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<document id="E497477278E8EE0D1DA8736F471FB14D" ID-CLB-Dataset="55730" ID-DOI="10.5852/ejt.2022.817.1767" ID-GBIF-Dataset="529dc516-47b9-424e-93ac-3787e407e100" ID-ISSN="2118-9773" ID-Zenodo-Dep="6517928" ID-ZooBank="FFB05CE7-3712-4FD0-91B4-FD84863CF9D7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651673830896" checkinUser="felipe" docAuthor="Yasuda, Kohei &amp; Yoshitomi, Hiroyuki" docDate="2022" docId="03BE8793BE48FFC1FE32B991822E069A" docLanguage="en" docName="ejt-817_111-142.pdf" docOrigin="European Journal of Taxonomy 817 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.817.1767" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Georissus (Nipponogeorissus) granulosus Sato 1972" docType="treatment" docVersion="9" lastPageNumber="123" masterDocId="FF87FFEBBE42FFCDFFE8BF5B80470455" masterDocTitle="Revision of the genus Georissus (Coleoptera, Hydrophiloidea, Georissidae) of Japan" masterLastPageNumber="142" masterPageNumber="111" pageNumber="121" updateTime="1699329566334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7980B1DE39AFEA10384B4D5D199DE966">Revision of the genus Georissus (Coleoptera, Hydrophiloidea, Georissidae) of Japan</mods:title>
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<mods:namePart id="DBAF01BFE39C97AB8645CD3E0566DB09">Yasuda, Kohei</mods:namePart>
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<mods:affiliation id="3D092B989133AC8979C7816D72B11EEC">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:namePart id="F192142468C684290A69546710A25EB6">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="8FC1C13E8553F2C4CBCFC9A0B5057EB5">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:title id="C5F1AA065E10AB04825F33028193FB50">European Journal of Taxonomy</mods:title>
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<treatment id="03BE8793BE48FFC1FE32B991822E069A" ID-DOI="http://doi.org/10.5281/zenodo.6518781" ID-GBIF-Taxon="195191638" ID-Zenodo-Dep="6518781" LSID="urn:lsid:plazi:treatment:03BE8793BE48FFC1FE32B991822E069A" httpUri="http://treatment.plazi.org/id/03BE8793BE48FFC1FE32B991822E069A" lastPageId="12" lastPageNumber="123" pageId="10" pageNumber="121">
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<heading id="D0E081E9BE48FFC7FE32B991841E02B0" box="[474,1113,1738,1765]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="121" reason="2">
<taxonomicName id="4C174D06BE48FFC7FE32B991841E02B0" ID-CoL="3FS3F" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[474,1113,1738,1765]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="121" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Nipponogeorissus">
<emphasis id="B963EA97BE48FFC7FE32B991839302B0" bold="true" box="[474,980,1738,1765]" pageId="10" pageNumber="121">
<emphasis id="B963EA97BE48FFC7FE32B991821602B1" bold="true" box="[474,593,1738,1764]" italics="true" pageId="10" pageNumber="121">Georissus</emphasis>
(
<emphasis id="B963EA97BE48FFC7FD8BB991837B02B1" bold="true" box="[611,828,1738,1764]" italics="true" pageId="10" pageNumber="121">Nipponogeorissus</emphasis>
)
<emphasis id="B963EA97BE48FFC7FCA5B991839302B0" bold="true" box="[845,980,1738,1765]" italics="true" pageId="10" pageNumber="121">granulosus</emphasis>
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<bibRefCitation id="EF864B74BE48FFC7FC33B990841E02B0" author="Sato M." box="[987,1113,1739,1765]" pageId="10" pageNumber="121" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
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<subSubSection id="C30D650EBE48FFC7FDC3B9B5844F035D" box="[555,1032,1773,1800]" pageId="10" pageNumber="121" type="vernacular_names">
<paragraph id="8BA83685BE48FFC7FDC3B9B5844F035D" blockId="10.[443,1144,1738,1836]" box="[555,1032,1773,1800]" pageId="10" pageNumber="121">[Japanese name: Sesuji-marudoromushi]</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE48FFC7FE53B84A843F037E" box="[443,1144,1809,1836]" pageId="10" pageNumber="121" type="description">
<paragraph id="8BA83685BE48FFC7FE53B84A843F037E" blockId="10.[443,1144,1738,1836]" box="[443,1144,1809,1836]" pageId="10" pageNumber="121">
<figureCitation id="132C2A00BE48FFC7FE53B84A825D037E" box="[443,538,1809,1835]" captionStart="Fig" captionStartId="7.[189,232,1886,1912]" captionTargetBox="[272,1331,265,1833]" captionTargetId="figure-11@7.[223,1365,265,1845]" captionTargetPageId="7" captionText="Fig. 2. Dorsal view of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F. G. (Ne.) takahashii sp. nov., holotype, ♂ (EUMJ). G. G. (Ne.) satoi sp. nov., holotype (EUMJ). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517952" httpUri="https://zenodo.org/record/6517952/files/figure.png" pageId="10" pageNumber="121">Figs 2C</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FDCEB84A82280379" box="[550,623,1809,1836]" captionStart="Fig" captionStartId="9.[189,232,1886,1912]" captionTargetBox="[196,1392,306,1839]" captionTargetId="figure-11@9.[189,1398,268,1842]" captionTargetPageId="9" captionText="Fig. 3. Holotypes of Georissus spp. A–F. Dorsal habitus. G–L. Labels. A, K. G. (G.) babai Satô, 1970, ♂ (EUMJ). B, L. G. (Ni.) granulosus Satô, 1972, ♀ (EUMJ). C, H. G. (Ne.) kurosawai Nakane, 1963, ♂ (SEHU). D, J. G. (Ne.) japonicus Satô, 1972, ♂ (EUMJ). E, G. G. (Ne.) sakaii Satô, 1972, ♂ (EUMJ). F, I. G. (Ne.) katsuoi Nakane, 1995 (SEHU). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517956" httpUri="https://zenodo.org/record/6517956/files/figure.png" pageId="10" pageNumber="121">3B, H</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FD93B84A83660379" box="[635,801,1809,1836]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="10" pageNumber="121">4B, E, H–I, N</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FCC5B84A83160379" box="[813,849,1809,1836]" captionStart="Fig" captionStartId="16.[189,232,1822,1848]" captionTargetBox="[192,1395,266,1778]" captionTargetId="figure-17@16.[189,1398,266,1783]" captionTargetPageId="16" captionText="Fig. 5. Dorsal habitus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517964" httpUri="https://zenodo.org/record/6517964/files/figure.png" pageId="10" pageNumber="121">5C</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FCB5B84A83C5037E" box="[861,898,1809,1835]" captionStart="Fig" captionStartId="18.[189,232,1882,1908]" captionTargetBox="[191,1394,563,1839]" captionTargetId="figure-90@18.[189,1398,562,1842]" captionTargetPageId="18" captionText="Fig. 6. Head of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517968" httpUri="https://zenodo.org/record/6517968/files/figure.png" pageId="10" pageNumber="121">6C</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FC66B84A83F4037E" box="[910,947,1809,1835]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="10" pageNumber="121">7C</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FC57B84A83A4037E" box="[959,995,1809,1835]" captionStart="Fig" captionStartId="21.[191,234,1890,1916]" captionTargetBox="[193,1394,569,1846]" captionTargetId="figure-107@21.[189,1398,569,1852]" captionTargetPageId="21" captionText="Fig. 8. Ventrites of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (F). I. G. (Ne.) satoi sp. nov. Abbreviation: vt = ventral teeth. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6517982" httpUri="https://zenodo.org/record/6517982/files/figure.png" pageId="10" pageNumber="121">8C</figureCitation>
,
<figureCitation id="132C2A00BE48FFC7FC18B84A8451037E" box="[1008,1046,1809,1835]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="10" pageNumber="121">9C</figureCitation>
;
<tableCitation id="C695033EBE48FFC7FBC9B84A843F037E" box="[1057,1144,1809,1835]" captionStart="Table 1" captionStartId="11.[189,264,265,293]" captionTargetBox="[352,1201,392,914]" captionTargetPageId="11" captionText="Table 1. Measurements of Georissus (G.) spp. and G. (Nipponogeorissus) granulosus Satô, 1972." httpUri="http://table.plazi.org/id/DF68660DBE49FFC6FF55BE5281470519" pageId="10" pageNumber="121" tableUuid="DF68660DBE49FFC6FF55BE5281470519">Table 1</tableCitation>
</paragraph>
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<treatmentCitationGroup id="AB0711ABBE48FFC7FF55B80883340338" box="[189,883,1874,1901]" pageId="10" pageNumber="121">
<treatmentCitation id="0AB61094BE48FFC7FF55B80883280338" author="Sato M." box="[189,879,1874,1901]" page="212" pageId="10" pageNumber="121" year="1972">
<taxonomicName id="4C174D06BE48FFC7FF55B80883280338" ID-CoL="3FS3F" authority="Sato, 1972: 212" authorityName="Sato" authorityPageNumber="212" authorityYear="1972" box="[189,879,1874,1901]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="121" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Nipponogeorissus">
<emphasis id="B963EA97BE48FFC7FF55B80881750338" box="[189,306,1875,1901]" italics="true" pageId="10" pageNumber="121">Georissus</emphasis>
(
<emphasis id="B963EA97BE48FFC7FEABB80882500339" box="[323,535,1875,1900]" italics="true" pageId="10" pageNumber="121">Nipponogeorissus</emphasis>
)
<emphasis id="B963EA97BE48FFC7FDC0B80982ED0339" box="[552,682,1874,1900]" italics="true" pageId="10" pageNumber="121">granulosus</emphasis>
<bibRefCitation id="EF864B74BE48FFC7FD5AB80883280338" author="Sato M." box="[690,879,1875,1901]" pageId="10" pageNumber="121" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972: 212</bibRefCitation>
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.
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<taxonomicName id="4C174D06BE48FFC7FF55B8CF82ED03FB" ID-CoL="3FS3F" authorityName="Sato" authorityYear="1972" box="[189,682,1940,1966]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="121" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Nipponogeorissus">
<emphasis id="B963EA97BE48FFC7FF55B8CF817503FB" box="[189,306,1940,1966]" italics="true" pageId="10" pageNumber="121">Georissus</emphasis>
(
<emphasis id="B963EA97BE48FFC7FEABB8CE825003FB" box="[323,535,1941,1966]" italics="true" pageId="10" pageNumber="121">Nipponogeorissus</emphasis>
)
<emphasis id="B963EA97BE48FFC7FDC0B8CF82ED03FB" box="[552,682,1940,1966]" italics="true" pageId="10" pageNumber="121">granulosus</emphasis>
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–
<treatmentCitation id="0AB61094BE48FFC7FD20B8CF832703FB" author="Sato M." box="[712,864,1940,1966]" page="4" pageId="10" pageNumber="121" year="1981">
<bibRefCitation id="EF864B74BE48FFC7FD20B8CF832703FB" author="Sato M." box="[712,864,1940,1966]" pageId="10" pageNumber="121" pagination="1 - 4" refId="ref14105" refString="Sato M. 1981. An outline of Japanese Georissidae. Coleopterist's News 53: 1 - 4. [In Japanese]" type="journal article" year="1981">Satô 1981: 4</bibRefCitation>
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;
<treatmentCitation id="0AB61094BE48FFC7FC84B8CF83A103FA" author="Sato M." box="[876,998,1940,1967]" page="205" pageId="10" pageNumber="121" year="1985">
<bibRefCitation id="EF864B74BE48FFC7FC84B8CF83A103FA" author="Sato M." box="[876,998,1940,1967]" pageId="10" pageNumber="121" refId="ref14128" refString="Sato M. 1985. Georissidae. In: Ueno S., Kurosawa Y. &amp; Sato M. (eds) The Coleoptera of Japan in Color Vol. II. Hoikusha, Osaka. [In Japanese]" type="book" year="1985">1985: 205</bibRefCitation>
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. —
<treatmentCitation id="0AB61094BE48FFC7FBFDB8CF84A703FB" author="Hansen M." box="[1045,1248,1940,1966]" page="49" pageId="10" pageNumber="121" year="1999">
<bibRefCitation id="EF864B74BE48FFC7FBFDB8CF84A703FB" author="Hansen M." box="[1045,1248,1940,1966]" pageId="10" pageNumber="121" refId="ref13193" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Amsterdam." type="book" year="1999">Hansen 1999: 49</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
<heading id="D0E081E9BE49FFC6FF55BC9981710789" bold="true" box="[189,310,962,988]" fontSize="11" level="3" pageId="11" pageNumber="122" reason="3">
<emphasis id="B963EA97BE49FFC6FF55BC9981710789" bold="true" box="[189,310,962,988]" pageId="11" pageNumber="122">Diagnosis</emphasis>
</heading>
</paragraph>
</subSubSection>
<caption id="DF68660DBE49FFC6FF55BE5281470519" ID-Table-UUID="DF68660DBE49FFC6FF55BE5281470519" httpUri="http://table.plazi.org/id/DF68660DBE49FFC6FF55BE5281470519" pageId="11" pageNumber="122" startId="11.[189,264,265,293]" targetBox="[352,1201,392,914]" targetIsTable="true" targetPageId="11">
<paragraph id="8BA83685BE49FFC6FF55BE5281470519" blockId="11.[189,1399,265,332]" pageId="11" pageNumber="122">
<emphasis id="B963EA97BE49FFC6FF55BE52816B0573" bold="true" box="[189,300,265,294]" pageId="11" pageNumber="122">Table 1.</emphasis>
Measurements of
<taxonomicName id="4C174D06BE49FFC6FDCEBE5282E10570" authorityName="Latreille" authorityYear="1809" box="[550,678,265,293]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B963EA97BE49FFC6FDCEBE5282E10570" box="[550,678,265,293]" italics="true" pageId="11" pageNumber="122">Georissus</emphasis>
</taxonomicName>
(
<emphasis id="B963EA97BE49FFC6FD54BE5282940570" box="[700,723,265,293]" italics="true" pageId="11" pageNumber="122">G</emphasis>
.) spp. and
<taxonomicName id="4C174D06BE49FFC6FC85BE5280BF051E" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Nipponogeorissus">
<emphasis id="B963EA97BE49FFC6FC85BE5283C30570" box="[877,900,265,293]" italics="true" pageId="11" pageNumber="122">G</emphasis>
. (
<emphasis id="B963EA97BE49FFC6FC4BBE5285680573" box="[931,1327,265,294]" italics="true" pageId="11" pageNumber="122">Nipponogeorissus) granulosus</emphasis>
<bibRefCitation id="EF864B74BE49FFC6FAD3BE5080BF051E" author="Sato M." pageId="11" pageNumber="122" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA83685BE49FFC6FE60BEFB84D907C7" pageId="11" pageNumber="122">
<table id="F917C425BE490032FE88BED384F607C7" box="[352,1201,392,914]" gridcols="4" gridrows="11" pageId="11" pageNumber="122">
<tr id="352734C7BE490032FE88BED384F60585" box="[352,1201,392,464]" gridrow="0" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE0ABED382EA0585" box="[482,685,392,464]" gridcol="1" gridrow="0" pageId="11" pageNumber="122">
<emphasis id="B963EA97BE49FFC6FE0ABED381B805F7" box="[482,511,392,418]" italics="true" pageId="11" pageNumber="122">G.</emphasis>
<taxonomicName id="4C174D06BE49FFC6FDE3BED382EA05F7" authorityName="Sharp" authorityYear="1888" box="[523,685,392,418]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="canalifer" subGenus="G .">
(
<emphasis id="B963EA97BE49FFC6FDFCBED3826E05F7" box="[532,553,392,418]" italics="true" pageId="11" pageNumber="122">G</emphasis>
.)
<emphasis id="B963EA97BE49FFC6FDAFBED382EA05F7" box="[583,685,392,418]" italics="true" pageId="11" pageNumber="122">canalifer</emphasis>
</taxonomicName>
(N = 53)
</th>
<th id="76F65DBBBE490032FD3FBED383DE0585" box="[727,921,392,464]" gridcol="2" gridrow="0" pageId="11" pageNumber="122">
<emphasis id="B963EA97BE49FFC6FD08BED382BA05F7" box="[736,765,392,418]" italics="true" pageId="11" pageNumber="122">G.</emphasis>
<taxonomicName id="4C174D06BE49FFC6FCE1BED383C105F7" authorityName="Sato" authorityYear="1970" box="[777,902,392,418]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="babai" subGenus="G .">
(
<emphasis id="B963EA97BE49FFC6FCFBBED3836F05F7" box="[787,808,392,418]" italics="true" pageId="11" pageNumber="122">G</emphasis>
.)
<emphasis id="B963EA97BE49FFC6FCADBED383C105F7" box="[837,902,392,418]" italics="true" pageId="11" pageNumber="122">babai</emphasis>
</taxonomicName>
(N = 20)
</th>
<th id="76F65DBBBE490032FC21BED384F60585" box="[969,1201,392,464]" gridcol="3" gridrow="0" pageId="11" pageNumber="122">
<emphasis id="B963EA97BE49FFC6FC21BED383A205F7" box="[969,997,392,418]" italics="true" pageId="11" pageNumber="122">G.</emphasis>
<taxonomicName id="4C174D06BE49FFC6FC1ABED384F605F7" authorityName="Sato" authorityYear="1972" box="[1010,1201,392,418]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Ni .">
(
<emphasis id="B963EA97BE49FFC6FC13BED2845005F7" box="[1019,1047,393,418]" italics="true" pageId="11" pageNumber="122">Ni</emphasis>
.)
<emphasis id="B963EA97BE49FFC6FBDBBED384F605F7" box="[1075,1201,392,418]" italics="true" pageId="11" pageNumber="122">granulosus</emphasis>
</taxonomicName>
(N = 5)
</th>
</tr>
<tr id="352734C7BE490032FE88BEB984F605A8" box="[352,1201,482,509]" gridrow="1" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BEB9818805A8" box="[352,463,482,509]" gridcol="0" gridrow="1" pageId="11" pageNumber="122">TL (mm)</th>
<td id="76F65DBBBE490032FE0ABEB982EA05A8" box="[482,685,482,509]" gridcol="1" gridrow="1" pageId="11" pageNumber="122">1.46–2.30 (1.91)</td>
<td id="76F65DBBBE490032FD3FBEB983DE05A8" box="[727,921,482,509]" gridcol="2" gridrow="1" pageId="11" pageNumber="122">1.34–1.90 (1.91)</td>
<td id="76F65DBBBE490032FC21BEB984F605A8" box="[969,1201,482,509]" gridcol="3" gridrow="1" pageId="11" pageNumber="122">1.28–1.54 (1.38)</td>
</tr>
<tr id="352734C7BE490032FE88BD5484F6067F" box="[352,1201,527,554]" gridrow="2" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BD548188067F" box="[352,463,527,554]" gridcol="0" gridrow="2" pageId="11" pageNumber="122">EL (mm)</th>
<td id="76F65DBBBE490032FE0ABD5482EA067F" box="[482,685,527,554]" gridcol="1" gridrow="2" pageId="11" pageNumber="122">0.86–1.62 (1.27)</td>
<td id="76F65DBBBE490032FD3FBD5483DE067F" box="[727,921,527,554]" gridcol="2" gridrow="2" pageId="11" pageNumber="122">0.82–1.26 (1.09)</td>
<td id="76F65DBBBE490032FC21BD5484F6067F" box="[969,1201,527,554]" gridcol="3" gridrow="2" pageId="11" pageNumber="122">0.86–1.02 (0.92)</td>
</tr>
<tr id="352734C7BE490032FE88BD6784F60602" box="[352,1201,572,599]" gridrow="3" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BD6781880602" box="[352,463,572,599]" gridcol="0" gridrow="3" pageId="11" pageNumber="122">EW (mm)</th>
<td id="76F65DBBBE490032FE0ABD6782EA0602" box="[482,685,572,599]" gridcol="1" gridrow="3" pageId="11" pageNumber="122">0.90–1.38 (1.17)</td>
<td id="76F65DBBBE490032FD3FBD6783DE0602" box="[727,921,572,599]" gridcol="2" gridrow="3" pageId="11" pageNumber="122">0.82–1.16 (1.05)</td>
<td id="76F65DBBBE490032FC21BD6784F60602" box="[969,1201,572,599]" gridcol="3" gridrow="3" pageId="11" pageNumber="122">0.70–0.88 (0.79)</td>
</tr>
<tr id="352734C7BE490032FE88BD3284F606D1" box="[352,1201,617,644]" gridrow="4" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BD32818806D1" box="[352,463,617,644]" gridcol="0" gridrow="4" pageId="11" pageNumber="122">PL (mm)</th>
<td id="76F65DBBBE490032FE0ABD3282EA06D1" box="[482,685,617,644]" gridcol="1" gridrow="4" pageId="11" pageNumber="122">0.52–0.78 (0.64)</td>
<td id="76F65DBBBE490032FD3FBD3283DE06D1" box="[727,921,617,644]" gridcol="2" gridrow="4" pageId="11" pageNumber="122">0.52–0.66 (0.60)</td>
<td id="76F65DBBBE490032FC21BD3284F606D1" box="[969,1201,617,644]" gridcol="3" gridrow="4" pageId="11" pageNumber="122">0.40–0.52 (0.46)</td>
</tr>
<tr id="352734C7BE490032FE88BDCD84F606E4" box="[352,1201,662,689]" gridrow="5" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BDCD818806E4" box="[352,463,662,689]" gridcol="0" gridrow="5" pageId="11" pageNumber="122">PW (mm)</th>
<td id="76F65DBBBE490032FE0ABDCD82EA06E4" box="[482,685,662,689]" gridcol="1" gridrow="5" pageId="11" pageNumber="122">0.70–1.04 (0.85)</td>
<td id="76F65DBBBE490032FD3FBDCD83DE06E4" box="[727,921,662,689]" gridcol="2" gridrow="5" pageId="11" pageNumber="122">0.68–0.88 (0.79)</td>
<td id="76F65DBBBE490032FC21BDCD84F606E4" box="[969,1201,662,689]" gridcol="3" gridrow="5" pageId="11" pageNumber="122">0.52–0.62 (0.58)</td>
</tr>
<tr id="352734C7BE490032FE88BD9884F6068B" box="[352,1201,707,734]" gridrow="6" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BD988188068B" box="[352,463,707,734]" gridcol="0" gridrow="6" pageId="11" pageNumber="122">PW/PL</th>
<td id="76F65DBBBE490032FE0ABD9882EA068B" box="[482,685,707,734]" gridcol="1" gridrow="6" pageId="11" pageNumber="122">1.08–1.66 (1.33)</td>
<td id="76F65DBBBE490032FD3FBD9883DE068B" box="[727,921,707,734]" gridcol="2" gridrow="6" pageId="11" pageNumber="122">1.15–1.41 (1.32)</td>
<td id="76F65DBBBE490032FC21BD9884F6068B" box="[969,1201,707,734]" gridcol="3" gridrow="6" pageId="11" pageNumber="122">1.15–1.35 (1.26)</td>
</tr>
<tr id="352734C7BE490032FE88BDAB84F6075E" box="[352,1201,752,779]" gridrow="7" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BDAB8188075E" box="[352,463,752,779]" gridcol="0" gridrow="7" pageId="11" pageNumber="122">EL/EW</th>
<td id="76F65DBBBE490032FE0ABDAB82EA075E" box="[482,685,752,779]" gridcol="1" gridrow="7" pageId="11" pageNumber="122">0.86–1.23 (1.08)</td>
<td id="76F65DBBBE490032FD3FBDAB83DE075E" box="[727,921,752,779]" gridcol="2" gridrow="7" pageId="11" pageNumber="122">0.83–1.24 (1.03)</td>
<td id="76F65DBBBE490032FC21BDAB84F6075E" box="[969,1201,752,779]" gridcol="3" gridrow="7" pageId="11" pageNumber="122">1.05–1.26 (1.16)</td>
</tr>
<tr id="352734C7BE490032FE88BC4684F6076D" box="[352,1201,797,824]" gridrow="8" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BC468188076D" box="[352,463,797,824]" gridcol="0" gridrow="8" pageId="11" pageNumber="122">EL/PL</th>
<td id="76F65DBBBE490032FE0ABC4682EA076D" box="[482,685,797,824]" gridcol="1" gridrow="8" pageId="11" pageNumber="122">1.43–2.48 (1.98)</td>
<td id="76F65DBBBE490032FD3FBC4683DE076D" box="[727,921,797,824]" gridcol="2" gridrow="8" pageId="11" pageNumber="122">1.58–2.07 (1.82)</td>
<td id="76F65DBBBE490032FC21BC4684F6076D" box="[969,1201,797,824]" gridcol="3" gridrow="8" pageId="11" pageNumber="122">1.87–2.20 (2.00)</td>
</tr>
<tr id="352734C7BE490032FE88BC1184F60730" box="[352,1201,842,869]" gridrow="9" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BC1181880730" box="[352,463,842,869]" gridcol="0" gridrow="9" pageId="11" pageNumber="122">EW/PW</th>
<td id="76F65DBBBE490032FE0ABC1182EA0730" box="[482,685,842,869]" gridcol="1" gridrow="9" pageId="11" pageNumber="122">1.10–1.56 (1.38)</td>
<td id="76F65DBBBE490032FD3FBC1183DE0730" box="[727,921,842,869]" gridcol="2" gridrow="9" pageId="11" pageNumber="122">1.21–1.53 (1.33)</td>
<td id="76F65DBBBE490032FC21BC1184F60730" box="[969,1201,842,869]" gridcol="3" gridrow="9" pageId="11" pageNumber="122">1.28–1.47 (1.37)</td>
</tr>
<tr id="352734C7BE490032FE88BC2C84F607C7" box="[352,1201,887,914]" gridrow="10" pageId="11" pageNumber="122">
<th id="76F65DBBBE490032FE88BC2C818807C7" box="[352,463,887,914]" gridcol="0" gridrow="10" pageId="11" pageNumber="122">TL/EW</th>
<td id="76F65DBBBE490032FE0ABC2C82EA07C7" box="[482,685,887,914]" gridcol="1" gridrow="10" pageId="11" pageNumber="122">1.45–1.84 (1.64)</td>
<td id="76F65DBBBE490032FD3FBC2C83DE07C7" box="[727,921,887,914]" gridcol="2" gridrow="10" pageId="11" pageNumber="122">1.34–1.86 (1.60)</td>
<td id="76F65DBBBE490032FC21BC2C84F607C7" box="[969,1201,887,914]" gridcol="3" gridrow="10" pageId="11" pageNumber="122">1.57–1.83 (1.74)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C30D650EBE49FFC6FF55BCAA845E0020" pageId="11" pageNumber="122" type="discussion">
<paragraph id="8BA83685BE49FFC6FF55BCAA845E0020" blockId="11.[189,1399,1009,1141]" pageId="11" pageNumber="122">
This species is easily distinguished from other Japanese species of
<taxonomicName id="4C174D06BE49FFC6FBF0BCAA84CA005E" authorityName="Latreille" authorityYear="1809" box="[1048,1165,1009,1035]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B963EA97BE49FFC6FBF0BCAA84CA005E" box="[1048,1165,1009,1035]" italics="true" pageId="11" pageNumber="122">Georissus</emphasis>
</taxonomicName>
by the granulated pronotum and shape of male aedeagus; shape elongated and narrow, phallobase widening. This species is distinguished from
<taxonomicName id="4C174D06BE49FFC6FE2CBB63823B0004" authorityName="Deleve" authorityYear="1969" box="[452,636,1079,1106]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="B963EA97BE49FFC6FE2CBB63823B0004" box="[452,636,1079,1106]" italics="true" pageId="11" pageNumber="122">Georissus biroi</emphasis>
</taxonomicName>
described from
<collectingCountry id="F3007615BE49FFC6FCABBB63846C0004" box="[835,1067,1079,1106]" name="Papua New Guinea" pageId="11" pageNumber="122">Papua New Guinea</collectingCountry>
by the carinae of frons;
<taxonomicName id="4C174D06BE49FFC6FAB2BB6381780021" authorityName="Sato" authorityYear="1972" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B963EA97BE49FFC6FAB2BB6385280007" box="[1370,1391,1080,1106]" italics="true" pageId="11" pageNumber="122">G</emphasis>
.
<emphasis id="B963EA97BE49FFC6FF55BB0181780021" box="[189,319,1114,1140]" italics="true" pageId="11" pageNumber="122">granulosus</emphasis>
</taxonomicName>
has transverse carinae, in contrast to their absence in
<taxonomicName id="4C174D06BE49FFC6FC5FBB0084550021" authorityName="Deleve" authorityYear="1969" box="[951,1042,1114,1141]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="B963EA97BE49FFC6FC5FBB00838B0020" box="[951,972,1115,1141]" italics="true" pageId="11" pageNumber="122">G</emphasis>
.
<emphasis id="B963EA97BE49FFC6FC32BB0184550021" box="[986,1042,1114,1140]" italics="true" pageId="11" pageNumber="122">biroi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE49FFC6FF55BBC085320138" pageId="11" pageNumber="122" type="materials_examined">
<paragraph id="8BA83685BE49FFC6FF55BBC081ED00E0" blockId="11.[189,426,1179,1205]" box="[189,426,1179,1205]" pageId="11" pageNumber="122">
<heading id="D0E081E9BE49FFC6FF55BBC081ED00E0" bold="true" box="[189,426,1179,1205]" fontSize="11" level="3" pageId="11" pageNumber="122" reason="3">
<emphasis id="B963EA97BE49FFC6FF55BBC081ED00E0" bold="true" box="[189,426,1179,1205]" pageId="11" pageNumber="122">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE49FFC6FF02BB918531015D" blockId="11.[189,1398,1226,1288]" pageId="11" pageNumber="122">
<materialsCitation id="3B7F3CD8BE49FFC6FF02BB918531015D" ID-GBIF-Occurrence="3762863301" collectingDate="1958-08-23" collectionCode="EUMJ" collectorName="H. Ichihashi" country="Japan" location="Honshu" pageId="11" pageNumber="122" specimenCount="1" specimenCount-female="1" stateProvince="Mie Pref." typeStatus="holotype">
<emphasis id="B963EA97BE49FFC6FF02BB91811B00B1" bold="true" box="[234,348,1226,1252]" pageId="11" pageNumber="122">
<typeStatus id="54AC8827BE49FFC6FF02BB91811B00B1" box="[234,348,1226,1252]" pageId="11" pageNumber="122" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3007615BE49FFC6FF55BBB58153015D" box="[189,276,1262,1288]" name="Japan" pageId="11" pageNumber="122">JAPAN</collectingCountry>
–
<emphasis id="B963EA97BE49FFC6FEC5BBB5824A0152" bold="true" box="[301,525,1261,1288]" pageId="11" pageNumber="122">
<location id="8EC8605EBE49FFC6FEC5BBB581D5015D" LSID="urn:lsid:plazi:treatment:03BE8793BE48FFC1FE32B991822E069A:8EC8605EBE49FFC6FEC5BBB581D5015D" box="[301,402,1262,1288]" country="Japan" name="Honshu" pageId="11" pageNumber="122" stateProvince="Mie Pref.">Honshu</location>
:
<collectingRegion id="49D3F867BE49FFC6FE74BBB6824A0152" box="[412,525,1261,1287]" country="Japan" name="Mie" pageId="11" pageNumber="122">Mie Pref.</collectingRegion>
</emphasis>
•
<specimenCount id="9D11FD0CBE49FFC6FDCBBBB5827D015D" box="[547,570,1262,1288]" pageId="11" pageNumber="122" type="female">♀</specimenCount>
;
<location id="8EC8605EBE49FFC6FDAEBBB682DA0152" LSID="urn:lsid:plazi:treatment:03BE8793BE48FFC1FE32B991822E069A:8EC8605EBE49FFC6FDAEBBB682DA0152" box="[582,669,1261,1287]" country="Japan" name="Tsu-shi" pageId="11" pageNumber="122" stateProvince="Mie Pref.">Tsu-shi</location>
,
<location id="8EC8605EBE49FFC6FD4EBBB6835A0152" LSID="urn:lsid:plazi:treatment:03BE8793BE48FFC1FE32B991822E069A:8EC8605EBE49FFC6FD4EBBB6835A0152" box="[678,797,1261,1287]" country="Japan" name="Furukawa" pageId="11" pageNumber="122" stateProvince="Mie Pref.">Furukawa</location>
;
<date id="FFA91045BE49FFC6FCCFBBB58383015D" box="[807,964,1261,1288]" pageId="11" pageNumber="122" value="1958-08-23">
<collectingDate id="EFEDE9ADBE49FFC6FCCFBBB58383015D" box="[807,964,1261,1288]" pageId="11" pageNumber="122" value="1958-08-23">23 Aug. 1958</collectingDate>
</date>
;
<collectorName id="26E25353BE49FFC6FC26BBB5841E0152" box="[974,1113,1261,1288]" pageId="11" pageNumber="122">H. Ichihashi</collectorName>
leg.;
<collectionCode id="ED06AE40BE49FFC6FB7EBBB584A5015D" box="[1174,1250,1262,1288]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="11" pageNumber="122">EUMJ</collectionCode>
(
<figureCitation id="132C2A00BE49FFC6FB18BBB6852E015D" box="[1264,1385,1261,1288]" captionStart="Fig" captionStartId="9.[189,232,1886,1912]" captionTargetBox="[196,1392,306,1839]" captionTargetId="figure-11@9.[189,1398,268,1842]" captionTargetPageId="9" captionText="Fig. 3. Holotypes of Georissus spp. A–F. Dorsal habitus. G–L. Labels. A, K. G. (G.) babai Satô, 1970, ♂ (EUMJ). B, L. G. (Ni.) granulosus Satô, 1972, ♀ (EUMJ). C, H. G. (Ne.) kurosawai Nakane, 1963, ♂ (SEHU). D, J. G. (Ne.) japonicus Satô, 1972, ♂ (EUMJ). E, G. G. (Ne.) sakaii Satô, 1972, ♂ (EUMJ). F, I. G. (Ne.) katsuoi Nakane, 1995 (SEHU). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517956" httpUri="https://zenodo.org/record/6517956/files/figure.png" pageId="11" pageNumber="122">Fig. 3B, L</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA83685BE49FFC6FF02BA7485320138" blockId="11.[189,1397,1327,1389]" pageId="11" pageNumber="122">
<materialsCitation id="3B7F3CD8BE49FFC6FF02BA7485370138" ID-GBIF-Occurrence="3762863374" collectingDate="2008-06-16" collectionCode="EUMJ" collectorName="K. Akita" country="Japan" county="Ise-shi" location="Honshu" municipality="Iso-cho" pageId="11" pageNumber="122" specimenCount="5" stateProvince="Mie Pref.">
<emphasis id="B963EA97BE49FFC6FF02BA7481E1011C" bold="true" box="[234,422,1327,1353]" pageId="11" pageNumber="122">Other material</emphasis>
<collectingCountry id="F3007615BE49FFC6FF55BA0981510139" box="[189,278,1362,1388]" name="Japan" pageId="11" pageNumber="122">JAPAN</collectingCountry>
–
<emphasis id="B963EA97BE49FFC6FEDBBA09825C0139" bold="true" box="[307,539,1362,1388]" pageId="11" pageNumber="122">
<location id="8EC8605EBE49FFC6FEDBBA0981DE0139" LSID="urn:lsid:plazi:treatment:03BE8793BE48FFC1FE32B991822E069A:8EC8605EBE49FFC6FEDBBA0981DE0139" box="[307,409,1362,1388]" country="Japan" county="Ise-shi" municipality="Iso-cho" name="Honshu" pageId="11" pageNumber="122" stateProvince="Mie Pref.">Honshu</location>
:
<collectingRegion id="49D3F867BE49FFC6FE4EBA09825C0139" box="[422,539,1362,1388]" country="Japan" name="Mie" pageId="11" pageNumber="122">Mie Pref.</collectingRegion>
</emphasis>
•
<specimenCount id="9D11FD0CBE49FFC6FDDBBA0982280138" box="[563,623,1362,1389]" pageId="11" pageNumber="122" type="generic">5 ex.</specimenCount>
;
<collectingCounty id="62C94E09BE49FFC6FD94BA09828A0139" box="[636,717,1362,1388]" pageId="11" pageNumber="122">Ise-shi</collectingCounty>
,
<collectingMunicipality id="6BCCACFFBE49FFC6FD31BA0983730139" box="[729,820,1362,1388]" pageId="11" pageNumber="122">Iso-cho</collectingMunicipality>
,
<location id="8EC8605EBE49FFC6FCA8BA09838E0139" LSID="urn:lsid:plazi:treatment:03BE8793BE48FFC1FE32B991822E069A:8EC8605EBE49FFC6FCA8BA09838E0139" box="[832,969,1362,1388]" country="Japan" county="Ise-shi" municipality="Iso-cho" name="Miya-gawa" pageId="11" pageNumber="122" stateProvince="Mie Pref.">Miya-gawa</location>
;
<date id="FFA91045BE49FFC6FC3DBA0984280139" box="[981,1135,1362,1389]" pageId="11" pageNumber="122" value="2008-06-16">
<collectingDate id="EFEDE9ADBE49FFC6FC3DBA0984280139" box="[981,1135,1362,1389]" pageId="11" pageNumber="122" value="2008-06-16">16 Jun. 2008</collectingDate>
</date>
;
<collectorName id="26E25353BE49FFC6FB93BA0884990139" box="[1147,1246,1362,1389]" pageId="11" pageNumber="122">K. Akita</collectorName>
leg.;
<collectionCode id="ED06AE40BE49FFC6FAC8BA0885370138" box="[1312,1392,1363,1389]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="11" pageNumber="122">EUMJ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE49FFC1FF55BAC882000598" lastPageId="12" lastPageNumber="123" pageId="11" pageNumber="122" type="description">
<paragraph id="8BA83685BE49FFC6FF55BAC8812C01F8" blockId="11.[189,363,1427,1453]" box="[189,363,1427,1453]" pageId="11" pageNumber="122">
<heading id="D0E081E9BE49FFC6FF55BAC8812C01F8" bold="true" box="[189,363,1427,1453]" fontSize="11" level="3" pageId="11" pageNumber="122" reason="3">
<emphasis id="B963EA97BE49FFC6FF55BAC8812C01F8" bold="true" box="[189,363,1427,1453]" pageId="11" pageNumber="122">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE49FFC6FF55BA99833D0189" blockId="11.[189,890,1474,1501]" box="[189,890,1474,1501]" pageId="11" pageNumber="122">
COLORATION (
<figureCitation id="132C2A00BE49FFC6FEB1BA9981F20189" box="[345,437,1474,1501]" captionStart="Fig" captionStartId="7.[189,232,1886,1912]" captionTargetBox="[272,1331,265,1833]" captionTargetId="figure-11@7.[223,1365,265,1845]" captionTargetPageId="7" captionText="Fig. 2. Dorsal view of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F. G. (Ne.) takahashii sp. nov., holotype, ♂ (EUMJ). G. G. (Ne.) satoi sp. nov., holotype (EUMJ). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517952" httpUri="https://zenodo.org/record/6517952/files/figure.png" pageId="11" pageNumber="122">Fig. 2C</figureCitation>
). Black, mat, legs dark reddish brown.
</paragraph>
<paragraph id="8BA83685BE49FFC6FF55B95F82820231" blockId="11.[187,1399,1539,1636]" pageId="11" pageNumber="122">
HEAD (
<figureCitation id="132C2A00BE49FFC6FEE4B9588121024B" box="[268,358,1539,1566]" captionStart="Fig" captionStartId="18.[189,232,1882,1908]" captionTargetBox="[191,1394,563,1839]" captionTargetId="figure-90@18.[189,1398,562,1842]" captionTargetPageId="18" captionText="Fig. 6. Head of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517968" httpUri="https://zenodo.org/record/6517968/files/figure.png" pageId="11" pageNumber="122">Fig. 6C</figureCitation>
). Clypeus uneven, provided with foveae and carinae throughout;
<emphasis id="B963EA97BE49FFC6FBA6B958842A0248" box="[1102,1133,1539,1565]" italics="true" pageId="11" pageNumber="122">slr</emphasis>
, consisted of granules, connecting to
<emphasis id="B963EA97BE49FFC6FE8EB97C81D20214" box="[358,405,1575,1601]" italics="true" pageId="11" pageNumber="122">smb</emphasis>
at median portion;
<emphasis id="B963EA97BE49FFC6FD68B97C82E80214" box="[640,687,1575,1601]" italics="true" pageId="11" pageNumber="122">smb</emphasis>
nonsequential, connecting each other, forming oblong oval;
<emphasis id="B963EA97BE49FFC6FF56B91180A70231" box="[190,224,1610,1636]" italics="true" pageId="11" pageNumber="122">lsa</emphasis>
connecting each other, forming U-shape.
</paragraph>
<paragraph id="8BA83685BE49FFC6FF55B9D78515035A" blockId="11.[189,1398,1675,1807]" pageId="11" pageNumber="122">
PRONOTUM (
<figureCitation id="132C2A00BE49FFC6FEA5B9D081E802F0" box="[333,431,1675,1702]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="11" pageNumber="122">Fig. 7C</figureCitation>
). Relatively large, widest at the middle. Pronotal disc provided with granules densely. Anterior portion provided with indistinct granules extensively; anterior margin smooth, not crenelated;
<emphasis id="B963EA97BE49FFC6FEAEB989813A02B9" box="[326,381,1746,1772]" italics="true" pageId="11" pageNumber="122">smlg</emphasis>
distinct, with longitudinal row of pits. Posterior portion slightly convex than anterior potion, bearing granulation regularly; lateral margins protruding;
<emphasis id="B963EA97BE49FFC6FC55B9AE8398035A" box="[957,991,1781,1807]" italics="true" pageId="11" pageNumber="122">slb</emphasis>
developed;
<emphasis id="B963EA97BE49FFC6FB87B9AD84CB035A" box="[1135,1164,1782,1807]" italics="true" pageId="11" pageNumber="122">pg</emphasis>
indistinct, short.
</paragraph>
<paragraph id="8BA83685BE49FFC6FF55B86C84280321" blockId="11.[187,1399,1846,1908]" pageId="11" pageNumber="122">ELYTRA. Entirely granulate. Elytral intervals 2, 4, 6 and 8 elevated, forming longitudinal carinae. Humeral bulge distinct, bearing granules. Lateral ridges indistinct. Hind wings developed.</paragraph>
<paragraph id="8BA83685BE49FFC6FF55B8C0842A038D" blockId="11.[189,1398,1947,2009]" pageId="11" pageNumber="122">
ABDOMEN (
<figureCitation id="132C2A00BE49FFC6FED7B8C081DB03E0" box="[319,412,1947,1973]" captionStart="Fig" captionStartId="21.[191,234,1890,1916]" captionTargetBox="[193,1394,569,1846]" captionTargetId="figure-107@21.[189,1398,569,1852]" captionTargetPageId="21" captionText="Fig. 8. Ventrites of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (F). I. G. (Ne.) satoi sp. nov. Abbreviation: vt = ventral teeth. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6517982" httpUri="https://zenodo.org/record/6517982/files/figure.png" pageId="11" pageNumber="122">Fig. 8C</figureCitation>
). Lateral margin of ventrites uniformly narrowing posteriorly. Ventrite 1 with large granules extensively. Granulation of ventrites 2–5 large and relatively indistinct.
</paragraph>
<paragraph id="8BA83685BE4EFFC1FF55BE5284AB05D9" blockId="12.[189,1401,265,396]" pageId="12" pageNumber="123">
MALE GENITALIA (
<figureCitation id="132C2A00BE4EFFC1FE62BE5281AF0576" box="[394,488,265,291]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="12" pageNumber="123">Fig. 9C</figureCitation>
). Aedeagus
<quantity id="4CEF9B60BE4EFFC1FD92BE5282A30576" box="[634,740,265,291]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="12" pageNumber="123" unit="mm" value="0.37">0.37 mm</quantity>
long. Parameres slightly shorter than phallobase, basal portions of parameres combined as wide as anterior portion of phallobase; lateral margins. Median lobe shorter than parameres, subparallel-sided, slightly angular at apex; basal struts short. Phallobase about 2.4 times as long as wide, widening posteriorly; posterior portion with large basal foramen.
</paragraph>
<paragraph id="8BA83685BE4EFFC1FF55BEEF82000598" blockId="12.[189,583,435,462]" box="[189,583,435,462]" pageId="12" pageNumber="123">FEMALE GENITALIA. Not examined.</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4EFFC1FF55BEAF81EF0635" pageId="12" pageNumber="123" type="discussion">
<paragraph id="8BA83685BE4EFFC1FF55BEAF81C6065B" blockId="12.[189,385,500,526]" box="[189,385,500,526]" pageId="12" pageNumber="123">
<heading id="D0E081E9BE4EFFC1FF55BEAF81C6065B" bold="true" box="[189,385,500,526]" fontSize="11" level="3" pageId="12" pageNumber="123" reason="3">
<emphasis id="B963EA97BE4EFFC1FF55BEAF81C6065B" bold="true" box="[189,385,500,526]" pageId="12" pageNumber="123">Biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4EFFC1FF55BD7881EF0635" blockId="12.[189,1399,547,608]" pageId="12" pageNumber="123">
This species inhabits tidal flats that are submerged at high tide (
<bibRefCitation id="EF864B74BE4EFFC1FC74BD7884610668" author="Akita K." box="[924,1062,547,573]" pageId="12" pageNumber="123" pagination="14 - 15" refId="ref12350" refString="Akita K. 2008. Rediscovery of Georissus granulosus. Coleopterist's News 164: 14 - 15 [in Japanese.]" type="journal article" year="2008">
<collectingRegion id="49D3F867BE4EFFC1FC74BD7883980668" box="[924,991,547,573]" country="Japan" name="Akita" pageId="12" pageNumber="123">Akita</collectingRegion>
2008
</bibRefCitation>
). The
<typeStatus id="54AC8827BE4EFFC1FB83BD7884950668" box="[1131,1234,547,573]" pageId="12" pageNumber="123" type="holotype">holotype</typeStatus>
was collected at light (
<bibRefCitation id="EF864B74BE4EFFC1FEF7BD1D81DC0635" author="Sato M." box="[287,411,582,608]" pageId="12" pageNumber="123" pagination="1 - 4" refId="ref14105" refString="Sato M. 1981. An outline of Japanese Georissidae. Coleopterist's News 53: 1 - 4. [In Japanese]" type="journal article" year="1981">Satô 1981</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4EFFC1FF55BDDC822E069A" box="[189,617,647,720]" pageId="12" pageNumber="123" type="distribution">
<paragraph id="8BA83685BE4EFFC1FF55BDDC811106F4" blockId="12.[189,342,647,673]" box="[189,342,647,673]" pageId="12" pageNumber="123">
<heading id="D0E081E9BE4EFFC1FF55BDDC811106F4" bold="true" box="[189,342,647,673]" fontSize="11" level="3" pageId="12" pageNumber="123" reason="3">
<emphasis id="B963EA97BE4EFFC1FF55BDDC811106F4" bold="true" box="[189,342,647,673]" pageId="12" pageNumber="123">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4EFFC1FF55BDED822E069A" blockId="12.[189,617,693,720]" box="[189,617,693,720]" pageId="12" pageNumber="123">
<collectingCountry id="F3007615BE4EFFC1FF55BDED81430685" box="[189,260,694,720]" name="Japan" pageId="12" pageNumber="123">Japan</collectingCountry>
: Honshu (
<collectingRegion id="49D3F867BE4EFFC1FE92BDEE81AA0685" box="[378,493,693,720]" country="Japan" name="Mie" pageId="12" pageNumber="123">Mie Pref.</collectingRegion>
) (
<figureCitation id="132C2A00BE4EFFC1FDECBDEE821C069A" box="[516,603,693,719]" captionStart="Fig" captionStartId="27.[189,232,1952,1978]" captionTargetBox="[206,1384,303,1881]" captionTargetId="figure-11@27.[189,1398,290,1888]" captionTargetPageId="27" captionText="Fig. 13. Distribution map of Georissus spp. Russian localities for Georissus (G.) canalifer Sharp, 1888 and G. (Ne.) japonicus Satô, 1972 are omitted." figureDoi="http://doi.org/10.5281/zenodo.6518004" httpUri="https://zenodo.org/record/6518004/files/figure.png" pageId="12" pageNumber="123">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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