treatments-xml/data/1B/4B/76/1B4B762AFFDEFFF3FD0E4228B771FDA0.xml
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<document id="4EBFB3FDBACA528FD0F0F2ACBF7FF3A4" ID-DOI="10.1206/0003-0090(2005)290&lt;0001:SBOTGA&gt;2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="5361751" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630237087708" checkinUser="felipe" docAuthor="WHEELER, QUENTIN D. &amp; MILLER, KELLY B." docDate="2005" docId="1B4B762AFFDEFFF3FD0E4228B771FDA0" docLanguage="en" docName="B290.pdf" docOrigin="Bulletin of the American Museum of Natural History 2005 (290)" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0090%282005%29290%3C0001%3ASBOTGA%3E2.0.CO%3B2" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Agathidium sexstriatum Horn 1880" docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="E7720E52FFF8FFDBFFCF4073B500FFF5" masterDocTitle="Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part I. Coleoptera: Leiodidae" masterLastPageNumber="95" masterPageNumber="1" pageNumber="39" updateTime="1699194440882" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4B7D54CE4EDE2C0AB85678ED22D43516">Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part I. Coleoptera: Leiodidae</mods:title>
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<subSubSection id="DBF894B7FFDEFFFDFD0E4228B6ECFD7A" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="935DC73CFFDEFFFDFD0E4228B6ECFD7A" blockId="38.[705,1204,603,655]" pageId="38" pageNumber="39">
<heading id="C8157050FFDEFFFDFD0E4228B6ECFD7A" centered="true" fontSize="10" level="2" pageId="38" pageNumber="39" reason="4">
<taxonomicName id="54E2BCBFFFDEFFFDFD0E4228B139FD87" ID-CoL="65QQJ" authorityName="Horn" authorityYear="1880" box="[705,1081,603,626]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="85" phylum="Arthropoda" rank="species" species="sexstriatum">
<emphasis id="A1961B2EFFDEFFFDFD0E4228B139FD87" box="[705,1081,603,626]" italics="true" pageId="38" pageNumber="39">AGATHIDIUM SEXSTRIATUM</emphasis>
</taxonomicName>
SPECIES GROUP
</heading>
</paragraph>
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<subSubSection id="DBF894B7FFDEFFFDFD1F42DFB61AFB95" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="935DC73CFFDEFFFDFD1F42DFB61AFB95" blockId="38.[691,1220,684,1741]" pageId="38" pageNumber="39">
DIAGNOSIS: The
<taxonomicName id="54E2BCBFFFDEFFFDFC6842DFB15BFD36" authorityName="Horn" authorityYear="1880" box="[935,1115,684,707]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sexstriatum">
<emphasis id="A1961B2EFFDEFFFDFC6842DFB15BFD36" box="[935,1115,684,707]" italics="true" pageId="38" pageNumber="39">A. sexstriatum</emphasis>
</taxonomicName>
species group is distinguished by the following combination of characters: head narrowed behind eye, without prolonged postocular temporum (fig. 44); body not strongly convex dorsally and not contractile (figs. 49); elytron with 2 8 serial series of punctures; aedeagus with ventral operculum divided into two plates; male tarsal formula 554, female 444; pronotum with evident posterolateral angles (fig. 52); clypeus large, separated from frons by impressed frontoclypeal suture, protruding anteriorly, from one­third to one­half of clypeus anterad of lateral margins of frons (fig. 52).
</paragraph>
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<subSubSection id="DBF894B7FFDEFFF3FD1F4415B46AFDEE" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="discussion">
<paragraph id="935DC73CFFDEFFFDFD1F4415B6BDFA36" blockId="38.[691,1220,684,1741]" pageId="38" pageNumber="39">
DISCUSSION: The
<taxonomicName id="54E2BCBFFFDEFFFDFC624414B15EFB8B" authorityName="Horn" authorityYear="1880" box="[941,1118,1127,1150]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sexstriatum">
<emphasis id="A1961B2EFFDEFFFDFC624414B15EFB8B" box="[941,1118,1127,1150]" italics="true" pageId="38" pageNumber="39">A. sexstriatum</emphasis>
</taxonomicName>
species group includes three similar species first described by
<bibRefCitation id="F773BACDFFDEFFFDFCFC44D1B6C8FB4C" author="Horn, G. H." box="[819,968,1186,1209]" pageId="38" pageNumber="39" pagination="219 - 322" refId="ref52484" refString="Horn, G. H. 1880. Synopsis of the Silphidae of the United States with reference to the genera of other countries. Transactions of the American Entomological Society 8: 219 - 322." type="journal article" year="1880">Horn (1880)</bibRefCitation>
. Hlisnikovský (1964) proposed the subgenus
<taxonomicName id="54E2BCBFFFDEFFFDFC1144CCB191FB23" authorityName="Hlisnikovsky" authorityYear="1964" box="[990,1169,1215,1238]" class="Insecta" family="Leiodidae" genus="Rhabdoelytrum" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="A1961B2EFFDEFFFDFC1144CCB191FB23" box="[990,1169,1215,1238]" italics="true" pageId="38" pageNumber="39">Rhabdoelytrum</emphasis>
</taxonomicName>
for these species. As discussed above, it is conceivable that these species are related to
<taxonomicName id="54E2BCBFFFDEFFFDFD7C456BB641FADA" box="[691,833,1304,1327]" class="Insecta" family="Leiodidae" genus="Stetholiodes" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="A1961B2EFFDEFFFDFD7C456BB641FADA" box="[691,833,1304,1327]" italics="true" pageId="38" pageNumber="39">Stetholiodes</emphasis>
</taxonomicName>
or other genera (
<bibRefCitation id="F773BACDFFDEFFFDFBC7456BB662FAB8" author="Angelini, F. &amp; L. de Marzo" pageId="38" pageNumber="39" pagination="3 - 15" refId="ref50159" refString="Angelini, F., and L. de Marzo. 1987 b. Notes on the genus Stetholiodes Fall with descriptions of four new species (Coleoptera, Leiodidae, Anisotomini). Revue Suisse de Zoologie 94: 3 - 15." type="journal article" year="1987">Angelini and de Marzo, 1987b</bibRefCitation>
;
<bibRefCitation id="F773BACDFFDEFFFDFCB74545B1B7FAB8" author="Angelini, F. &amp; S. Peck" box="[888,1207,1334,1357]" pageId="38" pageNumber="39" pagination="328 - 343" refId="ref50314" refString="Angelini, F., and S. Peck. 2000. Besuchetionella, new genus, with descriptions of 18 new species from Asia (Coleoptera: Leiodidae: Agathidiini). Zoological Studies 39: 328 - 343." type="journal article" year="2000">Angelini and Peck, 2000</bibRefCitation>
). They are conservatively maintained in
<taxonomicName id="54E2BCBFFFDEFFFDFD7C4502B63BFA7D" box="[691,827,1393,1416]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="A1961B2EFFDEFFFDFD7C4502B63BFA7D" box="[691,827,1393,1416]" italics="true" pageId="38" pageNumber="39">Agathidium</emphasis>
</taxonomicName>
until analyses are completed. Species of the group are distributed in the western
<collectingCountry id="EBF587ACFFDEFFFDFCD845DFB6B9FA36" box="[791,953,1452,1475]" name="United States of America" pageId="38" pageNumber="39">United States</collectingCountry>
.
</paragraph>
<paragraph id="935DC73CFFDEFFF3FD1F45B9B725FF00" blockId="38.[691,1220,684,1741]" lastBlockId="40.[115,644,193,539]" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
This group includes species that exhibit no observed unequivocal synapomorphies, although the rather short and broad form of the median lobe of the aedeagus, serial series of punctures, and prominently protruding clypeus make these species separable from other
<taxonomicName id="54E2BCBFFFDEFFFDFD7C4608B63BF967" box="[691,827,1659,1682]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="A1961B2EFFDEFFFDFD7C4608B63BF967" box="[691,827,1659,1682]" italics="true" pageId="38" pageNumber="39">Agathidium</emphasis>
</taxonomicName>
included in this study. The latter character refers to forms in which the clypeus protrudes clearly beyond the anterior an­ gles of the frons located dorsad of the antennal insertions and each side of the clypeus. It is particularly useful in distinguishing the
<taxonomicName id="54E2BCBFFFDFFFFCFFB4460FB423F966" authorityName="Horn" authorityYear="1880" box="[123,291,1660,1683]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sexstriatum">
<emphasis id="A1961B2EFFDFFFFCFFB4460FB423F966" box="[123,291,1660,1683]" italics="true" pageId="39" pageNumber="40">A. sexstriatum</emphasis>
</taxonomicName>
group species from most similarly small­sized
<taxonomicName id="54E2BCBFFFDFFFFCFE8246EAB4D5F945" box="[333,469,1689,1712]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="A1961B2EFFDFFFFCFE8246EAB4D5F945" box="[333,469,1689,1712]" italics="true" pageId="39" pageNumber="40">Agathidium</emphasis>
</taxonomicName>
with short postocular tempora. Due to a large number of symplesiomorphic similarities, these species share a general habitus and many specific characters with the related genus
<taxonomicName id="54E2BCBFFFDFFFFCFB80462CB1CBF983" authorityName="Panzer (Wheeler" authorityYear="1979" box="[1103,1227,1631,1654]" class="Magnoliopsida" family="Apocynaceae" genus="Anisotoma" kingdom="Plantae" order="Gentianales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="A1961B2EFFDFFFFCFB80462CB1CBF983" box="[1103,1227,1631,1654]" italics="true" pageId="39" pageNumber="40">Anisotoma</emphasis>
</taxonomicName>
(cf. fig. 41, see also
<bibRefCitation id="F773BACDFFDFFFFCFC6B460FB15CF966" author="Wheeler, Q. D." box="[932,1116,1660,1683]" pageId="39" pageNumber="40" pagination="251 - 309" refId="ref56916" refString="Wheeler, Q. D. 1979 b. Slime-mold beetles of the genus Anisotoma (Leiodidae): Evolution and classification. Systematic Entomology 4: 251 - 309." type="journal article" year="1979">Wheeler, 1979b</bibRefCitation>
) and oth­ er genera. However, the abrupt 3­segmented antennal club and distinct supraocular cari­ nae distinguish these
<taxonomicName id="54E2BCBFFFD0FFF3FE4940B2B70EFF2D" box="[390,526,193,216]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="A1961B2EFFD0FFF3FE4940B2B70EFF2D" box="[390,526,193,216]" italics="true" pageId="40" pageNumber="41">Agathidium</emphasis>
</taxonomicName>
from all
<taxonomicName id="54E2BCBFFFD0FFF3FFBC40ACB44DFF00" box="[115,333,222,246]" class="Magnoliopsida" family="Apocynaceae" genus="Anisotoma" kingdom="Plantae" order="Gentianales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A1961B2EFFD0FFF3FFBC40ACB5EFFF03" box="[115,239,223,246]" italics="true" pageId="40" pageNumber="41">Anisotoma</emphasis>
species
</taxonomicName>
and other genera.
</paragraph>
<caption id="C79D97B4FFDFFFFCFF5A45DCB153FA2A" ID-DOI="http://doi.org/10.5281/zenodo.5361791" ID-Zenodo-Dep="5361791" httpUri="https://zenodo.org/record/5361791/files/figure.png" pageId="39" pageNumber="40" startId="39.[149,187,1455,1476]" targetBox="[273,1080,190,1435]" targetPageId="39">
<paragraph id="935DC73CFFDFFFFCFF5A45DCB153FA2A" blockId="39.[123,1226,1455,1503]" pageId="39" pageNumber="40">
Fig. 52.
<taxonomicName id="54E2BCBFFFDFFFFCFECA45DCB747FA31" authority="Horn" authorityName="Horn" authorityYear="1880" box="[261,583,1455,1476]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sexstriatum">
<emphasis id="A1961B2EFFDFFFFCFECA45DCB704FA31" box="[261,516,1455,1476]" italics="true" pageId="39" pageNumber="40">Agathidium sexstriatum</emphasis>
Horn
</taxonomicName>
, habitus, dorsal (note: differences in angle of observation result in differences in shape and proportion of structures in this image and that in fig. 67).
</paragraph>
</caption>
<paragraph id="935DC73CFFD0FFF3FF5F408FB46AFDEE" blockId="40.[115,644,193,539]" pageId="40" pageNumber="41">Circumscription of these species is based on external characters, including color and punctation patterns, and male aedeagal shape. Although external characters are used extensively in the key, these features are somewhat less reliable than male aedeagal shape and, when possible, male specimens should be dissected and aedeagal shape compared with the illustrations for the most reliable identifications.</paragraph>
</subSubSection>
<subSubSection id="DBF894B7FFD0FFF3FF6E424DB771FDA0" box="[161,625,574,598]" pageId="40" pageNumber="41" type="key">
<paragraph id="935DC73CFFD0FFF3FF6E424DB771FDA0" blockId="40.[161,625,574,627]" box="[161,625,574,598]" pageId="40" pageNumber="41">
KEY TO
<taxonomicName id="54E2BCBFFFD0FFF3FED7424CB4F6FDA3" ID-CoL="65QQJ" authorityName="Horn" authorityYear="1880" box="[280,502,575,598]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="85" phylum="Arthropoda" rank="species" species="sexstriatum">
<emphasis id="A1961B2EFFD0FFF3FED7424CB4F6FDA3" box="[280,502,575,598]" italics="true" pageId="40" pageNumber="41">A. SEXSTRIATUM</emphasis>
</taxonomicName>
SPECIES
</paragraph>
</subSubSection>
</treatment>
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