220 lines
27 KiB
XML
220 lines
27 KiB
XML
<document id="B446FAF41CDF83533A9C2C475908C046" ID-DOI="10.5281/zenodo.13203792" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203792" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722658458551" checkinUser="felipe" docAuthor="Minkina, Ÿukasz, Anichtchenko, Alexander, Vasiljeva, Anastasija & Skelley, Paul E." docDate="2022" docId="03BD44335E52FFEBFC8C266800CDFE08" docLanguage="en" docName="BalticJColeopterol.22.1.81-101.pdf" docOrigin="Baltic Journal of Coleopterology 22 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.4:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="4" docTitle="Rhyparus buruensis Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="86" masterDocId="FF843C4B5E51FFEEFF8A22690417FFB0" masterDocTitle="Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females" masterLastPageNumber="101" masterPageNumber="81" pageNumber="84" updateTime="1722698086404" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="70BCECA3AFD96E12AAA908ED53F2B09A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A0A8B98D7980C55251D37356AD1C1000">
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<mods:title id="C64EB46C445D7275A961E0EFF6838A94">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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</mods:titleInfo>
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<mods:name id="CBD64F067FC78C343859CFBEB384FBA8" type="personal">
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<mods:role id="EE71FD0FF9AA4F29A94A7CEAB1EEC2A5">
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<mods:roleTerm id="664B1032CC959754083472BB6C31C876">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5716C1A20F287E1D5F97BF0C71CB56A9">Minkina, Ÿukasz</mods:namePart>
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</mods:name>
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<mods:name id="7D3800E79453F58717C0FA86799B2A8F" type="personal">
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<mods:role id="BC002AC4D5FCD686EFE04823EFB76008">
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<mods:roleTerm id="29667E6BF5F4DDCE248D1BC686E6E8BF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="742A02639B5B7F6E5BA5E5FF93EB4601">Anichtchenko, Alexander</mods:namePart>
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</mods:name>
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<mods:name id="89698DCB54D94F1539C8BA28BF2A2485" type="personal">
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<mods:roleTerm id="E00BB3263AA0640B3B23A535E6DF037C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9A204BE8DD9AC1BADF5D4F9D4913D03B">Vasiljeva, Anastasija</mods:namePart>
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</mods:name>
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<mods:name id="700376F181D288093D426E58E86AB70C" type="personal">
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<mods:role id="A97672301BAC7BDEE1B18F6FD5BFF21D">
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<mods:roleTerm id="EDEFFE2ACA75FA1214AE0D16E4D91F77">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="794F09641411793A1C66D1DAB5CA512B">Skelley, Paul E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A18E5A8379824AE5C449468119B18737">text</mods:typeOfResource>
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<mods:relatedItem id="04CD12FB68F8213562C7B83A3FAB2DCA" type="host">
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<mods:titleInfo id="8F417781C444C320B4490CD42CF333B5">
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<mods:title id="28DF26B46CD2A6C14670CA0EA16E2E31">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="42D1C9898893DE7EDAB44C50B035C847">
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<mods:date id="AD0654887D66E2FF1BEE884601C46D7C">2022</mods:date>
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<mods:detail id="147623E9F2FB49223E64DA3FEBECB896" type="volume">
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<mods:number id="4AE19805853F6078A664354549B8B094">22</mods:number>
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</mods:detail>
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<mods:detail id="01265ACD33BE0BF87D897B3C43CE4742" type="issue">
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<mods:number id="ED7A221F048FE3E6332FE69F080CDDE2">1</mods:number>
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</mods:detail>
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<mods:extent id="6F209632C3C4D52D40B63599C565CDED" unit="page">
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<mods:start id="A15CFE7E4264FF982E97B5811F3814E0">81</mods:start>
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<mods:end id="77C97AC4FA6789B9C36D6511B1B84B8D">101</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="EE63D933874C7E627A7822D32F070277">journal article</mods:classification>
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<mods:identifier id="8FBE8DB4F5288CFCDB30449B1325761A" type="DOI">10.5281/zenodo.13203792</mods:identifier>
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<mods:identifier id="945FE9EC825BB3270FFB040CEAE14F38" type="ISSN">1407-8619</mods:identifier>
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<mods:identifier id="A630788012446B6D2618E37CB055CC89" type="Zenodo-Dep">13203792</mods:identifier>
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</mods:mods>
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<treatment id="03BD44335E52FFEBFC8C266800CDFE08" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08" httpUri="http://treatment.plazi.org/id/03BD44335E52FFEBFC8C266800CDFE08" lastPageId="5" lastPageNumber="86" pageId="3" pageNumber="84">
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<subSubSection id="C30EA6AE5E52FFEDFC8C26680028FBA8" box="[774,1087,1025,1049]" pageId="3" pageNumber="84" type="nomenclature">
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<paragraph id="8BABF5255E52FFEDFC8C26680028FBA8" blockId="3.[774,1087,1025,1080]" box="[774,1087,1025,1049]" pageId="3" pageNumber="84">
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<heading id="D0E342495E52FFEDFC8C26680028FBA8" box="[774,1087,1025,1049]" fontSize="10" level="2" pageId="3" pageNumber="84" reason="7">
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<taxonomicName id="4C148EA65E52FFEDFC8C266807F2FBA9" ID-CoL="9WHZK" authorityName="Minkina & Anichtchenko & Vasiljeva & Skelley" authorityYear="2022" box="[774,997,1025,1049]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="buruensis" status="sp. nov.">
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<emphasis id="B96029375E52FFEDFC8C266807F2FBA9" bold="true" box="[774,997,1025,1049]" italics="true" pageId="3" pageNumber="84">Rhyparus buruensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A253944C5E52FFEDFC6626680028FBA8" box="[1004,1087,1025,1048]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30EA6AE5E52FFEDFC8C264807D8FB88" box="[774,975,1057,1080]" pageId="3" pageNumber="84" type="description">
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||
<paragraph id="8BABF5255E52FFEDFC8C264807D8FB88" blockId="3.[774,1087,1025,1080]" box="[774,975,1057,1080]" pageId="3" pageNumber="84">
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||
(
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<figureCitation id="132FE9A05E52FFEDFC8426480761FB88" box="[782,886,1057,1080]" captionStart="Figs" captionStartId="4.[302,352,1693,1716]" captionTargetBox="[483,1187,751,1656]" captionTargetId="figure-275@4.[478,1187,748,1659]" captionTargetPageId="4" captionText="Figs. 4–6. Rhyparus buruensis sp. nov.,, holotype: 4 – dorsal view; 5 – ventral view; 6 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203798" httpUri="https://zenodo.org/record/13203798/files/figure.png" pageId="3" pageNumber="84">Figs. 4–6</figureCitation>
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,
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<figureCitation id="132FE9A05E52FFEDFC0826480788FB88" box="[898,927,1057,1080]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 20–25. Heads: 20 – R. bacanensis sp. nov.,, holotype; 21 – R. buruensis sp. nov.,, holotype; 22 – R. fijiensis sp. nov.,, holotype; 23 – R. stebnickae sp. nov.,, holotype; 24 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 25 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203808" httpUri="https://zenodo.org/record/13203808/files/figure.png" pageId="3" pageNumber="84">21</figureCitation>
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,
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<figureCitation id="132FE9A05E52FFEDFC21264807D0FB88" box="[939,967,1057,1080]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 26–33.Abdominal apexes with blackwhite miniatures of pygidium on leftdown side: 26 – R. bacanensis sp. nov.,, holotype; 27 – R. buruensis sp. nov.,, holotype; 28 – R. fijiensis sp. nov.,, holotype; 29 – R. stebnickae sp. nov.,, holotype; 30 – R. approximans Fairmaire, 1893,; 31 – R. loebli Paulian, 1983,; 32 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 33 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13203810" httpUri="https://zenodo.org/record/13203810/files/figure.png" pageId="3" pageNumber="84">27</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30EA6AE5E52FFEDFC8C26090043FACA" pageId="3" pageNumber="84" type="materials_examined">
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<paragraph id="8BABF5255E52FFEDFC8C2609005DFB28" blockId="3.[774,1285,1120,1176]" pageId="3" pageNumber="84">
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<materialsCitation id="3B7CFF785E52FFEDFC8C26090051FB28" country="Indonesia" county="Ramaja Mount" location="Buru Island" municipality="Ilat" pageId="3" pageNumber="84" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B96029375E52FFEDFC8C2609078CFBC8" bold="true" box="[774,923,1120,1144]" pageId="3" pageNumber="84">
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<typeStatus id="54AF4B875E52FFEDFC8C26090729FBC8" box="[774,830,1120,1144]" pageId="3" pageNumber="84">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F303B5B55E52FFEDFC152608001EFBC8" box="[927,1033,1121,1144]" name="Indonesia" pageId="3" pageNumber="84">Indonesia</collectingCountry>
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, Moluccas,
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<location id="8ECBA3FE5E52FFEDFB0E26080116FBC8" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08:8ECBA3FE5E52FFEDFB0E26080116FBC8" box="[1156,1281,1121,1144]" country="Indonesia" county="Ramaja Mount" municipality="Ilat" name="Buru Island" pageId="3" pageNumber="84">Buru Island</location>
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,
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<collectingMunicipality id="6BCF6F5F5E52FFEDFC8C26E8073DFB28" box="[774,810,1153,1176]" pageId="3" pageNumber="84">Ilat</collectingMunicipality>
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vill., env.,
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<collectingCounty id="62CA8DA95E52FFEDFC2826E80051FB28" box="[930,1094,1153,1176]" pageId="3" pageNumber="84">Ramaja Mount</collectingCounty>
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</materialsCitation>
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.
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</paragraph>
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||
<paragraph id="8BABF5255E52FFEDFC8C26A80043FACA" blockId="3.[774,1286,1217,1403]" pageId="3" pageNumber="84">
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||
<emphasis id="B96029375E52FFEDFC8C26A807BBFB69" bold="true" box="[774,940,1217,1241]" pageId="3" pageNumber="84">Type material.</emphasis>
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||
<materialsCitation id="3B7CFF785E52FFEDFC3A26A80116FAAA" collectingDate="2012-12" collectionCode="Y, MCN" collectorName="S. Jakl" country="Indonesia" county="Ramaja Mt." elevation="275" location="Buru Island" municipality="Ilat" pageId="3" pageNumber="84" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54AF4B875E52FFEDFC3A26A80003FB68" box="[944,1044,1217,1240]" pageId="3" pageNumber="84" type="holotype">Holotype</typeStatus>
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||
():
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||
<collectingCountry id="F303B5B55E52FFEDFBC226A800A7FB68" box="[1096,1200,1217,1240]" name="Indonesia" pageId="3" pageNumber="84">Indonesia</collectingCountry>
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| C Moluccas: see
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<location id="8ECBA3FE5E52FFEDFC0F268A0012FB4A" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08:8ECBA3FE5E52FFEDFC0F268A0012FB4A" box="[901,1029,1251,1274]" country="Indonesia" county="Ramaja Mt." municipality="Ilat" name="Buru Island" pageId="3" pageNumber="84">Buru Island</location>
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|
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||
<quantity id="4CEC58C05E52FFEDFB91268A009AFB4A" box="[1051,1165,1251,1274]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="3" pageNumber="84" unit="m" value="275.0" valueMax="350.0" valueMin="200.0">
|
||
<elevation id="003912165E52FFEDFB91268A009AFB4A" box="[1051,1165,1251,1274]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="3" pageNumber="84" unit="m" value="275.0" valueMax="350.0" valueMin="200.0">200 350m</elevation>
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</quantity>
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||
. |
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<collectingMunicipality id="6BCF6F5F5E52FFEDFB2D268A00DCFB4A" box="[1191,1227,1251,1274]" pageId="3" pageNumber="84">Ilat</collectingMunicipality>
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vill., env.,
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<collectingCounty id="62CA8DA95E52FFEDFCB1276A07A3FAAA" box="[827,948,1283,1306]" pageId="3" pageNumber="84">Ramaja Mt.</collectingCounty>
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||
|
|
||
<date id="FFAAD3E55E52FFEDFC4B276A0001FAAA" box="[961,1046,1283,1306]" pageId="3" pageNumber="84" value="2012-12">
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||
<collectingDate id="EFEE2A0D5E52FFEDFC4B276A0001FAAA" box="[961,1046,1283,1306]" pageId="3" pageNumber="84" value="2012-12">xii.2012</collectingDate>
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</date>
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| leg.
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<collectorName id="26E190F35E52FFEDFBC7276A0087FAAA" box="[1101,1168,1283,1306]" pageId="3" pageNumber="84">S. Jakl</collectorName>
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|| (
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||
<emphasis id="B96029375E52FFEDFB21276900ACFAAA" bold="true" box="[1195,1211,1280,1306]" pageId="3" pageNumber="84">
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||
<collectionCode id="ED056DE05E52FFEDFB21276900ACFAAA" box="[1195,1211,1280,1306]" pageId="3" pageNumber="84">Ÿ</collectionCode>
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||
</emphasis>
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||
<collectionCode id="ED056DE05E52FFEDFB31276A00ECFAAA" box="[1211,1275,1283,1306]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="3" pageNumber="84" type="Herbarium">MCN</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B7CFF785E52FFEDFC8C274A0047FACA" collectingDate="2013-06" collectionCode="Y, MCN" country="Indonesia" county="Ramaja Mt." elevation="200" location="Buru Island" municipality="Ilat" pageId="3" pageNumber="84" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54AF4B875E52FFEDFC8C274A0773FA8A" box="[774,868,1315,1338]" pageId="3" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
():
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||
<collectingCountry id="F303B5B55E52FFEDFC16274A0012FA8A" box="[924,1029,1315,1338]" name="Indonesia" pageId="3" pageNumber="84">Indonesia</collectingCountry>
|
||
| C Moluccas: see
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||
<location id="8ECBA3FE5E52FFEDFB45274A075FFAEA" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08:8ECBA3FE5E52FFEDFB45274A075FFAEA" country="Indonesia" county="Ramaja Mt." municipality="Ilat" name="Buru Island" pageId="3" pageNumber="84">Buru Island</location>
|
||
|
|
||
<quantity id="4CEC58C05E52FFEDFCD7272A07D6FAEA" box="[861,961,1347,1370]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="3" pageNumber="84" unit="m" value="200.0" valueMax="350.0" valueMin="50.0">
|
||
<elevation id="003912165E52FFEDFCD7272A07D6FAEA" box="[861,961,1347,1370]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="3" pageNumber="84" unit="m" value="200.0" valueMax="350.0" valueMin="50.0">50 350m</elevation>
|
||
</quantity>
|
||
. |
|
||
<collectingMunicipality id="6BCF6F5F5E52FFEDFC53272A07EAFAEA" box="[985,1021,1347,1370]" pageId="3" pageNumber="84">Ilat</collectingMunicipality>
|
||
vill., env.,
|
||
<collectingCounty id="62CA8DA95E52FFEDFBF2272A00EEFAEA" box="[1144,1273,1347,1370]" pageId="3" pageNumber="84">Ramaja Mt.</collectingCounty>
|
||
|
|
||
<date id="FFAAD3E55E52FFEDFC8C270A074FFACA" box="[774,856,1379,1402]" pageId="3" pageNumber="84" value="2013-06">
|
||
<collectingDate id="EFEE2A0D5E52FFEDFC8C270A074FFACA" box="[774,856,1379,1402]" pageId="3" pageNumber="84" value="2013-06">vi.2013</collectingDate>
|
||
</date>
|
||
| local coll. || (
|
||
<emphasis id="B96029375E52FFEDFC7C27080011FACB" bold="true" box="[1014,1030,1377,1403]" pageId="3" pageNumber="84">
|
||
<collectionCode id="ED056DE05E52FFEDFC7C27080011FACB" box="[1014,1030,1377,1403]" pageId="3" pageNumber="84">Ÿ</collectionCode>
|
||
</emphasis>
|
||
<collectionCode id="ED056DE05E52FFEDFB8C270A005FFACA" box="[1030,1096,1379,1402]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="3" pageNumber="84" type="Herbarium">MCN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30EA6AE5E52FFEBFC8C27CA0038FF48" lastPageId="5" lastPageNumber="86" pageId="3" pageNumber="84" type="description">
|
||
<paragraph id="8BABF5255E52FFEDFC8C27CA0097F90A" blockId="3.[774,1285,1443,1722]" pageId="3" pageNumber="84">
|
||
<emphasis id="B96029375E52FFEDFC8C27CA0050FA0B" bold="true" box="[774,1095,1443,1467]" pageId="3" pageNumber="84">
|
||
Description of the
|
||
<typeStatus id="54AF4B875E52FFEDFC5727CA0055FA0B" box="[989,1090,1443,1467]" pageId="3" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Dorsum (
|
||
<figureCitation id="132FE9A05E52FFEDFB3E27CA00EFFA0A" box="[1204,1272,1443,1466]" captionStart="Figs" captionStartId="4.[302,352,1693,1716]" captionTargetBox="[483,1187,751,1656]" captionTargetId="figure-275@4.[478,1187,748,1659]" captionTargetPageId="4" captionText="Figs. 4–6. Rhyparus buruensis sp. nov.,, holotype: 4 – dorsal view; 5 – ventral view; 6 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203798" httpUri="https://zenodo.org/record/13203798/files/figure.png" pageId="3" pageNumber="84">Fig. 4</figureCitation>
|
||
). Length:
|
||
<quantity id="4CEC58C05E52FFEDFCD127AA07BCFA6A" box="[859,939,1475,1498]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="84" unit="mm" value="7.3">7.3 mm</quantity>
|
||
; maximum width:
|
||
<quantity id="4CEC58C05E52FFEDFBED27AA00D2FA6A" box="[1127,1221,1475,1498]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="84" unit="mm" value="2.45">2.45 mm</quantity>
|
||
. Body largesized for members of this genus, elongate, not so distinctly convex, flattened in central part; shiny; apparently almost glabrous, though partly clothed with very small yellowish macrosetae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E55FFEAFEA422A805F8FDC8" blockId="4.[302,813,193,632]" pageId="4" pageNumber="85">
|
||
Head (
|
||
<figureCitation id="132FE9A05E55FFEAFEFC22A805D1FF68" box="[374,454,193,216]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 20–25. Heads: 20 – R. bacanensis sp. nov.,, holotype; 21 – R. buruensis sp. nov.,, holotype; 22 – R. fijiensis sp. nov.,, holotype; 23 – R. stebnickae sp. nov.,, holotype; 24 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 25 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203808" httpUri="https://zenodo.org/record/13203808/files/figure.png" pageId="4" pageNumber="85">Fig. 21</figureCitation>
|
||
) weakly shiny; transversely subhexagonal; clypeus trapezoidal in outline, anteriorly weakly sinuate, on sides weakly upturned as obtuse, quite distinct tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of quite distinct, very short, convergent ridges, on whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E55FFEAFCD822A80146FD08" blockId="4.[850,1361,193,696]" pageId="4" pageNumber="85">Pronotum quite distinctly shiny; with eight distinct costae and seven intercostae, with two lateral, rounded lobes on each side.Anterior lobes somewhat higherand distinctly narrower than posterior, on the top are the widest part of pronotum. Costae of third and fourth pair not interrupted in basal part of apical half, very gently convergent, distinctly convergent in the middle of apical third; median and second pair of costae distinctly interrupted in basal part of apical half; costae on each side with very small punctures bearing very small macrosetae. All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, weakly concentrated around median part into longitudinal line, all intercostae in basal part with distinct, dense punctation.</paragraph>
|
||
<caption id="DF6BA5AD5E55FFEAFEA424F4065FF964" ID-DOI="http://doi.org/10.5281/zenodo.13203798" ID-Zenodo-Dep="13203798" httpUri="https://zenodo.org/record/13203798/files/figure.png" pageId="4" pageNumber="85" startId="4.[302,352,1693,1716]" targetBox="[483,1187,751,1656]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BABF5255E55FFEAFEA424F4065FF964" blockId="4.[302,1361,1693,1748]" pageId="4" pageNumber="85">
|
||
Figs. 4–6.
|
||
<taxonomicName id="4C148EA65E55FFEAFE2B24F7066EF905" authorityName="Minkina & Anichtchenko & Vasiljeva & Skelley" authorityYear="2022" box="[417,633,1694,1717]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="buruensis" status="sp. nov.">
|
||
<emphasis id="B96029375E55FFEAFE2B24F7066EF905" box="[417,633,1694,1717]" italics="true" pageId="4" pageNumber="85">Rhyparus buruensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A253944C5E55FFEAFD0A24F406C4F904" box="[640,723,1693,1716]" pageId="4" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,, holotype: 4 – dorsal view; 5 – ventral view; 6 – lateral view. Scale lines: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BABF5255E54FFEBFD8A22F807FFFF14" blockId="5.[512,1000,145,164]" box="[512,1000,145,164]" pageId="5" pageNumber="86">
|
||
<emphasis id="B96029375E54FFEBFD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="5" pageNumber="86">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF6922A80628FF68" blockId="5.[227,575,193,216]" box="[227,575,193,216]" pageId="5" pageNumber="86">Scutellum almost imperceptible.</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF69236806B9FD48" blockId="5.[227,737,257,760]" pageId="5" pageNumber="86">Elytra shiny. Each elytron with six elevated costae, and five flat intercostae. Costae on sides with very small punctures bearing very small macrosetae. Preapical glandular area relatively small. Intercostae first to fourth with two distinct rows of punctures; on third intercostae there is short additional costae with additional row of punctures untill half of length of elytra. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremely short additional costa, with some additional punctures here. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and mediointernal caudal bulbs not divided; mediointernal caudal bulb shortened, transversally rounded, internally stretched.</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF6921480582FC08" blockId="5.[227,737,801,952]" pageId="5" pageNumber="86">
|
||
Pygidium (
|
||
<figureCitation id="132FE9A05E54FFEBFEDF21480588FC88" box="[341,415,801,824]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 26–33.Abdominal apexes with blackwhite miniatures of pygidium on leftdown side: 26 – R. bacanensis sp. nov.,, holotype; 27 – R. buruensis sp. nov.,, holotype; 28 – R. fijiensis sp. nov.,, holotype; 29 – R. stebnickae sp. nov.,, holotype; 30 – R. approximans Fairmaire, 1893,; 31 – R. loebli Paulian, 1983,; 32 – R. paraphilippinensis Ochi & Kakizoe, 2021,; 33 – R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13203810" httpUri="https://zenodo.org/record/13203810/files/figure.png" pageId="5" pageNumber="86">Fig. 27</figureCitation>
|
||
) with dense, irregularly spaced punctation, with weak longitudinal rib in the middle, and deep excision on its sides; with distinctly longitudinal apex in the shape of volcano with sinuate top.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF69218806C2FAC8" blockId="5.[227,738,993,1400]" pageId="5" pageNumber="86">
|
||
Venter (
|
||
<figureCitation id="132FE9A05E54FFEBFEB121880597FC48" box="[315,384,993,1016]" captionStart="Figs" captionStartId="4.[302,352,1693,1716]" captionTargetBox="[483,1187,751,1656]" captionTargetId="figure-275@4.[478,1187,748,1659]" captionTargetPageId="4" captionText="Figs. 4–6. Rhyparus buruensis sp. nov.,, holotype: 4 – dorsal view; 5 – ventral view; 6 – lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203798" httpUri="https://zenodo.org/record/13203798/files/figure.png" pageId="5" pageNumber="86">Fig. 5</figureCitation>
|
||
) shiny. Mesometaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of mesometaventral plate dense, regularly spaced, irregular in size; all punctures bearing short macrosetae. Abdominal ventrites moderately shiny, on sides with rows of punctures; with an additional punctured furrow in basal part. Last abdominal ventrite with dense punctures, which are similar size as on last but one ventrite; in basal half in the middle part with dense, very deep, longitudinal rows; in apical part in the middle with very deep groove, clearly located much below apex of pygidium.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF6927C8069CF988" blockId="5.[227,737,1441,1624]" pageId="5" pageNumber="86">Mesofemora with two very indistinct tubercles on lower border; metafemora without any tubercles on lower border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF69242905DBF9E8" blockId="5.[227,737,1441,1624]" box="[227,460,1600,1624]" pageId="5" pageNumber="86">
|
||
<emphasis id="B96029375E54FFEBFF6924290541F9E8" bold="true" box="[227,342,1600,1624]" pageId="5" pageNumber="86">Variation.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFF6924E90038FF48" blockId="5.[227,737,1664,1752]" lastBlockId="5.[774,1285,193,248]" pageId="5" pageNumber="86">
|
||
<emphasis id="B96029375E54FFEBFF6924E905DEF928" bold="true" box="[227,457,1664,1688]" pageId="5" pageNumber="86">Sexual dimorphism.</emphasis>
|
||
Unknown. When we look at shape of mesometaventral plate and apex of mesotibiae we can suppose that sexual dimorphism it is like typical for a whole genus (except pygidium and last ventrite).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30EA6AE5E54FFEBFC8C234900CDFE08" pageId="5" pageNumber="86" type="etymology">
|
||
<paragraph id="8BABF5255E54FFEBFC8C23490788FEC8" blockId="5.[774,1285,288,376]" pageId="5" pageNumber="86">
|
||
<emphasis id="B96029375E54FFEBFC8C23490794FE88" bold="true" box="[774,899,288,312]" pageId="5" pageNumber="86">Etymology.</emphasis>
|
||
Toponymic; an adjective derived from the name of Buru Island – where the
|
||
<typeStatus id="54AF4B875E54FFEBFB2C23280113FEE8" box="[1190,1284,321,344]" pageId="5" pageNumber="86" type="holotype">holotype</typeStatus>
|
||
was collected.
|
||
</paragraph>
|
||
<paragraph id="8BABF5255E54FFEBFC8C23C900CDFE08" blockId="5.[774,1242,416,440]" box="[774,1242,416,440]" pageId="5" pageNumber="86">
|
||
<emphasis id="B96029375E54FFEBFC8C23C90771FE08" bold="true" box="[774,870,416,440]" pageId="5" pageNumber="86">Affinity.</emphasis>
|
||
See discussion and tables 1 and 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |