127 lines
15 KiB
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127 lines
15 KiB
XML
<document id="B02BD894EC85BEC0E1F994996D382E11" ID-CLB-Dataset="33931" ID-DOI="http://dx.doi.org/10.3897/zookeys.414.7671" ID-GBIF-Dataset="ee145c8e-3cd6-4146-ab36-33f3d406bd94" ID-PMC="PMC4086050" ID-Pensoft-Pub="1313-2970-414-19" ID-PubMed="25009416" ID-ZooBank="C1E83718BC8544A69F00ED4C93D758B5" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-414-19" ModsDocOrigin="ZooKeys 414" ModsDocTitle="Review of the millipede family Trichopolydesmidae in the Oriental realm (Diplopoda, Polydesmida), with descriptions of new genera and species" checkinTime="1451245813408" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques & VandenSpiegel, Didier" docDate="2014" docId="D95C39C50F825C66B7F4C5DEC36BEDB9" docLanguage="en" docName="ZooKeys 414: 19-65" docOrigin="ZooKeys 414" docSource="http://dx.doi.org/10.3897/zookeys.414.7671" docTitle="Trichopolydesmidae Verhoeff 1910" docType="treatment" docVersion="6" lastPageNumber="27" masterDocId="FFE351740A7A365EFF8EE113FF99FFAD" masterDocTitle="Review of the millipede family Trichopolydesmidae in the Oriental realm (Diplopoda, Polydesmida), with descriptions of new genera and species" masterLastPageNumber="65" masterPageNumber="19" pageNumber="26" updateTime="1732596600966" updateUser="ExternalLinkService">
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<mods:mods id="A64495E2EE2E773B374D6D05F5959F02" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EAD57CA2A8E80EBB8DA27860A0DCB7C7">
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<mods:title id="EC4FD4A8E1178C0048E9055C82928F2C">Review of the millipede family Trichopolydesmidae in the Oriental realm (Diplopoda, Polydesmida), with descriptions of new genera and species</mods:title>
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<mods:name id="F22EC27EC894000B7317238ECF7A9594" type="personal">
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<mods:namePart id="0842A86D9478B5750069784D8D2B677C">Golovatch, Sergei I.</mods:namePart>
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<mods:roleTerm id="BEB516DC4AA792A57F23990B93C78793">Author</mods:roleTerm>
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<mods:namePart id="8656D43224E688F94D86529E4237C795">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:roleTerm id="4C99233A3C66DFA5BC5A24292743549E">Author</mods:roleTerm>
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<mods:namePart id="E6463C8261DBC1B5226493DC0195622E">VandenSpiegel, Didier</mods:namePart>
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<mods:typeOfResource id="6F5CD023B91CE65DF257EB1C5FDE4D68">text</mods:typeOfResource>
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<mods:titleInfo id="7F85CCF3EFB128B1CE7E057AB75D99D7">
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<mods:title id="586DB5B269E5F04508A7A564A5CDCC0B">ZooKeys</mods:title>
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<mods:part id="8DB8A833C351D7BAA42A90AE0097D23F">
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<mods:date id="46067B5B16CBC2918561C35D93961C81">2014</mods:date>
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<mods:detail id="D2A6213EF44D1D928C75543218C4EAE7" type="volume">
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<mods:number id="0BC7D3576A2B047415056EF072409FDD">414</mods:number>
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</mods:detail>
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<mods:start id="C114F50E3CA2247E91EF31DDE50F9B00">19</mods:start>
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<mods:end id="EE897AC8B9CC8B7BFFB6EA7E0DA6D9B6">65</mods:end>
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<mods:url id="ED696166EDDB77793C80F32588F9B81B">http://dx.doi.org/10.3897/zookeys.414.7671</mods:url>
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<mods:classification id="EC92F8A92D513D383D051EFDEBBF631A">journal article</mods:classification>
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<mods:identifier id="A3D65EF0A9D30A8B10A5A3CE7A99BA0A" type="DOI">http://dx.doi.org/10.3897/zookeys.414.7671</mods:identifier>
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<mods:identifier id="E1E2CA3629527267B2FD133E3D819406" type="Pensoft-Pub">1313-2970-414-19</mods:identifier>
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<mods:identifier id="EDFD52D2794D19BC721FE8C06C484DD8" type="ZooBank">C1E83718BC8544A69F00ED4C93D758B5</mods:identifier>
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<treatment id="D95C39C50F825C66B7F4C5DEC36BEDB9" ID-GBIF-Taxon="152053374" LSID="urn:lsid:plazi:treatment:D95C39C50F825C66B7F4C5DEC36BEDB9" httpUri="http://treatment.plazi.org/id/D95C39C50F825C66B7F4C5DEC36BEDB9" lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26" scope_class="Diplopoda" scope_order="Polydesmida">
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<subSubSection id="459FE2F342016E465D9D995BBF8FFFD6" pageId="7" pageNumber="26" type="nomenclature">
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<paragraph id="01BA86AC8AA7ADA0B65F4F47C2514AA5" pageId="7" pageNumber="26">
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Family
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<taxonomicName id="333BC8A2EBC6743D0B3ACC33C703F53D" ID-CoL="87CC6" LSID="http://species-id.net/wiki/Trichopolydesmidae" authority="Verhoeff, 1910" authorityName="Verhoeff" authorityYear="1910" class="Diplopoda" family="Trichopolydesmidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="7" pageNumber="26" phylum="Arthropoda" rank="family">Trichopolydesmidae Verhoeff, 1910</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="530989113DBCE1769187C215ED7217D6" lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26" type="diagnosis">
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<paragraph id="7BFE127EFF2891AA970CCB93297DE297" pageId="7" pageNumber="26">Diagnosis.</paragraph>
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<paragraph id="FB336B2DD224FDEB737A62E849C48921" lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26">
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A family of the superfamily
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<taxonomicName id="B87BC81E0C471FE0FAA5A5785AF3B60A" genus="Trichopolydesmoidea" lsidName="Trichopolydesmoidea" pageId="7" pageNumber="26" rank="genus">Trichopolydesmoidea</taxonomicName>
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, suborder
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<taxonomicName id="BDC52833090E99727F114C7AFEE05255" genus="Polydesmidea" lsidName="Polydesmidea" pageId="7" pageNumber="26" rank="genus">Polydesmidea</taxonomicName>
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with 18, 19 or 20 segments, sometimes varying between sexes. Body very small to
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<pageBreakToken id="BE53FD64CA4BC6BEC9E7D6798F5C924C" pageId="8" pageNumber="27" start="start">small</pageBreakToken>
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(ca 2-20 mm long). Tegument microalveolate, limbus mostly microspiculate. ♂ head with or without vertigial modifications. Antennae often geniculate between segments 5 and 6, antennomeres 5 and 6 each usually with a compact group of bacilliform sensilla apicodorsally, rarely 6th in ♂ with a dorsoparabasal stump. Metaterga usually with 3 regular, more rarely with more and/or irregular, transverse rows of sharp (= simple), bacilliform or clavate setae sometimes borne on knobs; side margin of paraterga incised or tuberculate, with 2 or 3 setae. Paraterga from absent to strongly developed, usually only slightly indented laterally, only exceptionally deeply trilobate (
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<taxonomicName id="1CA9D5D1CE8579233418572DC16639F5" class="Diplopoda" family="Trichopolydesmidae" genus="Trilobodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trilobodesmus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Trilobodesmus</taxonomicName>
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Golovatch &
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<normalizedToken id="EF4B084FA57A4170A218E013CA5F7B5C" originalValue="Mauriès">Mauries</normalizedToken>
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, 2007). Pore formula usually normal: 5, 7, 9, 10, 12, 13, 15-17 (18,19), but sometimes ozopores completely wanting, rarely unusually large, normally opening flush on dorsal or dorsolateral surface either near penultimate lateral incision or at caudal corner of paraterga. Legs rather short to long, ♂ ones often stouter and elongate, rarely modified, inflated, only sometimes with peculiar ventral setae, including sphaerotrichomes.
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</paragraph>
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<paragraph id="842BA1A0824EC85E95BA2AA4CDE1261A" pageId="8" pageNumber="27">
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Gonopods with subglobose, medially fused coxae, these being rather small to quite large (with correspondingly large gonocoel), micropapillate and at most only slightly setose laterally, sometimes with a frontal (= anterior) process, each supporting a cannula medially (exception:
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<taxonomicName id="6B7072B2F18BA6BA009708C064B166E6" class="Diplopoda" family="Macrosternodesmidae" genus="Caucasodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasodesmus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Caucasodesmus</taxonomicName>
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Golovatch, 1987, in which there is no cannula) and a sack-shaped to elongate telopodite. The latter ranging from stout and short, deeply sunken inside a deep gonocoel, to slender and long, nearly fully exposed, strongly to modestly curved caudad or mesad, only exceptionally geniculate (distal half directed abruptly laterad:
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<taxonomicName id="144FD32D9DB525597AEC360CEF577914" class="Diplopoda" family="Trichopolydesmidae" genus="Gonatodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonatodesmus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Gonatodesmus</taxonomicName>
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gen. n.) or perforating coxal wall with a prominent process (
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<taxonomicName id="DE960FD69E04958B2F63CEE3E7FE6AB3" class="Diplopoda" family="Trichopolydesmidae" genus="Schizotelopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizotelopus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Schizotelopus</taxonomicName>
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Verhoeff, 1941), usually complex, with various processes or outgrowths, sometimes fringed; seminal groove normally terminating distally or apically on a separate branch or tooth (= solenomere), rarely on a tooth inside gonocoel, exceptionally absent (
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<taxonomicName id="028013466E9BF43FC73969E749F1F87C" class="Diplopoda" family="Macrosternodesmidae" genus="Caucasodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasodesmus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Caucasodesmus</taxonomicName>
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). Typically neither an accessory seminal chamber nor a hairy pulvillus (a few exceptions, e.g.
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<taxonomicName id="07D3AEEB139E4C82E3A08ED4609AF0C0" class="Diplopoda" family="Trichopolydesmidae" genus="Gonatodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonatodesmus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Gonatodesmus</taxonomicName>
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gen. n.).
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</paragraph>
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</subSubSection>
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<subSubSection id="8EBD6EE4753FFE14EB52BEA379E268E8" pageId="8" pageNumber="27" type="type genus">
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<paragraph id="3F5F55AA19BF1516A15309FC410C9287" pageId="8" pageNumber="27">Type genus.</paragraph>
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<paragraph id="F50B1C96A2671FC539EE7307C7093D03" pageId="8" pageNumber="27">
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<taxonomicName id="7A3ED630B06B2F4BE2E597847D3950D0" class="Diplopoda" family="Trichopolydesmidae" genus="Trichopolydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichopolydesmus" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">Trichopolydesmus</taxonomicName>
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Verhoeff, 1910.
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</paragraph>
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</subSubSection>
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<subSubSection id="6A420F5C16CFF0A6F3CE6045A7A1D143" pageId="8" pageNumber="27" type="remarks">
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<paragraph id="0D8ECC567E320AA33BB0B2F10B9631EF" pageId="8" pageNumber="27">Remarks.</paragraph>
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<paragraph id="45D4FE81DF942DEB20C8631CDFE00CA5" pageId="8" pageNumber="27">
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Most of the above somatic and gonopod features of
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<taxonomicName id="DEC9500E7B148422878BDFBE434B84FA" class="Diplopoda" family="Trichopolydesmidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
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are in no way unique to the family, sometimes being also encountered, in various combinations, in the micropolydesmoid family
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<taxonomicName id="D89962742F986B22B5D459ECBEADE667" class="Diplopoda" family="Opisotretidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Opisotretidae</taxonomicName>
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of the same superfamily
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<taxonomicName id="8117DCBB5872A940A75061F797644A02" genus="Trichopolydesmoidea" lsidName="Trichopolydesmoidea" pageId="8" pageNumber="27" rank="genus">Trichopolydesmoidea</taxonomicName>
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, as well as in certain macropolydesmoid members of the family
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<taxonomicName id="7099E6A295F287AC881337D1F74150C0" class="Diplopoda" family="Polydesmidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
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, superfamily
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<taxonomicName id="BF92ED8C25F37FE54A694BD12EF23F53" genus="Polydesmoidea" lsidName="Polydesmoidea" pageId="8" pageNumber="27" rank="genus">Polydesmoidea</taxonomicName>
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(
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<bibRefCitation id="09D5D051B15569398CD9ACC197D4089B" author="Golovatch, SI" journalOrPublisher="ZooKeys" pageId="44" pageNumber="63" pagination="13 - 77" title="Review of the millipede family Opisotretidae (Diplopoda, Polydesmida), with descriptions of new species." url="10.3897/zookeys.302.5357" volume="302" year="2013">Golovatch et al. 2013</bibRefCitation>
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,
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<bibRefCitation id="1494CED5B371536545596047DA4013A8" author="Golovatch, SI" journalOrPublisher="ZooKeys" pageId="44" pageNumber="63" pagination="63 - 78" title="A reclassification of the millipede superfamily Trichopolydesmoidea, with descriptions of two new species from the Aegean region (Diplopoda, Polydesmida)." url="10.3897/zookeys.340.6295" volume="340" year="2013">Golovatch 2013</bibRefCitation>
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). The finely microspiculate limbus is also characteristic of most of the
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<taxonomicName id="2C11DA7BE07C7B40E6BAE2A09D4ACA54" genus="Polydesmidea" lsidName="Polydesmidea" pageId="8" pageNumber="27" rank="genus">Polydesmidea</taxonomicName>
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, and is another feature whose importance was obviously overestimated by
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<bibRefCitation id="8DFF24692FE0067AF5454AB0E4F33D6A" author="Simonsen, A" journalOrPublisher="Museum of Zoology, University of Bergen, Bergen" pageId="45" pageNumber="64" title="Phylogeny and biogeography of the millipede order Polydesmida, with special emphasis on the suborder Polydesmidea." year="1990">Simonsen (1990)</bibRefCitation>
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. It is only the gonopod structure that seems to be characteristic of
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<taxonomicName id="759F06A46F8F73793F8FCF60A3D66B6C" class="Diplopoda" family="Trichopolydesmidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
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, at least so to a certain extent. Superficially, female and/or juvenile trichopolydesmids from the Oriental realm are not or only barely distinguishable from the often sympatric or even syntopic female or juvenile
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<taxonomicName id="9E88A4F7853CEA3B307FAB5B453AF6E3" class="Diplopoda" family="Opisotretidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Opisotretidae</taxonomicName>
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or smaller
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<taxonomicName id="5328E2B9C295DB5AEF879C1634AB5948" class="Diplopoda" family="Polydesmidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="5BFE8A5FF16E9A194412792CA7F6DF80" pageId="8" pageNumber="27">
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The following 13 nominate Oriental genera of
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<taxonomicName id="6478C3E5FCD8DE8E7129A11630A4FCEA" class="Diplopoda" family="Trichopolydesmidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="8" pageNumber="27" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
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seem to be valid, and are arranged and defined below in alphabetic order.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |