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<document id="4D52FDCB977357438C870D2F41FB707E" ID-CLB-Dataset="46473" ID-DOI="10.11646/zootaxa.3764.3.8" ID-GBIF-Dataset="8a7a9770-4717-40c3-8c19-85dee45429c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="227547" ID-ZooBank="5DA1C9F3-F0FD-4798-BBEC-AF8EE8FD5917" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460382641431" checkinUser="plazi" docAuthor="Mahnert, Volker, Sharaf, Mostafa &amp; Aldawood, Abdulrahman S." docDate="2014" docId="0D31EF27FFECFFCCBEB1B9DEFD2B2D8E" docLanguage="en" docName="zt03764p393.pdf" docOrigin="Zootaxa 3764 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudochthonius arabicus Mahnert, n. sp." docType="treatment" docUuid="4728F304-236C-4D93-B06F-9D4035936CA1" docUuidSource="ZooBank" docVersion="11" lastPageNumber="391" masterDocId="F108975FFFEDFFC8BE26BB50FFF72F0A" masterDocTitle="Further records of pseudoscorpions (Arachnida, Pseudoscorpiones) from Saudi Arabia" masterLastPageNumber="393" masterPageNumber="387" pageNumber="388" updateTime="1698297006382" updateUser="ExternalLinkService">
<mods:mods id="6212E13DB503423B6BB6FC7093752561" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9B7C0139A1BC0BAD9FF95EE4C58FA00B">Further records of pseudoscorpions (Arachnida, Pseudoscorpiones) from Saudi Arabia</mods:title>
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<mods:namePart id="6AFA67DB3103107EACE1110E6DDEBD3E">Mahnert, Volker</mods:namePart>
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<mods:namePart id="1720C9FE86F84338A528EC3064429BD5">Sharaf, Mostafa</mods:namePart>
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<mods:namePart id="DC69D8D21F97AF2D6BCEC1E1F64153EA">Aldawood, Abdulrahman S.</mods:namePart>
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<mods:title id="4567B151E519094E13C4C23927DF3DAA">Zootaxa</mods:title>
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<mods:date id="0E1DFC3C6F373A37C53DEA09997358D5">2014</mods:date>
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<mods:number id="87DFDC5363A56B91B4AB2AFF37E7C0D0">3764</mods:number>
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<mods:identifier id="89DDFFFF66F5FCAE169593A029123C23" type="DOI">10.11646/zootaxa.3764.3.8</mods:identifier>
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<treatment id="0D31EF27FFECFFCCBEB1B9DEFD2B2D8E" ID-DOI="http://doi.org/10.5281/zenodo.6138955" ID-GBIF-Taxon="119414445" ID-Zenodo-Dep="6138955" LSID="urn:lsid:zoobank.org:act:4728F304-236C-4D93-B06F-9D4035936CA1" httpUri="http://treatment.plazi.org/id/0D31EF27FFECFFCCBEB1B9DEFD2B2D8E" lastPageId="4" lastPageNumber="391" pageId="1" pageNumber="388">
<subSubSection id="CD820DBAFFECFFC9BEB1B9DEFEFC2DC1" pageId="1" pageNumber="388" type="nomenclature">
<paragraph id="85275E31FFECFFC9BEB1B9DEFD882DAD" blockId="1.[151,639,654,715]" box="[151,639,654,679]" pageId="1" pageNumber="388">
<heading id="DE6FE95DFFECFFC9BEB1B9DEFD882DAD" bold="true" box="[151,639,654,679]" fontSize="10" level="2" pageId="1" pageNumber="388" reason="2">
<emphasis id="B7EC8223FFECFFC9BEB1B9DEFD882DAD" bold="true" box="[151,639,654,679]" pageId="1" pageNumber="388">
<taxonomicName id="429825B2FFECFFC9BEB1B9DEFDCE2DAD" ID-CoL="4NTP9" authority="Mahnert" authorityName="Mahnert" box="[151,569,654,679]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B7EC8223FFECFFC9BEB1B9DEFE3E2DAD" bold="true" box="[151,457,654,679]" italics="true" pageId="1" pageNumber="388">Pseudochthonius arabicus</emphasis>
Mahnert
</taxonomicName>
<taxonomicNameLabel id="ACDF3F58FFECFFC9BC66B9DEFD882DAD" box="[576,639,654,679]" pageId="1" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="85275E31FFECFFC9BEB1B9E2FEFC2DC1" blockId="1.[151,639,654,715]" box="[151,267,690,715]" pageId="1" pageNumber="388">
(
<figureCitation id="1DA342B4FFECFFC9BEB9B9E2FEF62DC1" box="[159,257,690,715]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1814,1836]" captionStartId-1="3.[151,250,1939,1961]" captionTargetBox-0="[187,1405,454,1787]" captionTargetBox-1="[186,1410,193,1912]" captionTargetId-0="figure@2.[181,1405,447,1790]" captionTargetId-1="figure@3.[186,1410,193,1914]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." captionText-1="FIGURE 2. Type locality of Pseudochthonius arabicus Mahnert n. sp. a, Map of Saudi Arabia, showing location of site; b, Al Bahah, Al Mukwah Dhi Ayn Archeological Village (type locality), (photograph by M. Abdel-Dayem); c, Habitat (photograph by M. Sharaf)." httpUri-0="https://zenodo.org/record/227548/files/figure.png" httpUri-1="https://zenodo.org/record/227549/files/figure.png" pageId="1" pageNumber="388">Figs 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CD820DBAFFECFFC9BEB1B9AAFBD72C3D" pageId="1" pageNumber="388" type="materials_examined">
<paragraph id="85275E31FFECFFC9BEB1B9AAFBD72C3D" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="388">
<emphasis id="B7EC8223FFECFFC9BEB1B9AAFE772C19" bold="true" box="[151,384,762,787]" pageId="1" pageNumber="388">Material examined.</emphasis>
<materialsCitation id="35F0546CFFECFFC9BFA1B9AAFD5B2C3C" ID-GBIF-Occurrence="3886338308" collectingDate="2011-05-11" collectionCode="MHNG" country="Saudi Arabia" elevation="741" latitude="20.21103" location="Dhi Ayn Archeological Village" longLatPrecision="1" longitude="41.441723" municipality="Al Mukwah" pageId="1" pageNumber="388" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahah" typeStatus="holotype">
<typeStatus id="5A23E093FFECFFC9BFA1B9AAFE182C19" box="[391,495,762,787]" pageId="1" pageNumber="388" type="holotype">Holotype</typeStatus>
<specimenCount id="939E95B8FFECFFC9BFD1B9ABFDFB2C18" box="[503,524,763,786]" pageId="1" pageNumber="388" type="male">♂</specimenCount>
(
<collectionCode id="E389C6F4FFECFFC9BC3DB9ABFD802C19" box="[539,631,763,787]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="388">MHNG</collectionCode>
),
<collectingCountry id="FD8F1EA1FFECFFC9BCAFB9AAFCE82C19" box="[649,799,762,787]" name="Saudi Arabia" pageId="1" pageNumber="388">Saudi Arabia</collectingCountry>
,
<collectingRegion id="475C90D3FFECFFC9BD0DB9AAFC6E2C19" box="[811,921,762,787]" country="Saudi Arabia" name="Al Bahah" pageId="1" pageNumber="388">Al Bahah</collectingRegion>
,
<collectingMunicipality id="6543C44BFFECFFC9BD82B9AAFBDA2C19" box="[932,1069,762,787]" pageId="1" pageNumber="388">Al Mukwah</collectingMunicipality>
,
<location id="804708EAFFECFFC9BA1EB9AAFA6F2C19" LSID="urn:lsid:plazi:treatment:0D31EF27FFECFFCCBEB1B9DEFD2B2D8E:804708EAFFECFFC9BA1EB9AAFA6F2C19" box="[1080,1432,762,787]" country="Saudi Arabia" latitude="20.21103" longLatPrecision="1" longitude="41.441723" municipality="Al Mukwah" name="Dhi Ayn Archeological Village" pageId="1" pageNumber="388" stateProvince="Al Bahah">Dhi Ayn Archeological Village</location>
,
<date id="F12678F1FFECFFC9BEB1B84FFEF22C3D" box="[151,261,799,823]" pageId="1" pageNumber="388" value="2011-05-11">
<collectingDate id="E1628119FFECFFC9BEB1B84FFEF22C3D" box="[151,261,799,823]" pageId="1" pageNumber="388" value="2011-05-11">11.v.2011</collectingDate>
</date>
,
<geoCoordinate id="E0AC38F6FFECFFC9BF37B84EFE592C3D" box="[273,430,798,823]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="388" precision="1" seconds="39.7" value="20.21103">20°12′39.7″N</geoCoordinate>
,
<geoCoordinate id="E0AC38F6FFECFFC9BF9FB84EFDA42C3D" box="[441,595,798,823]" degrees="41" direction="east" minutes="26" orientation="longitude" pageId="1" pageNumber="388" precision="1" seconds="30.2" value="41.441723">41°26′30.2″E</geoCoordinate>
,
<quantity id="4260F3D4FFECFFC9BC78B84FFD5F2C3C" box="[606,680,799,823]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="1" pageNumber="388" unit="m" value="741.0">
<elevation id="0EB5B902FFECFFC9BC78B84FFD5F2C3C" box="[606,680,799,823]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="1" pageNumber="388" unit="m" value="741.0">741 m</elevation>
</quantity>
.
</materialsCitation>
<materialsCitation id="35F0546CFFECFFC9BC95B84FFBEB2C3D" ID-GBIF-Occurrence="3886338306" box="[691,1052,798,823]" collectionCode="KSMA" pageId="1" pageNumber="388" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="5A23E093FFECFFC9BC95B84FFCE22C3D" box="[691,789,799,823]" pageId="1" pageNumber="388" type="paratype">Paratype</typeStatus>
<specimenCount id="939E95B8FFECFFC9BD3AB84FFCC62C3C" box="[796,817,799,822]" pageId="1" pageNumber="388" type="female">♀</specimenCount>
(
<collectionCode id="E389C6F4FFECFFC9BD66B84FFC612C3D" box="[832,918,799,823]" pageId="1" pageNumber="388">KSMA</collectionCode>
), same data
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CD820DBAFFECFFCABEE1B812FD9A2E82" lastPageId="2" lastPageNumber="389" pageId="1" pageNumber="388" type="description">
<paragraph id="85275E31FFECFFC9BEE1B812FE812BC9" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="388">
<emphasis id="B7EC8223FFECFFC9BEE1B812FEAE2C51" bold="true" box="[199,345,834,859]" pageId="1" pageNumber="388">Description.</emphasis>
General coloration yellowish brown, tergites III–VI with a dark median mark. Carapace 1.1 × longer than broad, slightly constricted posteriorly and there reticulated (
<figureCitation id="1DA342B4FFECFFC9BDFAB836FBD52C75" box="[988,1058,870,895]" captionStart="FIGURE 1" captionStartId="2.[151,250,1814,1836]" captionTargetBox="[187,1405,454,1787]" captionTargetId="figure@2.[181,1405,447,1790]" captionTargetPageId="2" captionText="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
a); two small eyes with distinct lenses present, about one diameter from anterior margin; epistome (
<figureCitation id="1DA342B4FFECFFC9BDB7B8DAFC242CA9" box="[913,979,906,931]" captionStart="FIGURE 1" captionStartId="2.[151,250,1814,1836]" captionTargetBox="[187,1405,454,1787]" captionTargetId="figure@2.[181,1405,447,1790]" captionTargetPageId="2" captionText="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
b) broad and dentate; 16 long setae (4 + 1 pair preocular seta/4/4/2/2); tergal chaetotaxy:
<date id="F12678F1FFECFFC9BC9CB8FFFCF12CCD" box="[698,774,943,967]" pageId="1" pageNumber="388">I–IV 4</date>
,
<date id="F12678F1FFECFFC9BD35B8FFFC842CCD" box="[787,883,942,967]" pageId="1" pageNumber="388">VI–IX 6</date>
,
<date id="F12678F1FFECFFC9BD59B8FFFC5C2CCD" box="[895,939,943,967]" pageId="1" pageNumber="388">X 4</date>
,
<date id="F12678F1FFECFFC9BD91B8FFFC1D2CCD" box="[951,1002,942,967]" pageId="1" pageNumber="388">XI 6</date>
(2 tactile setae). Manducatory process acute, with 2 setae; pedipalpal coxa 3 (one discal seta in basal third), coxa
<date id="F12678F1FFECFFC9BA20B883FBD02CE1" box="[1030,1063,979,1003]" pageId="1" pageNumber="388">I 3</date>
long setae and one tiny seta on anterior margin + 4–5 dentate (median one apparently smooth) coxal spines in single row,
<date id="F12678F1FFECFFC9BAB5B8A7FB4C2B05" box="[1171,1211,1015,1039]" pageId="1" pageNumber="388">II 4</date>
setae + 5–6 dentate (median one apparently smooth) coxal spines (
<figureCitation id="1DA342B4FFECFFC9BC82BF4AFD102B39" box="[676,743,1050,1075]" captionStart="FIGURE 1" captionStartId="2.[151,250,1814,1836]" captionTargetBox="[187,1405,454,1787]" captionTargetId="figure@2.[181,1405,447,1790]" captionTargetPageId="2" captionText="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
f),
<date id="F12678F1FFECFFC9BD29BF4BFCB72B38" box="[783,832,1051,1074]" pageId="1" pageNumber="388">III 7</date>
setae,
<date id="F12678F1FFECFFC9BDA8BF4BFC362B39" box="[910,961,1051,1075]" pageId="1" pageNumber="388">IV 8</date>
setae, intercoxal tubercle absent. Anterior genital operculum of ♂ (not observable in ♀) with 9 marginal and discal setae, genital opening of ♂ slit-like triangular (
<figureCitation id="1DA342B4FFECFFC9BF33BF32FEA72B71" box="[277,336,1122,1147]" captionStart="FIGURE 1" captionStartId="2.[151,250,1814,1836]" captionTargetBox="[187,1405,454,1787]" captionTargetId="figure@2.[181,1405,447,1790]" captionTargetPageId="2" captionText="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="1" pageNumber="388">
Fig.
<quantity id="4260F3D4FFECFFC9BF60BF32FEA72B71" box="[326,336,1122,1147]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="1" pageNumber="388" unit="g" value="1.0">1</quantity>
</figureCitation>
g), with 7–8 unmodified marginal setae on each side; sternal chaetotaxy III–XI (♂): 8 + 3 pairs suprastigmal setae/8 + 3 pairs suprastigmal setae/9 (2 lateral setae on each side tiny)/8 (lateral setae tiny)/8 (lateral setae short)/8/8/9/0.
</paragraph>
<paragraph id="85275E31FFECFFC9BEE1BF9EFDB12A59" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="388">
Chelicera (
<figureCitation id="1DA342B4FFECFFC9BF65BF9EFE7F2BED" box="[323,392,1230,1255]" captionStart="FIGURE 1" captionStartId="2.[151,250,1814,1836]" captionTargetBox="[187,1405,454,1787]" captionTargetId="figure@2.[181,1405,447,1790]" captionTargetPageId="2" captionText="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
c): hand with 4 long and 2–3 shorter, basal, accessory setae, fixed finger with 10–11 acute teeth, movable finger with 11–14 small acute teeth and one slightly isolated distal tooth; subgaleal seta in basal third of movable finger; galea tubercle-like (slightly larger in ♀ than in ♂), serrula exterior 14–17 lamellae, rallum with about 9 setae with long denticles.
</paragraph>
<paragraph id="85275E31FFECFFC9BEE1BE0EFF2D290C" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="388">
Pedipalps (
<figureCitation id="1DA342B4FFECFFC9BF65BE0EFE712A7D" box="[323,390,1374,1399]" captionStart="FIGURE 1" captionStartId="2.[151,250,1814,1836]" captionTargetBox="[187,1405,454,1787]" captionTargetId="figure@2.[181,1405,447,1790]" captionTargetPageId="2" captionText="FIGURE 1. Pseudochthonius arabicus Mahnert n. sp., holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm." httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
d, e): trochanter 1.6 (♀ 1.6) × longer than broad, femur 4.9 (♀ 5.0) ×, patella 2.3 (♀ 2.2) × longer than broad, hand 2.0 (♀ 1.9) ×, chela 6.4 (♀ 6.2) × longer than deep, fixed finger 2.2 (♀ 2.3) × longer than hand; fixed finger with 33–35 acute, erect teeth, in distal half, of alternating length and in saw-like arrangement, movable finger shorter than the fixed one, with 37–39 low retrorse teeth; trichobothrium
<emphasis id="B7EC8223FFECFFC9BA52BE9CFB782AE9" box="[1140,1167,1484,1507]" italics="true" pageId="1" pageNumber="388">ist</emphasis>
twice nearer to
<emphasis id="B7EC8223FFECFFC9BB66BE9AFA922AE9" box="[1344,1381,1482,1507]" italics="true" pageId="1" pageNumber="388">esb</emphasis>
than to
<emphasis id="B7EC8223FFECFFC9BE92BEBFFF2D290C" box="[180,218,1519,1542]" italics="true" pageId="1" pageNumber="388">est.</emphasis>
</paragraph>
<paragraph id="85275E31FFECFFC9BEE1BD43FB802979" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="388">Leg I: femur 5.1 (♀ 4.8.) × longer than deep and 1.6 × longer than patella, patella 3.4 (♀ 3.7) ×, tibia 5.0 (♀ 4.4) ×, tarsus 9.9 (♀ 9.4) × longer than deep; leg IV: femur+patella 2.4 (2.4) ×, tibia 4.0 (♀ 4.1) ×, basitarsus 3.1 (♀ 3.2) ×, telotarsus 9.9 × (♀: broken) longer than deep and 1.5 × longer than basitarsus.</paragraph>
<paragraph id="85275E31FFECFFC9BEE1BD2FFCFA2841" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="388">Measurements of ♂ (♀) in mm: Total length 1.27 (1.91). Carapace (length/breadth) 0.44/0.40 (0.48/0.44). Pedipalps (length/breadth): trochanter 0.17/0.11 (0.19/0.12), femur 0.55/0.11 (0.61/0.12), patella 0.23/0.10 (0.25/ 0.12), hand length/depth 0.24/0.12 (0.26/0.14), length of fixed finger 0.54 (0.59), of movable finger 0.50 (0.53), of chela 0.77 (0.85). Leg I (length/depth): femur 0.30/0.06 (0.32/0.07), patella 0.19/0.05 (0.20/0.05), tibia 0.19/0.04 (0.17/0.04), tarsus 0.31/0.03 (0.31/0.03); leg IV: femur+patella 0.49/0.21 (0.50/0.21), tibia 0.31/0.08 (0.32/0.08), basitarsus 0.17/0.06 (0.18/0.06), telotarsus 0.33/0.03 (-).</paragraph>
<paragraph id="85275E31FFECFFCABEE1BC06FADF2E4A" blockId="1.[151,1436,762,2011]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="389" pageId="1" pageNumber="388">
<emphasis id="B7EC8223FFECFFC9BEE1BC06FECA2865" bold="true" box="[199,317,1878,1903]" pageId="1" pageNumber="388">Affinities.</emphasis>
The known geographical distribution of the 29 species of
<taxonomicName id="429825B2FFECFFC9BDECBC06FAD32865" authority="Balzan, 1892" authorityName="Balzan" authorityYear="1892" box="[970,1316,1878,1903]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
<emphasis id="B7EC8223FFECFFC9BDECBC06FB7E2865" box="[970,1161,1878,1903]" italics="true" pageId="1" pageNumber="388">Pseudochthonius</emphasis>
Balzan, 1892
</taxonomicName>
(28 extant and one fossil species) was previously restricted to sub-Saharan Africa and South
<collectingCountry id="FD8F1EA1FFECFFC9BA1ABC2AFB6B2899" box="[1084,1180,1914,1939]" name="United States of America" pageId="1" pageNumber="388">America</collectingCountry>
north to
<collectingCountry id="FD8F1EA1FFECFFC9BB23BC2AFAA92899" box="[1285,1374,1914,1939]" name="Mexico" pageId="1" pageNumber="388">Mexico</collectingCountry>
. Five species have been reported from Africa, and the new species seems to be morphologically similar to those from central and eastern Africa. Whereas trichobothrium
<emphasis id="B7EC8223FFECFFC9BCC1BC94FCF528D1" box="[743,770,1988,2011]" italics="true" pageId="1" pageNumber="388">ist</emphasis>
is approximately halfway between
<emphasis id="B7EC8223FFECFFC9BABFBC92FB4928D1" box="[1177,1214,1986,2011]" italics="true" pageId="1" pageNumber="388">esb</emphasis>
and
<emphasis id="B7EC8223FFECFFC9BADFBC94FAEF28D1" box="[1273,1304,1988,2011]" italics="true" pageId="1" pageNumber="388">est</emphasis>
in
<taxonomicName id="429825B2FFECFFCABB19BC94FEC32FBA" authority="Vachon, 1941" authorityName="Vachon" authorityYear="1941" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" lastPageId="2" lastPageNumber="389" order="Pseudoscorpiones" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="billae">
<emphasis id="B7EC8223FFECFFC9BB19BC94FA6B28D1" box="[1343,1436,1986,2011]" italics="true" pageId="1" pageNumber="388">P. billae</emphasis>
Vachon, 1941
</taxonomicName>
(
<collectingCountry id="FD8F1EA1FFEFFFCABF63BBC8FE382FBA" box="[325,463,152,176]" name="Ivory Coast" pageId="2" pageNumber="389">Ivory Coast</collectingCountry>
) and
<taxonomicName id="429825B2FFEFFFCABC35BBC9FC912FBA" authority="Beier, 1959" authorityName="Beier" authorityYear="1959" box="[531,870,151,176]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="B7EC8223FFEFFFCABC35BBC9FD8A2FBA" box="[531,637,151,176]" italics="true" pageId="2" pageNumber="389">P. leleupi</emphasis>
Beier, 1959 (
<collectingCountry id="FD8F1EA1FFEFFFCABD38BBC8FCA92FBA" box="[798,862,152,176]" name="Democratic Republic of the Congo" pageId="2" pageNumber="389">Zaïre</collectingCountry>
)
</taxonomicName>
,
<emphasis id="B7EC8223FFEFFFCABD52BBC9FC782FBA" box="[884,911,153,176]" italics="true" pageId="2" pageNumber="389">ist</emphasis>
originates&gt;2× closer to
<emphasis id="B7EC8223FFEFFFCABA93BBC7FB2D2FBA" box="[1205,1242,151,176]" italics="true" pageId="2" pageNumber="389">esb</emphasis>
than to
<emphasis id="B7EC8223FFEFFFCABB18BBC9FAAA2FBA" box="[1342,1373,153,176]" italics="true" pageId="2" pageNumber="389">est</emphasis>
in
<taxonomicName id="429825B2FFEFFFCABBA1BBC9FE2A2FDE" authority="Beier, 1959" authorityName="Beier" authorityYear="1959" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="congicus">
<emphasis id="B7EC8223FFEFFFCABBA1BBC9FF0C2FDE" italics="true" pageId="2" pageNumber="389">P. congicus</emphasis>
Beier, 1959 (
<collectingCountry id="FD8F1EA1FFEFFFCABFB1BBECFE222FDE" box="[407,469,188,212]" name="Democratic Republic of the Congo" pageId="2" pageNumber="389">Zaïre</collectingCountry>
)
</taxonomicName>
and
<taxonomicName id="429825B2FFEFFFCABC31BBEDFC602FDE" authority="Mahnert, 1986" authorityName="Mahnert" authorityYear="1986" box="[535,919,188,213]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="perreti">
<emphasis id="B7EC8223FFEFFFCABC31BBEDFD882FDE" box="[535,639,189,212]" italics="true" pageId="2" pageNumber="389">P. perreti</emphasis>
Mahnert, 1986 (
<collectingCountry id="FD8F1EA1FFEFFFCABD66BBECFC782FDE" box="[832,911,188,212]" name="Kenya" pageId="2" pageNumber="389">Kenya</collectingCountry>
)
</taxonomicName>
.
<taxonomicName id="429825B2FFEFFFCABD82BBECFB392FDF" box="[932,1230,188,213]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="B7EC8223FFEFFFCABD82BBECFB392FDF" box="[932,1230,188,213]" italics="true" pageId="2" pageNumber="389">Pseudochthonius arabicus</emphasis>
</taxonomicName>
is distinguishable from the latter two species by its much larger size (chela length
<quantity id="4260F3D4FFEFFFCABD7BBB8FFBF62FFD" box="[861,1025,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.5" metricValueMin="7.7" pageId="2" pageNumber="389" unit="mm" value="0.81" valueMax="0.85" valueMin="0.77">0.77–0.85 mm</quantity>
, versus
<quantity id="4260F3D4FFEFFFCABA7CBBB0FB492FFD" box="[1114,1214,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="389" unit="mm" value="0.38">0.38 mm</quantity>
in
<taxonomicName id="429825B2FFEFFFCABAC6BBB1FAB02FF2" box="[1248,1351,224,248]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="perreti">
<emphasis id="B7EC8223FFEFFFCABAC6BBB1FB192FF2" box="[1248,1262,225,248]" italics="true" pageId="2" pageNumber="389">P</emphasis>
.
<emphasis id="B7EC8223FFEFFFCABADABBB1FAB02FF2" box="[1276,1351,225,248]" italics="true" pageId="2" pageNumber="389">perreti</emphasis>
</taxonomicName>
or
<quantity id="4260F3D4FFEFFFCABB4DBB8FFF342E16" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="389" unit="mm" value="0.53">0.53 mm</quantity>
in
<taxonomicName id="429825B2FFEFFFCABECEBA55FE9E2E16" box="[232,361,261,284]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="congicus">
<emphasis id="B7EC8223FFEFFFCABECEBA55FF012E16" box="[232,246,261,284]" italics="true" pageId="2" pageNumber="389">P</emphasis>
.
<emphasis id="B7EC8223FFEFFFCABF23BA55FE9E2E16" box="[261,361,261,284]" italics="true" pageId="2" pageNumber="389">congicus</emphasis>
</taxonomicName>
). Furthermore it differs from
<taxonomicName id="429825B2FFEFFFCABC9DBA55FCCB2E16" box="[699,828,261,284]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="congicus">
<emphasis id="B7EC8223FFEFFFCABC9DBA55FD3E2E16" box="[699,713,261,284]" italics="true" pageId="2" pageNumber="389">P</emphasis>
.
<emphasis id="B7EC8223FFEFFFCABCFEBA55FCCB2E16" box="[728,828,261,284]" italics="true" pageId="2" pageNumber="389">congicus</emphasis>
</taxonomicName>
in having a distinct epistome (lacking in
<taxonomicName id="429825B2FFEFFFCABB34BA55FA642E16" box="[1298,1427,261,284]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="congicus">
<emphasis id="B7EC8223FFEFFFCABB34BA55FAD72E16" box="[1298,1312,261,284]" italics="true" pageId="2" pageNumber="389">P</emphasis>
.
<emphasis id="B7EC8223FFEFFFCABB09BA55FA642E16" box="[1327,1427,261,284]" italics="true" pageId="2" pageNumber="389">congicus</emphasis>
</taxonomicName>
) and from
<taxonomicName id="429825B2FFEFFFCABF22BA79FE9C2E4A" box="[260,363,296,320]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="perreti">
<emphasis id="B7EC8223FFEFFFCABF22BA79FEE52E4A" box="[260,274,297,320]" italics="true" pageId="2" pageNumber="389">P</emphasis>
.
<emphasis id="B7EC8223FFEFFFCABF06BA79FE9C2E4A" box="[288,363,297,320]" italics="true" pageId="2" pageNumber="389">perreti</emphasis>
</taxonomicName>
by the presence of distinct, lensed eyes (eyes indistinct, without lenses, in
<taxonomicName id="429825B2FFEFFFCABA97BA79FAEF2E4A" box="[1201,1304,296,320]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="perreti">
<emphasis id="B7EC8223FFEFFFCABA97BA79FB482E4A" box="[1201,1215,297,320]" italics="true" pageId="2" pageNumber="389">P</emphasis>
.
<emphasis id="B7EC8223FFEFFFCABAEBBA79FAEF2E4A" box="[1229,1304,297,320]" italics="true" pageId="2" pageNumber="389">perreti</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="85275E31FFEFFFCABEE1BA1CFD9A2E82" blockId="2.[151,1436,151,392]" pageId="2" pageNumber="389">The presence of this genus in the southwestern mountains of the Arabian Peninsula confirms the affinity of this region with the Afrotropical faunal realm.</paragraph>
</subSubSection>
<caption id="D1E70EB9FFEFFFCABEB1BC46FDD42860" httpUri="https://zenodo.org/record/227548/files/figure.png" pageId="2" pageNumber="389" targetBox="[187,1405,454,1787]" targetPageId="2">
<paragraph id="85275E31FFEFFFCABEB1BC46FDD42860" blockId="2.[151,1436,1814,1898]" pageId="2" pageNumber="389">
<emphasis id="B7EC8223FFEFFFCABEB1BC46FEE12826" bold="true" box="[151,278,1814,1836]" pageId="2" pageNumber="389">FIGURE 1.</emphasis>
<taxonomicName id="429825B2FFEFFFCABF06BC46FD672826" authority="Mahnert" authorityName="Mahnert" box="[288,656,1814,1836]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B7EC8223FFEFFFCABF06BC46FDC72826" box="[288,560,1814,1836]" italics="true" pageId="2" pageNumber="389">Pseudochthonius arabicus</emphasis>
Mahnert
</taxonomicName>
<emphasis id="B7EC8223FFEFFFCABCBFBC47FD212826" bold="true" box="[665,726,1815,1836]" pageId="2" pageNumber="389">
<taxonomicNameLabel id="ACDF3F58FFEFFFCABCBFBC47FD212826" box="[665,726,1815,1836]" pageId="2" pageNumber="389" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype. a, carapace (most setae fallen off); b, epistome; c, right chelicera; d, left pedipalp, minus chela, dorsal view; e, left chela, lateral view; f, left coxae I and II with details of coxal spines; g, genital opening. Scale units 0.1 mm.
</paragraph>
</caption>
<caption id="D1E70EB9FFEEFFCBBEB1BCC3FEDC28EC" httpUri="https://zenodo.org/record/227549/files/figure.png" pageId="3" pageNumber="390" targetBox="[186,1410,193,1912]" targetPageId="3">
<paragraph id="85275E31FFEEFFCBBEB1BCC3FEDC28EC" blockId="3.[151,1435,1939,2022]" pageId="3" pageNumber="390">
<emphasis id="B7EC8223FFEEFFCBBEB1BCC3FEE528A2" bold="true" box="[151,274,1939,1961]" pageId="3" pageNumber="390">FIGURE 2.</emphasis>
Type locality of
<taxonomicName id="429825B2FFEEFFCBBFE6BCC3FCDF28A3" authority="Mahnert" authorityName="Mahnert" box="[448,808,1939,1961]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="B7EC8223FFEEFFCBBFE6BCC3FD3B28A3" box="[448,716,1939,1961]" italics="true" pageId="3" pageNumber="390">Pseudochthonius arabicus</emphasis>
Mahnert
</taxonomicName>
<emphasis id="B7EC8223FFEEFFCBBD09BCC4FC9F28A3" bold="true" box="[815,872,1940,1961]" pageId="3" pageNumber="390">
<taxonomicNameLabel id="ACDF3F58FFEEFFCBBD09BCC4FC9F28A3" box="[815,872,1940,1961]" pageId="3" pageNumber="390" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
a, Map of Saudi Arabia, showing location of site; b, Al Bahah, Al Mukwah Dhi Ayn Archeological Village (type locality), (photograph by M. Abdel-Dayem); c, Habitat (photograph by M. Sharaf).
</paragraph>
</caption>
<subSubSection id="CD820DBAFFE9FFCCBEE1BBC7FD2B2D8E" pageId="4" pageNumber="391" type="discussion">
<paragraph id="85275E31FFE9FFCCBEE1BBC7FDD52E4A" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="391">
<emphasis id="B7EC8223FFE9FFCCBEE1BBC7FECC2FBA" bold="true" box="[199,315,151,176]" pageId="4" pageNumber="391">Remarks.</emphasis>
The
<typeStatus id="5A23E093FFE9FFCCBF5DBBC8FE5C2FBA" box="[379,427,152,176]" pageId="4" pageNumber="391">type</typeStatus>
locality (
<figureCitation id="1DA342B4FFE9FFCCBC3DBBC7FD962FBA" box="[539,609,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1939,1961]" captionTargetBox="[186,1410,193,1912]" captionTargetId="figure@3.[186,1410,193,1914]" captionTargetPageId="3" captionText="FIGURE 2. Type locality of Pseudochthonius arabicus Mahnert n. sp. a, Map of Saudi Arabia, showing location of site; b, Al Bahah, Al Mukwah Dhi Ayn Archeological Village (type locality), (photograph by M. Abdel-Dayem); c, Habitat (photograph by M. Sharaf)." httpUri="https://zenodo.org/record/227549/files/figure.png" pageId="4" pageNumber="391">Fig. 2</figureCitation>
a, b) is a national archaeological site that is completely surrounded by mountains. It forms part of the upper Tihama Territory, belonging to Al Bahah Province. The locality has a high diversity of native vegetation, but numerous fruits are cultivated (e.g. banana, date palm and
<taxonomicName id="429825B2FFE9FFCCBAECBBB1FAFF2FF2" box="[1226,1288,225,248]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="391" phylum="Mollusca" rank="genus">
<emphasis id="B7EC8223FFE9FFCCBAECBBB1FAFF2FF2" box="[1226,1288,225,248]" italics="true" pageId="4" pageNumber="391">Ficus</emphasis>
</taxonomicName>
), along with alfalfa, and lemon orchards have been planted. There are several permanent drainages and localized soil moisture is higher than in surrounding regions.
</paragraph>
<paragraph id="85275E31FFE9FFCCBEE1BA1CFD2B2D8E" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="391">
The
<typeStatus id="5A23E093FFE9FFCCBEDDBA1CFEAA2E6F" box="[251,349,332,357]" pageId="4" pageNumber="391" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="429825B2FFE9FFCCBFE0BA1DFDB12E6F" box="[454,582,332,357]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="B7EC8223FFE9FFCCBFE0BA1DFDB12E6F" box="[454,582,332,357]" italics="true" pageId="4" pageNumber="391">P. arabicus</emphasis>
</taxonomicName>
was found under a stone next to a large, older
<taxonomicName id="429825B2FFE9FFCCBA41BA1DFB522E6E" box="[1127,1189,333,356]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="391" phylum="Mollusca" rank="genus">
<emphasis id="B7EC8223FFE9FFCCBA41BA1DFB522E6E" box="[1127,1189,333,356]" italics="true" pageId="4" pageNumber="391">Ficus</emphasis>
</taxonomicName>
tree, close to a small stream that is part of an agricultural irrigation ditch system (
<figureCitation id="1DA342B4FFE9FFCCBD6BBA3FFC652E82" box="[845,914,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1939,1961]" captionTargetBox="[186,1410,193,1912]" captionTargetId="figure@3.[186,1410,193,1914]" captionTargetPageId="3" captionText="FIGURE 2. Type locality of Pseudochthonius arabicus Mahnert n. sp. a, Map of Saudi Arabia, showing location of site; b, Al Bahah, Al Mukwah Dhi Ayn Archeological Village (type locality), (photograph by M. Abdel-Dayem); c, Habitat (photograph by M. Sharaf)." httpUri="https://zenodo.org/record/227549/files/figure.png" pageId="4" pageNumber="391">Fig. 2</figureCitation>
c). Two ant (
<taxonomicName id="429825B2FFE9FFCCBA08BA3FFB462E82" box="[1070,1201,367,392]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
) species were found with the new pseudoscorpion:
<taxonomicName id="429825B2FFE9FFCCBFD1BAC4FC632EA6" authority="Nylander, 1849" authorityName="Nylander" authorityYear="1849" box="[503,916,404,429]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidula">
<emphasis id="B7EC8223FFE9FFCCBFD1BAC4FD3E2EA7" box="[503,713,404,429]" italics="true" pageId="4" pageNumber="391">Pheidole pallidula</emphasis>
(Nylander, 1849)
</taxonomicName>
and
<taxonomicName id="429825B2FFE9FFCCBDF6BAC5FA6F2EA6" authority="Emery, 1877" authorityName="Emery" authorityYear="1877" box="[976,1432,404,429]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="sericeiventre">
<emphasis id="B7EC8223FFE9FFCCBDF6BAC5FB0C2EA6" box="[976,1275,405,428]" italics="true" pageId="4" pageNumber="391">Tetramorium sericeiventre</emphasis>
Emery, 1877
</taxonomicName>
. Additionally, several other myrmecophilous arthropods were collected in the same area, including:
<taxonomicName id="429825B2FFE9FFCCBEB1BA8CFD622EFE" authority="Hlavac, 2013" authorityName="Hlavac" authorityYear="2013" box="[151,661,476,501]" class="Insecta" family="Staphylinidae" genus="Centrophthalmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="subtilissimus">
<emphasis id="B7EC8223FFE9FFCCBEB1BA8CFE042EFF" box="[151,499,476,501]" italics="true" pageId="4" pageNumber="391">Centrophthalmus subtilissimus</emphasis>
Hlaváč, 2013
</taxonomicName>
(
<taxonomicName id="429825B2FFE9FFCCBC80BA8CFCD02EFF" box="[678,807,476,501]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="429825B2FFE9FFCCBD12BA8CFC482EFF" box="[820,959,476,501]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
);
<taxonomicName id="429825B2FFE9FFCCBDF3BA8CFA842EFE" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[981,1395,476,501]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B7EC8223FFE9FFCCBDF3BA8CFB382EFE" box="[981,1231,476,501]" italics="true" pageId="4" pageNumber="391">Pinophilinus spinosus</emphasis>
Assing, 2013
</taxonomicName>
; an undescribed
<taxonomicName id="429825B2FFE9FFCCBF09BAAFFE5D2D12" box="[303,426,511,536]" class="Insecta" family="Staphylinidae" genus="Peltodonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B7EC8223FFE9FFCCBF09BAAFFE5D2D12" box="[303,426,511,536]" italics="true" pageId="4" pageNumber="391">Peltodonia</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="429825B2FFE9FFCCBFD4BAAFFD842D12" box="[498,627,511,536]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="429825B2FFE9FFCCBCA1BAAFFCD52D12" box="[647,802,511,536]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
) and unidentified species of the family
<taxonomicName id="429825B2FFE9FFCCBB07BAAFFEDF2D37" authority="(Coleoptera)" baseAuthorityName="Coleoptera" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="family">Anthicidae (Coleoptera)</taxonomicName>
. The
<typeStatus id="5A23E093FFE9FFCCBF48B975FE392D37" box="[366,462,549,573]" pageId="4" pageNumber="391" type="paratype">paratype</typeStatus>
female was collected by sifting leaf litter under banana in company with the ants
<taxonomicName id="429825B2FFE9FFCCBEB1B917FD112D6A" authority="Collingwood &amp; Van Harten, 2001" authorityName="Collingwood &amp; Van Harten" authorityYear="2001" box="[151,742,583,608]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="B7EC8223FFE9FFCCBEB1B917FE962D6A" box="[151,353,583,608]" italics="true" pageId="4" pageNumber="391">Carebara arabica</emphasis>
Collingwood &amp; Van Harten, 2001
</taxonomicName>
,
<taxonomicName id="429825B2FFE9FFCCBCD7B919FB1D2D6A" authority="Sharaf &amp; Aldawood, 2012" authorityName="Sharaf &amp; Aldawood" authorityYear="2012" box="[753,1258,583,608]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B7EC8223FFE9FFCCBCD7B919FC4F2D6A" box="[753,952,583,608]" italics="true" pageId="4" pageNumber="391">Tapinoma wilsoni</emphasis>
Sharaf &amp; Aldawood, 2012
</taxonomicName>
,
<taxonomicName id="429825B2FFE9FFCCBAD3B917FE9A2D8E" authority="Forel, 1881" authorityName="Forel" authorityYear="1881" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B7EC8223FFE9FFCCBAD3B917FF142D8E" italics="true" pageId="4" pageNumber="391">Cardiocondyla emeryi</emphasis>
Forel, 1881
</taxonomicName>
,
<taxonomicName id="429825B2FFE9FFCCBF5FB93DFDF22D8E" box="[377,517,621,644]" class="Insecta" family="Formicidae" genus="Hypoponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B7EC8223FFE9FFCCBF5FB93DFDF22D8E" box="[377,517,621,644]" italics="true" pageId="4" pageNumber="391">Hypoponera</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="429825B2FFE9FFCCBC44B93DFD422D8E" box="[610,693,621,644]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B7EC8223FFE9FFCCBC44B93DFD422D8E" box="[610,693,621,644]" italics="true" pageId="4" pageNumber="391">Messor</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</treatment>
</document>
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