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<document id="6EB496230054EB1307DEC3BCB74F9291" ID-DOI="10.5852/ejt.2024.967.2699" ID-ISSN="2118-9773" ID-Zenodo-Dep="13990819" ID-ZooBank="C08B8027-50CC-417E-BCD4-5183B9FF6738" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1729849508920" checkinUser="plazi" docAuthor="Zonstein, Sergei L." docDate="2024" docId="03A8B7235519FF8FFDD9E318FD84C9B2" docLanguage="en" docName="EJT.2024.967.1-185.pdf" docOrigin="European Journal of Taxonomy 967" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola fedotovi" docType="treatment" docVersion="4" lastPageNumber="54" masterDocId="FF91CF5B552AFFBAFFEFE72AFFB2CE5B" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia" masterLastPageNumber="185" masterPageNumber="1" pageNumber="52" updateTime="1730386529905" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A7E6BC370653DF7479AD11999FE0FBC7">A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia</mods:title>
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<paragraph id="8BBE06355519FF89FDD9E318FC4FCA17" blockId="51.[262,1325,1074,1173]" box="[566,1021,1074,1101]" pageId="51" pageNumber="52">
<heading id="D0F6B1595519FF89FDD9E318FC4FCA17" box="[566,1021,1074,1101]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="4C017DB65519FF89FDD9E318FC4FCA17" ID-CoL="4RMTR" authority="(Charitonov, 1946)" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[566,1021,1074,1101]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA275519FF89FDD9E318FCA5CA16" bold="true" box="[566,791,1074,1101]" italics="true" pageId="51" pageNumber="52">Raveniola fedotovi</emphasis>
(
<bibRefCitation id="EF907BC45519FF89FCC8E318FC47CA17" author="Charitonov D. E." box="[807,1013,1074,1100]" pageId="51" pageNumber="52" pagination="19 - 32" refId="ref77775" refString="Charitonov D. E. 1946. New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo 12: 19 - 32. [In Russian and Latin.]" type="journal article" year="1946">Charitonov, 1946</bibRefCitation>
)
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<subSubSection id="C31B55BE5519FF89FEE9E37CFCD8CACE" pageId="51" pageNumber="52" type="description">
<paragraph id="8BBE06355519FF89FEE9E37CFCD8CACE" blockId="51.[262,1325,1074,1173]" pageId="51" pageNumber="52">
<figureCitation id="133A1AB05519FF89FEE9E37CFED2CA2B" box="[262,352,1110,1136]" captionStart="Figs 10–18" captionStartId="108.[189,241,1826,1852]" captionTargetBox="[189,1398,265,1785]" captionTargetId="figure-83@108.[189,1398,265,1785]" captionTargetPageId="108" captionText="Figs 10–18. Raveniola spp., ♂♂, holotypes (except 14, 16), habitus, dorsal views. 10. R. insolita sp. nov. (ZISP). 11. R. ornata sp. nov. (SMNH). 12. R. ornatula sp. nov. (SMNH). 13. R. pamira sp. nov. (SMNH). 14. R. pamira sp. nov., paratype (SMNH). 15. R. diluta sp. nov. (SMNH). 16. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 17. R. pallens sp. nov. (SMNH). 18. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990826" httpUri="https://zenodo.org/record/13990826/files/figure.png" pageId="51" pageNumber="52">Figs 16</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FE83E37CFE3ECA2A" box="[364,396,1110,1137]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="51" pageNumber="52">51</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FE77E37CFE0ACA2B" box="[408,440,1110,1136]" captionStart="Figs 91–99" captionStartId="118.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-80@118.[189,1398,265,1816]" captionTargetPageId="118" captionText="Figs 91–99. Raveniola spp., ♂♂, holotypes (except 96, 98), cephalothorax, dorsal views. 91. R. ignobilis sp. nov. (SMNH). 92. R. insolita sp. nov. (ZISP). 93. R. ornata sp. nov. (SMNH). 94. R. ornatula sp. nov. (SMNH). 95. R. pamira sp. nov. (SMNH). 96. R. pamira, paratype (SMNH). 97. R. diluta sp. nov. (SMNH). 98. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 99. R. pallens sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990852" httpUri="https://zenodo.org/record/13990852/files/figure.png" pageId="51" pageNumber="52">98</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FE2BE37CFE40CA2A" box="[452,498,1110,1137]" captionStart="Figs 118–126" captionStartId="121.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-90@121.[189,1398,265,1816]" captionTargetPageId="121" captionText="Figs 118–126. Raveniola spp., ♀♀, paratypes (except 119, 123, 125), cephalothorax, dorsal views. 118. R. ignobilis sp. nov. (SMNH). 119. R. karategensis sp. nov., holotype (SMNH). 120. R. ornata sp. nov., from Sanglok (SMNH). 121. R. ornatula sp. nov. (SMNH). 122. R. pamira sp. nov. (SMNH). 123. R. sororculasp. nov., holotype (SMNH). 124. R. dilutasp. nov.(SMNH). 125. R. fedotovi(Charitonov, 1946), ♀ from Hojakurgan (SMNH). 126. R. pallens sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990860" httpUri="https://zenodo.org/record/13990860/files/figure.png" pageId="51" pageNumber="52">125</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FE10E37CFD9FCA2A" box="[511,557,1110,1137]" captionStart="Figs 148–159" captionStartId="124.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1691]" captionTargetId="figure-104@124.[189,1399,265,1691]" captionTargetPageId="124" captionText="Figs 148–159. Raveniola spp., ♂♂, holotypes (except 150, 152–153), clypeus and eye tubercle, dorsal views. 148. R. ornatula sp. nov. (SMNH). 149. R. pamira sp. nov. (SMNH). 150. R. pamira, paratype (SMNH). 151. R. dilutasp. nov. (SMNH). 152. R. diluta, paratype (SMNH).153. R. fedotovi(Charitonov, 1946), paralectotype (PSU). 154. R. pallens sp. nov. (SMNH). 155. R. zyuzini sp. nov. (SMNH). 156. R. ferghanensis (Zonstein, 1984) (ZISP). 157. R. kirgizica sp. nov. (SMNH).158. R. kopetdaghensis (Fet, 1984) (ZISP). 159. R. mikhailovi Zonstein, 2021 (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990868" httpUri="https://zenodo.org/record/13990868/files/figure.png" pageId="51" pageNumber="52">153</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FDD5E37CFDDACA2B" box="[570,616,1110,1136]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="51" pageNumber="52">183</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FD9BE37CFD11CA2B" box="[628,675,1110,1136]" captionStart="Figs 196–201" captionStartId="128.[189,241,1850,1876]" captionTargetBox="[189,1398,276,1794]" captionTargetId="figure-68@128.[189,1398,265,1816]" captionTargetPageId="128" captionText="Figs 196–201. Raveniola spp., ♂♂, chelicera, prolateral views. 196. R. redikorzevi (Spassky, 1937), ♂ from Kepele (SMNH). 197. R. cucullata sp. nov., paratype from Shikildara (SMNH). 198. R. ignobilis sp. nov., paratype (SMNH). 199. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 200. R. kopetdaghensis (Fet, 1984), paratype (SMNH). 201. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990876" httpUri="https://zenodo.org/record/13990876/files/figure.png" pageId="51" pageNumber="52">199</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FD40E37CFD6CCA2B" box="[687,734,1110,1136]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="51" pageNumber="52">217</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FD05E37CFCABCA2B" box="[746,793,1110,1136]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="51" pageNumber="52">244</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FCCBE37CFC3CCA2A" box="[804,910,1110,1137]" captionStart="Figs 274–281" captionStartId="137.[189,241,1743,1769]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure-78@137.[189,1398,265,1703]" captionTargetPageId="137" captionText="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990897" httpUri="https://zenodo.org/record/13990897/files/figure.png" pageId="51" pageNumber="52">274–275</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FC75E37CFC7BCA2B" box="[922,969,1110,1136]" captionStart="Figs 364–378" captionStartId="144.[189,241,1714,1740]" captionTargetBox="[189,1398,271,1690]" captionTargetId="figure-142@144.[189,1398,265,1690]" captionTargetPageId="144" captionText="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990915" httpUri="https://zenodo.org/record/13990915/files/figure.png" pageId="51" pageNumber="52">366</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FC3BE37CFB8CCA2B" box="[980,1086,1110,1136]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="51" pageNumber="52">432–433</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FBA5E37CFB01CA2A" box="[1098,1203,1110,1137]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="51" pageNumber="52">528–529</figureCitation>
, 
<figureCitation id="133A1AB05519FF89FB50E37CFA9ACA2A" box="[1215,1320,1110,1137]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="51" pageNumber="52">591–592</figureCitation>
, 707–710, 755
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5519FF89FF52E396FE80CBD8" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="8BBE06355519FF89FF52E396FC81CA8D" blockId="51.[189,819,1212,1238]" box="[189,819,1212,1238]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB11211B5519FF89FF52E396FC81CA8D" box="[189,819,1212,1238]" pageId="51" pageNumber="52">
<treatmentCitation id="0AA020245519FF89FF52E396FD00CA8D" author="Charitonov D. E." box="[189,690,1212,1238]" page="19" pageId="51" pageNumber="52" year="1946">
<taxonomicName id="4C017DB65519FF89FF52E396FD00CA8D" authority="Charitonov, 1946: 19" authorityName="Charitonov" authorityPageNumber="19" authorityYear="1946" box="[189,690,1212,1238]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA275519FF89FF52E396FE03CA8D" box="[189,433,1212,1238]" italics="true" pageId="51" pageNumber="52">Brachythele fedotovi</emphasis>
<bibRefCitation id="EF907BC45519FF89FE57E396FD00CA8D" author="Charitonov D. E." box="[440,690,1212,1238]" pageId="51" pageNumber="52" pagination="19 - 32" refId="ref77775" refString="Charitonov D. E. 1946. New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo 12: 19 - 32. [In Russian and Latin.]" type="journal article" year="1946">Charitonov, 1946: 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 2 (
<specimenCount id="9D07CDBC5519FF89FCE3E396FC96CA8D" box="[780,804,1212,1238]" count="1" pageId="51" pageNumber="52" type="male">♂</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBE06355519FF89FF52E3D4FB40CB42" blockId="51.[189,1399,1278,1411]" box="[189,1266,1278,1305]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB11211B5519FF89FF52E3D4FB40CB42" box="[189,1266,1278,1305]" pageId="51" pageNumber="52">
<taxonomicName id="4C017DB65519FF89FF52E3D4FE03CB43" authorityName="Charitonov" authorityYear="1946" box="[189,433,1278,1304]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA275519FF89FF52E3D4FE03CB43" box="[189,433,1278,1304]" italics="true" pageId="51" pageNumber="52">Brachythele fedotovi</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AA020245519FF89FE21E3D4FD76CB42" author="Charitonov D. E." box="[462,708,1278,1305]" page="65" pageId="51" pageNumber="52" year="1969">
<bibRefCitation id="EF907BC45519FF89FE21E3D4FD76CB42" author="Charitonov D. E." box="[462,708,1278,1305]" pageId="51" pageNumber="52" pagination="59 - 133" refId="ref77847" refString="Charitonov D. E. 1969. Materialy po faune paukov SSSR [Materials to the spider fauna of the USSR]. Uchenye Zapiski Permskogo Universiteta 179: 59 - 133. [In Russian.]" type="journal article" year="1969">Charitonov 1969: 65</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AA020245519FF89FD1BE3D4FC64CB43" author="Brignoli P. M." box="[756,982,1278,1304]" page="123" pageId="51" pageNumber="52" year="1983">
<bibRefCitation id="EF907BC45519FF89FD1BE3D4FC64CB43" author="Brignoli P. M." box="[756,982,1278,1304]" pageId="51" pageNumber="52" refId="ref77457" refString="Brignoli P. M. 1983. A Catalogue of Araneae Described between 1940 and 1981. Manchester University Press, Manchester." type="book" year="1983">Brignoli 1983: 123</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AA020245519FF89FBE8E3D4FB5CCB42" author="Zonstein S. L." box="[1031,1262,1278,1305]" page="158" pageId="51" pageNumber="52" year="1985">
<bibRefCitation id="EF907BC45519FF89FBE8E3D4FB5CCB42" author="Zonstein S. L." box="[1031,1262,1278,1305]" pageId="51" pageNumber="52" pagination="156 - 161" refId="ref83012" refString="Zonstein S. L. 1985. Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR. Trudy zoologicheskogo Instituta AN SSSR 139: 156 - 161. [In Russian with English summary.]" type="journal article" year="1985">Zonstein 1985: 158</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBE06355519FF89FF52E20BFE80CBD8" blockId="51.[189,1399,1278,1411]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB11211B5519FF89FF52E20BFE80CBD8" pageId="51" pageNumber="52">
<taxonomicName id="4C017DB65519FF89FF52E20BFE2FCB60" ID-CoL="4RMTR" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[189,413,1313,1339]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA275519FF89FF52E20BFE2FCB60" box="[189,413,1313,1339]" italics="true" pageId="51" pageNumber="52">Raveniola fedotovi</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AA020245519FF89FE2FE20BFD0FCB67" author="Zonstein S. L." box="[448,701,1313,1340]" page="1015" pageId="51" pageNumber="52" year="1987">
<bibRefCitation id="EF907BC45519FF89FE2FE20BFD0FCB67" author="Zonstein S. L." box="[448,701,1313,1340]" pageId="51" pageNumber="52" pagination="1013 - 1019" refId="ref83052" refString="Zonstein S. L. 1987. A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna. Zoologicheskii Zhurnal 66 (7): 1013 - 1019. [In Russian with English summary.]" type="journal article" year="1987">Zonstein 1987: 1015</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AA020245519FF89FD1CE20BFC7ECB67" author="Platnick N. I." box="[755,972,1313,1340]" page="90" pageId="51" pageNumber="52" year="1989">
<bibRefCitation id="EF907BC45519FF89FD1CE20BFC7ECB67" author="Platnick N. I." box="[755,972,1313,1340]" pageId="51" pageNumber="52" refId="ref81453" refString="Platnick N. I. 1989. Advances in Spider Taxonomy 1981 - 1987. A Supplement to Brignoli's &quot; A Catalogue of Araneae Described between 1940 and 1981 &quot;. New York / Manchester." type="book" year="1989">Platnick 1989: 90</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AA020245519FF89FBEDE20BFB47CB67" author="Mikhailov K. G." box="[1026,1269,1313,1340]" page="77" pageId="51" pageNumber="52" year="1996">
<bibRefCitation id="EF907BC45519FF89FBEDE20BFB47CB67" author="Mikhailov K. G." box="[1026,1269,1313,1340]" pageId="51" pageNumber="52" pagination="75 - 137" refId="ref80535" refString="Mikhailov K. G. 1996. A checklist of the spiders of Russia and other territories of the former USSR. Arthropoda Selecta 5 (1 - 2): 75 - 137." type="journal article" year="1996">Mikhailov 1996: 77</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AA020245519FF89FAEBE208FAC0CB67" author="Mikhailov K. G." box="[1284,1394,1314,1340]" page="20" pageId="51" pageNumber="52" year="1997">
<bibRefCitation id="EF907BC45519FF89FAEBE208FAC0CB67" author="Mikhailov K. G." box="[1284,1394,1314,1340]" pageId="51" pageNumber="52" refId="ref80570" refString="Mikhailov K. G. 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum of the Moscow State University." type="book" year="1997">1997: 20</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AA020245519FF89FF05E26FFEE5CB04" author="Mikhailov K. G." box="[234,343,1349,1375]" page="12" pageId="51" pageNumber="52" year="2013">
<bibRefCitation id="EF907BC45519FF89FF05E26FFEE5CB04" author="Mikhailov K. G." box="[234,343,1349,1375]" pageId="51" pageNumber="52" pagination="1 - 262" refId="ref80603" refString="Mikhailov K. G. 2013. The spiders (Arachnida: Aranei) of Russia and the adjacement countries: a nonannotated checklist. Arthropoda Selecta, Supplement 3: 1 - 262." type="journal article" year="2013">2013: 12</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AA020245519FF89FE63E26FFD0CCB04" author="Zonstein S." box="[396,702,1349,1376]" page="66" pageId="51" pageNumber="52" year="2018">
<bibRefCitation id="EF907BC45519FF89FE63E26FFD0CCB04" author="Zonstein S." box="[396,702,1349,1376]" pageId="51" pageNumber="52" pagination="1 - 100" refId="ref83568" refString="Zonstein S. 2018 b. A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae). European Journal of Taxonomy 485: 1 - 100. https: // doi. org / 10.5852 / ejt. 2018.485" type="journal article" year="2018">
Zonstein 
<emphasis id="B975DA275519FF89FE11E26CFD80CB04" box="[510,562,1349,1375]" italics="true" pageId="51" pageNumber="52">et al</emphasis>
. 2018b: 66
</bibRefCitation>
</treatmentCitation>
, fig. 100 (
<specimenCount id="9D07CDBC5519FF89FCD4E26FFCE1CB04" box="[827,851,1349,1375]" count="1" pageId="51" pageNumber="52" type="male">♂</specimenCount>
). — 
<treatmentCitation id="0AA020245519FF89FC7CE26FFB48CB04" author="Zonstein S. L. &amp; Esyunin S. L." box="[915,1274,1349,1375]" page="76" pageId="51" pageNumber="52" year="2023">
<bibRefCitation id="EF907BC45519FF89FC7CE26FFB48CB04" author="Zonstein S. L. &amp; Esyunin S. L." box="[915,1274,1349,1375]" pageId="51" pageNumber="52" pagination="75 - 79" refId="ref83697" refString="Zonstein S. L. &amp; Esyunin S. L. 2023. A redescription of Raveniola fedotovi (Charitonov, 1946), with first description of the female (Aranei: Nemesiidae). Arthropoda Selecta 32 (1): 75 - 79. https: // doi. org / 10.15298 / arthsel. 32.1.06" type="journal article" year="2023">Zonstein &amp; Esyunin 2023: 76</bibRefCitation>
</treatmentCitation>
, figs 1–14 (
<specimenCount id="9D07CDBC5519FF89FF1AE243FEBDCBD8" box="[245,271,1385,1411]" count="1" pageId="51" pageNumber="52" type="male">♂</specimenCount>
<specimenCount id="9D07CDBC5519FF89FEE0E243FE93CBD8" box="[271,289,1385,1411]" count="1" pageId="51" pageNumber="52" type="female">♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5519FF89FF52E281FBA5C890" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BBE06355519FF89FF52E281FE84CB9E" blockId="51.[189,310,1451,1477]" box="[189,310,1451,1477]" pageId="51" pageNumber="52">
<heading id="D0F6B1595519FF89FF52E281FE84CB9E" bold="true" box="[189,310,1451,1477]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B975DA275519FF89FF52E281FE84CB9E" bold="true" box="[189,310,1451,1477]" pageId="51" pageNumber="52">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355519FF89FF52E2F0FBA5C890" blockId="51.[188,1399,1498,1739]" pageId="51" pageNumber="52">
Males of 
<taxonomicName id="4C017DB65519FF89FEC5E2F0FDB5CBAF" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[298,519,1498,1524]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA275519FF89FEC5E2F0FDB5CBAF" box="[298,519,1498,1524]" italics="true" pageId="51" pageNumber="52">Raveniola fedotovi</emphasis>
</taxonomicName>
differ from other male congeners in having a characteristic triangular (very short, but high and acute) keel, demarcating proximal and distal sections of the embolus (vs either the presence of a lengthened keel in 
<taxonomicName id="4C017DB65519FF89FDD0E108FD16C860" authorityName="Zonstein" authorityYear="2024" box="[575,676,1569,1596]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="diluta" status="sp. nov.">
<emphasis id="B975DA275519FF89FDD0E108FDE3C860" box="[575,593,1570,1595]" italics="true" pageId="51" pageNumber="52">R</emphasis>
. 
<emphasis id="B975DA275519FF89FD8FE10BFD16C860" box="[608,676,1569,1595]" italics="true" pageId="51" pageNumber="52">diluta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5519FF89FD43E108FCBBC867" box="[684,777,1570,1596]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
, or the absence of a raised keel in 
<taxonomicName id="4C017DB65519FF89FB70E108FAA7C860" authorityName="Zonstein" authorityYear="2024" box="[1183,1301,1569,1596]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pallens" status="sp. nov.">
<emphasis id="B975DA275519FF89FB70E108FB03C860" box="[1183,1201,1570,1595]" italics="true" pageId="51" pageNumber="52">R</emphasis>
. 
<emphasis id="B975DA275519FF89FB2FE10BFAA7C860" box="[1216,1301,1569,1595]" italics="true" pageId="51" pageNumber="52">pallens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5519FF89FAF3E108FAC5C867" box="[1308,1399,1570,1596]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C017DB65519FF89FF01E16CFEEDC804" authorityName="Zonstein" authorityYear="2024" box="[238,351,1606,1632]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="B975DA275519FF89FF01E16CFEB2C804" box="[238,256,1606,1631]" italics="true" pageId="51" pageNumber="52">R</emphasis>
. 
<emphasis id="B975DA275519FF89FEE1E16CFEEDC804" box="[270,351,1606,1631]" italics="true" pageId="51" pageNumber="52">zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5519FF89FE88E16CFE76C83B" box="[359,452,1606,1632]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
; 
<figureCitation id="133A1AB05519FF89FE20E16FFDC0C804" box="[463,626,1605,1631]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="51" pageNumber="52">Figs 432–433</figureCitation>
, cf. 
<figureCitation id="133A1AB05519FF89FD4DE16FFC0FC804" box="[674,957,1605,1631]" captionStart-0="Figs 418–428" captionStart-1="Figs 429–438" captionStart-2="Figs 319–333" captionStart-3="Figs 334–348" captionStart-4="Figs 349–363" captionStart-5="Figs 364–378" captionStart-6="Figs 379–388" captionStart-7="Figs 389–399" captionStart-8="Figs 400–408" captionStart-9="Figs 409–417" captionStartId-0="149.[189,241,1845,1871]" captionStartId-1="150.[189,241,1855,1881]" captionStartId-2="141.[189,241,1712,1738]" captionStartId-3="142.[189,241,1720,1746]" captionStartId-4="143.[189,241,1727,1753]" captionStartId-5="144.[189,241,1714,1740]" captionStartId-6="145.[189,241,1844,1870]" captionStartId-7="146.[189,241,1854,1880]" captionStartId-8="147.[189,241,1853,1879]" captionStartId-9="148.[189,241,1847,1873]" captionTargetBox-0="[189,1398,265,1794]" captionTargetBox-1="[189,1398,265,1818]" captionTargetBox-2="[189,1398,265,1676]" captionTargetBox-3="[189,1398,265,1679]" captionTargetBox-4="[189,1390,270,1690]" captionTargetBox-5="[189,1398,271,1690]" captionTargetBox-6="[189,1398,265,1806]" captionTargetBox-7="[189,1398,265,1814]" captionTargetBox-8="[198,1398,265,1817]" captionTargetBox-9="[189,1398,265,1801]" captionTargetId-0="figure-44@149.[189,1398,265,1794]" captionTargetId-1="figure-75@150.[189,1398,265,1818]" captionTargetId-2="figure-93@141.[189,1398,265,1676]" captionTargetId-3="figure-88@142.[189,1398,265,1679]" captionTargetId-4="figure-139@143.[189,1398,265,1690]" captionTargetId-5="figure-142@144.[189,1398,265,1690]" captionTargetId-6="figure-75@145.[189,1398,265,1806]" captionTargetId-7="figure-68@146.[189,1398,265,1814]" captionTargetId-8="figure-50@147.[189,1398,265,1817]" captionTargetId-9="figure-49@148.[189,1398,265,1801]" captionTargetPageId-0="149" captionTargetPageId-1="150" captionTargetPageId-2="141" captionTargetPageId-3="142" captionTargetPageId-4="143" captionTargetPageId-5="144" captionTargetPageId-6="145" captionTargetPageId-7="146" captionTargetPageId-8="147" captionTargetPageId-9="148" captionText-0="Figs 418–428. Raveniola pamira sp. nov, ♂♂, holotype (418–422) and two paratypes (423–426 and 427–428, respectively), details of copulatoty bulb; retrolateral (418, 427), retroventral (419, 428), ventral (420, 423, 426), proventral (421, 425) and prolateral (422, 424) views (all SMNH). Scale bars: 0.5 mm." captionText-1="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." captionText-2="Figs 319–333. Raveniola spp., ♂ (319–324) and ♀♀ (325–333); apical edge of maxilla (319); tarsus I, tarsal organ, (320, 325, 331); same, zigzag row of trichobothria (321, 326, dorsal and retrodorsal views, respectively); same, bases of trichobotria (322–323, 327, 332); tarsal claws of leg IV (324) and leg I (333); entire chelicera (328) and its probasal corner (329–330); ventroapical (319), dorsal (320–323, 325, 327, 331–332), retrolateral (324, 333) and proventro-caudal (328–330) views. 319–324. R. redikorzevi (Spassky, 1937) (SMNH). 325–327. R. alajensis sp. nov. (SMNH). 328–333. R. cucullata sp. nov. (SMNH)." captionText-3="Figs 334–348. Raveniola spp., ♂♂ (334, 337–348) and ♀♀ (335–336); chelicera, probasal corner (334) and entire structure (343); apical edge of maxilla (335, 344); tarsus I (336); same, tarsal organ (337, 340–341, 345–346); same, bases of trichobotria (338–339, 342, 347); same, claws (348); proventrocaudal (334, 343), ventroapical (335, 344), dorsal (336–342, 345–347) and anterior (348) views. 334– 336. R. pamira sp. nov. (SMNH). 337–339. R. zyuzini sp. nov. (SMNH). 340–342. R. ferghanensis (Zonstein, 1984) (SMNH). 343–348. R. virgata (Simon, 1891) (SMNH)." captionText-4="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." captionText-5="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." captionText-6="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." captionText-7="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." captionText-8="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." captionText-9="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990925" figureDoi-1="http://doi.org/10.5281/zenodo.13990927" figureDoi-2="http://doi.org/10.5281/zenodo.13990908" figureDoi-3="http://doi.org/10.5281/zenodo.13990911" figureDoi-4="http://doi.org/10.5281/zenodo.13990913" figureDoi-5="http://doi.org/10.5281/zenodo.13990915" figureDoi-6="http://doi.org/10.5281/zenodo.13990917" figureDoi-7="http://doi.org/10.5281/zenodo.13990919" figureDoi-8="http://doi.org/10.5281/zenodo.13990921" figureDoi-9="http://doi.org/10.5281/zenodo.13990923" httpUri-0="https://zenodo.org/record/13990925/files/figure.png" httpUri-1="https://zenodo.org/record/13990927/files/figure.png" httpUri-2="https://zenodo.org/record/13990908/files/figure.png" httpUri-3="https://zenodo.org/record/13990911/files/figure.png" httpUri-4="https://zenodo.org/record/13990913/files/figure.png" httpUri-5="https://zenodo.org/record/13990915/files/figure.png" httpUri-6="https://zenodo.org/record/13990917/files/figure.png" httpUri-7="https://zenodo.org/record/13990919/files/figure.png" httpUri-8="https://zenodo.org/record/13990921/files/figure.png" httpUri-9="https://zenodo.org/record/13990923/files/figure.png" pageId="51" pageNumber="52">Figs 428–431, 324–438</figureCitation>
). The sole adult female of 
<taxonomicName id="4C017DB65519FF89FB1FE16CFADDC804" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[1264,1391,1605,1632]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA275519FF89FB1FE16CFAB0C804" box="[1264,1282,1606,1631]" italics="true" pageId="51" pageNumber="52">R</emphasis>
. 
<emphasis id="B975DA275519FF89FAFEE16FFADDC804" box="[1297,1391,1605,1631]" italics="true" pageId="51" pageNumber="52">fedotovi</emphasis>
</taxonomicName>
, known to date, is distinguishable owing to a specific structure of the spermathecal trunks, which are considerably longer, and relatively more closely spaced to each other than in females of other species representing the same group (
<figureCitation id="133A1AB05519FF89FDF6E19AFD0CC890" box="[537,702,1712,1739]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="51" pageNumber="52">Figs 528–529</figureCitation>
, cf. 
<figureCitation id="133A1AB05519FF89FD1FE19AFBB9C890" box="[752,1035,1712,1739]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="51" pageNumber="52">Figs 526–527, 530–533</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5519FF8EFF52E1D8FEA8CFAB" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BBE06355519FF89FF52E1D8FE18C957" blockId="51.[189,426,1778,1804]" box="[189,426,1778,1804]" pageId="51" pageNumber="52">
<heading id="D0F6B1595519FF89FF52E1D8FE18C957" bold="true" box="[189,426,1778,1804]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B975DA275519FF89FF52E1D8FE18C957" bold="true" box="[189,426,1778,1804]" pageId="51" pageNumber="52">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355519FF89FF05E00BFED4C960" blockId="51.[189,1399,1825,1923]" box="[234,358,1825,1851]" pageId="51" pageNumber="52">
<emphasis id="B975DA275519FF89FF05E00BFED4C960" bold="true" box="[234,358,1825,1851]" pageId="51" pageNumber="52">
<typeStatus id="54BAB8975519FF89FF05E00BFED4C960" box="[234,358,1825,1851]" pageId="51" pageNumber="52" type="lectotype">Lectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBE06355519FF89FF52E06FFBF6C9D8" blockId="51.[189,1399,1825,1923]" pageId="51" pageNumber="52">
<materialsCitation id="3B690C685519FF89FF52E06FFBF6C9D8" collectingDate="1942-03-25" collectingDateMax="1942-03-28" collectingDateMin="1942-03-25" collectionCode="ZMMU" collectorName="D. M. Fedotov" country="Uzbekistan" elevation="1200" latitude="38.816666" location="Hissar Mts" longLatPrecision="1173" longitude="66.96667" municipality="Ishkent Village" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F31646A55519FF89FF52E06FFEC1C904" box="[189,371,1861,1887]" name="Uzbekistan" pageId="51" pageNumber="52">UZBEKISTAN</collectingCountry>
• 
<specimenCount id="9D07CDBC5519FF89FE77E06FFE03C904" box="[408,433,1861,1887]" count="1" pageId="51" pageNumber="52" type="male">♂</specimenCount>
; 
<location id="8EDE50EE5519FF89FE2BE06FFDF9C904" LSID="urn:lsid:plazi:treatment:03A8B7235519FF8FFDD9E318FD84C9B2:8EDE50EE5519FF89FE2BE06FFDF9C904" box="[452,587,1861,1887]" country="Uzbekistan" latitude="38.816666" longLatPrecision="1173" longitude="66.96667" municipality="Ishkent Village" name="Hissar Mts" pageId="51" pageNumber="52">Hissar Mts</location>
(northwestern slope), surroundings of 
<collectingMunicipality id="6BDA9C4F5519FF89FBDDE06FFB5FC904" box="[1074,1261,1861,1887]" pageId="51" pageNumber="52">Ishkent Village</collectingMunicipality>
; 
<geoCoordinate id="EE3560F25519FF89FAEFE069FAC0C904" box="[1280,1394,1859,1887]" degrees="38" direction="north" minutes="49" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="38.816666">38°49′ N</geoCoordinate>
, 
<geoCoordinate id="EE3560F25519FF89FF52E04DFE94C9D8" box="[189,294,1895,1923]" degrees="66" direction="east" minutes="58" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="66.96667">66°58′ E</geoCoordinate>
; 
<elevation id="002CE1065519FF89FEDCE043FDA6C9D8" box="[307,532,1897,1923]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="51" pageNumber="52" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">
<quantity id="4CF9ABD05519FF89FEDCE043FE66C9D8" box="[307,468,1897,1923]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="51" pageNumber="52" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">1100–1300 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F55519FF89FDCEE042FD5CC9D8" box="[545,750,1896,1923]" pageId="51" pageNumber="52" value="1942-03-25" valueMax="1942-03-28" valueMin="1942-03-25">
<collectingDate id="EFFBD91D5519FF89FDCEE042FD5CC9D8" box="[545,750,1896,1923]" pageId="51" pageNumber="52" value="1942-03-25" valueMax="1942-03-28" valueMin="1942-03-25">25–28 Mar. 1942</collectingDate>
</date>
; 
<collectorName id="26F463E35519FF89FD15E043FC2DC9D8" box="[762,927,1897,1923]" pageId="51" pageNumber="52">D.M. Fedotov</collectorName>
leg.; 
<collectionCode id="ED109EF05519FF89FC0EE043FBF2C9D8" box="[993,1088,1897,1923]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="51" pageNumber="52" type="Museum">ZMMU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBE06355519FF89FF05E081FE10C99E" blockId="51.[189,1288,1963,2025]" box="[234,418,1963,1989]" pageId="51" pageNumber="52">
<emphasis id="B975DA275519FF89FF05E081FE10C99E" bold="true" box="[234,418,1963,1989]" pageId="51" pageNumber="52">
<typeStatus id="54BAB8975519FF89FF05E081FE10C99E" box="[234,418,1963,1989]" pageId="51" pageNumber="52" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBE06355519FF89FF52E0E4FABAC9B3" blockId="51.[189,1288,1963,2025]" box="[189,1288,1998,2025]" pageId="51" pageNumber="52">
<materialsCitation id="3B690C685519FF89FF52E0E4FAB6C9B3" box="[189,1284,1998,2025]" collectionCode="PSU" country="Uzbekistan" pageId="51" pageNumber="52" specimenCount="3" specimenCount-male="3" typeStatus="paralectotype">
<collectingCountry id="F31646A55519FF89FF52E0E4FEC1C9B3" box="[189,371,1998,2024]" name="Uzbekistan" pageId="51" pageNumber="52">UZBEKISTAN</collectingCountry>
• 
<specimenCount id="9D07CDBC5519FF89FE63E0E4FE63C9B2" box="[396,465,1998,2025]" count="3" pageId="51" pageNumber="52" type="male">3 ♂♂</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED109EF05519FF89FC7CE0E4FC7EC9B3" box="[915,972,1998,2024]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="51" pageNumber="52" type="Museum">PSU</collectionCode>
, deposited as preparations
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF05E623FD0ACF78" blockId="52.[188,1399,265,496]" box="[234,696,265,291]" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFF05E623FE53CF78" bold="true" box="[234,481,265,291]" pageId="52" pageNumber="53">Additional material</emphasis>
(
<specimenCount id="9D07CDBC551EFF8EFE1BE623FD92CF78" box="[500,544,265,291]" count="1" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D07CDBC551EFF8EFDC2E623FDEBCF78" box="[557,601,265,291]" count="1" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
, 2 juvs)
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E601FEA8CFAB" blockId="52.[188,1399,265,496]" pageId="52" pageNumber="53">
<materialsCitation id="3B690C68551EFF8EFF52E601FBEACF3C" collectingDate="1989-04-08" collectionCode="SMNH" collectorName="S. Zonstein" country="Uzbekistan" elevation="1350" latitude="37.966667" location="Kugitang (Koitendagh) Mts" longLatPrecision="1178" longitude="66.71667" municipality="Baglydara Canyon" pageId="52" pageNumber="53" specimenCount="1">
<collectingCountry id="F31646A5551EFF8EFF52E601FEC1CF1E" box="[189,371,299,325]" name="Uzbekistan" pageId="52" pageNumber="53">UZBEKISTAN</collectingCountry>
• 
<specimenCount id="9D07CDBC551EFF8EFE65E601FE7CCF1E" box="[394,462,299,325]" count="1" pageId="52" pageNumber="53" type="generic">1 juv.</specimenCount>
; 
<location id="8EDE50EE551EFF8EFE36E601FCADCF1D" LSID="urn:lsid:plazi:treatment:03A8B7235519FF8FFDD9E318FD84C9B2:8EDE50EE551EFF8EFE36E601FCADCF1D" box="[473,799,299,326]" country="Uzbekistan" latitude="37.966667" longLatPrecision="1178" longitude="66.71667" municipality="Baglydara Canyon" name="Kugitang (Koitendagh) Mts" pageId="52" pageNumber="53">Kugitang (Koitendagh) Mts</location>
, 
<collectingMunicipality id="6BDA9C4F551EFF8EFCC4E601FBBACF1E" box="[811,1032,299,325]" pageId="52" pageNumber="53">Baglydara Canyon</collectingMunicipality>
, 
<locationDeviation id="9A0C1F17551EFF8EFBFCE601FAC0CF1E" box="[1043,1394,299,326]" location="Hatak Village" pageId="52" pageNumber="53">
<quantity id="4CF9ABD0551EFF8EFBFCE601FB35CF1E" box="[1043,1159,299,325]" metricMagnitude="4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="52" pageNumber="53" unit="km" value="10.5" valueMax="11.0" valueMin="10.0">10–11 km</quantity>
W of Hatak Village
</locationDeviation>
; 
<geoCoordinate id="EE3560F2551EFF8EFF53E666FE9BCF33" box="[188,297,332,360]" degrees="37" direction="north" minutes="58" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="37.966667">37°58′ N</geoCoordinate>
, 
<geoCoordinate id="EE3560F2551EFF8EFED9E666FE12CF33" box="[310,416,332,360]" degrees="66" direction="east" minutes="43" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="66.71667">66°43′ E</geoCoordinate>
; 
<elevation id="002CE106551EFF8EFE42E667FD23CF3C" box="[429,657,333,360]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="52" pageNumber="53" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
<quantity id="4CF9ABD0551EFF8EFE42E667FDE2CF33" box="[429,592,333,360]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="52" pageNumber="53" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F5551EFF8EFD70E667FC9FCF3C" box="[671,813,333,359]" pageId="52" pageNumber="53" value="1989-04-08">
<collectingDate id="EFFBD91D551EFF8EFD70E667FC9FCF3C" box="[671,813,333,359]" pageId="52" pageNumber="53" value="1989-04-08">8 Apr. 1989</collectingDate>
</date>
; 
<collectorName id="26F463E3551EFF8EFCD5E667FC72CF3C" box="[826,960,333,359]" pageId="52" pageNumber="53">S. Zonstein</collectorName>
leg.; 
<collectionCode id="ED109EF0551EFF8EFBECE667FBEACF3C" box="[1027,1112,333,359]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="52" pageNumber="53">SMNH</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B690C68551EFF8EFB9DE667FB1FCFD1" collectingDate="1987-04-17" collectionCode="SMNH" collectorName="S. Zonstein" country="Uzbekistan" elevation="1500" latitude="38.25" location="Baisuntau Mts" longLatPrecision="1176" longitude="66.833336" municipality="Akrabat Pass" pageId="52" pageNumber="53" specimenCount="1">
<specimenCount id="9D07CDBC551EFF8EFB9DE667FB0ACF3C" box="[1138,1208,333,359]" count="1" pageId="52" pageNumber="53" type="generic">1 juv.</specimenCount>
; 
<location id="8EDE50EE551EFF8EFB2BE667FAC3CF33" LSID="urn:lsid:plazi:treatment:03A8B7235519FF8FFDD9E318FD84C9B2:8EDE50EE551EFF8EFB2BE667FAC3CF33" box="[1220,1393,333,360]" country="Uzbekistan" latitude="38.25" longLatPrecision="1176" longitude="66.833336" municipality="Akrabat Pass" name="Baisuntau Mts" pageId="52" pageNumber="53">Baisuntau Mts</location>
, 
<collectingMunicipality id="6BDA9C4F551EFF8EFF52E645FEE8CFD1" box="[189,346,367,394]" pageId="52" pageNumber="53">Akrabat Pass</collectingMunicipality>
; 
<geoCoordinate id="EE3560F2551EFF8EFE8BE644FE7DCFD1" box="[356,463,366,394]" degrees="38" direction="north" minutes="15" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="38.25">38°15′ N</geoCoordinate>
, 
<geoCoordinate id="EE3560F2551EFF8EFE35E644FDF0CFD1" box="[474,578,366,394]" degrees="66" direction="east" minutes="50" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="66.833336">66°50′ E</geoCoordinate>
; 
<elevation id="002CE106551EFF8EFDA2E645FD51CFD2" box="[589,739,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="52" pageNumber="53" unit="m" value="1500.0">
<quantity id="4CF9ABD0551EFF8EFDA2E645FD17CFD1" box="[589,677,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="52" pageNumber="53" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F5551EFF8EFD1FE65AFC3BCFD1" box="[752,905,368,394]" pageId="52" pageNumber="53" value="1987-04-17">
<collectingDate id="EFFBD91D551EFF8EFD1FE65AFC3BCFD1" box="[752,905,368,394]" pageId="52" pageNumber="53" value="1987-04-17">17 Apr. 1987</collectingDate>
</date>
; 
<collectorName id="26F463E3551EFF8EFC7BE65AFBABCFD2" box="[916,1049,367,394]" pageId="52" pageNumber="53">S. Zonstein</collectorName>
leg.; 
<collectionCode id="ED109EF0551EFF8EFBB7E65AFB1FCFD1" box="[1112,1197,368,394]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="52" pageNumber="53">SMNH</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B690C68551EFF8EFB2CE65AFE50CF95" collectingDate="1992-04-29" collectionCode="SMNH" collectorName="S. Zonstein" country="Uzbekistan" county="Hojakurgan Gorge" elevation="1500" latitude="39.183334" location="Zeravshan Mts" longLatPrecision="1170" longitude="67.28333" municipality="Jindy-Daria Canyon" pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D07CDBC551EFF8EFB2CE65AFB43CFD1" box="[1219,1265,368,394]" count="1" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
; 
<location id="8EDE50EE551EFF8EFB13E645FF5FCFF7" LSID="urn:lsid:plazi:treatment:03A8B7235519FF8FFDD9E318FD84C9B2:8EDE50EE551EFF8EFB13E645FF5FCFF7" country="Uzbekistan" county="Hojakurgan Gorge" latitude="39.183334" longLatPrecision="1170" longitude="67.28333" municipality="Jindy-Daria Canyon" name="Zeravshan Mts" pageId="52" pageNumber="53">Zeravshan Mts</location>
, 
<collectingMunicipality id="6BDA9C4F551EFF8EFF14E6BBFE5FCFF7" box="[251,493,401,428]" pageId="52" pageNumber="53">Jindy-Daria Canyon</collectingMunicipality>
, 
<collectingCounty id="62DF7EB9551EFF8EFE14E6BBFD69CFF7" box="[507,731,401,428]" pageId="52" pageNumber="53">Hojakurgan Gorge</collectingCounty>
; 
<geoCoordinate id="EE3560F2551EFF8EFD07E6BAFCE7CFF7" box="[744,853,400,428]" degrees="39" direction="north" minutes="11" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="39.183334">39°11′ N</geoCoordinate>
, 
<geoCoordinate id="EE3560F2551EFF8EFC8CE6BAFC7CCFF7" box="[867,974,400,428]" degrees="67" direction="east" minutes="17" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="67.28333">67°17′ E</geoCoordinate>
; 
<elevation id="002CE106551EFF8EFC32E6BBFB71CFF0" box="[989,1219,401,428]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="52" pageNumber="53" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">
<quantity id="4CF9ABD0551EFF8EFC32E6BBFB33CFF7" box="[989,1153,401,428]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="52" pageNumber="53" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">1400–1600 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F5551EFF8EFB3DE6B8FAC0CFF7" box="[1234,1394,402,428]" pageId="52" pageNumber="53" value="1992-04-29">
<collectingDate id="EFFBD91D551EFF8EFB3DE6B8FAC0CFF7" box="[1234,1394,402,428]" pageId="52" pageNumber="53" value="1992-04-29">29 Apr. 1992</collectingDate>
</date>
; 
<collectorName id="26F463E3551EFF8EFF52E69EFEF7CF95" box="[189,325,436,462]" pageId="52" pageNumber="53">S. Zonstein</collectorName>
leg.; 
<collectionCode id="ED109EF0551EFF8EFE62E69EFE50CF95" box="[397,482,436,462]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="52" pageNumber="53">SMNH</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B690C68551EFF8EFE12E69EFEA4CFAB" collectingDate="2022-05-04" collectionCode="SMNH" collectorName="S. Zonstein" country="Uzbekistan" county="Hojakurgan Gorge" elevation="1500" latitude="39.183334" location="Zeravshan Mts" longLatPrecision="1170" longitude="67.28333" municipality="Jindy-Daria Canyon" pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D07CDBC551EFF8EFE12E69EFD9FCF95" box="[509,557,436,462]" count="1" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
; same collection data as for preceding; 
<date id="FFBF20F5551EFF8EFBE1E69EFB11CF95" box="[1038,1187,436,462]" pageId="52" pageNumber="53" value="2022-05-04">
<collectingDate id="EFFBD91D551EFF8EFBE1E69EFB11CF95" box="[1038,1187,436,462]" pageId="52" pageNumber="53" value="2022-05-04">4 May 2022</collectingDate>
</date>
; 
<collectorName id="26F463E3551EFF8EFB5DE69EFA88CF95" box="[1202,1338,436,462]" pageId="52" pageNumber="53">S. Zonstein</collectorName>
leg.; 
<collectionCode id="ED109EF0551EFF8EFF52E6FCFEA4CFAB" box="[189,278,470,496]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="52" pageNumber="53">SMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE551EFF8FFF52E53CFDD6C8CF" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="8BBE0635551EFF8EFF52E53CFEFCCC6B" blockId="52.[189,334,534,560]" box="[189,334,534,560]" pageId="52" pageNumber="53">
<heading id="D0F6B159551EFF8EFF52E53CFEFCCC6B" bold="true" box="[189,334,534,560]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="B975DA27551EFF8EFF52E53CFEFCCC6B" bold="true" box="[189,334,534,560]" pageId="52" pageNumber="53">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551EFF8EFF05E56EFE02CC05" blockId="52.[189,1398,580,674]" box="[234,432,580,606]" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFF05E56EFE98CC05" bold="true" box="[234,298,580,606]" pageId="52" pageNumber="53">Male</emphasis>
(
<typeStatus id="54BAB897551EFF8EFED6E56EFE1ACC05" box="[313,424,580,606]" pageId="52" pageNumber="53" type="lectotype">lectotype</typeStatus>
)
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E54CFE97CCF9" blockId="52.[189,1398,580,674]" pageId="52" pageNumber="53">
<figureCitation id="133A1AB0551EFF8EFF52E54CFE8ECCDB" box="[189,316,614,640]" captionStart="Figs 10–18" captionStartId="108.[189,241,1826,1852]" captionTargetBox="[189,1398,265,1785]" captionTargetId="figure-83@108.[189,1398,265,1785]" captionTargetPageId="108" captionText="Figs 10–18. Raveniola spp., ♂♂, holotypes (except 14, 16), habitus, dorsal views. 10. R. insolita sp. nov. (ZISP). 11. R. ornata sp. nov. (SMNH). 12. R. ornatula sp. nov. (SMNH). 13. R. pamira sp. nov. (SMNH). 14. R. pamira sp. nov., paratype (SMNH). 15. R. diluta sp. nov. (SMNH). 16. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 17. R. pallens sp. nov. (SMNH). 18. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990826" httpUri="https://zenodo.org/record/13990826/files/figure.png" pageId="52" pageNumber="53">Figures 16</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFEA8E54CFEC7CCDA" box="[327,373,614,641]" captionStart="Figs 148–159" captionStartId="124.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1691]" captionTargetId="figure-104@124.[189,1399,265,1691]" captionTargetPageId="124" captionText="Figs 148–159. Raveniola spp., ♂♂, holotypes (except 150, 152–153), clypeus and eye tubercle, dorsal views. 148. R. ornatula sp. nov. (SMNH). 149. R. pamira sp. nov. (SMNH). 150. R. pamira, paratype (SMNH). 151. R. dilutasp. nov. (SMNH). 152. R. diluta, paratype (SMNH).153. R. fedotovi(Charitonov, 1946), paralectotype (PSU). 154. R. pallens sp. nov. (SMNH). 155. R. zyuzini sp. nov. (SMNH). 156. R. ferghanensis (Zonstein, 1984) (ZISP). 157. R. kirgizica sp. nov. (SMNH).158. R. kopetdaghensis (Fet, 1984) (ZISP). 159. R. mikhailovi Zonstein, 2021 (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990868" httpUri="https://zenodo.org/record/13990868/files/figure.png" pageId="52" pageNumber="53">153</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFE6DE54CFE03CCDB" box="[386,433,614,640]" captionStart="Figs 196–201" captionStartId="128.[189,241,1850,1876]" captionTargetBox="[189,1398,276,1794]" captionTargetId="figure-68@128.[189,1398,265,1816]" captionTargetPageId="128" captionText="Figs 196–201. Raveniola spp., ♂♂, chelicera, prolateral views. 196. R. redikorzevi (Spassky, 1937), ♂ from Kepele (SMNH). 197. R. cucullata sp. nov., paratype from Shikildara (SMNH). 198. R. ignobilis sp. nov., paratype (SMNH). 199. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 200. R. kopetdaghensis (Fet, 1984), paratype (SMNH). 201. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990876" httpUri="https://zenodo.org/record/13990876/files/figure.png" pageId="52" pageNumber="53">199</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFE53E54CFE59CCDB" box="[444,491,614,640]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="52" pageNumber="53">217</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFE19E54CFDEDCCDA" box="[502,607,614,641]" captionStart="Figs 274–281" captionStartId="137.[189,241,1743,1769]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure-78@137.[189,1398,265,1703]" captionTargetPageId="137" captionText="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990897" httpUri="https://zenodo.org/record/13990897/files/figure.png" pageId="52" pageNumber="53">274–275</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFD84E54CFD28CCDB" box="[619,666,614,640]" captionStart="Figs 364–378" captionStartId="144.[189,241,1714,1740]" captionTargetBox="[189,1398,271,1690]" captionTargetId="figure-142@144.[189,1398,265,1690]" captionTargetPageId="144" captionText="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990915" httpUri="https://zenodo.org/record/13990915/files/figure.png" pageId="52" pageNumber="53">366</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFD4AE54CFCBDCCDB" box="[677,783,614,640]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="52" pageNumber="53">432–433</figureCitation>
, 
<figureCitation id="133A1AB0551EFF8EFCF5E54CFCF4CCDA" box="[794,838,614,641]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="52" pageNumber="53">591</figureCitation>
are based on the 
<typeStatus id="54BAB897551EFF8EFBFEE54CFB08CCDB" box="[1041,1210,614,640]" pageId="52" pageNumber="53" type="paralectotype">paralectotypes</typeStatus>
and conspecific material.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E5E3FE05CCB9" blockId="52.[189,439,712,739]" box="[189,439,712,739]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E5E3FEAECCBA" baselines="732,732" box="[189,284,713,739]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="52" pageNumber="53">HABITUS</smallCapsWord>
. See 
<figureCitation id="133A1AB0551EFF8EFEB2E5E2FE05CCB9" box="[349,439,712,738]" captionStart="Figs 10–18" captionStartId="108.[189,241,1826,1852]" captionTargetBox="[189,1398,265,1785]" captionTargetId="figure-83@108.[189,1398,265,1785]" captionTargetPageId="108" captionText="Figs 10–18. Raveniola spp., ♂♂, holotypes (except 14, 16), habitus, dorsal views. 10. R. insolita sp. nov. (ZISP). 11. R. ornata sp. nov. (SMNH). 12. R. ornatula sp. nov. (SMNH). 13. R. pamira sp. nov. (SMNH). 14. R. pamira sp. nov., paratype (SMNH). 15. R. diluta sp. nov. (SMNH). 16. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 17. R. pallens sp. nov. (SMNH). 18. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990826" httpUri="https://zenodo.org/record/13990826/files/figure.png" pageId="52" pageNumber="53">Fig. 16.</figureCitation>
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E423FB2ACD7F" blockId="52.[189,1176,777,804]" box="[189,1176,777,804]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E423FEDECD7A" baselines="796,796" box="[189,364,777,803]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="52" pageNumber="53">MEASUREMENTS</smallCapsWord>
. TBL 6.90, CL 3.38, CW 2.85, LL 0.28, LW 0.60, SL 1.78, SW 1.51.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E463FED5CDF3" blockId="52.[189,1399,841,936]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E463FEAACD39" baselines="860,861" box="[189,280,841,867]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="52" pageNumber="53">COLOUR</smallCapsWord>
. Carapace, palps and legs light brownish orange; eye tubercle blackish brown; chelicerae light reddish brown; sternum, labium and maxillae yellow; abdomen uniformly greyish white without clear dorsal pattern.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E4E4FD9ACAC9" blockId="52.[187,1399,974,1171]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E4E4FECACDBD" baselines="993,993" box="[189,376,974,1000]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Cephalothorax" pageId="52" pageNumber="53">CEPHALOTHORAX</smallCapsWord>
. Carapace and chelicerae as shown in 
<figureCitation id="133A1AB0551EFF8EFCDDE4E4FC35CDB3" box="[818,903,974,1000]" captionStart="Figs 91–99" captionStartId="118.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-80@118.[189,1398,265,1816]" captionTargetPageId="118" captionText="Figs 91–99. Raveniola spp., ♂♂, holotypes (except 96, 98), cephalothorax, dorsal views. 91. R. ignobilis sp. nov. (SMNH). 92. R. insolita sp. nov. (ZISP). 93. R. ornata sp. nov. (SMNH). 94. R. ornatula sp. nov. (SMNH). 95. R. pamira sp. nov. (SMNH). 96. R. pamira, paratype (SMNH). 97. R. diluta sp. nov. (SMNH). 98. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 99. R. pallens sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990852" httpUri="https://zenodo.org/record/13990852/files/figure.png" pageId="52" pageNumber="53">Fig. 98</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="133A1AB0551EFF8EFB34E4E4FA8CCDB2" box="[1243,1342,974,1001]" captionStart="Figs 148–159" captionStartId="124.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1691]" captionTargetId="figure-104@124.[189,1399,265,1691]" captionTargetPageId="124" captionText="Figs 148–159. Raveniola spp., ♂♂, holotypes (except 150, 152–153), clypeus and eye tubercle, dorsal views. 148. R. ornatula sp. nov. (SMNH). 149. R. pamira sp. nov. (SMNH). 150. R. pamira, paratype (SMNH). 151. R. dilutasp. nov. (SMNH). 152. R. diluta, paratype (SMNH).153. R. fedotovi(Charitonov, 1946), paralectotype (PSU). 154. R. pallens sp. nov. (SMNH). 155. R. zyuzini sp. nov. (SMNH). 156. R. ferghanensis (Zonstein, 1984) (ZISP). 157. R. kirgizica sp. nov. (SMNH).158. R. kopetdaghensis (Fet, 1984) (ZISP). 159. R. mikhailovi Zonstein, 2021 (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990868" httpUri="https://zenodo.org/record/13990868/files/figure.png" pageId="52" pageNumber="53">Fig. 153</figureCitation>
. Eye diameters and interdistances: AME 0.08(0.12), ALE 0.13, PLE 0.09, PME 0.06; AME–AME 0.08(0.05), ALE–AME 0.05(0.04), ALE–PLE 0.02, PLE–PME 0.02, PME–PME 0.16. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 7 promarginal teeth and 4–5 mesobasal denticles. MIT indiscernible (
<figureCitation id="133A1AB0551EFF8EFD76E37CFCB2CA2A" box="[665,768,1110,1137]" captionStart="Figs 196–201" captionStartId="128.[189,241,1850,1876]" captionTargetBox="[189,1398,276,1794]" captionTargetId="figure-68@128.[189,1398,265,1816]" captionTargetPageId="128" captionText="Figs 196–201. Raveniola spp., ♂♂, chelicera, prolateral views. 196. R. redikorzevi (Spassky, 1937), ♂ from Kepele (SMNH). 197. R. cucullata sp. nov., paratype from Shikildara (SMNH). 198. R. ignobilis sp. nov., paratype (SMNH). 199. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 200. R. kopetdaghensis (Fet, 1984), paratype (SMNH). 201. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990876" httpUri="https://zenodo.org/record/13990876/files/figure.png" pageId="52" pageNumber="53">Fig. 199</figureCitation>
). Sternum, labium and maxillae as shown in 
<figureCitation id="133A1AB0551EFF8EFAE2E37CFAC1CA2A" box="[1293,1395,1110,1137]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="52" pageNumber="53">Fig. 217</figureCitation>
. Maxillae with 7 cuspules each.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E393FC20CB62" blockId="52.[189,1399,1209,1338]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E393FF44CA8A" baselines="1228,1228" box="[189,246,1209,1235]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="52" pageNumber="53">LEGS</smallCapsWord>
. Tibia and metatarsus I as in 
<figureCitation id="133A1AB0551EFF8EFDA1E393FD06CA88" box="[590,692,1209,1235]" captionStart="Figs 274–281" captionStartId="137.[189,241,1743,1769]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure-78@137.[189,1398,265,1703]" captionTargetPageId="137" captionText="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990897" httpUri="https://zenodo.org/record/13990897/files/figure.png" pageId="52" pageNumber="53">Fig. 274</figureCitation>
. Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided with setae on tarsus II; widely divided on tarsus III; rudimentary on tarsus IV. Trichobothria: 2 rows of 7 each on tibiae, 10–12 on metatarsi, 7–10 on tarsi, 6 on cymbium. PTC I–IV with 6–7 and 5 teeth on outer and inner margins, respectively.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E24AFE99C859" blockId="52.[189,1399,1376,1538]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E24AFE9ECB23" baselines="1395,1395" box="[189,300,1376,1402]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spination" pageId="52" pageNumber="53">SPINATION</smallCapsWord>
. Palp: femur d3, pd2; tibia d2, p3, r1, v4; cymbium d4. Leg I: femur d4, pd3, rd3; patella p1; tibia p2, pv2, rv2+2M; metatarsus v1. Leg II: femur d4, pd3; patella p1; tibia p3, v6; metatarsus v6. Leg III: femur d4, pd3, rd3; patella p1, r1; tibia d2, p2, r2, v6; metatarsus d1, p3, r3, v7. Leg IV: femur d4, pd3, rd3; patella p1; tibia d2, p3, r3, v7; metatarsus d1, p3, r2, v9(8). Palpal patella and tarsi I–IV aspinose.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E103FAB2C83E" blockId="52.[189,1398,1577,1637]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E103FF46C81A" baselines="1596,1596" box="[189,244,1577,1603]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Palp" pageId="52" pageNumber="53">PALP</smallCapsWord>
. Tibia, cymbium and copulatory bulb as shown in 
<figureCitation id="133A1AB0551EFF8EFCB9E103FC0CC818" box="[854,958,1577,1603]" captionStart="Figs 364–378" captionStartId="144.[189,241,1714,1740]" captionTargetBox="[189,1398,271,1690]" captionTargetId="figure-142@144.[189,1398,265,1690]" captionTargetPageId="144" captionText="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990915" httpUri="https://zenodo.org/record/13990915/files/figure.png" pageId="52" pageNumber="53">Fig. 366</figureCitation>
. Embolus with widely tapering basal portion ending with triangular keel, and with twisted apical part (as shown in 
<figureCitation id="133A1AB0551EFF8EFBA2E161FB41C83E" box="[1101,1267,1611,1637]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="52" pageNumber="53">Figs 432–433</figureCitation>
).
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E1A1FC30C89C" blockId="52.[189,1398,1675,1736]" pageId="52" pageNumber="53">
<smallCapsWord id="8D5890E9551EFF8EFF52E1A1FE8EC8FF" baselines="1694,1695" box="[189,316,1675,1701]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spinnerets" pageId="52" pageNumber="53">SPINNERETS</smallCapsWord>
. See 
<figureCitation id="133A1AB0551EFF8EFE6DE1A1FE58C8FD" box="[386,490,1675,1702]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="52" pageNumber="53">Fig. 591</figureCitation>
. PLS: maximal diameter 0.37; length of basal, medial and apical segments 0.60, 0.42, 0.27; total length 1.29; apical segment triangular.
</paragraph>
<paragraph id="8BBE0635551EFF8EFF52E1C4FE55C953" blockId="52.[189,487,1774,1800]" box="[189,487,1774,1800]" pageId="52" pageNumber="53">
<tableNote id="76E707BB551EFF8EFF52E1C4FE55C953" box="[189,487,1774,1800]" pageId="52" pageNumber="53" targetBox="[197,1259,1812,2017]" targetPageId="52">
<smallCapsWord id="8D5890E9551EFF8EFF52E1C4FF58C95D" baselines="1793,1793" box="[189,234,1774,1800]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="52" pageNumber="53">LEG</smallCapsWord>
<smallCapsWord id="8D5890E9551EFF8EFF1DE1D9FE28C95D" baselines="1793" box="[242,410,1779,1798]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="52" pageNumber="53">MEASUREMENTS</smallCapsWord>
. ♂(♀)
</tableNote>
</paragraph>
<paragraph id="8BBE0635551EFF8EFEBCE03FFB59C9BB" pageId="52" pageNumber="53">
<table id="F901F495551E0045FF2AE03EFB59C9BA" box="[197,1259,1812,2017]" gridcols="7" gridrows="6" pageId="52" pageNumber="53">
<tr id="35310477551E0045FF2AE03EFB59C974" box="[197,1259,1812,1839]" gridrow="0" pageId="52" pageNumber="53" rowspan-0="1">
<th id="76E06D0B551E0045FED3E03EFE0CC974" box="[316,446,1812,1839]" gridcol="1" gridrow="0" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFEBCE03FFE1AC974" bold="true" box="[339,424,1813,1839]" pageId="52" pageNumber="53">Femur</emphasis>
</th>
<th id="76E06D0B551E0045FE34E03EFDEEC974" box="[475,604,1812,1839]" gridcol="2" gridrow="0" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFE1FE03FFDF4C974" bold="true" box="[496,582,1813,1839]" pageId="52" pageNumber="53">Patella</emphasis>
</th>
<th id="76E06D0B551E0045FD96E03EFD49C974" box="[633,763,1812,1839]" gridcol="3" gridrow="0" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFD76E03EFD69C975" bold="true" box="[665,731,1812,1838]" pageId="52" pageNumber="53">Tibia</emphasis>
</th>
<th id="76E06D0B551E0045FCFEE03EFC2DC974" box="[785,927,1812,1839]" gridcol="4" gridrow="0" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFCFEE03FFC2DC974" bold="true" box="[785,927,1813,1839]" pageId="52" pageNumber="53">Metatarsus</emphasis>
</th>
<th id="76E06D0B551E0045FC59E03EFB8AC974" box="[950,1080,1812,1839]" gridcol="5" gridrow="0" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFC22E03FFB93C974" bold="true" box="[973,1057,1813,1839]" pageId="52" pageNumber="53">Tarsus</emphasis>
</th>
<th id="76E06D0B551E0045FBA2E03EFB59C974" box="[1101,1259,1812,1839]" gridcol="6" gridrow="0" pageId="52" pageNumber="53">
<emphasis id="B975DA27551EFF8EFB93E03FFB0EC974" bold="true" box="[1148,1212,1813,1839]" pageId="52" pageNumber="53">Total</emphasis>
</th>
</tr>
<tr id="35310477551E0045FF2AE012FB59C908" box="[197,1259,1848,1875]" gridrow="1" pageId="52" pageNumber="53">
<th id="76E06D0B551E0045FF2AE012FEAAC908" box="[197,280,1848,1875]" gridcol="0" gridrow="1" pageId="52" pageNumber="53">Palp</th>
<td id="76E06D0B551E0045FED3E012FE0CC908" box="[316,446,1848,1875]" gridcol="1" gridrow="1" pageId="52" pageNumber="53">1.93 (2.68)</td>
<td id="76E06D0B551E0045FE34E012FDEEC908" box="[475,604,1848,1875]" gridcol="2" gridrow="1" pageId="52" pageNumber="53">1.07 (1.45)</td>
<td id="76E06D0B551E0045FD96E012FD49C908" box="[633,763,1848,1875]" gridcol="3" gridrow="1" pageId="52" pageNumber="53">1.43 (1.69)</td>
<td id="76E06D0B551E0045FCFEE012FC2DC908" box="[785,927,1848,1875]" gridcol="4" gridrow="1" pageId="52" pageNumber="53">–</td>
<td id="76E06D0B551E0045FC59E012FB8AC908" box="[950,1080,1848,1875]" gridcol="5" gridrow="1" pageId="52" pageNumber="53">0.65 (1.42)</td>
<td id="76E06D0B551E0045FBA2E012FB59C908" box="[1101,1259,1848,1875]" gridcol="6" gridrow="1" pageId="52" pageNumber="53">5.08 (7.24)</td>
</tr>
<tr id="35310477551E0045FF2AE076FB59C92C" box="[197,1259,1884,1911]" gridrow="2" pageId="52" pageNumber="53">
<th id="76E06D0B551E0045FF2AE076FEAAC92C" box="[197,280,1884,1911]" gridcol="0" gridrow="2" pageId="52" pageNumber="53">Leg I</th>
<td id="76E06D0B551E0045FED3E076FE0CC92C" box="[316,446,1884,1911]" gridcol="1" gridrow="2" pageId="52" pageNumber="53">3.20 (3.23)</td>
<td id="76E06D0B551E0045FE34E076FDEEC92C" box="[475,604,1884,1911]" gridcol="2" gridrow="2" pageId="52" pageNumber="53">1.78 (2.09)</td>
<td id="76E06D0B551E0045FD96E076FD49C92C" box="[633,763,1884,1911]" gridcol="3" gridrow="2" pageId="52" pageNumber="53">2.43 (2.35)</td>
<td id="76E06D0B551E0045FCFEE076FC2DC92C" box="[785,927,1884,1911]" gridcol="4" gridrow="2" pageId="52" pageNumber="53">2.54 (1.77)</td>
<td id="76E06D0B551E0045FC59E076FB8AC92C" box="[950,1080,1884,1911]" gridcol="5" gridrow="2" pageId="52" pageNumber="53">1.45 (1.34)</td>
<td id="76E06D0B551E0045FBA2E076FB59C92C" box="[1101,1259,1884,1911]" gridcol="6" gridrow="2" pageId="52" pageNumber="53">11.40 (10.78)</td>
</tr>
<tr id="35310477551E0045FF2AE055FB59C9C1" box="[197,1259,1919,1946]" gridrow="3" pageId="52" pageNumber="53">
<th id="76E06D0B551E0045FF2AE055FEAAC9C1" box="[197,280,1919,1946]" gridcol="0" gridrow="3" pageId="52" pageNumber="53">Leg II</th>
<td id="76E06D0B551E0045FED3E055FE0CC9C1" box="[316,446,1919,1946]" gridcol="1" gridrow="3" pageId="52" pageNumber="53">2.98 (2.96)</td>
<td id="76E06D0B551E0045FE34E055FDEEC9C1" box="[475,604,1919,1946]" gridcol="2" gridrow="3" pageId="52" pageNumber="53">1.50 (1.89)</td>
<td id="76E06D0B551E0045FD96E055FD49C9C1" box="[633,763,1919,1946]" gridcol="3" gridrow="3" pageId="52" pageNumber="53">2.18 (2.08)</td>
<td id="76E06D0B551E0045FCFEE055FC2DC9C1" box="[785,927,1919,1946]" gridcol="4" gridrow="3" pageId="52" pageNumber="53">2.14 (1.84)</td>
<td id="76E06D0B551E0045FC59E055FB8AC9C1" box="[950,1080,1919,1946]" gridcol="5" gridrow="3" pageId="52" pageNumber="53">1.43 (1.36)</td>
<td id="76E06D0B551E0045FBA2E055FB59C9C1" box="[1101,1259,1919,1946]" gridcol="6" gridrow="3" pageId="52" pageNumber="53">10.23 (10.13)</td>
</tr>
<tr id="35310477551E0045FF2AE088FB59C9E6" box="[197,1259,1954,1981]" gridrow="4" pageId="52" pageNumber="53">
<th id="76E06D0B551E0045FF2AE088FEAAC9E6" box="[197,280,1954,1981]" gridcol="0" gridrow="4" pageId="52" pageNumber="53">Leg III</th>
<td id="76E06D0B551E0045FED3E088FE0CC9E6" box="[316,446,1954,1981]" gridcol="1" gridrow="4" pageId="52" pageNumber="53">2.57 (2.63)</td>
<td id="76E06D0B551E0045FE34E088FDEEC9E6" box="[475,604,1954,1981]" gridcol="2" gridrow="4" pageId="52" pageNumber="53">1.43 (1.54)</td>
<td id="76E06D0B551E0045FD96E088FD49C9E6" box="[633,763,1954,1981]" gridcol="3" gridrow="4" pageId="52" pageNumber="53">1.89 (1.78)</td>
<td id="76E06D0B551E0045FCFEE088FC2DC9E6" box="[785,927,1954,1981]" gridcol="4" gridrow="4" pageId="52" pageNumber="53">2.60 (2.25)</td>
<td id="76E06D0B551E0045FC59E088FB8AC9E6" box="[950,1080,1954,1981]" gridcol="5" gridrow="4" pageId="52" pageNumber="53">1.46 (1.39)</td>
<td id="76E06D0B551E0045FBA2E088FB59C9E6" box="[1101,1259,1954,1981]" gridcol="6" gridrow="4" pageId="52" pageNumber="53">9.95 (9.59)</td>
</tr>
<tr id="35310477551E0045FF2AE0ECFB59C9BA" box="[197,1259,1990,2017]" gridrow="5" pageId="52" pageNumber="53">
<th id="76E06D0B551E0045FF2AE0ECFEAAC9BA" box="[197,280,1990,2017]" gridcol="0" gridrow="5" pageId="52" pageNumber="53">Leg IV</th>
<td id="76E06D0B551E0045FED3E0ECFE0CC9BA" box="[316,446,1990,2017]" gridcol="1" gridrow="5" pageId="52" pageNumber="53">3.51 (3.54)</td>
<td id="76E06D0B551E0045FE34E0ECFDEEC9BA" box="[475,604,1990,2017]" gridcol="2" gridrow="5" pageId="52" pageNumber="53">1.48 (1.92)</td>
<td id="76E06D0B551E0045FD96E0ECFD49C9BA" box="[633,763,1990,2017]" gridcol="3" gridrow="5" pageId="52" pageNumber="53">2.76 (2.66)</td>
<td id="76E06D0B551E0045FCFEE0ECFC2DC9BA" box="[785,927,1990,2017]" gridcol="4" gridrow="5" pageId="52" pageNumber="53">3.75 (3.31)</td>
<td id="76E06D0B551E0045FC59E0ECFB8AC9BA" box="[950,1080,1990,2017]" gridcol="5" gridrow="5" pageId="52" pageNumber="53">2.07 (1.61)</td>
<td id="76E06D0B551E0045FBA2E0ECFB59C9BA" box="[1101,1259,1990,2017]" gridcol="6" gridrow="5" pageId="52" pageNumber="53">13.57 (13.04)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBE0635551FFF8FFF05E623FE77CF78" blockId="53.[189,453,265,329]" box="[234,453,265,291]" pageId="53" pageNumber="54">
<emphasis id="B975DA27551FFF8FFF05E623FEF7CF78" bold="true" box="[234,325,265,291]" pageId="53" pageNumber="54">Female</emphasis>
(
<typeStatus id="54BAB897551FFF8FFEBAE623FE0CCF78" box="[341,446,265,291]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
)
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E604FE05CF12" blockId="53.[189,453,265,329]" box="[189,439,302,329]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E604FEAECF1D" baselines="321,321" box="[189,284,302,328]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="53" pageNumber="54">HABITUS</smallCapsWord>
. See 
<figureCitation id="133A1AB0551FFF8FFEB2E604FE05CF12" box="[349,439,302,329]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="53" pageNumber="54">Fig. 51.</figureCitation>
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E65BFB15CFD0" blockId="53.[189,1191,369,396]" box="[189,1191,369,396]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E65BFEDECFD2" baselines="388,388" box="[189,364,369,395]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="53" pageNumber="54">MEASUREMENTS</smallCapsWord>
. TBL 15.05, CL 4.22, CW 3.91, LL 0.41, LW 0.87, SL 2.15, SW 2.06.
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E69EFDD1CFA8" blockId="53.[189,1399,436,499]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E69EFEAACF97" baselines="455,455" box="[189,280,436,462]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="53" pageNumber="54">COLOUR</smallCapsWord>
. Mostly as in male, but chelicerae light scarlet red and legs more uniformly coloured, without difference between legs I and II–IV.
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E536FC66CC93" blockId="53.[187,1399,539,713]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E536FECACC6F" baselines="559,559" box="[189,376,540,566]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Cephalothorax" pageId="53" pageNumber="54">CEPHALOTHORAX</smallCapsWord>
. Carapace and chelicerae as shown in 
<figureCitation id="133A1AB0551FFF8FFCC3E531FC3CCC6D" box="[812,910,539,566]" captionStart="Figs 118–126" captionStartId="121.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-90@121.[189,1398,265,1816]" captionTargetPageId="121" captionText="Figs 118–126. Raveniola spp., ♀♀, paratypes (except 119, 123, 125), cephalothorax, dorsal views. 118. R. ignobilis sp. nov. (SMNH). 119. R. karategensis sp. nov., holotype (SMNH). 120. R. ornata sp. nov., from Sanglok (SMNH). 121. R. ornatula sp. nov. (SMNH). 122. R. pamira sp. nov. (SMNH). 123. R. sororculasp. nov., holotype (SMNH). 124. R. dilutasp. nov.(SMNH). 125. R. fedotovi(Charitonov, 1946), ♀ from Hojakurgan (SMNH). 126. R. pallens sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990860" httpUri="https://zenodo.org/record/13990860/files/figure.png" pageId="53" pageNumber="54">Fig. 125</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="133A1AB0551FFF8FFB32E531FA8DCC6D" box="[1245,1343,539,566]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="53" pageNumber="54">Fig. 183</figureCitation>
. Eye diameters and interdistances: AME 0.13(0.17), ALE 0.24, PLE 0.15, PME 0.07; AME–AME 0.13(0.09), ALE–AME 0.09(0.07), ALE–PLE 0.09, PLE–PME 0.06, PME–PME 0.40. Cheliceral rastellum absent. Each cheliceral furrow with 10 promarginal teeth and 5–6 mesobasal denticles. Sternum, labium and maxillae as shown in 
<figureCitation id="133A1AB0551FFF8FFE52E584FD91CC92" box="[445,547,686,713]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="53" pageNumber="54">Fig. 244</figureCitation>
. Maxillae with 13–14 cuspules each.
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E5D8FE03CDC5" blockId="53.[188,1401,753,926]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E5D8FF44CD51" baselines="773,773" box="[189,246,754,780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="53" pageNumber="54">LEGS</smallCapsWord>
. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus; narrowly divided on tarsus I; widely divided by setae on tarsus II; absent on metatarsi and tarsi III and IV. Trichobothria: 2 rows of 9–10 each on tibiae, 13–15 on metatarsi, 11–13 on tarsi, 9–10 on palpal tarsus. Palpal claw with 4 promarginal teeth widely separated from each other. PTC I–II and III–IV with 6–7 and 6–8 teeth on each margin, respectively.
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E4E2FAB0CA2E" blockId="53.[189,1400,967,1141]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E4E2FE9ECDBB" baselines="987,987" box="[189,300,968,994]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spination" pageId="53" pageNumber="54">SPINATION</smallCapsWord>
. Palpal femur and femora I–II with 4 dorsal bristles; femora III–IV with one thin basodorsal spine and 3 dorsal bristles alongside midline; palpal patella, patellae I–II and tarsi I–IV aspinose. Palp: femur pd1; tibia v10(7); tarsus v4(3). Leg I: femur pd1; tibia v5; metatarsus v6. Leg II: femur pd1; tibia p3, v7; metatarsus p1, v7(6). Leg III: femur pd3, rd2; patella p1, r1; tibia d1, p2, r2, v7; metatarsus d2, p3, r3(2), v7. Leg IV: femur rd1; patella r1; tibia d1, p2, r3, v7; metatarsus d1, p3, r3, v10(9).
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E3B4FE62CA86" blockId="53.[189,1398,1182,1245]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E3B4FED0CAED" baselines="1201,1201" box="[189,354,1182,1208]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spermathecae" pageId="53" pageNumber="54">SPERMATHECAE</smallCapsWord>
. Each of paired spermathecae with wide cone-shaped trunk and short fusiform outer branch (
<figureCitation id="133A1AB0551FFF8FFEF3E3E8FE76CA86" box="[284,452,1218,1245]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="53" pageNumber="54">Figs 528–529</figureCitation>
).
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E22CFC44CB1F" blockId="53.[189,1398,1285,1349]" pageId="53" pageNumber="54">
<smallCapsWord id="8D5890E9551FFF8FFF52E22CFE8ECB45" baselines="1305,1305" box="[189,316,1286,1312]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spinnerets" pageId="53" pageNumber="54">SPINNERETS</smallCapsWord>
. See 
<figureCitation id="133A1AB0551FFF8FFE93E22FFE53CB7B" box="[380,481,1285,1312]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="53" pageNumber="54">Fig. 592</figureCitation>
. PMS: absent. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.84, 0.55, 0.53; total length 1.92; apical segment triangular.
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E247FE81CBDC" blockId="53.[189,307,1389,1415]" box="[189,307,1389,1415]" pageId="53" pageNumber="54">
<heading id="D0F6B159551FFF8FFF52E247FE81CBDC" bold="true" box="[189,307,1389,1415]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B975DA27551FFF8FFF52E247FE81CBDC" bold="true" box="[189,307,1389,1415]" pageId="53" pageNumber="54">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E2B7FDD6C8CF" blockId="53.[189,1401,1437,1684]" pageId="53" pageNumber="54">
Carapace length in males (n= 3) varies from 3.30 to 3.95. One of the fragmented 
<typeStatus id="54BAB897551FFF8FFB64E2B7FA9BCBEC" box="[1163,1321,1437,1463]" pageId="53" pageNumber="54" type="paralectotype">paralectotype</typeStatus>
males possesses 8 (vs 
<quantity id="4CF9ABD0551FFF8FFE90E2E9FE1CCB87" box="[383,430,1474,1501]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="53" pageNumber="54" unit="in" value="7.0">7 in</quantity>
the 
<typeStatus id="54BAB897551FFF8FFE0AE2E8FDE0CB87" box="[485,594,1474,1500]" pageId="53" pageNumber="54" type="lectotype">lectotype</typeStatus>
) teeth on the cheliceral furrow promargin (
<figureCitation id="133A1AB0551FFF8FFBB2E2E8FB74CB87" box="[1117,1222,1474,1500]" captionStart="Figs 196–201" captionStartId="128.[189,241,1850,1876]" captionTargetBox="[189,1398,276,1794]" captionTargetId="figure-68@128.[189,1398,265,1816]" captionTargetPageId="128" captionText="Figs 196–201. Raveniola spp., ♂♂, chelicera, prolateral views. 196. R. redikorzevi (Spassky, 1937), ♂ from Kepele (SMNH). 197. R. cucullata sp. nov., paratype from Shikildara (SMNH). 198. R. ignobilis sp. nov., paratype (SMNH). 199. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 200. R. kopetdaghensis (Fet, 1984), paratype (SMNH). 201. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990876" httpUri="https://zenodo.org/record/13990876/files/figure.png" pageId="53" pageNumber="54">Fig. 199</figureCitation>
). According to 
<bibRefCitation id="EF907BC4551FFF8FFF50E2CDFE2DC85A" author="Charitonov D. E." box="[191,415,1511,1537]" pageId="53" pageNumber="54" pagination="19 - 32" refId="ref77775" refString="Charitonov D. E. 1946. New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo 12: 19 - 32. [In Russian and Latin.]" type="journal article" year="1946">Charitonov (1946)</bibRefCitation>
, within the type series, the number of the maxillary cuspules ranges from 5 to 8 (some type specimens were used by D.E. Charitonov as preparations, and only a few of their parts have survived to date). A weak difference in the structure of the male tibia and metatarsus I is shown in 
<figureCitation id="133A1AB0551FFF8FFF52E17FFE90C82B" box="[189,290,1621,1648]" captionStart="Figs 274–281" captionStartId="137.[189,241,1743,1769]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure-78@137.[189,1398,265,1703]" captionTargetPageId="137" captionText="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990897" httpUri="https://zenodo.org/record/13990897/files/figure.png" pageId="53" pageNumber="54">Fig. 275</figureCitation>
. Most other features, including a specific shape of the embolic keel, were found to be consistent throughout the available specimens.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE551FFF8FFF52E197FAD9C92E" pageId="53" pageNumber="54" type="biology_ecology">
<paragraph id="8BBE0635551FFF8FFF52E197FE92C88C" blockId="53.[189,288,1725,1751]" box="[189,288,1725,1751]" pageId="53" pageNumber="54">
<heading id="D0F6B159551FFF8FFF52E197FE92C88C" bold="true" box="[189,288,1725,1751]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B975DA27551FFF8FFF52E197FE92C88C" bold="true" box="[189,288,1725,1751]" pageId="53" pageNumber="54">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E1C7FAD9C92E" blockId="53.[189,1401,1773,1910]" pageId="53" pageNumber="54">
<taxonomicName id="4C017DB6551FFF8FFF52E1C7FE2EC95C" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[189,412,1773,1799]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA27551FFF8FFF52E1C7FE2EC95C" box="[189,412,1773,1799]" italics="true" pageId="53" pageNumber="54">Raveniola fedotovi</emphasis>
</taxonomicName>
inhabits the rocky midland montane slopes between 1100 and 
<quantity id="4CF9ABD0551FFF8FFB7DE1C7FB5FC95C" box="[1170,1261,1773,1799]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="53" pageNumber="54" unit="m" value="1600.0">1600 m</quantity>
a.s.l., from the steppe shrubland zone in high foothills (
<figureCitation id="133A1AB0551FFF8FFD3AE038FC8FC977" box="[725,829,1810,1836]" captionStart="Figs 707–714" captionStartId="178.[189,241,1822,1848]" captionTargetBox="[189,1399,265,1784]" captionTargetId="figure-85@178.[189,1399,265,1784]" captionTargetPageId="178" captionText="Figs 707–714. Raveniola spp., habitats of R. fedotovi (Charitonov, 1946) (707–710) and R. zyuzini sp. nov. (711–714), Uzbekistan. 707. Hissar Mts (northwestern slope), montane slopes south of Ishkent Village. 708. Zeravshan Mts, Kitab Reserve, Hojakurgan Canyon. 709–710. Same, a rocky substrate under the tree canopy. 711. Babatagh Mts, as seen from foothills. 712. Same, as seen from the midland zone (ca 1200 m a.s.l.). 713–714. Same, the collection sites at 1400 m and 1800 m a.s.l., respectively." figureDoi="http://doi.org/10.5281/zenodo.13990973" httpUri="https://zenodo.org/record/13990973/files/figure.png" pageId="53" pageNumber="54">Fig. 707</figureCitation>
) to the higher-located zone of mosaic riverside woodland dominated by 
<taxonomicName id="4C017DB6551FFF8FFE08E01CFD3AC90B" box="[487,648,1846,1872]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="regia">
<emphasis id="B975DA27551FFF8FFE08E01CFD3AC90B" box="[487,648,1846,1872]" italics="true" pageId="53" pageNumber="54">Juglans regia</emphasis>
</taxonomicName>
(
<figureCitation id="133A1AB0551FFF8FFD75E01CFCB1C90A" box="[666,771,1846,1873]" captionStart="Figs 707–714" captionStartId="178.[189,241,1822,1848]" captionTargetBox="[189,1399,265,1784]" captionTargetId="figure-85@178.[189,1399,265,1784]" captionTargetPageId="178" captionText="Figs 707–714. Raveniola spp., habitats of R. fedotovi (Charitonov, 1946) (707–710) and R. zyuzini sp. nov. (711–714), Uzbekistan. 707. Hissar Mts (northwestern slope), montane slopes south of Ishkent Village. 708. Zeravshan Mts, Kitab Reserve, Hojakurgan Canyon. 709–710. Same, a rocky substrate under the tree canopy. 711. Babatagh Mts, as seen from foothills. 712. Same, as seen from the midland zone (ca 1200 m a.s.l.). 713–714. Same, the collection sites at 1400 m and 1800 m a.s.l., respectively." figureDoi="http://doi.org/10.5281/zenodo.13990973" httpUri="https://zenodo.org/record/13990973/files/figure.png" pageId="53" pageNumber="54">Fig. 708</figureCitation>
). The collected conspecific specimens were found in overgrown screes under shrubs and tree canopies hiding deeply inside the rocky layer (
<figureCitation id="133A1AB0551FFF8FFB57E071FAEDC92E" box="[1208,1375,1883,1909]" captionStart="Figs 707–714" captionStartId="178.[189,241,1822,1848]" captionTargetBox="[189,1399,265,1784]" captionTargetId="figure-85@178.[189,1399,265,1784]" captionTargetPageId="178" captionText="Figs 707–714. Raveniola spp., habitats of R. fedotovi (Charitonov, 1946) (707–710) and R. zyuzini sp. nov. (711–714), Uzbekistan. 707. Hissar Mts (northwestern slope), montane slopes south of Ishkent Village. 708. Zeravshan Mts, Kitab Reserve, Hojakurgan Canyon. 709–710. Same, a rocky substrate under the tree canopy. 711. Babatagh Mts, as seen from foothills. 712. Same, as seen from the midland zone (ca 1200 m a.s.l.). 713–714. Same, the collection sites at 1400 m and 1800 m a.s.l., respectively." figureDoi="http://doi.org/10.5281/zenodo.13990973" httpUri="https://zenodo.org/record/13990973/files/figure.png" pageId="53" pageNumber="54">Figs 709–710</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE551FFF8FFF52E0B4FD84C9B2" box="[189,566,1950,2025]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="8BBE0635551FFF8FFF52E0B4FEE4C9E3" blockId="53.[189,342,1950,1976]" box="[189,342,1950,1976]" pageId="53" pageNumber="54">
<heading id="D0F6B159551FFF8FFF52E0B4FEE4C9E3" bold="true" box="[189,342,1950,1976]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B975DA27551FFF8FFF52E0B4FEE4C9E3" bold="true" box="[189,342,1950,1976]" pageId="53" pageNumber="54">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551FFF8FFF52E0E4FD84C9B2" blockId="53.[189,566,1998,2025]" box="[189,566,1998,2025]" pageId="53" pageNumber="54">
South 
<collectingCountry id="F31646A5551FFF8FFEE6E0E4FE23C9B3" box="[265,401,1998,2024]" name="Uzbekistan" pageId="53" pageNumber="54">Uzbekistan</collectingCountry>
. See 
<figureCitation id="133A1AB0551FFF8FFE22E0E4FD81C9B2" box="[461,563,1998,2025]" captionStart="Figs 751–760" captionStartId="184.[189,241,1662,1688]" captionTargetBox="[258,1323,265,1629]" captionTargetId="figure-247@184.[258,1323,265,1629]" captionTargetPageId="184" captionText="Figs 751–760. Distribution records of Raveniola spp. 751. R. alajensis sp. nov. (), R. cucullata sp. nov. () and R. dolosa sp. nov. (). 752. R. hirta sp. nov. (), R. ignobilis sp. nov. () and R. insolita sp. nov. (). 753. R. karategensis sp. nov. (), R. ornata sp. nov. () and R. ornatula sp. nov. (). 754. R. pamira sp. nov. (), R. sororcula sp. nov. () and R. diluta sp. nov. (). 755. R. fedotovi (Charitonov, 1946) (), R. pallens sp. nov. () and R. zyuzini sp. nov. (). 756. R. ferghanensis (Zonstein, 1984) () and R. kirgizica sp. nov. (). 757. R. kopetdaghensis (Fet, 1984) (). 758. R. mikhailovi Zonstein, 2021 () and R. nenilini sp. nov. (). 759. R. ovchinnikovi sp. nov. () and R. tarabaevi sp. nov. (). 760. R. virgata (Simon, 1891) (;? = the questionable type locality) and R. vulpina sp. nov. (). Country abbreviations as in Figs 747–750." figureDoi="http://doi.org/10.5281/zenodo.14018423" httpUri="https://zenodo.org/record/14018423/files/figure.png" pageId="53" pageNumber="54">Fig. 755</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>