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<document id="4F9BCB6EC49A5BB491041D4F195F7153" ID-DOI="10.11646/zootaxa.3915.3.10" ID-GBIF-Dataset="c0300ee8-f2c6-46cd-8cb2-ff2d945c2270" ID-ISSN="1175-5326" ID-Zenodo-Dep="239095" ID-ZooBank="79599AEE-8F41-4ABE-B918-320148B62161" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460973348811" checkinUser="plazi" docAuthor="Jiao, Meng &amp; Yang, Maofa" docDate="2015" docId="03BE87B0706FFF9CFF1DFC288726C7DB" docLanguage="en" docName="zt03915p450.pdf" docOrigin="Zootaxa 3915 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Karachiota recurva Jiao &amp; Yang, sp. nov." docType="treatment" docVersion="8" lastPageNumber="447" masterDocId="FF87FFC8706EFF9DFF8AFF908402C11E" masterDocTitle="Description of two new species of the leafhopper genus Karachiota (Hemiptera: Cicadellidae: Typhlocybinae: Dikraneurini) from China" masterLastPageNumber="450" masterPageNumber="446" pageNumber="447" updateTime="1698559416952" updateUser="plazi">
<mods:mods id="B0A5FF7853B2DB904AA447497330BB03" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CB3EA6E5CAC94D7F291CFAF311DCA9C3">
<mods:title id="2F03F3DA6098C3B3EAD403630941E90E">Description of two new species of the leafhopper genus Karachiota (Hemiptera: Cicadellidae: Typhlocybinae: Dikraneurini) from China</mods:title>
</mods:titleInfo>
<mods:name id="4C153177DD41ADF08BB3E63EA017E5F4" type="personal">
<mods:role id="DC1D8FC500F0AF5CCBEB6B73CED5A919">
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</mods:role>
<mods:namePart id="9D3E5608E7C5159E0DC39C215A3F633B">Jiao, Meng</mods:namePart>
</mods:name>
<mods:name id="C0C55195B801C2F4D845B94C2AFECB03" type="personal">
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<mods:namePart id="ED090469F085D5FB47CD62D2FEA60863">Yang, Maofa</mods:namePart>
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<mods:typeOfResource id="2A3BC861CFD1FA7B79D875CEE9835F39">text</mods:typeOfResource>
<mods:relatedItem id="66130C008DFFA9CDEBCF240E4421FD9F" type="host">
<mods:titleInfo id="D136EF7D2DB79B7D8B559A123DA7BFEE">
<mods:title id="4B4689A026CA92DF1B1DEAD769BB8D2A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6F334E0F7FD99112C57063391DF35FB4">
<mods:date id="F5DF422F96845B8744749B42C387EE8D">2015</mods:date>
<mods:detail id="57BDCCF0A7C7FF3A298435FF5084B083" type="volume">
<mods:number id="415D582FE7980D10EC150E8E6D350AD2">3915</mods:number>
</mods:detail>
<mods:detail id="42046C921BEAA8103000E766038B5D0B" type="issue">
<mods:number id="16C66AF63C9D226422328F28179D445E">3</mods:number>
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<mods:start id="98663F541144FCBCAD13904C931E1D07">446</mods:start>
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<mods:classification id="F2FA1296DFE923553BE4D249F3D4D373">journal article</mods:classification>
<mods:identifier id="C41304EC802D63678B97312DF6216775" type="DOI">10.11646/zootaxa.3915.3.10</mods:identifier>
<mods:identifier id="1955F8EF78A617263742646B6CA350B5" type="GBIF-Dataset">c0300ee8-f2c6-46cd-8cb2-ff2d945c2270</mods:identifier>
<mods:identifier id="08028853B89A2B2BC4C2A7E52385C764" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EF79E844970BC39B0475358B3E11FF37" type="Zenodo-Dep">239095</mods:identifier>
<mods:identifier id="D9889D5F83872A09F8A18588DF0D5B4C" type="ZooBank">79599AEE-8F41-4ABE-B918-320148B62161</mods:identifier>
</mods:mods>
<treatment id="03BE87B0706FFF9CFF1DFC288726C7DB" ID-DOI="http://doi.org/10.5281/zenodo.6106232" ID-GBIF-Taxon="119594040" ID-Zenodo-Dep="6106232" LSID="urn:lsid:plazi:treatment:03BE87B0706FFF9CFF1DFC288726C7DB" httpUri="http://treatment.plazi.org/id/03BE87B0706FFF9CFF1DFC288726C7DB" lastPageNumber="447" pageId="1" pageNumber="447">
<subSubSection id="C30D652D706FFF9CFF1DFC28851BC2EB" pageId="1" pageNumber="447" type="nomenclature">
<paragraph id="8BA836A6706FFF9CFF1DFC28868CC2CD" blockId="1.[151,654,952,1013]" box="[151,654,952,979]" pageId="1" pageNumber="447">
<heading id="D0E081CA706FFF9CFF1DFC28868CC2CD" bold="true" box="[151,654,952,979]" fontSize="11" level="1" pageId="1" pageNumber="447" reason="1">
<emphasis id="B963EAB4706FFF9CFF1DFC28868CC2CD" bold="true" box="[151,654,952,979]" pageId="1" pageNumber="447">
<taxonomicName id="4C174D25706FFF9CFF1DFC288621C2CD" ID-CoL="8KLH4" authority="Jiao &amp; Yang" authorityName="Jiao &amp; Yang" box="[151,547,952,979]" class="Insecta" family="Cicadellidae" genus="Karachiota" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="B963EAB4706FFF9CFF1DFC288580C2CC" bold="true" box="[151,386,952,978]" italics="true" pageId="1" pageNumber="447">Karachiota recurva</emphasis>
Jiao
<emphasis id="B963EAB4706FFF9CFE4EFC2885D9C2CC" bold="true" box="[452,475,952,978]" italics="true" pageId="1" pageNumber="447">&amp;</emphasis>
Yang
</taxonomicName>
,
<taxonomicNameLabel id="A25057CF706FFF9CFDBAFC29868CC2CD" box="[560,654,953,979]" pageId="1" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA836A6706FFF9CFF1DFC4C851BC2EB" blockId="1.[151,654,952,1013]" box="[151,281,988,1013]" pageId="1" pageNumber="447">
(
<figureCitation id="132C2A23706FFF9CFF15FC4C8512C2EB" box="[159,272,988,1013]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Figs 1–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30D652D706FFF9CFF1DFBB480D5C70E" pageId="1" pageNumber="447" type="description">
<paragraph id="8BA836A6706FFF9CFF1DFBB48743C522" blockId="1.[151,1437,1060,1733]" box="[151,833,1060,1085]" pageId="1" pageNumber="447">
<emphasis id="B963EAB4706FFF9CFF1DFBB48523C523" bold="true" box="[151,289,1060,1085]" pageId="1" pageNumber="447">Description</emphasis>
. Length: male 3.2–3.3 mm, female 2.8–3.1 mm.
</paragraph>
<paragraph id="8BA836A6706FFF9CFF4DFBD78701C5EE" blockId="1.[151,1437,1060,1733]" pageId="1" pageNumber="447">
Body (
<figureCitation id="132C2A23706FFF9CFE98FBD78576C57E" box="[274,372,1095,1120]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Figs 1, 2</figureCitation>
) slightly yellowish. Head (
<figureCitation id="132C2A23706FFF9CFD28FBD786E6C57E" box="[674,740,1095,1120]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 3</figureCitation>
) triangular. Vertex median length nearly the same as the width between the apex of eyes, coronal suture distinct, to the 1/3 of vertex. Crown slightly yellowish, front margin white, two white patches mesad of eyes. Pronotum yellowish, with several irregular white patches near front margin. Scutellum yellowish, two white longitudinal stripes centrally, side angle white. Face (
<figureCitation id="132C2A23706FFF9CFB21FB2480EFC5D2" box="[1195,1261,1204,1229]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 4</figureCitation>
) compressed in lateral view. Anteclypeus (
<figureCitation id="132C2A23706FFF9CFE48FB478606C5EE" box="[450,516,1239,1264]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 5</figureCitation>
) slender, lorum broad.
</paragraph>
<paragraph id="8BA836A6706FFF9CFF4DFB6C8701C443" blockId="1.[151,1437,1060,1733]" pageId="1" pageNumber="447">
Forewing (
<figureCitation id="132C2A23706FFF9CFECBFB6C8587C40B" box="[321,389,1276,1301]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 6</figureCitation>
) slightly yellowish, with a small black spot on the basal part of the first apical cell, the fourth apical cell with a gray patch beside outer margin, the first apical cell nearly parallel with the second apical cell. Hindwing (
<figureCitation id="132C2A23706FFF9CFE9CFAD4855BC442" box="[278,345,1348,1373]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 7</figureCitation>
) semitransparent, anal vein branched.
</paragraph>
<paragraph id="8BA836A6706FFF9CFF4DFAF780D5C70E" blockId="1.[151,1437,1060,1733]" pageId="1" pageNumber="447">
Male abdomen (
<figureCitation id="132C2A23706FFF9CFE08FAF785C5C49E" box="[386,455,1383,1408]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 8</figureCitation>
) with apodemes of sternum II well developed. Pygofer (
<figureCitation id="132C2A23706FFF9CFBDFFAF780FAC49E" box="[1109,1272,1383,1408]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Figs 9, 10, 15</figureCitation>
) very slender, sclerotised, with short dorsal appendage, distal part of pygofer with transverse striations. Subgenital (
<figureCitation id="132C2A23706FFF9CFAB0FA1C8196C4BB" box="[1338,1428,1420,1445]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Figs 15</figureCitation>
) triangular, greatly reduced and without setae. Paramere (
<figureCitation id="132C2A23706FFF9CFC92FA3F876AC4D6" box="[792,872,1455,1480]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Fig. 11</figureCitation>
) strongly curved, two arms as long as each other. Aedeagus (
<figureCitation id="132C2A23706FFF9CFE93FA4485D1C4F2" box="[281,467,1492,1517]" captionStart="FIGURES 1 – 16" captionStartId="2.[151,264,1885,1908]" captionTargetBox="[266,1294,152,1856]" captionTargetId="figure@2.[266,1295,151,1861]" captionTargetPageId="2" captionText="FIGURES 1 – 16. Karachiota recurva Jiao &amp; Yang, sp. nov., 1 whole body, dorsal view. 2 whole body, lateral view. 3 head and thorax, dorsal view. 4 head and thorax, lateral view. 5 face. 6 forewing. 7 hindwing. 8 male abdominal apodemes. 9 male terminalia, lateral view. 10 male pygofer, dorsal view. 11 Paramere, lateral view. 12 aedeagus and connective, lateral view. 13 aedeagus and connective, dorsal view. 14 aedeagus and connective, ventral view. 15 genital capsule, ventral view. 16 genital capsule, lateral view." httpUri="https://zenodo.org/record/239096/files/figure.png" pageId="1" pageNumber="447">Figs 12, 13, 14</figureCitation>
) with shaft moderately long, broader at base and narrowed to apex, with pair of processes arising subapically and extended distad but not surpassing shaft apex, gonopore apical.
</paragraph>
</subSubSection>
<subSubSection id="C30D652D706FFF9CFF4DF98C8557C746" pageId="1" pageNumber="447" type="materials_examined">
<paragraph id="8BA836A6706FFF9CFF4DF98C8557C746" blockId="1.[151,1437,1060,1733]" pageId="1" pageNumber="447">
<emphasis id="B963EAB4706FFF9CFF4DF98C8576C72B" bold="true" box="[199,372,1564,1589]" pageId="1" pageNumber="447">
<typeStatus id="54AC8804706FFF9CFF4DF98C8500C72B" box="[199,258,1564,1589]" pageId="1" pageNumber="447">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AC8804706FFF9CFEF0F98C85E7C72B" box="[378,485,1564,1589]" pageId="1" pageNumber="447" type="holotype">Holotype</typeStatus>
, ♂, Luodian, Guizhou, April. 1980, coll. by Zizhong Li.
<typeStatus id="54AC8804706FFF9CFBD7F98D80CFC72B" box="[1117,1229,1565,1589]" pageId="1" pageNumber="447" type="paratype">Paratypes</typeStatus>
, 2♂♂, 3♀♀, same data as
<typeStatus id="54AC8804706FFF9CFF61F9AF8553C746" box="[235,337,1599,1624]" pageId="1" pageNumber="447" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D652D706FFF9CFF4DF9F48095C763" box="[199,1175,1636,1661]" pageId="1" pageNumber="447" type="etymology">
<paragraph id="8BA836A6706FFF9CFF4DF9F48095C763" blockId="1.[151,1437,1060,1733]" box="[199,1175,1636,1661]" pageId="1" pageNumber="447">
<emphasis id="B963EAB4706FFF9CFF4DF9F4854CC763" bold="true" box="[199,334,1636,1661]" pageId="1" pageNumber="447">Etymology.</emphasis>
The species name refers to the pair of upturned processes on the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C30D652D706FFF9CFF4DF9178726C7DB" pageId="1" pageNumber="447" type="discussion">
<paragraph id="8BA836A6706FFF9CFF4DF9178726C7DB" blockId="1.[151,1437,1060,1733]" pageId="1" pageNumber="447">
<emphasis id="B963EAB4706FFF9CFF4DF9178539C7BE" bold="true" box="[199,315,1671,1696]" pageId="1" pageNumber="447">Remarks.</emphasis>
The new species differs from all other species of this genus by the absence of teeth on the processes of the aedeagus, and the presence of a pygofer appendage.
</paragraph>
</subSubSection>
</treatment>
</document>
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