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<mods:title id="C861BD1F677729D0C18DF9FB7969F66D">Taxonomic studies on the ant genus &lt;i&gt; Lepisiota &lt;/ i&gt; Santschi 1926 (Hymenoptera: Formicidae: Formicinae) in India, with description of four new species</mods:title>
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<emphasis id="B963EADE1F3EFFA48982FE16964CCC82" bold="true" box="[462,545,391,418]" pageId="4" pageNumber="4">Genus</emphasis>
<taxonomicName id="4C174D4F1F3EFFA48A64FE169738CC82" ID-CoL="5CN6" authority="Santschi, 1926" authorityName="Santschi" authorityYear="1926" box="[552,853,391,418]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EADE1F3EFFA48A64FE1696F1CC82" bold="true" box="[552,668,391,418]" italics="true" pageId="4" pageNumber="4">Lepisiota</emphasis>
<bibRefCitation id="EF864B3D1F3EFFA48AE9FE169738CC82" author="Santschi F." box="[677,853,391,418]" journalOrPublisher="Annales de la Societe Entomologique de France." pageId="4" pageNumber="4" pagination="13 - 28" part="95" refId="ref13237" refString="Santschi F. 1926. Trois notes myrmecologiques. Annales de la Societe Entomologique de France. 95: 13 - 28." title="Trois notes myrmecologiques" type="journal article" year="1926">Santschi, 1926</bibRefCitation>
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<typeStatus id="54AC886E1F3EFFA4888CFE519493CCFA" box="[192,254,448,474]" pageId="4" pageNumber="4">Type</typeStatus>
species:
<taxonomicName id="4C174D4F1F3EFFA48920FE2E9637CCF9" authorityName="Menozzi" authorityYear="1939" box="[364,602,447,473]" class="Insecta" family="Formicidae" genus="Plagiolepis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B963EADE1F3EFFA48920FE2E9637CCF9" box="[364,602,447,473]" italics="true" pageId="4" pageNumber="4">Plagiolepis rothneyi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF864B3D1F3EFFA48A26FE2E9698CCF9" author="Forel A." box="[618,757,447,473]" journalOrPublisher="Journal of the Bombay Natural History Society." pageId="4" pageNumber="4" pagination="396 - 420" part="8" refId="ref12813" refString="Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. Journal of the Bombay Natural History Society. 8: 396 - 420." title="Les Formicides de l'Empire des Indes et de Ceylan. Part IV" type="journal article" year="1894">Forel, 1894</bibRefCitation>
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<paragraph id="8BA836CC1F3EFFA4888CFD9A95E3CF05" blockId="4.[192,1123,523,660]" box="[192,398,523,549]" pageId="4" pageNumber="4">
<heading id="D0E081A01F3EFFA4888CFD9A95E3CF05" bold="true" box="[192,398,523,549]" fontSize="11" level="4" pageId="4" pageNumber="4" reason="6">
<emphasis id="B963EADE1F3EFFA4888CFD9A95E3CF05" bold="true" box="[192,398,523,549]" italics="true" pageId="4" pageNumber="4">Worker diagnosis</emphasis>
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Antennae 11 segmented; propodeum armed with a pair of spines, teeth, or tubercles; the dorsal edge of petiole armed with a pair of teeth or spines but sometimes only emarginated; acidopore well-developed (
<bibRefCitation id="EF864B3D1F3EFFA48BCFFDEB9071CFB4" author="Bolton B." bookContentInfo="222 pp." box="[899,1052,634,660]" journalOrPublisher="Cambridge (MA) and London: Harvard University Press" pageId="4" pageNumber="4" refId="ref12376" refString="Bolton B. 1994. Identification guide to the ant genera of the world. Cambridge (MA) and London: Harvard University Press. 222 pp." title="Identification guide to the ant genera of the world" type="book" year="1994">Bolton 1994</bibRefCitation>
).
</paragraph>
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<emphasis id="B963EADE1F3EFFA4888CFD49971DCFD3" bold="true" box="[192,880,728,755]" pageId="4" pageNumber="4">
List of the Indian species of
<taxonomicName id="4C174D4F1F3EFFA48A78FD49971DCFD3" authority="Santschi, 1926" authorityName="Santschi" authorityYear="1926" box="[564,880,728,755]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EADE1F3EFFA48A78FD4996C5CFD3" bold="true" box="[564,680,728,755]" italics="true" pageId="4" pageNumber="4">Lepisiota</emphasis>
<bibRefCitation id="EF864B3D1F3EFFA48AFCFD49971DCFD3" author="Santschi F." box="[688,880,728,755]" journalOrPublisher="Annales de la Societe Entomologique de France." pageId="4" pageNumber="4" pagination="13 - 28" part="95" refId="ref13237" refString="Santschi F. 1926. Trois notes myrmecologiques. Annales de la Societe Entomologique de France. 95: 13 - 28." title="Trois notes myrmecologiques" type="journal article" year="1926">Santschi, 1926</bibRefCitation>
</taxonomicName>
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<paragraph id="8BA836CC1F3EFFA4888CFC8196F3CE0A" blockId="4.[192,793,783,1407]" box="[192,670,783,810]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFC8196F3CE0A" authority="(Mukerjee, 1930)" baseAuthorityName="Mukerjee" baseAuthorityYear="1930" box="[192,670,783,810]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="annandalei">
<emphasis id="B963EADE1F3EFFA4888CFC8195ADCE09" box="[192,448,783,809]" italics="true" pageId="4" pageNumber="4">Lepisiota annandalei</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA4899CFC9E96FECE09" author="Mukerjee D." box="[464,659,783,809]" journalOrPublisher="Journal of the Bombay Natural History Society." pageId="4" pageNumber="4" pagination="149 - 163" part="34" refId="ref13093" refString="Mukerjee D. 1930. Report on a collection of ants in the Indian Museum, Calcutta. Journal of the Bombay Natural History Society. 34: 149 - 163." title="Report on a collection of ants in the Indian Museum, Calcutta" type="journal article" year="1930">Mukerjee, 1930</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFCA4967ACE6F" blockId="4.[192,793,783,1407]" box="[192,535,821,847]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFCA495C7CE6F" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[192,426,821,847]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="binghami" status="sp. nov.">
<emphasis id="B963EADE1F3EFFA4888CFCA495C7CE6F" box="[192,426,821,847]" italics="true" pageId="4" pageNumber="4">Lepisiota binghami</emphasis>
</taxonomicName>
<emphasis id="B963EADE1F3EFFA489FEFCA4967ACE6F" bold="true" box="[434,535,821,847]" pageId="4" pageNumber="4">
<taxonomicNameLabel id="A25057A51F3EFFA489FEFCA4967ACE6F" box="[434,535,821,847]" pageId="4" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFCCA963ECE54" blockId="4.[192,793,783,1407]" box="[192,595,858,884]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFCCA963ECE54" authority="(Smith, 1861)" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[192,595,858,884]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="bipartita">
<emphasis id="B963EADE1F3EFFA4888CFCCA95CCCE54" box="[192,417,858,884]" italics="true" pageId="4" pageNumber="4">Lepisiota bipartita</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489FEFCCB9625CE54" author="Smith F." box="[434,584,858,884]" journalOrPublisher="Journal and Proceedings of the Linnean Society of London, Zoology." pageId="4" pageNumber="4" pagination="31 - 35" part="6" refId="ref13333" refString="Smith F. 1861. Descriptions of some new species of ants from the Holy Land, with a synonymic list of others previously described. Journal and Proceedings of the Linnean Society of London, Zoology. 6 (21): 31 - 35. doi: 10.1111 / j. 1096 - 3642.1861. tb 00926. x." title="Descriptions of some new species of ants from the Holy Land, with a synonymic list of others previously described" type="journal article" year="1861">Smith, 1861</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFC119620CEBA" blockId="4.[192,793,783,1407]" box="[192,589,895,922]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFC119620CEBA" authority="(Forel, 1895)" baseAuthorityName="Forel" baseAuthorityYear="1895" box="[192,589,895,922]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="fergusoni">
<emphasis id="B963EADE1F3EFFA4888CFC1195C9CEB9" box="[192,420,895,921]" italics="true" pageId="4" pageNumber="4">Lepisiota fergusoni</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489FAFCEE962FCEBA" author="Forel A." box="[438,578,895,922]" journalOrPublisher="Journal of the Bombay Natural History Society." pageId="4" pageNumber="4" pagination="453 - 472" part="9" refId="ref12845" refString="Forel A. 1895. Les Formicides de l'Empire des Indes et de Ceylan. Part V. Journal of the Bombay Natural History Society. 9: 453 - 472." title="Les Formicides de l'Empire des Indes et de Ceylan. Part V" type="journal article" year="1895">Forel, 1895</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFC349659CE9F" blockId="4.[192,793,783,1407]" box="[192,564,933,959]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFC349659CE9F" authority="(Forel, 1894)" baseAuthorityName="Forel" baseAuthorityYear="1894" box="[192,564,933,959]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="B963EADE1F3EFFA4888CFC3495E6CE9E" box="[192,395,933,958]" italics="true" pageId="4" pageNumber="4">Lepisiota integra</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489D1FC349644CE9F" author="Forel A." box="[413,553,933,959]" journalOrPublisher="Journal of the Bombay Natural History Society." pageId="4" pageNumber="4" pagination="396 - 420" part="8" refId="ref12813" refString="Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. Journal of the Bombay Natural History Society. 8: 396 - 420." title="Les Formicides de l'Empire des Indes et de Ceylan. Part IV" type="journal article" year="1894">Forel, 1894</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFC5A9766CEC4" blockId="4.[192,793,783,1407]" box="[192,779,970,996]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFC5A9766CEC4" authority="Wachkoo, Bharti &amp; Akbar, 2021" authorityName="Wachkoo, Bharti &amp; Akbar" authorityYear="2021" box="[192,779,970,996]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="layla">
<emphasis id="B963EADE1F3EFFA4888CFC5A951CCEC4" box="[192,369,970,996]" italics="true" pageId="4" pageNumber="4">Lepisiota layla</emphasis>
<bibRefCitation id="EF864B3D1F3EFFA48935FC5B9766CEC4" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" box="[377,779,970,996]" journalOrPublisher="Bonn Zoological Bulletin." pageId="4" pageNumber="4" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo, Bharti &amp; Akbar, 2021</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFC619628C92A" blockId="4.[192,793,783,1407]" box="[192,581,1007,1034]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFC619628C92A" authority="(Emery, 1893)" baseAuthorityName="Emery" baseAuthorityYear="1893" box="[192,581,1007,1034]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="lunaris">
<emphasis id="B963EADE1F3EFFA4888CFC6195E6C929" box="[192,395,1007,1033]" italics="true" pageId="4" pageNumber="4">Lepisiota lunaris</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489D0FC7E9657C929" author="Emery C." box="[412,570,1007,1033]" journalOrPublisher="Annales de la Societe Entomologique de France." pageId="4" pageNumber="4" pagination="239 - 258" part="62" refId="ref12601" refString="Emery C. 1893 a. Voyage de M. E. Simon a l'ile de Ceylan (janvier-fevrier 1892). Formicides. Annales de la Societe Entomologique de France. 62: 239 - 258." title="Voyage de M. E. Simon a l'ile de Ceylan (janvier-fevrier 1892). Formicides" type="journal article" year="1893">Emery, 1893</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFB849774C90F" blockId="4.[192,793,783,1407]" box="[192,793,1045,1071]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFB849774C90F" authority="Wachkoo, Bharti &amp; Akbar, 2021" authorityName="Wachkoo, Bharti &amp; Akbar" authorityYear="2021" box="[192,793,1045,1071]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B963EADE1F3EFFA4888CFB849513C90E" box="[192,382,1045,1070]" italics="true" pageId="4" pageNumber="4">Lepisiota mayri</emphasis>
<bibRefCitation id="EF864B3D1F3EFFA489CBFB849774C90F" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" box="[391,793,1045,1071]" journalOrPublisher="Bonn Zoological Bulletin." pageId="4" pageNumber="4" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo, Bharti &amp; Akbar, 2021</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFBAA9629C975" blockId="4.[192,793,783,1407]" box="[192,580,1082,1109]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFBAA9629C975" authority="(Forel, 1894)" baseAuthorityName="Forel" baseAuthorityYear="1894" box="[192,580,1082,1109]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B963EADE1F3EFFA4888CFBAA95F6C974" box="[192,411,1082,1108]" italics="true" pageId="4" pageNumber="4">Lepisiota modesta</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489E0FBAB9654C974" author="Forel A." box="[428,569,1082,1108]" journalOrPublisher="Journal of the Bombay Natural History Society." pageId="4" pageNumber="4" pagination="396 - 420" part="8" refId="ref12813" refString="Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. Journal of the Bombay Natural History Society. 8: 396 - 420." title="Les Formicides de l'Empire des Indes et de Ceylan. Part IV" type="journal article" year="1894">Forel, 1894</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFBF1964BC95A" blockId="4.[192,793,783,1407]" box="[192,550,1119,1146]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFBF1964BC95A" authority="(Forel, 1892)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[192,550,1119,1146]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B963EADE1F3EFFA4888CFBF19510C959" box="[192,381,1120,1145]" italics="true" pageId="4" pageNumber="4">Lepisiota opaca</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489C2FBCE9676C959" author="Forel A." box="[398,539,1119,1145]" journalOrPublisher="Annales de la Societe Entomologique de Belgique." pageId="4" pageNumber="4" pagination="1 - 6" part="36" refId="ref12791" refString="Forel A. 1892. Notes myrmecologiques. Annales de la Societe Entomologique de Belgique. 36: 1 - 6." title="Notes myrmecologiques" type="journal article" year="1892">Forel, 1892</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFB149621C9BF" blockId="4.[192,793,783,1407]" box="[192,588,1157,1183]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFB149621C9BF" authority="(Forel, 1892)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[192,588,1157,1183]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="B963EADE1F3EFFA4888CFB1495CEC9BF" box="[192,419,1157,1183]" italics="true" pageId="4" pageNumber="4">Lepisiota pulchella</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489F8FB14962CC9BF" author="Forel A." box="[436,577,1157,1183]" journalOrPublisher="Annales de la Societe Entomologique de Belgique." pageId="4" pageNumber="4" pagination="1 - 6" part="36" refId="ref12791" refString="Forel A. 1892. Notes myrmecologiques. Annales de la Societe Entomologique de Belgique. 36: 1 - 6." title="Notes myrmecologiques" type="journal article" year="1892">Forel, 1892</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFB3A967BC9E4" blockId="4.[192,793,783,1407]" box="[192,534,1194,1220]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFB3A95C5C9E4" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[192,424,1195,1220]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="pusaensis" status="sp. nov.">
<emphasis id="B963EADE1F3EFFA4888CFB3A95C5C9E4" box="[192,424,1195,1220]" italics="true" pageId="4" pageNumber="4">Lepisiota pusaensis</emphasis>
</taxonomicName>
<emphasis id="B963EADE1F3EFFA489FDFB3B967BC9E4" bold="true" box="[433,534,1194,1220]" pageId="4" pageNumber="4">
<taxonomicNameLabel id="A25057A51F3EFFA489FDFB3B967BC9E4" box="[433,534,1194,1220]" pageId="4" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFB419629C9CA" blockId="4.[192,793,783,1407]" box="[192,580,1231,1258]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFB419629C9CA" authority="(Forel, 1894)" baseAuthorityName="Forel" baseAuthorityYear="1894" box="[192,580,1231,1258]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B963EADE1F3EFFA4888CFB4195F6C9C9" box="[192,411,1231,1257]" italics="true" pageId="4" pageNumber="4">Lepisiota rothneyi</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489E0FB5E9654C9C9" author="Forel A." box="[428,569,1231,1257]" journalOrPublisher="Journal of the Bombay Natural History Society." pageId="4" pageNumber="4" pagination="396 - 420" part="8" refId="ref12813" refString="Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. Journal of the Bombay Natural History Society. 8: 396 - 420." title="Les Formicides de l'Empire des Indes et de Ceylan. Part IV" type="journal article" year="1894">Forel, 1894</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFB649654C82F" blockId="4.[192,793,783,1407]" box="[192,569,1269,1295]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFB6495A1C82E" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[192,460,1269,1294]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="satpuraensis" status="sp. nov.">
<emphasis id="B963EADE1F3EFFA4888CFB6495A1C82E" box="[192,460,1269,1294]" italics="true" pageId="4" pageNumber="4">Lepisiota satpuraensis</emphasis>
</taxonomicName>
<emphasis id="B963EADE1F3EFFA48999FB649654C82F" bold="true" box="[469,569,1269,1295]" pageId="4" pageNumber="4">
<taxonomicNameLabel id="A25057A51F3EFFA48999FB649654C82F" box="[469,569,1269,1295]" pageId="4" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFA8A965DC814" blockId="4.[192,793,783,1407]" box="[192,560,1306,1332]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFA8A965DC814" authority="(Forel, 1892)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[192,560,1306,1332]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="sericea">
<emphasis id="B963EADE1F3EFFA4888CFA8A95EAC814" box="[192,391,1307,1332]" italics="true" pageId="4" pageNumber="4">Lepisiota sericea</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA489D4FA8B9648C814" author="Forel A." box="[408,549,1306,1332]" journalOrPublisher="Annales de la Societe Entomologique de Belgique." pageId="4" pageNumber="4" pagination="1 - 6" part="36" refId="ref12791" refString="Forel A. 1892. Notes myrmecologiques. Annales de la Societe Entomologique de Belgique. 36: 1 - 6." title="Notes myrmecologiques" type="journal article" year="1892">Forel, 1892</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFAD19594C879" blockId="4.[192,793,783,1407]" box="[192,505,1343,1369]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFAD195E1C879" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[192,396,1343,1369]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
<emphasis id="B963EADE1F3EFFA4888CFAD195E1C879" box="[192,396,1343,1369]" italics="true" pageId="4" pageNumber="4">Lepisiota wilsoni</emphasis>
</taxonomicName>
<emphasis id="B963EADE1F3EFFA489D9FAAE9594C879" bold="true" box="[405,505,1343,1369]" pageId="4" pageNumber="4">
<taxonomicNameLabel id="A25057A51F3EFFA489D9FAAE9594C879" box="[405,505,1343,1369]" pageId="4" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BA836CC1F3EFFA4888CFAF4961EC85F" blockId="4.[192,793,783,1407]" box="[192,627,1380,1407]" pageId="4" pageNumber="4">
<taxonomicName id="4C174D4F1F3EFFA4888CFAF4961EC85F" authority="(Forel, 1902)" baseAuthorityName="Forel" baseAuthorityYear="1902" box="[192,627,1380,1407]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="B963EADE1F3EFFA4888CFAF495A7C85E" box="[192,458,1380,1406]" italics="true" pageId="4" pageNumber="4">Lepisiota wroughtonii</emphasis>
(
<bibRefCitation id="EF864B3D1F3EFFA48997FAF59605C85F" author="Forel A." box="[475,616,1380,1407]" journalOrPublisher="Annales de la Societe Entomologique de Belgique." pageId="4" pageNumber="4" pagination="284 - 296" part="46" refId="ref12877" refString="Forel A. 1902. Varietes myrmecologiques. Annales de la Societe Entomologique de Belgique. 46: 284 - 296." title="Varietes myrmecologiques" type="journal article" year="1902">Forel, 1902</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30D65471F3EFFA4888CFA2195A4C8CF" pageId="4" pageNumber="4" type="multiple">
<paragraph id="8BA836CC1F3EFFA4888CFA2195A4C8CF" blockId="4.[192,1123,1455,1519]" pageId="4" pageNumber="4">
Key to the workers of Indian species of
<taxonomicName id="4C174D4F1F3EFFA48AFBFA21978DC8E9" authority="Santschi, 1926" authorityName="Santschi" authorityYear="1926" box="[695,992,1455,1481]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EADE1F3EFFA48AFBFA219749C8E9" box="[695,804,1456,1481]" italics="true" pageId="4" pageNumber="4">Lepisiota</emphasis>
<bibRefCitation id="EF864B3D1F3EFFA48B62FA3E978DC8E9" author="Santschi F." box="[814,992,1455,1481]" journalOrPublisher="Annales de la Societe Entomologique de France." pageId="4" pageNumber="4" pagination="13 - 28" part="95" refId="ref13237" refString="Santschi F. 1926. Trois notes myrmecologiques. Annales de la Societe Entomologique de France. 95: 13 - 28." title="Trois notes myrmecologiques" type="journal article" year="1926">Santschi, 1926</bibRefCitation>
</taxonomicName>
(based on
<bibRefCitation id="EF864B3D1F3EFFA4888CFA4495D3C8CF" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" box="[192,446,1493,1519]" journalOrPublisher="Bonn Zoological Bulletin." pageId="4" pageNumber="4" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30D65471F3EFFA68893F98B900ECEB1" lastPageId="6" lastPageNumber="6" pageId="4" pageNumber="4" type="key">
<key id="FF8661FD1F3EFFA68893F98B900ECEB1" lastPageId="6" lastPageNumber="6" pageId="4" pageNumber="4">
<keyStep id="30E32E691F3EFFA48893F98B9009CB84" pageId="4" pageNumber="4">
<paragraph id="8BA836CC1F3EFFA48893F98B9009CB79" blockId="4.[223,1124,1562,1701]" pageId="4" pageNumber="4">
<keyLead id="30E695F91F3EFFA48893F98B9009CB79" pageId="4" pageNumber="4">
1. Antennal scape long, surpassing posterior margin of head by about half of its length or more (
<figureCitation id="132C2A491F3EFFA48A58F9AE961DCB7A" box="[532,624,1599,1626]" captionStart="Figure 5" captionStartId="19.[192,257,1090,1113]" captionTargetBox="[194,1119,141,1057]" captionTargetId="figure-136@19.[193,1123,140,1059]" captionTargetPageId="19" captionText="Figure 5AF. Workers of Lepisiota annandalei (AC), Lepisiota bipartita (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152944" httpUri="https://zenodo.org/record/7152944/files/figure.png" pageId="4" pageNumber="4">Figs 5E</figureCitation>
,
<figureCitation id="132C2A491F3EFFA48A32F9D196F2CB7A" box="[638,671,1600,1626]" captionStart="Figure 7" captionStartId="26.[192,257,996,1019]" captionTargetBox="[194,1122,141,962]" captionTargetId="figure-156@26.[193,1123,140,964]" captionTargetPageId="26" captionText="Figure 7AF. Workers of Lepisiota integra (AC), Lepisiota layla (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152948" httpUri="https://zenodo.org/record/7152948/files/figure.png" pageId="4" pageNumber="4">7B</figureCitation>
)........................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3EFFA48893F9F49009CB84" blockId="4.[223,1124,1562,1701]" pageId="4" pageNumber="4">
<keyLead id="30E695F91F3EFFA48893F9F49009CB84" pageId="4" pageNumber="4">
. Antennal scape short, surpassing posterior margin of head by a third of its length or less (
<figureCitation id="132C2A491F3EFFA48990F91B965BCB84" box="[476,566,1674,1700]" captionStart="Figure 8" captionStartId="30.[193,258,1000,1023]" captionTargetBox="[194,1121,143,962]" captionTargetId="figure-154@30.[193,1123,140,968]" captionTargetPageId="30" captionText="Figure 8AF. Syntype worker (AntWeb: CASENT0909893, photographed by Zach Lieberman) of Lepisiota opaca (AC), worker of Lepisiota pulchella (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152950" httpUri="https://zenodo.org/record/7152950/files/figure.png" pageId="4" pageNumber="4">Fig. 8C</figureCitation>
).......................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FF1C9009CDD1" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FF1C9009CDED" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FF1C9009CDED" pageId="5" pageNumber="5">
2. Bicoloured species; head and gaster dark brown to black while mesosoma reddish-brown (
<figureCitation id="132C2A491F3FFFA58A58FF2396FDCDED" box="[532,656,178,205]" captionStart="Figure 5" captionStartId="19.[192,257,1090,1113]" captionTargetBox="[194,1119,141,1057]" captionTargetId="figure-136@19.[193,1123,140,1059]" captionTargetPageId="19" captionText="Figure 5AF. Workers of Lepisiota annandalei (AC), Lepisiota bipartita (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152944" httpUri="https://zenodo.org/record/7152944/files/figure.png" pageId="5" pageNumber="5">Fig. 5DF</figureCitation>
)............................
<taxonomicName id="4C174D4F1F3FFFA58B26FF239009CDED" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[874,1124,177,205]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="bipartita">
<emphasis id="B963EADE1F3FFFA58B26FF239791CDEC" bold="true" box="[874,1020,177,204]" italics="true" pageId="5" pageNumber="5">L. bipartita</emphasis>
(Smith)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58890FF499009CDD1" blockId="5.[220,1124,141,1736]" box="[220,1124,215,242]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58890FF499009CDD1" box="[220,1124,215,242]" pageId="5" pageNumber="5">. Uniformly dark-brown or black-brown species..................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FF6C900ECCF2" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FF6C900ECC42" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FF6C900ECC42" pageId="5" pageNumber="5">
3. Dark brown species; petiolar scale with distinctly angular sides, dorsally emarginate with teeth-like apical corners (
<figureCitation id="132C2A491F3FFFA58BC0FEB39787CC1C" box="[908,1002,290,316]" captionStart="Figure 7" captionStartId="26.[192,257,996,1019]" captionTargetBox="[194,1122,141,962]" captionTargetId="figure-156@26.[193,1123,140,964]" captionTargetPageId="26" captionText="Figure 7AF. Workers of Lepisiota integra (AC), Lepisiota layla (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152948" httpUri="https://zenodo.org/record/7152948/files/figure.png" pageId="5" pageNumber="5">Fig. 7A</figureCitation>
)............... ..........................................................................................
<taxonomicName id="4C174D4F1F3FFFA58BCFFED9900ECC42" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1894" box="[899,1123,327,354]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="B963EADE1F3FFFA58BCFFED9906FCC41" bold="true" box="[899,1026,327,354]" italics="true" pageId="5" pageNumber="5">L. integra</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FEFC900ECCF2" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FEFC900ECCF2" pageId="5" pageNumber="5">
. Black-brown species; petiolar scale with smoothly curved sides, dorsally rounded and narrow, without teeth (
<bibRefCitation id="EF864B3D1F3FFFA58A89FE0397A8CC8C" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" box="[709,965,402,429]" journalOrPublisher="Bonn Zoological Bulletin." pageId="5" pageNumber="5" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
, Figs 3536) ............................................................................................
<taxonomicName id="4C174D4F1F3FFFA58BC2FE29900ECCF2" ID-CoL="3TFGN" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[910,1123,439,466]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="sericea">
<emphasis id="B963EADE1F3FFFA58BC2FE29906BCCF1" bold="true" box="[910,1030,439,466]" italics="true" pageId="5" pageNumber="5">L. sericea</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FE4C900ECF3C" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FE4C9009CCD7" blockId="5.[220,1124,141,1736]" box="[223,1124,476,503]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FE4C9009CCD7" box="[223,1124,476,503]" pageId="5" pageNumber="5">
4. Bicoloured species (
<figureCitation id="132C2A491F3FFFA589BAFE4C9633CCD7" box="[502,606,477,503]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="7.[192,257,1643,1666]" captionStartId-1="10.[192,257,1643,1666]" captionStartId-2="14.[192,257,1759,1782]" captionTargetBox-0="[243,1072,191,1610]" captionTargetBox-1="[196,1120,141,1610]" captionTargetBox-2="[218,1096,142,1726]" captionTargetId-0="figure-3@7.[241,1074,140,1612]" captionTargetId-1="figure-3@10.[193,1123,140,1612]" captionTargetId-2="figure-3@14.[217,1098,140,1728]" captionTargetPageId-0="7" captionTargetPageId-1="10" captionTargetPageId-2="14" captionText-0="Figure 1AC. Lepisiota binghami sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." captionText-1="Figure 2AC. Lepisiota pusaensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." captionText-2="Figure 3AC. Lepisiota satpuraensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi-0="http://doi.org/10.5281/zenodo.7152935" figureDoi-1="http://doi.org/10.5281/zenodo.7152938" figureDoi-2="http://doi.org/10.5281/zenodo.7152940" httpUri-0="https://zenodo.org/record/7152935/files/figure.png" httpUri-1="https://zenodo.org/record/7152938/files/figure.png" httpUri-2="https://zenodo.org/record/7152940/files/figure.png" pageId="5" pageNumber="5">Figs 13</figureCitation>
,
<figureCitation id="132C2A491F3FFFA58A21FE4C9611CCD7" box="[621,636,477,503]" captionStart="Figure 8" captionStartId="30.[193,258,1000,1023]" captionTargetBox="[194,1121,143,962]" captionTargetId="figure-154@30.[193,1123,140,968]" captionTargetPageId="30" captionText="Figure 8AF. Syntype worker (AntWeb: CASENT0909893, photographed by Zach Lieberman) of Lepisiota opaca (AC), worker of Lepisiota pulchella (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152950" httpUri="https://zenodo.org/record/7152950/files/figure.png" pageId="5" pageNumber="5">8</figureCitation>
)................................................................ 5
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FD92900ECF3C" blockId="5.[220,1124,141,1736]" box="[223,1123,514,541]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FD92900ECF3C" box="[223,1123,514,541]" pageId="5" pageNumber="5">
. Uniformly dark brown to black species (
<figureCitation id="132C2A491F3FFFA58ABBFD939711CF3D" box="[759,892,514,541]" captionStart="Figure 5" captionStartId="19.[192,257,1090,1113]" captionTargetBox="[194,1119,141,1057]" captionTargetId="figure-136@19.[193,1123,140,1059]" captionTargetPageId="19" captionText="Figure 5AF. Workers of Lepisiota annandalei (AC), Lepisiota bipartita (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152944" httpUri="https://zenodo.org/record/7152944/files/figure.png" pageId="5" pageNumber="5">Figs 5AC</figureCitation>
,
<figureCitation id="132C2A491F3FFFA58BC6FD9297A2CF3D" box="[906,975,515,541]" captionStart="Figure 7" captionStartId="26.[192,257,996,1019]" captionTargetBox="[194,1122,141,962]" captionTargetId="figure-156@26.[193,1123,140,964]" captionTargetPageId="26" captionText="Figure 7AF. Workers of Lepisiota integra (AC), Lepisiota layla (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152948" httpUri="https://zenodo.org/record/7152948/files/figure.png" pageId="5" pageNumber="5">7DF</figureCitation>
) ............. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FDB69009CF47" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FDB69009CF61" blockId="5.[220,1124,141,1736]" box="[223,1124,551,578]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FDB69009CF61" box="[223,1124,551,578]" pageId="5" pageNumber="5">
5. Body shiny to subopaque; weakly sculptured (
<figureCitation id="132C2A491F3FFFA58B76FDB697CFCF61" box="[826,930,551,577]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="7.[192,257,1643,1666]" captionStartId-1="10.[192,257,1643,1666]" captionStartId-2="14.[192,257,1759,1782]" captionTargetBox-0="[243,1072,191,1610]" captionTargetBox-1="[196,1120,141,1610]" captionTargetBox-2="[218,1096,142,1726]" captionTargetId-0="figure-3@7.[241,1074,140,1612]" captionTargetId-1="figure-3@10.[193,1123,140,1612]" captionTargetId-2="figure-3@14.[217,1098,140,1728]" captionTargetPageId-0="7" captionTargetPageId-1="10" captionTargetPageId-2="14" captionText-0="Figure 1AC. Lepisiota binghami sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." captionText-1="Figure 2AC. Lepisiota pusaensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." captionText-2="Figure 3AC. Lepisiota satpuraensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi-0="http://doi.org/10.5281/zenodo.7152935" figureDoi-1="http://doi.org/10.5281/zenodo.7152938" figureDoi-2="http://doi.org/10.5281/zenodo.7152940" httpUri-0="https://zenodo.org/record/7152935/files/figure.png" httpUri-1="https://zenodo.org/record/7152938/files/figure.png" httpUri-2="https://zenodo.org/record/7152940/files/figure.png" pageId="5" pageNumber="5">Figs 13</figureCitation>
)...................... 6
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FDDC9009CF47" blockId="5.[220,1124,141,1736]" box="[223,1124,589,615]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FDDC9009CF47" box="[223,1124,589,615]" pageId="5" pageNumber="5">
. Body dull; coarsely sculptured (
<figureCitation id="132C2A491F3FFFA58AC4FDDC96AACF47" box="[648,711,589,615]" captionStart="Figure 8" captionStartId="30.[193,258,1000,1023]" captionTargetBox="[194,1121,143,962]" captionTargetId="figure-154@30.[193,1123,140,968]" captionTargetPageId="30" captionText="Figure 8AF. Syntype worker (AntWeb: CASENT0909893, photographed by Zach Lieberman) of Lepisiota opaca (AC), worker of Lepisiota pulchella (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152950" httpUri="https://zenodo.org/record/7152950/files/figure.png" pageId="5" pageNumber="5">Fig 8</figureCitation>
) ..................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FDE39009CE97" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FDE3900ECFDC" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FDE3900ECFDC" pageId="5" pageNumber="5">
6. Body with scanty pilosity; posterior margin of head with 23 erect setae, dorsum of pronotum with 24 long erect setae, mesonotum with 12 erect setae, metanotum lacks erect setae and propodeum with 01 erect setae (
<figureCitation id="132C2A491F3FFFA589DEFD739581CFDC" box="[402,492,738,764]" captionStart="Figure 1" captionStartId="7.[192,257,1643,1666]" captionTargetBox="[243,1072,191,1610]" captionTargetId="figure-3@7.[241,1074,140,1612]" captionTargetPageId="7" captionText="Figure 1AC. Lepisiota binghami sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152935" httpUri="https://zenodo.org/record/7152935/files/figure.png" pageId="5" pageNumber="5">Fig. 1A</figureCitation>
) .................................................
<taxonomicName id="4C174D4F1F3FFFA58B2CFD739797CFDC" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[864,1018,737,764]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="binghami" status="sp. nov.">
<emphasis id="B963EADE1F3FFFA58B2CFD739797CFDC" bold="true" box="[864,1018,737,764]" italics="true" pageId="5" pageNumber="5">L. binghami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057A51F3FFFA58C4FFD73900ECFDC" box="[1027,1123,738,764]" pageId="5" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FC999009CE97" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FC999009CE97" pageId="5" pageNumber="5">
. Body with abundant pilosity; posterior margin of head with 45 erect setae or more, dorsum of pronotum with more than 10 long erect setae, mesonotum with four or more erect setae, metanotum with two or more erect setae, and propodeum with 46 or more erect setae (
<figureCitation id="132C2A491F3FFFA58C7DFCE69548CE97" captionStart="Figure 2" captionStartId="10.[192,257,1643,1666]" captionTargetBox="[196,1120,141,1610]" captionTargetId="figure-3@10.[193,1123,140,1612]" captionTargetPageId="10" captionText="Figure 2AC. Lepisiota pusaensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152938" httpUri="https://zenodo.org/record/7152938/files/figure.png" pageId="5" pageNumber="5">Figs 2A</figureCitation>
,
<figureCitation id="132C2A491F3FFFA5897EFC0C953BCE97" box="[306,342,925,951]" captionStart="Figure 3" captionStartId="14.[192,257,1759,1782]" captionTargetBox="[218,1096,142,1726]" captionTargetId="figure-3@14.[217,1098,140,1728]" captionTargetPageId="14" captionText="Figure 3AC. Lepisiota satpuraensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152940" httpUri="https://zenodo.org/record/7152940/files/figure.png" pageId="5" pageNumber="5">3A</figureCitation>
).......................................................................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FC53900EC96D" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FC53900EC922" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FC53900EC922" pageId="5" pageNumber="5">
7. Smaller species (HL 0.580.59, WL 0.830.85); body with appressed pubescence (
<figureCitation id="132C2A491F3FFFA589D3FC769589C921" box="[415,484,999,1025]" captionStart="Figure 2" captionStartId="10.[192,257,1643,1666]" captionTargetBox="[196,1120,141,1610]" captionTargetId="figure-3@10.[193,1123,140,1612]" captionTargetPageId="10" captionText="Figure 2AC. Lepisiota pusaensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152938" httpUri="https://zenodo.org/record/7152938/files/figure.png" pageId="5" pageNumber="5">Fig. 2</figureCitation>
)......................... .........................
<taxonomicName id="4C174D4F1F3FFFA58B2DFC799796C921" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[865,1019,999,1026]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="pusaensis" status="sp. nov.">
<emphasis id="B963EADE1F3FFFA58B2DFC799796C921" bold="true" box="[865,1019,999,1026]" italics="true" pageId="5" pageNumber="5">L. pusaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057A51F3FFFA58C4FFC79900EC922" box="[1027,1123,1000,1026]" pageId="5" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FB9C900EC96D" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FB9C900EC96D" pageId="5" pageNumber="5">
. Larger species (HL 0.800.83, WL 1.121.21); body with semi-erect pubescence (
<figureCitation id="132C2A491F3FFFA589D3FBA39589C96C" box="[415,484,1074,1100]" captionStart="Figure 3" captionStartId="14.[192,257,1759,1782]" captionTargetBox="[218,1096,142,1726]" captionTargetId="figure-3@14.[217,1098,140,1728]" captionTargetPageId="14" captionText="Figure 3AC. Lepisiota satpuraensis sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152940" httpUri="https://zenodo.org/record/7152940/files/figure.png" pageId="5" pageNumber="5">Fig. 3</figureCitation>
)..............................................
<taxonomicName id="4C174D4F1F3FFFA58B77FBA39797C96C" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[827,1018,1074,1100]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="satpuraensis" status="sp. nov.">
<emphasis id="B963EADE1F3FFFA58B77FBA39797C96C" bold="true" box="[827,1018,1074,1100]" italics="true" pageId="5" pageNumber="5">L. satpuraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057A51F3FFFA58C4FFBA2900EC96D" box="[1027,1123,1075,1101]" pageId="5" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FBC69009C827" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FBC6900EC99D" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FBC6900EC99D" pageId="5" pageNumber="5">
8. Head reticulate-striate; propodeal spines blunt, directed backward; petiole dorsally emarginated (
<bibRefCitation id="EF864B3D1F3FFFA58A3CFBEC971CC9B7" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" box="[624,881,1149,1175]" journalOrPublisher="Bonn Zoological Bulletin." pageId="5" pageNumber="5" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
, Fig. 39)................... .......................................................................................
<taxonomicName id="4C174D4F1F3FFFA58B3CFB33900EC99D" ID-CoL="3TFFN" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1895" box="[880,1123,1186,1213]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="fergusoni">
<emphasis id="B963EADE1F3FFFA58B3CFB339068C99C" bold="true" box="[880,1029,1186,1212]" italics="true" pageId="5" pageNumber="5">L. fergusoni</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA588A8FB599009C827" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA588A8FB599009C827" pageId="5" pageNumber="5">. Head reticulate-punctate; propodeal spines pointed, directed upward; petiole dorsally bispinose........................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FA839009C857" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FA83900EC80D" blockId="5.[220,1124,141,1736]" box="[223,1123,1298,1325]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FA83900EC80D" box="[223,1123,1298,1325]" pageId="5" pageNumber="5">
9. Gaster smooth and shiny (
<figureCitation id="132C2A491F3FFFA58A05FA8396CDC80C" box="[585,672,1298,1324]" captionStart="Figure 8" captionStartId="30.[193,258,1000,1023]" captionTargetBox="[194,1121,143,962]" captionTargetId="figure-154@30.[193,1123,140,968]" captionTargetPageId="30" captionText="Figure 8AF. Syntype worker (AntWeb: CASENT0909893, photographed by Zach Lieberman) of Lepisiota opaca (AC), worker of Lepisiota pulchella (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152950" httpUri="https://zenodo.org/record/7152950/files/figure.png" pageId="5" pageNumber="5">Fig. 8B</figureCitation>
)................................
<taxonomicName id="4C174D4F1F3FFFA58BD5FA83900EC80D" ID-CoL="3TFGD" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[921,1123,1298,1325]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B963EADE1F3FFFA58BD5FA83906BC80C" bold="true" box="[921,1030,1298,1324]" italics="true" pageId="5" pageNumber="5">L. opaca</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FAA99009C857" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FAA99009C857" pageId="5" pageNumber="5">
. Gaster microreticulate and subopaque (
<figureCitation id="132C2A491F3FFFA58B42FAA69701C871" box="[782,876,1335,1361]" captionStart="Figure 8" captionStartId="30.[193,258,1000,1023]" captionTargetBox="[194,1121,143,962]" captionTargetId="figure-154@30.[193,1123,140,968]" captionTargetPageId="30" captionText="Figure 8AF. Syntype worker (AntWeb: CASENT0909893, photographed by Zach Lieberman) of Lepisiota opaca (AC), worker of Lepisiota pulchella (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152950" httpUri="https://zenodo.org/record/7152950/files/figure.png" pageId="5" pageNumber="5">Fig. 8E</figureCitation>
)............................... ..............................................................................
<taxonomicName id="4C174D4F1F3FFFA58B2CFACC9009C857" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[864,1124,1372,1399]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="B963EADE1F3FFFA58B2CFACC9793C857" bold="true" box="[864,1022,1372,1399]" italics="true" pageId="5" pageNumber="5">L. pulchella</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893FA13900ECB11" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893FA13900EC8E1" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FA13900EC8E1" pageId="5" pageNumber="5">10. Propodeal spines indistinct; gastral pilosity restricted to few pairs of black setae on the posterior margin of tergites .................................... 11</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893FA5C900ECB11" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893FA5C900ECB11" pageId="5" pageNumber="5">. Propodeal spines well-developed as two broad-based blunt tubercles, teeth or spines; whole gastral dorsum covered with abundant white to yellowish setae............................................................................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3FFFA58893F9AC900ECBE8" pageId="5" pageNumber="5">
<paragraph id="8BA836CC1F3FFFA58893F9AC900ECB5D" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893F9AC900ECB5D" pageId="5" pageNumber="5">
11. Pronotum without setae; body sparsely pubescent, shiny (
<bibRefCitation id="EF864B3D1F3FFFA58BA6F9AC9510CB5C" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" journalOrPublisher="Bonn Zoological Bulletin." pageId="5" pageNumber="5" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
, Fig. 32)......................................................
<taxonomicName id="4C174D4F1F3FFFA58B36F9F3900ECB5D" ID-CoL="3TFGH" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1894" box="[890,1123,1633,1661]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B963EADE1F3FFFA58B36F9F39068CB5C" bold="true" box="[890,1029,1633,1660]" italics="true" pageId="5" pageNumber="5">L. rothneyi</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3FFFA58893F919900ECBE8" blockId="5.[220,1124,141,1736]" pageId="5" pageNumber="5">
<keyLead id="30E695F91F3FFFA58893F919900ECBE8" pageId="5" pageNumber="5">
. Pronotum with few short setae; body fairly pubescent, opaque (
<bibRefCitation id="EF864B3D1F3FFFA58BA7F9169517CBE7" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" journalOrPublisher="Bonn Zoological Bulletin." pageId="5" pageNumber="5" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
, Fig. 38)....................................................
<taxonomicName id="4C174D4F1F3FFFA58B1CF93C900ECBE8" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1902" box="[848,1123,1708,1736]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="5" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="B963EADE1F3FFFA58B1CF93C9065CBE7" bold="true" box="[848,1032,1708,1735]" italics="true" pageId="5" pageNumber="5">L. wroughtonii</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3CFFA68893FF1C900ECDD1" pageId="6" pageNumber="6">
<paragraph id="8BA836CC1F3CFFA68893FF1C900ECDEC" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FF1C900ECDEC" pageId="6" pageNumber="6">
12. Body abundantly covered with long, erect white setae (
<figureCitation id="132C2A491F3CFFA68BFBFF1C905ECD87" box="[951,1075,141,167]" captionStart="Figure 7" captionStartId="26.[192,257,996,1019]" captionTargetBox="[194,1122,141,962]" captionTargetId="figure-156@26.[193,1123,140,964]" captionTargetPageId="26" captionText="Figure 7AF. Workers of Lepisiota integra (AC), Lepisiota layla (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152948" httpUri="https://zenodo.org/record/7152948/files/figure.png" pageId="6" pageNumber="6">Fig. 7DF</figureCitation>
)..... ............................................................
<taxonomicName id="4C174D4F1F3CFFA68AFFFF23900ECDEC" authority="Wachkoo, Bharti &amp; Akbar" authorityName="Wachkoo, Bharti &amp; Akbar" authorityYear="2021" box="[691,1123,177,204]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="layla">
<emphasis id="B963EADE1F3CFFA68AFFFF23977ECDEC" bold="true" box="[691,787,177,204]" italics="true" pageId="6" pageNumber="6">L. layla</emphasis>
Wachkoo, Bharti &amp; Akbar
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3CFFA68893FF49900ECDD1" blockId="6.[223,1123,141,914]" box="[223,1123,215,242]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FF49900ECDD1" box="[223,1123,215,242]" pageId="6" pageNumber="6">
. Body covered with usual short erect setae (
<figureCitation id="132C2A491F3CFFA68B59FF46970CCDD1" box="[789,865,215,241]" captionStart="Figure 4" captionStartId="17.[192,257,1682,1705]" captionTargetBox="[194,1121,204,1648]" captionTargetId="figure-3@17.[193,1123,140,1650]" captionTargetPageId="17" captionText="Figure 4AC. Lepisiota wilsoni sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152942" httpUri="https://zenodo.org/record/7152942/files/figure.png" pageId="6" pageNumber="6">Figs 4</figureCitation>
,
<figureCitation id="132C2A491F3CFFA68B22FF4697FFCDD2" box="[878,914,215,242]" captionStart="Figure 5" captionStartId="19.[192,257,1090,1113]" captionTargetBox="[194,1119,141,1057]" captionTargetId="figure-136@19.[193,1123,140,1059]" captionTargetPageId="19" captionText="Figure 5AF. Workers of Lepisiota annandalei (AC), Lepisiota bipartita (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152944" httpUri="https://zenodo.org/record/7152944/files/figure.png" pageId="6" pageNumber="6">5A</figureCitation>
)...................... 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3CFFA68893FF6C900ECCA7" pageId="6" pageNumber="6">
<paragraph id="8BA836CC1F3CFFA68893FF6C900ECC1D" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FF6C900ECC1D" pageId="6" pageNumber="6">
13. Petiole dorsally emarginated without distinct teeth or spines; scanty mesosomal setae (
<figureCitation id="132C2A491F3CFFA689ADFEB39656CC1D" box="[481,571,290,317]" captionStart="Figure 5" captionStartId="19.[192,257,1090,1113]" captionTargetBox="[194,1119,141,1057]" captionTargetId="figure-136@19.[193,1123,140,1059]" captionTargetPageId="19" captionText="Figure 5AF. Workers of Lepisiota annandalei (AC), Lepisiota bipartita (DF). A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152944" httpUri="https://zenodo.org/record/7152944/files/figure.png" pageId="6" pageNumber="6">Fig. 5A</figureCitation>
)............................
<taxonomicName id="4C174D4F1F3CFFA68B52FEB3900ECC1D" ID-CoL="6PD48" authority="(Mukerjee)" baseAuthorityName="Mukerjee" baseAuthorityYear="1930" box="[798,1123,289,317]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="annandalei">
<emphasis id="B963EADE1F3CFFA68B52FEB397BDCC1C" bold="true" box="[798,976,289,316]" italics="true" pageId="6" pageNumber="6">L. annandalei</emphasis>
(Mukerjee)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3CFFA688A9FED9900ECCA7" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA688A9FED9900ECCA7" pageId="6" pageNumber="6">. Petiole dorsally distinctly bispinose; setae present across entire mesosomal dorsum............................................................................................. 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3CFFA68893FE03900ECF62" pageId="6" pageNumber="6">
<paragraph id="8BA836CC1F3CFFA68893FE03900ECCD8" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FE03900ECCD8" pageId="6" pageNumber="6">
14. Head and pronotum smooth and shiny; propodeal spines as two broad-based blunt tubercles (
<bibRefCitation id="EF864B3D1F3CFFA68A26FE269706CCF1" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" box="[618,875,439,466]" journalOrPublisher="Bonn Zoological Bulletin." pageId="6" pageNumber="6" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
, Figs 2223)............ .........................................................................................
<taxonomicName id="4C174D4F1F3CFFA68B36FE4C900ECCD8" ID-CoL="3TFG4" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1894" box="[890,1123,476,504]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B963EADE1F3CFFA68B36FE4C9068CCD7" bold="true" box="[890,1029,476,503]" italics="true" pageId="6" pageNumber="6">L. modesta</emphasis>
(Forel)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3CFFA68893FD92900ECF62" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FD92900ECF62" pageId="6" pageNumber="6">. Head and pronotum either microreticulate or microreticulate with feeble striations; propodeal spines pointed or blunt........................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3CFFA68893FDDD900ECF91" pageId="6" pageNumber="6">
<paragraph id="8BA836CC1F3CFFA68893FDDD900ECFAC" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FDDD900ECFAC" pageId="6" pageNumber="6">
15. Head and mesosoma microreticulate with feeble striations (
<figureCitation id="132C2A491F3CFFA68BB7FDDC957ECFAC" captionStart="Figure 4" captionStartId="17.[192,257,1682,1705]" captionTargetBox="[194,1121,204,1648]" captionTargetId="figure-3@17.[193,1123,140,1650]" captionTargetPageId="17" captionText="Figure 4AC. Lepisiota wilsoni sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152942" httpUri="https://zenodo.org/record/7152942/files/figure.png" pageId="6" pageNumber="6">Fig. 4B C</figureCitation>
); propodeal spines blunt (
<figureCitation id="132C2A491F3CFFA68A11FDE396D5CFAC" box="[605,696,626,652]" captionStart="Figure 4" captionStartId="17.[192,257,1682,1705]" captionTargetBox="[194,1121,204,1648]" captionTargetId="figure-3@17.[193,1123,140,1650]" captionTargetPageId="17" captionText="Figure 4AC. Lepisiota wilsoni sp. nov. (Holotype worker). A, body in profile view; B, body in dorsal view; C, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7152942" httpUri="https://zenodo.org/record/7152942/files/figure.png" pageId="6" pageNumber="6">Fig. 4A</figureCitation>
)............ ............
<taxonomicName id="4C174D4F1F3CFFA68B33FDE39797CFAC" authority="Harshana &amp; Dey, 2022" authorityName="Harshana &amp; Dey" authorityYear="2022" box="[895,1018,625,652]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
<emphasis id="B963EADE1F3CFFA68B33FDE39797CFAC" bold="true" box="[895,1018,625,652]" italics="true" pageId="6" pageNumber="6">L. wilsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057A51F3CFFA68C4FFDE3900ECFAC" box="[1027,1123,626,652]" pageId="6" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3CFFA68893FD09900ECF91" blockId="6.[223,1123,141,914]" box="[223,1123,663,690]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FD09900ECF91" box="[223,1123,663,690]" pageId="6" pageNumber="6">. Head and mesosoma microreticulate; propodeal spines pointed...... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E691F3CFFA68893FD2D900ECEB1" pageId="6" pageNumber="6">
<paragraph id="8BA836CC1F3CFFA68893FD2D900ECE01" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FD2D900ECE01" pageId="6" pageNumber="6">
16. Antennal scape with sparse appressed to decumbent pubescence; mesosomal setae sparse; smaller species (HW 0.510.57)....................... .........................................................................................
<taxonomicName id="4C174D4F1F3CFFA68B35FC99900ECE01" authority="(Emery)" baseAuthorityName="Emery" baseAuthorityYear="1893" box="[889,1123,775,802]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="lunaris">
<emphasis id="B963EADE1F3CFFA68B35FC999798CE02" bold="true" box="[889,1013,775,802]" italics="true" pageId="6" pageNumber="6">L. lunaris</emphasis>
(Emery)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836CC1F3CFFA68893FCBC900ECEB1" blockId="6.[223,1123,141,914]" pageId="6" pageNumber="6">
<keyLead id="30E695F91F3CFFA68893FCBC900ECEB1" pageId="6" pageNumber="6">
. Antennal scape with dense subdecumbent to suberect pubescence; mesosomal setae abundant; larger species (HW 0.600.69) (
<bibRefCitation id="EF864B3D1F3CFFA68BA6FCC39510CEB1" author="Wachkoo AA &amp; Bharti H &amp; Akbar SA" journalOrPublisher="Bonn Zoological Bulletin." pageId="6" pageNumber="6" pagination="227 - 245" part="70" refId="ref13395" refString="Wachkoo AA, Bharti H, Akbar SA. 2021. Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India. Bonn Zoological Bulletin. 70 (2): 227 - 245." title="Taxonomic review of the ant genus Lepisiota Santschi, 1926 (Hymenoptera: Formicidae: Formicinae) from India" type="journal article" year="2021">Wachkoo et al. 2021</bibRefCitation>
, Figs 1921)..................
<taxonomicName id="4C174D4F1F3CFFA68AEAFCE9900ECEB1" authority="Wachkoo, Bharti &amp; Akbar" authorityName="Wachkoo, Bharti &amp; Akbar" authorityYear="2021" box="[678,1123,887,914]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B963EADE1F3CFFA68AEAFCE9977ECEB1" bold="true" box="[678,787,887,914]" italics="true" pageId="6" pageNumber="6">L. mayri</emphasis>
Wachkoo, Bharti &amp; Akbar
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
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