275 lines
30 KiB
XML
275 lines
30 KiB
XML
<document id="0E59B391925E08D9AFA87876F11E8823" ID-CLB-Dataset="28924" ID-DOI="http://dx.doi.org/10.3897/zookeys.790.26894" ID-GBIF-Dataset="a0fc65ba-3abf-4f9a-8b0e-875828e59b69" ID-PMC="PMC6198030" ID-Pensoft-Pub="1313-2970-790-35" ID-PubMed="30364707" ID-ZooBank="EF3342CA4E9C4EF2ACDD4768026D1967" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-790-35" ModsDocOrigin="ZooKeys 790" ModsDocTitle="A morphological re-evaluation of Pachyseiushumeralis Berlese, 1910 (Acari, Mesostigmata, Pachylaelapidae)" checkinTime="1539655234588" checkinUser="pensoft" docAuthor="Masan, Peter" docDate="2018" docId="E772BDFE6DD9C38ECCE8DD5A486371E3" docLanguage="en" docName="ZooKeys 790: 35-44" docOrigin="ZooKeys 790" docSource="http://dx.doi.org/10.3897/zookeys.790.26894" docTitle="Pachyseius subhumeralis Masan, 2018, sp. n." docType="treatment" docUuid="3B4BDFFF-BE84-4FF9-B2E0-91E053317BC8" docUuidSource="ZooBank" docVersion="7" lastPageNumber="41" masterDocId="8A23FF9A2679FFD63662FFD77C28FFE9" masterDocTitle="A morphological re-evaluation of Pachyseiushumeralis Berlese, 1910 (Acari, Mesostigmata, Pachylaelapidae)" masterLastPageNumber="44" masterPageNumber="35" pageNumber="38" updateTime="1701384504147" updateUser="ExternalLinkService">
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<mods:title id="942FB6B399FE414813DCE164535D7B27">A morphological re-evaluation of Pachyseiushumeralis Berlese, 1910 (Acari, Mesostigmata, Pachylaelapidae)</mods:title>
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<mods:namePart id="537856E1AEE237EED35E3022A0D23E40">Masan, Peter</mods:namePart>
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<mods:title id="3C8FD5D43D07D111347F08D8BC002766">ZooKeys</mods:title>
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<mods:part id="B655E9717B11EF6EA53A00462D103B7C">
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<mods:date id="3371E3A990F2BA613E507D122367A70B">2018</mods:date>
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<mods:number id="D19BADF6DFA005C2920584E3FC93BD39">790</mods:number>
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<mods:url id="3F8EE19C3F3F328AFD5077141FD7314D">http://dx.doi.org/10.3897/zookeys.790.26894</mods:url>
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<mods:classification id="635A18FF4A05AB25AB3309B0CD665A26">journal article</mods:classification>
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<mods:identifier id="749D8EFF08EA7E53CDEF56733F93A753" type="DOI">http://dx.doi.org/10.3897/zookeys.790.26894</mods:identifier>
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<treatment id="E772BDFE6DD9C38ECCE8DD5A486371E3" ID-GBIF-Taxon="148771559" LSID="urn:lsid:zoobank.org:act:3B4BDFFF-BE84-4FF9-B2E0-91E053317BC8" httpUri="http://treatment.plazi.org/id/E772BDFE6DD9C38ECCE8DD5A486371E3" lastPageId="6" lastPageNumber="41" pageId="3" pageNumber="38">
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<subSubSection id="D3EABE9BF27963602B87A14F5B0F8FDC" pageId="3" pageNumber="38" type="nomenclature">
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<paragraph id="1EC0B4687A2681C17E108040FF8B40CB" pageId="3" pageNumber="38">
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<taxonomicName id="DF8FDF19E9D526BB35D76C493C4FADB1" ID-CoL="8Y7NV" LSID="http://zoobank.org/3B4BDFFF-BE84-4FF9-B2E0-91E053317BC8" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius subhumeralis" order="Mesostigmata" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="subhumeralis">
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<pageBreakToken id="FDEE97F875273A1AE365730DC8CC8CA7" pageId="3" pageNumber="38" start="start">Pachyseius</pageBreakToken>
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subhumeralis
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</taxonomicName>
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<taxonomicNameLabel id="297E91651C952015F7A49013054AF5F3" pageId="3" pageNumber="38">sp. n.</taxonomicNameLabel>
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Figures 2A, 2B, 3B, 4B, 5B, 6B, 7B
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</paragraph>
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</subSubSection>
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<subSubSection id="6E6B307E7FC5B7C66D2BC03208006A65" pageId="4" pageNumber="39" type="reference_group">
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<paragraph id="E939938F28520FE40F168BA5314FB39F" pageId="4" pageNumber="39">
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<taxonomicName id="BE9965BFD6F8A2129F8E7644FF62ADD4" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius humeralis" order="Mesostigmata" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="humeralis">
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<pageBreakToken id="6D08200F62695D488DA4CF3D5BA44199" pageId="4" pageNumber="39" start="start">Pachyseius</pageBreakToken>
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humeralis
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</taxonomicName>
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:
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<bibRefCitation id="60AB1A28031ED50EE6FCAEEC9F13F8AD" author="Hyatt, KH" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="7" pageNumber="42" url="https://doi.org/10.1080/00222935608655716" year="1956">Hyatt 1956</bibRefCitation>
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: 4;
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<bibRefCitation id="9D60149ECDE04C21803BAE2D6103D271" author="Karg, W" journalOrPublisher="Die Tierwelt Deutschlands" pageId="7" pageNumber="42" pagination="1 - 475" title="Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben." volume="59" year="1971">Karg 1971</bibRefCitation>
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: 141;
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<bibRefCitation id="D0787D80F2DA4916C50F320D62CFA8AC" author="Solomon, L" journalOrPublisher="Revue Roumaine de Biologie, Serie de Biologie Animale" pageId="7" pageNumber="42" pagination="99 - 102" title="A new Pachyseius species (Acari: Mesostigmata) and a new one for the Romanian fauna." volume="27" year="1982">Solomon 1982</bibRefCitation>
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: 102;
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<bibRefCitation id="88CBF15C70DE18651F5BD47E7A21A707" author="Lapina, IM" journalOrPublisher="Zinatne Publishing House, Riga" pageId="7" pageNumber="42" title="Gamasina mites of Latvia." year="1988">Lapina 1988</bibRefCitation>
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: 178;
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<bibRefCitation id="63F49EE52347F9579310A423B132634D" author="Karg, W" journalOrPublisher="Die Tierwelt Deutschlands" pageId="7" pageNumber="42" pagination="1 - 523" title="Acari (Acarina), Milben. Parasitiformes (Anactinochaeta). Cohors Gamasina Leach. Raubmilben. 2. Ueberarbeitete Auflage." volume="59" year="1993">Karg 1993</bibRefCitation>
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: 116;
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<bibRefCitation id="B8132B7DEB068E68E6E3E162A477E4F9" author="Masan, P" journalOrPublisher="NOI Press, Bratislava" pageId="7" pageNumber="42" title="A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea)." year="2007">
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<normalizedToken id="B98863E0DBFF17B85BC52599F3AA1B28" originalValue="Mašán">Masan</normalizedToken>
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2007
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</bibRefCitation>
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||
: 32, 210.
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</paragraph>
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</subSubSection>
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<subSubSection id="60B810187A257748A2B36A50DD3BF4DE" lastPageId="5" lastPageNumber="40" pageId="4" pageNumber="39" type="materials_examined">
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<paragraph id="E4032B6646BEA166B8A4326D4E30E1DE" pageId="4" pageNumber="39">Material examined.</paragraph>
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<paragraph id="64FF86656A6B1C18EDF6633456F727BC" pageId="4" pageNumber="39">
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Type material. Holotype female: Slovakia,
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<normalizedToken id="98C6573527F82D7557A2EF15E9A82E07" originalValue="Trnavská">Trnavska</normalizedToken>
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Pahorkatina Highland,
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<normalizedToken id="224358DD49FF93415B5385078244A49B" originalValue="Štefanová">Stefanova</normalizedToken>
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Village,
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<normalizedToken id="4D3E3A32156E4D606960BF5C09C6B6C7" originalValue="Dubník">Dubnik</normalizedToken>
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Forest, oak forest (with
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<taxonomicName id="18298833731F7D50587F027EF397A4F9" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
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spp.), leaf litter and soil detritus, altitude 250 m, June 18, 1997, leg. P.
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<normalizedToken id="25A8FF49034B3575A63836BA7B1ADB8B" originalValue="Mašán">Masan</normalizedToken>
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. Paratypes: 2 females, with the same collection data as in holotype; seven females, Slovakia,
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<normalizedToken id="C7C3185DEB352DC92ECA127189570F7A" originalValue="Malé">Male</normalizedToken>
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Karpaty Mts., Bratislava Capital,
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<normalizedToken id="B6E90B9416C7F625FCCC7D475E41DB3B" originalValue="Železná">Zelezna</normalizedToken>
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Studienka Forest, beech forest (with
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<taxonomicName id="70E55DD3989AE48A21AEA9119F068233" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
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), leaf litter and soil detritus, altitude 370 m, April 27, 1993, leg. P.
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<normalizedToken id="C4944C0734742F42ADB41E1332B44728" originalValue="Mašán">Masan</normalizedToken>
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; 17 females, Bratislava Capital,
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<normalizedToken id="FC903B35EECC59D0416C3131261228F7" originalValue="Devín">Devin</normalizedToken>
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Settlement,
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<normalizedToken id="4B6B91B22D3FDE0925B71FC3F993787F" originalValue="Devínska">Devinska</normalizedToken>
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Kobyla Mt., oak forest (with
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<taxonomicName id="B7FA68C65472E241A775F75BC37167B8" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
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spp.), leaf litter and soil detritus; altitude 270 m, July 30, 1997, leg. P.
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<normalizedToken id="D40DAC40435068E12343B1CC9499988D" originalValue="Mašán">Masan</normalizedToken>
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. All these specimens were previously published as
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<taxonomicName id="4E10EE7E79927AD621B39DA27C223259" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius humeralis" order="Mesostigmata" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="humeralis">Pachyseius humeralis</taxonomicName>
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by
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<bibRefCitation id="9BC37DDBF333584D21DDA428C3359018" author="Masan, P" journalOrPublisher="NOI Press, Bratislava" pageId="7" pageNumber="42" title="A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea)." year="2007">
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<normalizedToken id="298D7A2B933B2BC50180381B76EFC0CB" originalValue="Mašán">Masan</normalizedToken>
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(2007)
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</bibRefCitation>
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, and are deposited at the Institute of Zoology, Slovak Academy of Sciences, Bratislava.
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</paragraph>
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<paragraph id="6E7744BEE755201313745EE50F8FD689" lastPageId="5" lastPageNumber="40" pageId="4" pageNumber="39">
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Non-type material. Bulgaria: one female, Shumen Plateau Natural Park, Shumen City, Bukaka Reserve, old beech forest (
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<taxonomicName id="7E2E5C9E98E3C414FBC0EE6893F42BCC" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
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) with admixed hornbeam (
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<taxonomicName id="3FE0EF94B213C4C7F8590C4D192D0F0C" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="betulus">Carpinus betulus</taxonomicName>
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), leaf litter and soil detritus, altitude 500 m, October 23, 2007, leg. I.
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<normalizedToken id="E2390A026912CBEBF6144B0F7DFCF5C4" originalValue="Mihál">Mihal</normalizedToken>
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. Czech Republic: two females, with unknown collection data. France: two females, Alpes Cottiennes Mts., Arvieux Village, Gorges du Guil Canyon, pine forest (
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<taxonomicName id="67927D6B038CBBFE9A2A36226B97367D" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="genus">Pinus</taxonomicName>
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sp.) on a slope with loose rock debris (limestone), humid needle litter with tussocks of grass and mushrooms, altitude 1,180 m, June 11, 2007, leg. P.
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<normalizedToken id="38CEF1A4134C0E403518FB28A1F23BF1" originalValue="Fenďa">Fenďa</normalizedToken>
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. Germany: three females, Bavaria, Bavarian Prealps Mts., Flintsbach am Inn Village, St.
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<normalizedToken id="B0E70E9A7DC58CDD370090748FE5F2DE" originalValue="Peter’s">Peter's</normalizedToken>
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Abbey on the Madron (
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<normalizedToken id="F39F5BCED43B031E893035C645E18568" originalValue="“Peterskirchlein”">"Peterskirchlein"</normalizedToken>
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), broadleaved deciduous forest predominated by beech (
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<taxonomicName id="ECB626EBFEA72864F30EAEB7AE7AF2D5" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
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), humid soil detritus under a deep layer of leaf-fall, altitude 600 m, April 25, 2007, leg. P.
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<normalizedToken id="5969D991F135FD413E562CCD15ECDC08" originalValue="Mašán">Masan</normalizedToken>
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. Hungary: six females, with unknown collection data. Italy: four females, Lombardy Region, Bergamo Province, Bergamasque Alps and Prealps Mts., Zambla Alta Village, near to Zambla Pass, spruce forest (
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<taxonomicName id="BB66CEDC4D9F62C06F8273E143602033" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea abies" order="Pinales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="abies">Picea abies</taxonomicName>
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) with admixed beech (
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<taxonomicName id="BE73779123BDA43D9B480CEF26CBDDFF" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
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), needle litter and soil detritus with decomposed wood substrate, altitude 1,170 m, May 13, 2015, leg. P.
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<normalizedToken id="D619C4F39E65F89D536867F6B5043E36" originalValue="Mašán">Masan</normalizedToken>
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; three females, Tuscany Region, Florence City, Villa Camerata Hostel, park, broadleaved deciduous wood, leaf litter, altitude 75 m, April 23, 2009, leg. P.
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<normalizedToken id="3C1F9B78EB7741739E5CA7243B058FDC" originalValue="Mašán">Masan</normalizedToken>
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; two females, Florence City, Parco delle Cascine Gardens, broad-leaved deciduous wood, leaf litter with soil and wood detritus, altitude 45 m, May 23, 2006, leg. P.
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<normalizedToken id="0C01A812205F1B491B9A743E389117E5" originalValue="Mašán">Masan</normalizedToken>
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. Poland: one female, with unknown collection data. Romania: one female, Transylvania Region, Apuseni Mts. (Gilău Mts. in Bihor Massif), Cluj County, Turda Town, Turda Gorge, near to
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<normalizedToken id="44FA167F100E00DEAF159C8F47E33CD7" originalValue="Peştera">Pestera</normalizedToken>
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<normalizedToken id="EBD9C9F3D0A508B7AE2503FCBD97E483" originalValue="Ungurească">Ungurească</normalizedToken>
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Cave, oak-hornbeam forest, leaf litter, altitude 530 m, July 13, 2006, leg. P.
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<normalizedToken id="3D0D74131E17BB34C70DEF0D5E6863D2" originalValue="Fenďa">Fenďa</normalizedToken>
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. Serbia: three females,
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<normalizedToken id="BEE6A94AEC42E7441B3720C73EE32B96" originalValue="Kučajske">Kucajske</normalizedToken>
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Planine Mts., Pomoravlje District, Troglan Bara Settlement, Velika Brezovica Forest, beech forest (
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<taxonomicName id="182836B8CAC251D188906823E2EB0D2B" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
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), leaf litter and soil detritus, altitude 900 m, April 23, 2009, leg. I.
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<normalizedToken id="A911EF71ED7742A7A1256960320C9F7F" originalValue="Mihál">Mihal</normalizedToken>
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. Switzerland: four females, Basel-Stadt
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<pageBreakToken id="5375D1D2F8470F3E89E6B073DAA831C5" pageId="5" pageNumber="40" start="start">Canton</pageBreakToken>
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, Riehen Town, Wenken Settlement, old beech forest with limes (
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<taxonomicName id="13EC685DFF6FD755F92AFDB27D235D8D" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia" order="Malvales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="genus">Tilia</taxonomicName>
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sp.) and maples (
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<taxonomicName id="D8060CF56AF8C814FC950E40940B4FD3" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acer" order="Sapindales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
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sp.), leaf litter and soil detritus, altitude 380 m, June 1, 2016, leg. P.
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<normalizedToken id="563C03C8127BE2C8231682C26BF2C41A" originalValue="Mašán">Masan</normalizedToken>
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. United Kingdom, Wales: two females, Anglesey Island, Llangefni Town, The Dingle Forest, alluvium of Cefni River, mixed broadleaved deciduous forest, leaf litter and soil detritus, altitude 35 m, July 31, 2010, leg. P.
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<normalizedToken id="B4F6D54648FDDB266F5C57E63D46B0E4" originalValue="Mašán">Masan</normalizedToken>
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; four females, Anglesey Island, Llandegfan Village, alluvium of Cadnant River, mixed broadleaved deciduous wood, leaf litter and soil detritus, altitude 50 m, July 25, 2010, leg. P.
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<normalizedToken id="ECC4F03BFD778E558F6B13FE0CD4F37A" originalValue="Mašán">Masan</normalizedToken>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C17068646E56B21FD401A9C83041A6E4" pageId="5" pageNumber="40" type="diagnosis">
|
||
<paragraph id="44651FB23B2A8E2D2796A0C64C825829" pageId="5" pageNumber="40">Diagnosis.</paragraph>
|
||
<paragraph id="142FF53D1DAC98B6CB072151B9EEF46F" pageId="5" pageNumber="40">The species may be distinguished from the other congeners especially by combination of the following female characters: (1) dorsal shield setae simple, needle-like; (2) dorsal shield between setae z1 and z2 and peritrematal shields close to stigma with enlarged and cavity-like poroid structure; (3) presternal platelets well sclerotized in anterior part, transversely striate, separate each other but connected to anterior margin of sternal shield; (4) exopodal platelets II-III and III-IV free, not fused to peritrematal shields; (5) ventrianal shield with three pairs of preanal setae (JV1‒JV3); (6) lateral and opisthogastric soft integument with eight pairs of setae: r6, R2‒R5, ZV2, JV4, and JV5; (7) tarsus II with two subdistal posterolateral setae thickened, of which pl1 with obtuse apex, spur-like, and pl2 terminally attenuate and sharply pointed (but the tip of pl2 is fragile and can be very often broken in slide-mounted specimens); (8) tarsus IV with 17 setae.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F02BC766E14F63DCDA3080119514B278" pageId="5" pageNumber="40" type="description">
|
||
<paragraph id="7536EF4115C9DC91D61C2F3BEDF99433" pageId="5" pageNumber="40">Description.</paragraph>
|
||
<paragraph id="61B9E3351FFF429A5B14E4B430A050EA" pageId="5" pageNumber="40">
|
||
The morphological attributes of the species were described and illustrated in detail by
|
||
<bibRefCitation id="42F807F893576020B25EE51FF0765D7C" author="Masan, P" journalOrPublisher="NOI Press, Bratislava" pageId="7" pageNumber="42" title="A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea)." year="2007">
|
||
<normalizedToken id="AC65584725917E931A969676F3FC50D0" originalValue="Mašán">Masan</normalizedToken>
|
||
(2007)
|
||
</bibRefCitation>
|
||
, and the description does not need to be repeated here. The original illustrations given by the cited author are based on the type specimens from
|
||
<normalizedToken id="D64C0DA6D7EBAAEFF98A28CBA31A918A" originalValue="Štefanová">Stefanova</normalizedToken>
|
||
Village (see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4084ECD5D88F0DD994AF50A5BBC6BFB0" pageId="5" pageNumber="40" type="etymology">
|
||
<paragraph id="FAEE704D66FE50C2B25A8EA7DC0C999B" pageId="5" pageNumber="40">Etymology.</paragraph>
|
||
<paragraph id="B2AA8A0A154B9E72D6243535284F9A1A" pageId="5" pageNumber="40">
|
||
Many epithets beginning with sub-, a Latin name-forming prefix meaning
|
||
<normalizedToken id="8050BACCB660C6032F6C933472E44659" originalValue="“approaching”">"approaching"</normalizedToken>
|
||
, are intended to distinguish a species from that with which it was previously confused. The new epithet is proposed as an alternative name for
|
||
<taxonomicName id="2C23B725469714FDB7DA09654E61BF61" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius humeralis" order="Mesostigmata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="humeralis">Pachyseius humeralis</taxonomicName>
|
||
in the broad sense understood to date.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A626DE32CC42C7A193701D75B50123B0" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="discussion">
|
||
<paragraph id="1880FC86ECCE688C3C75048902584E81" pageId="5" pageNumber="40">Discussion.</paragraph>
|
||
<paragraph id="DF72A7E5A5E24C94C8B94EC7E5C76BF7" pageId="5" pageNumber="40">
|
||
The re-evaluation of the lectotype and newly collected material of
|
||
<taxonomicName id="866B6832897148C789669653D3FAE647" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius humeralis" order="Mesostigmata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="humeralis">Pachyseius humeralis</taxonomicName>
|
||
sensu Berlese, 1910 showed that there are several apparent and constant morphological differences between this species and its congener widely reported from Europe under the same name, and here established as
|
||
<taxonomicName id="E033F6C019116305D73C3BB056FC119E" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius subhumeralis" order="Mesostigmata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="subhumeralis">Pachyseius subhumeralis</taxonomicName>
|
||
sp. n. According to these findings,
|
||
<taxonomicName id="6ADDAB30CCFBE12448FEFDC2CE244179" lsidName="P. humeralis" pageId="5" pageNumber="40" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
may be most reliably distinguished from
|
||
<taxonomicName id="08A22922552A343538D89056CCDB2B5D" lsidName="P. subhumeralis" pageId="5" pageNumber="40" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
by the following characters:
|
||
</paragraph>
|
||
<paragraph id="327856480944F61F823EB58D218C9DFB" pageId="5" pageNumber="40">
|
||
(1) the placement of genital pores (placed outside the epigynal shield in
|
||
<taxonomicName id="8E4D4B518B8E531AC1AE15FBBFD67776" lsidName="P. humeralis" pageId="5" pageNumber="40" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
, or on posterolateral corners of the shield in
|
||
<taxonomicName id="C949655C759309D57C12CB539275DA82" lsidName="P. subhumeralis" pageId="5" pageNumber="40" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
; Figs 1 and 2),
|
||
</paragraph>
|
||
<paragraph id="A82F515FC8C7E27F1037E92303DC2E58" pageId="5" pageNumber="40">
|
||
(2) the form and placement of presternal platelets (the platelets evenly sclerotized and free on soft integument in
|
||
<taxonomicName id="361317A42DF356BFA0A5B26806349C37" lsidName="P. humeralis" pageId="5" pageNumber="40" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
, or separately connected to sternal shield with their weakly sclerotized posterior portions in
|
||
<taxonomicName id="B9CA3BBA253CE171D0B0504D69F71D13" lsidName="P. subhumeralis" pageId="5" pageNumber="40" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
; Figure 3),
|
||
</paragraph>
|
||
<paragraph id="ECE67D4D9D2F7976C3803FBFCF45BF0C" pageId="5" pageNumber="40">
|
||
(3) the number of dorsomarginal setae on soft integument (four pairs in
|
||
<taxonomicName id="3E294015F542A03C0E20E0765EDE211D" lsidName="P. humeralis" pageId="5" pageNumber="40" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
‒ setae R5 absent, or five pairs in
|
||
<taxonomicName id="4BC86691CCEFDDE38E7377ABEE205D57" lsidName="P. subhumeralis" pageId="5" pageNumber="40" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
‒ setae R5 present; Figure 4),
|
||
</paragraph>
|
||
<paragraph id="2044D0821896F57B86E25D0D14C53E24" pageId="6" pageNumber="41">
|
||
<pageBreakToken id="FB1DE30E56AA84B5F6BABBF7F591E428" pageId="6" pageNumber="41" start="start">(</pageBreakToken>
|
||
4) the form of posterolateral seta pl2 on tarsus II (pl2 robust, spur-like, and apically rounded in
|
||
<taxonomicName id="38D6BB94B34D69C6F77398E41FC814EB" lsidName="P. humeralis" pageId="6" pageNumber="41" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
, or regularly tapered and apically pointed in
|
||
<taxonomicName id="13106CCF8C2307BF8F361064AC313E87" lsidName="P. subhumeralis" pageId="6" pageNumber="41" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
; Figure 5),
|
||
</paragraph>
|
||
<paragraph id="BA6E61F6B589F7330766502EFD283BC1" pageId="6" pageNumber="41">
|
||
(5) the relative length of peritremes (the peritreme with anterior tip never reaching beyond the longitudinal axis of gland pore gdj3 in
|
||
<taxonomicName id="3A98296623A05B2A4C411B1B92462F39" lsidName="P. humeralis" pageId="6" pageNumber="41" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
, or reaching slightly beyond this point in
|
||
<taxonomicName id="DB50DE3DD43CD6F4795DB9CA51172FE7" lsidName="P. subhumeralis" pageId="6" pageNumber="41" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
; Figure 6),
|
||
</paragraph>
|
||
<paragraph id="DD5F974C6B2D6C689E5478E918DF6FE9" pageId="6" pageNumber="41">
|
||
(6) the form of epistome (the epistome with narrow base and large central cusp in
|
||
<taxonomicName id="B396AE80FEEA43E00ECE17564EA93CD1" lsidName="P. humeralis" pageId="6" pageNumber="41" rank="species" species="humeralis">P. humeralis</taxonomicName>
|
||
, or with widened base and uniformly spinate anteriormost margin in
|
||
<taxonomicName id="3211915C6C419013AF5C7FA86D268C95" lsidName="P. subhumeralis" pageId="6" pageNumber="41" rank="species" species="subhumeralis">P. subhumeralis</taxonomicName>
|
||
; Figure 7).
|
||
</paragraph>
|
||
<paragraph id="713C094E2F4C7042F3CD3EB008BBCBAC" pageId="6" pageNumber="41">
|
||
There are also less serious differences recognizable between the two compared species, having partly transitional character, for instance in size and proportions of the dorsal and ventroanal shield, and relative length of some idiosomal setae. Generally, when compared with
|
||
<taxonomicName id="60E95EBBEF23933ADEBF192C2FC276E9" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius subhumeralis" order="Mesostigmata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="subhumeralis">Pachyseius subhumeralis</taxonomicName>
|
||
(based on metric data given by
|
||
<normalizedToken id="7B8D475BAFC1D3477DBFED1AC7BDB17F" originalValue="Mašán">Masan</normalizedToken>
|
||
in 2007),
|
||
<taxonomicName id="088ACC7DB27CD85D1870C4C77E230AB7" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius humeralis" order="Mesostigmata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="humeralis">Pachyseius humeralis</taxonomicName>
|
||
is smaller species (optimum length: 565-595
|
||
<normalizedToken id="2336AE1485C88C72DB9586D1C60B4FEE" originalValue="μm">μm</normalizedToken>
|
||
versus 595-640
|
||
<normalizedToken id="07F4E6107A583BA983246D2AF39185C0" originalValue="μm">μm</normalizedToken>
|
||
), possessing relatively narrower ventrianal shield (L/W ratio: 1.2‒1.37 versus 1.08‒1.28; Figs 1 and 2), relatively longer setae (j5: 19-25
|
||
<normalizedToken id="3B82AA2996FD80FC0F5575094501C031" originalValue="μm">μm</normalizedToken>
|
||
versus 15-23
|
||
<normalizedToken id="D6ABE3F3B5257710AD331258A0217DC6" originalValue="μm">μm</normalizedToken>
|
||
, J5: 31-39
|
||
<normalizedToken id="F65BD8CF01C557DBF02E4E606AB01049" originalValue="μm">μm</normalizedToken>
|
||
versus 24-34
|
||
<normalizedToken id="05A3EC0D9E070657936AF325D656E3C4" originalValue="μm">μm</normalizedToken>
|
||
), and shallower medial concavity of posterior margin of sternal shield.
|
||
</paragraph>
|
||
<paragraph id="681674343437F855FE05690B4C33B1F5" pageId="6" pageNumber="41">
|
||
The specimens of
|
||
<taxonomicName id="66C36D454667564150B5B8D503B88199" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachyseius humeralis" order="Mesostigmata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="humeralis">Pachyseius humeralis</taxonomicName>
|
||
available in the Berlese Collection in Florence from several localities of Central Italy (Monte Giovi, Filettino, Vallombrosa, and Florence), and those from other European regions widely published by various authors (especially from the Mediterranean areas), should be carefully re-examined in the next studies to define their correct identity, and to re-evaluate spatial distribution of the species treated in this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |