treatments-xml/data/E7/05/CC/E705CC50FFA2FF84DE93FB6E5F961D42.xml

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<mods:title id="AFF8903E8C94FD2FE63978B8E94F779C">A new Physoschistura from a Salween affluent in western Yunnan (Teleostei: Nemacheilidae)</mods:title>
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<mods:namePart id="38BF89F94612256F2DE8499A6365508C">Endruweit, Marco</mods:namePart>
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<mods:date id="E30B41C1E8527FDE5C6B9ADED018688F">2017</mods:date>
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<paragraph id="6F137D46FFA2FF82DE93FB6E5C631BBB" blockId="1.[151,621,1194,1254]" box="[151,621,1194,1221]" pageId="1" pageNumber="379">
<heading id="345BCA2AFFA2FF82DE93FB6E5C631BBB" bold="true" box="[151,621,1194,1221]" fontSize="11" level="1" pageId="1" pageNumber="379" reason="1">
<emphasis id="5DD8A154FFA2FF82DE93FB6E5C631BBB" bold="true" box="[151,621,1194,1221]" pageId="1" pageNumber="379">
<taxonomicName id="A8AC06C5FFA2FF82DE93FB6E5FC01BBA" ID-CoL="4HJFY" box="[151,462,1194,1220]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="379" phylum="Chordata" rank="species" species="absumbra" status="sp. nov.">
<emphasis id="5DD8A154FFA2FF82DE93FB6E5FC01BBA" bold="true" box="[151,462,1194,1220]" italics="true" pageId="1" pageNumber="379">Physoschistura absumbra</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="46EB1C2FFFA2FF82DFD9FB6F5C631BBB" box="[477,621,1195,1221]" pageId="1" pageNumber="379" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="6F137D46FFA2FF82DE93FB095F0E1B98" blockId="1.[151,621,1194,1254]" box="[151,256,1229,1254]" pageId="1" pageNumber="379">
<figureCitation id="F79761C3FFA2FF82DE93FB095F0E1B98" box="[151,256,1229,1254]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1205,1227]" captionStartId-1="4.[151,250,1912,1934]" captionStartId-2="5.[151,250,935,957]" captionTargetBox-0="[150,1436,409,1183]" captionTargetBox-1="[300,1287,481,1890]" captionTargetBox-2="[155,1426,202,904]" captionTargetId-0="figure@3.[150,1436,409,1183]" captionTargetId-1="figure@4.[300,1287,481,1890]" captionTargetId-2="figure@5.[150,1436,193,913]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Physoschistura absumbra, KIZ 2014006018, holotype, 39.0 mm SL; Yunnan: Cangyuan; Nangunhe River." captionText-1="FIGURE 2. Physoschistura absumbra, ZFMK-ICH 103627, paratype, 37.6 mm SL; head in ventral view." captionText-2="FIGURE 3. Physoschistura absumbra, ZFMK-ICH 103627, paratype, 37.6 mm SL; gas bladder." httpUri-0="https://zenodo.org/record/573194/files/figure.png" httpUri-1="https://zenodo.org/record/573195/files/figure.png" httpUri-2="https://zenodo.org/record/573197/files/figure.png" pageId="1" pageNumber="379">Figs. 1–3</figureCitation>
</paragraph>
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<subSubSection id="27B62ECDFFA2FF82DE93FAD15D231AE4" pageId="1" pageNumber="379" type="materials_examined">
<paragraph id="6F137D46FFA2FF82DE93FAD15B501A2C" blockId="1.[151,1437,1301,2010]" pageId="1" pageNumber="379">
<materialsCitation id="DFC4771BFFA2FF82DE93FAD15B991A53" ID-GBIF-Occurrence="1500393282" box="[151,1431,1301,1326]" collectionCode="KIZ" country="China" location="Banlao town" pageId="1" pageNumber="379" specimenCode="KIZ 2014006018" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="5DD8A154FFA2FF82DE93FAD15F051A50" bold="true" box="[151,267,1301,1326]" pageId="1" pageNumber="379">
<typeStatus id="B017C3E4FFA2FF82DE93FAD15F081A50" box="[151,262,1301,1326]" pageId="1" pageNumber="379" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="3F0AD53DFFA2FF82DF12FAD25FD01A50" box="[278,478,1301,1326]" collectionCode="KIZ" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="379">
<collectionCode id="09BDE583FFA2FF82DF12FAD25F4A1A53" box="[278,324,1302,1325]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="379">KIZ</collectionCode>
2014006018
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, 39.0 mm SL; 
<collectingCountry id="17BB3DD6FFA2FF82DC97FAD15CD41A50" box="[659,730,1301,1326]" name="China" pageId="1" pageNumber="379">China</collectingCountry>
: 
<collectingRegion id="AD68B3A4FFA2FF82DCEDFAD25D301A53" box="[745,830,1302,1325]" country="China" name="Yunnan" pageId="1" pageNumber="379">Yunnan</collectingRegion>
Prov.: Lincang Pref.: Cangyuan Cty.: 
<location id="6A732B9DFFA2FF82DB06FAD15B991A53" LSID="urn:lsid:plazi:treatment:E705CC50FFA2FF84DE93FB6E5F961D42:6A732B9DFFA2FF82DB06FAD15B991A53" box="[1282,1431,1301,1326]" country="China" name="Banlao town" pageId="1" pageNumber="379" stateProvince="Yunnan">Banlao town</location>
</materialsCitation>
; 
<materialsCitation id="DFC4771BFFA2FF82DE93FAFD5B501A2C" ID-GBIF-Occurrence="1500393279" box="[151,1374,1337,1362]" collectingDate="2014-08-23" collectorName="Endruweit, M. &amp; Qin, T." country="China" elevation="523" latitude="23.1822" location="Nangunhe River" longLatPrecision="1" longitude="98.94292" pageId="1" pageNumber="379" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<location id="6A732B9DFFA2FF82DE93FAFD5F5D1A2C" LSID="urn:lsid:plazi:treatment:E705CC50FFA2FF84DE93FB6E5F961D42:6A732B9DFFA2FF82DE93FAFD5F5D1A2C" box="[151,339,1337,1362]" country="China" latitude="23.1822" longLatPrecision="1" longitude="98.94292" name="Nangunhe River" pageId="1" pageNumber="379" stateProvince="Yunnan">Nangunhe River</location>
; 
<geoCoordinate id="0A981B81FFA2FF82DF5AFAFE5FFB1A2F" box="[350,501,1338,1362]" direction="north" orientation="latitude" pageId="1" pageNumber="379" precision="1" value="23.1822">23°10.932' N</geoCoordinate>
, 
<geoCoordinate id="0A981B81FFA2FF82DC04FAFD5C9A1A2F" box="[512,660,1337,1362]" direction="east" orientation="longitude" pageId="1" pageNumber="379" precision="1" value="98.94292">98°56.575' E</geoCoordinate>
, 
<elevation id="E4819A75FFA2FF82DCA4FAFD5D591A2F" box="[672,855,1337,1362]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" pageId="1" pageNumber="379" unit="m" value="523.0">
elevation 
<quantity id="A854D0A3FFA2FF82DD09FAFD5D591A2F" box="[781,855,1337,1362]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" pageId="1" pageNumber="379" unit="m" value="523.0">523 m</quantity>
</elevation>
; coll. 
<collectorName id="C2591890FFA2FF82DD9FFAFD5A301A2F" box="[923,1086,1337,1362]" pageId="1" pageNumber="379">Endruweit, M.</collectorName>
&amp; 
<collectorName id="C2591890FFA2FF82DA65FAFD5ABC1A2F" box="[1121,1202,1337,1362]" pageId="1" pageNumber="379">Qin, T.</collectorName>
, 
<date id="1B125B86FFA2FF82DAB9FAFE5B551A2C" box="[1213,1371,1338,1362]" pageId="1" pageNumber="379" value="2014-08-23">
<collectingDate id="0B56A26EFFA2FF82DAB9FAFE5B551A2C" box="[1213,1371,1338,1362]" pageId="1" pageNumber="379" value="2014-08-23">23 Aug. 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="6F137D46FFA2FF82DEC3FA995D231AE4" blockId="1.[151,1437,1301,2010]" pageId="1" pageNumber="379">
<materialsCitation id="DFC4771BFFA2FF82DEC3FA995D231AE4" ID-GBIF-Occurrence="1500393286" collectionCode="KIZ" country="China" location="Banlao town" pageId="1" pageNumber="379" specimenCode="KIZ 2014006017, 6019" specimenCount="6" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="5DD8A154FFA2FF82DEC3FA995F481A08" bold="true" box="[199,326,1373,1398]" pageId="1" pageNumber="379">
<typeStatus id="B017C3E4FFA2FF82DEC3FA995F4F1A08" box="[199,321,1373,1398]" pageId="1" pageNumber="379" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="3F0AD53DFFA2FF82DF54FA9A5C6F1A08" box="[336,609,1373,1398]" pageId="1" pageNumber="379">
<collectionCode id="09BDE583FFA2FF82DF54FA9A5F711A0B" box="[336,383,1374,1397]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="379">KIZ</collectionCode>
2014006017, 6019
</specimenCode>
, 6022, 6024, 6026, 6028, 
<specimenCount id="79AAB6CFFFA2FF82DD90FA995A2D1A08" box="[916,1059,1373,1398]" pageId="1" pageNumber="379" type="generic">6 specimens</specimenCount>
, 
<quantity id="A854D0A3FFA2FF82DA36FA995AD61A0B" box="[1074,1240,1373,1398]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" metricValueMax="4.35" metricValueMin="2.63" pageId="1" pageNumber="379" unit="mm" value="34.9" valueMax="43.5" valueMin="26.3">26.3–43.5 mm</quantity>
SL; ZFMK-ICH 103626-3630, 5, 
<quantity id="A854D0A3FFA2FF82DF52FA465FF71AE7" box="[342,505,1410,1434]" metricMagnitude="-2" metricUnit="m" metricValue="3.6049999999999995" metricValueMax="3.9299999999999997" metricValueMin="3.28" pageId="1" pageNumber="379" unit="mm" value="36.05" valueMax="39.3" valueMin="32.8">32.8–39.3 mm</quantity>
SL; same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="27B62ECDFFA2FF82DEC3FA615F541930" pageId="1" pageNumber="379" type="diagnosis">
<paragraph id="6F137D46FFA2FF82DEC3FA615F541930" blockId="1.[151,1437,1301,2010]" pageId="1" pageNumber="379">
<emphasis id="5DD8A154FFA2FF82DEC3FA615F4F1AC0" bold="true" box="[199,321,1445,1470]" pageId="1" pageNumber="379">Diagnosis.</emphasis>
<taxonomicName id="A8AC06C5FFA2FF82DF4FFA615C611AC3" box="[331,623,1445,1469]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="379" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA2FF82DF4FFA615C611AC3" box="[331,623,1445,1469]" italics="true" pageId="1" pageNumber="379">Physoschistura absumbra</emphasis>
</taxonomicName>
is easily distinguished from all congeners in lacking a suborbital lobe and in having a complete lateral line and a continuous lower jaw. Further diagnostic characters are head depth at nape 13–14% SL; head length dorsally 23–25% SL, laterally 26–28% SL; body depth 15–17% SL; caudalpeduncle depth 13–14% SL; 9–12 rather regular flank bars; basicaudal bar complete; and a maximum adult size of about 
<quantity id="A854D0A3FFA2FF82DED8F9F25F251933" box="[220,299,1590,1614]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="379" unit="mm" value="44.0">44 mm</quantity>
SL.
</paragraph>
</subSubSection>
<subSubSection id="27B62ECDFFA2FF80DEC3F99D5FE61E1A" lastPageId="3" lastPageNumber="381" pageId="1" pageNumber="379" type="description">
<paragraph id="6F137D46FFA2FF82DEC3F99D5C5E18A4" blockId="1.[151,1437,1301,2010]" pageId="1" pageNumber="379">
<emphasis id="5DD8A154FFA2FF82DEC3F99D5F57190C" bold="true" box="[199,345,1625,1650]" pageId="1" pageNumber="379">Description.</emphasis>
See 
<figureCitation id="F79761C3FFA2FF82DF95F99D5FFF190C" box="[401,497,1625,1650]" captionStart="FIGURE 1" captionStartId="3.[151,250,1205,1227]" captionTargetBox="[150,1436,409,1183]" captionTargetId="figure@3.[150,1436,409,1183]" captionTargetPageId="3" captionText="FIGURE 1. Physoschistura absumbra, KIZ 2014006018, holotype, 39.0 mm SL; Yunnan: Cangyuan; Nangunhe River." httpUri="https://zenodo.org/record/573194/files/figure.png" pageId="1" pageNumber="379">Figure 1</figureCitation>
for general appearance and 
<tableCitation id="222E48FDFFA2FF82DD31F99D5D84190C" box="[821,906,1625,1650]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Morphometrics of the holotype and 10 paratypes of Physoschistura absumbra; SD = standard deviation." pageId="1" pageNumber="379">Table 1</tableCitation>
for morphometric data of the 
<typeStatus id="B017C3E4FFA2FF82DAE0F99D5B48190C" box="[1252,1350,1625,1650]" pageId="1" pageNumber="379" type="holotype">holotype</typeStatus>
and 
<specimenCount id="79AAB6CFFFA2FF82DB84F99E5F0819E8" pageId="1" pageNumber="379" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
. Body compact, predorsally cylindrical, postdorsally laterally compressed. Body depth 5.8–6.7 times in SL, maximum immediately in front of dorsal fin. Caudal peduncle deep, depth 1.0–1.2 times in its length. Axillary pelvic lobe rudimentary, tip free. Anus located 0.9–1.7 times eye diameter in front of anal-fin origin. Snout pointed. Cheek not inflated. Head width constantly increasing from nostril to about edge of preoperculum. Eye large; located dorsolaterally, not reaching dorsal profile when viewed laterally. Interorbital space flat, narrow. Anterior nostril a flap-like tube, not pierced, not reaching orbit. Mouth (
<figureCitation id="F79761C3FFA2FF82DD68F8F55DBF1834" box="[876,945,1841,1866]" captionStart="FIGURE 2" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[300,1287,481,1890]" captionTargetId="figure@4.[300,1287,481,1890]" captionTargetPageId="4" captionText="FIGURE 2. Physoschistura absumbra, ZFMK-ICH 103627, paratype, 37.6 mm SL; head in ventral view." httpUri="https://zenodo.org/record/573195/files/figure.png" pageId="1" pageNumber="379">Fig. 2</figureCitation>
) inferior, arched, 1.5–1.8 times wider than long. Lips thick, furrowed close to axis; upper lip not notched, lower lip slightly notched medianly, lateral sides slightly triangular, not forming pads. Upper jaw with pointed processus dentiformis; lower jaw spoon-like, not notched. Inner rostral barbel failing to reach rictus, outer reaching vertical through anterior rim of orbit, maxillary reaching vertical through posterior rim.
</paragraph>
<caption id="3BD32DCEFFA1FF81DE93FF5C5B661FD1" box="[151,1384,151,175]" pageId="2" pageNumber="380">
<paragraph id="6F137D46FFA1FF81DE93FF5C5B661FD1" blockId="2.[151,1384,151,175]" box="[151,1384,151,175]" pageId="2" pageNumber="380">
<emphasis id="5DD8A154FFA1FF81DE93FF5C5F041FD1" bold="true" box="[151,266,152,175]" pageId="2" pageNumber="380">TABLE 1.</emphasis>
Morphometrics of the holotype and 10 paratypes of 
<taxonomicName id="A8AC06C5FFA1FF81DD38FF535A5E1FD1" box="[828,1104,151,175]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="380" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA1FF81DD38FF535A5E1FD1" box="[828,1104,151,175]" italics="true" pageId="2" pageNumber="380">Physoschistura absumbra</emphasis>
</taxonomicName>
; SD = standard deviation.
</paragraph>
</caption>
<paragraph id="6F137D46FFA1FF81DCC3FF0E5B4D19F9" pageId="2" pageNumber="380">
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<tr id="D19C7F04FFA1007CDEC3FF0E5B4D1F9E" box="[199,1347,202,224]" gridrow="0" pageId="2" pageNumber="380" rowspan-0="1">
<th id="924D1678FFA1007CDCC3FF0E5D2E1F9E" box="[711,800,202,224]" gridcol="1" gridrow="0" pageId="2" pageNumber="380">holotype</th>
<th id="924D1678FFA1007CDDCBFF0E5A061F9E" box="[975,1032,202,224]" gridcol="2" gridrow="0" pageId="2" pageNumber="380">mean</th>
<th id="924D1678FFA1007CDA55FF0E5ADE1F9E" box="[1105,1232,202,224]" gridcol="3" gridrow="0" pageId="2" pageNumber="380">range</th>
<th id="924D1678FFA1007CDB27FF0E5B4D1F9E" box="[1315,1347,202,224]" gridcol="4" gridrow="0" pageId="2" pageNumber="380">SD</th>
</tr>
<tr id="D19C7F04FFA1007CDEC3FF305B4D1E74" box="[199,1347,244,266]" gridrow="1" pageId="2" pageNumber="380" rowspan-2="1" rowspan-4="1">
<th id="924D1678FFA1007CDEC3FF305C321E74" box="[199,572,244,266]" gridcol="0" gridrow="1" pageId="2" pageNumber="380">SL in mm</th>
<td id="924D1678FFA1007CDCC3FF305D2E1E74" box="[711,800,244,266]" gridcol="1" gridrow="1" pageId="2" pageNumber="380">39.0</td>
<td id="924D1678FFA1007CDA55FF305ADE1E74" box="[1105,1232,244,266]" gridcol="3" gridrow="1" pageId="2" pageNumber="380">26.3–43.5</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FE825B4D1E22" box="[199,1347,326,348]" gridrow="2" pageId="2" pageNumber="380" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="924D1678FFA1007CDEC3FE825C321E22" box="[199,572,326,348]" gridcol="0" gridrow="2" pageId="2" pageNumber="380">In percent of SL</th>
</tr>
<tr id="D19C7F04FFA1007CDEC3FEB45B4D1EF8" box="[199,1347,368,390]" gridrow="3" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FEB45C321EF8" box="[199,572,368,390]" gridcol="0" gridrow="3" pageId="2" pageNumber="380">Total length</th>
<td id="924D1678FFA1007CDCC3FEB45D2E1EF8" box="[711,800,368,390]" gridcol="1" gridrow="3" pageId="2" pageNumber="380">122.1</td>
<td id="924D1678FFA1007CDDCBFEB45A061EF8" box="[975,1032,368,390]" gridcol="2" gridrow="3" pageId="2" pageNumber="380">124.4</td>
<td id="924D1678FFA1007CDA55FEB45ADE1EF8" box="[1105,1232,368,390]" gridcol="3" gridrow="3" pageId="2" pageNumber="380">122.1–127.1</td>
<td id="924D1678FFA1007CDB27FEB45B4D1EF8" box="[1315,1347,368,390]" gridcol="4" gridrow="3" pageId="2" pageNumber="380">1.8</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FE5D5B4D1ED1" box="[199,1347,409,431]" gridrow="4" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FE5D5C321ED1" box="[199,572,409,431]" gridcol="0" gridrow="4" pageId="2" pageNumber="380">Dorsal head length</th>
<td id="924D1678FFA1007CDCC3FE5D5D2E1ED1" box="[711,800,409,431]" gridcol="1" gridrow="4" pageId="2" pageNumber="380">24.6</td>
<td id="924D1678FFA1007CDDCBFE5D5A061ED1" box="[975,1032,409,431]" gridcol="2" gridrow="4" pageId="2" pageNumber="380">23.4</td>
<td id="924D1678FFA1007CDA55FE5D5ADE1ED1" box="[1105,1232,409,431]" gridcol="3" gridrow="4" pageId="2" pageNumber="380">22.7–24.6</td>
<td id="924D1678FFA1007CDB27FE5D5B4D1ED1" box="[1315,1347,409,431]" gridcol="4" gridrow="4" pageId="2" pageNumber="380">0.7</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FE065B4D1EA6" box="[199,1347,450,472]" gridrow="5" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FE065C321EA6" box="[199,572,450,472]" gridcol="0" gridrow="5" pageId="2" pageNumber="380">Lateral head length</th>
<td id="924D1678FFA1007CDCC3FE065D2E1EA6" box="[711,800,450,472]" gridcol="1" gridrow="5" pageId="2" pageNumber="380">26.4</td>
<td id="924D1678FFA1007CDDCBFE065A061EA6" box="[975,1032,450,472]" gridcol="2" gridrow="5" pageId="2" pageNumber="380">26.9</td>
<td id="924D1678FFA1007CDA55FE065ADE1EA6" box="[1105,1232,450,472]" gridcol="3" gridrow="5" pageId="2" pageNumber="380">25.5–28.3</td>
<td id="924D1678FFA1007CDB27FE065B4D1EA6" box="[1315,1347,450,472]" gridcol="4" gridrow="5" pageId="2" pageNumber="380">0.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FE285B4D1D7C" box="[199,1347,492,514]" gridrow="6" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FE285C321D7C" box="[199,572,492,514]" gridcol="0" gridrow="6" pageId="2" pageNumber="380">Predorsal length</th>
<td id="924D1678FFA1007CDCC3FE285D2E1D7C" box="[711,800,492,514]" gridcol="1" gridrow="6" pageId="2" pageNumber="380">54.1</td>
<td id="924D1678FFA1007CDDCBFE285A061D7C" box="[975,1032,492,514]" gridcol="2" gridrow="6" pageId="2" pageNumber="380">51.8</td>
<td id="924D1678FFA1007CDA55FE285ADE1D7C" box="[1105,1232,492,514]" gridcol="3" gridrow="6" pageId="2" pageNumber="380">50.0–54.1</td>
<td id="924D1678FFA1007CDB27FE285B4D1D7C" box="[1315,1347,492,514]" gridcol="4" gridrow="6" pageId="2" pageNumber="380">1.5</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FDD15B4D1D55" box="[199,1347,533,555]" gridrow="7" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FDD15C321D55" box="[199,572,533,555]" gridcol="0" gridrow="7" pageId="2" pageNumber="380">Prepelvic length</th>
<td id="924D1678FFA1007CDCC3FDD15D2E1D55" box="[711,800,533,555]" gridcol="1" gridrow="7" pageId="2" pageNumber="380">53.1</td>
<td id="924D1678FFA1007CDDCBFDD15A061D55" box="[975,1032,533,555]" gridcol="2" gridrow="7" pageId="2" pageNumber="380">53.7</td>
<td id="924D1678FFA1007CDA55FDD15ADE1D55" box="[1105,1232,533,555]" gridcol="3" gridrow="7" pageId="2" pageNumber="380">52.4–54.8</td>
<td id="924D1678FFA1007CDB27FDD15B4D1D55" box="[1315,1347,533,555]" gridcol="4" gridrow="7" pageId="2" pageNumber="380">0.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FDFA5B4D1D2A" box="[199,1347,574,596]" gridrow="8" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FDFA5C321D2A" box="[199,572,574,596]" gridcol="0" gridrow="8" pageId="2" pageNumber="380">Preanal length</th>
<td id="924D1678FFA1007CDCC3FDFA5D2E1D2A" box="[711,800,574,596]" gridcol="1" gridrow="8" pageId="2" pageNumber="380">76.2</td>
<td id="924D1678FFA1007CDDCBFDFA5A061D2A" box="[975,1032,574,596]" gridcol="2" gridrow="8" pageId="2" pageNumber="380">77.9</td>
<td id="924D1678FFA1007CDA55FDFA5ADE1D2A" box="[1105,1232,574,596]" gridcol="3" gridrow="8" pageId="2" pageNumber="380">76.2–79.5</td>
<td id="924D1678FFA1007CDB27FDFA5B4D1D2A" box="[1315,1347,574,596]" gridcol="4" gridrow="8" pageId="2" pageNumber="380">1.1</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FDAC5B4D1D00" box="[199,1347,616,638]" gridrow="9" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FDAC5C321D00" box="[199,572,616,638]" gridcol="0" gridrow="9" pageId="2" pageNumber="380">Pre-anus length</th>
<td id="924D1678FFA1007CDCC3FDAC5D2E1D00" box="[711,800,616,638]" gridcol="1" gridrow="9" pageId="2" pageNumber="380">68.7</td>
<td id="924D1678FFA1007CDDCBFDAC5A061D00" box="[975,1032,616,638]" gridcol="2" gridrow="9" pageId="2" pageNumber="380">70.4</td>
<td id="924D1678FFA1007CDA55FDAC5ADE1D00" box="[1105,1232,616,638]" gridcol="3" gridrow="9" pageId="2" pageNumber="380">68.7–72.2</td>
<td id="924D1678FFA1007CDB27FDAC5B4D1D00" box="[1315,1347,616,638]" gridcol="4" gridrow="9" pageId="2" pageNumber="380">1.3</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FD555B4D1DD9" box="[199,1347,657,679]" gridrow="10" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FD555C321DD9" box="[199,572,657,679]" gridcol="0" gridrow="10" pageId="2" pageNumber="380">Head depth (at eye)</th>
<td id="924D1678FFA1007CDCC3FD555D2E1DD9" box="[711,800,657,679]" gridcol="1" gridrow="10" pageId="2" pageNumber="380">12.3</td>
<td id="924D1678FFA1007CDDCBFD555A061DD9" box="[975,1032,657,679]" gridcol="2" gridrow="10" pageId="2" pageNumber="380">12.2</td>
<td id="924D1678FFA1007CDA55FD555ADE1DD9" box="[1105,1232,657,679]" gridcol="3" gridrow="10" pageId="2" pageNumber="380">11.7–12.6</td>
<td id="924D1678FFA1007CDB27FD555B4D1DD9" box="[1315,1347,657,679]" gridcol="4" gridrow="10" pageId="2" pageNumber="380">0.3</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FD7E5B4D1DAE" box="[199,1347,698,720]" gridrow="11" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FD7E5C321DAE" box="[199,572,698,720]" gridcol="0" gridrow="11" pageId="2" pageNumber="380">Head depth (at nape)</th>
<td id="924D1678FFA1007CDCC3FD7E5D2E1DAE" box="[711,800,698,720]" gridcol="1" gridrow="11" pageId="2" pageNumber="380">14.4</td>
<td id="924D1678FFA1007CDDCBFD7E5A061DAE" box="[975,1032,698,720]" gridcol="2" gridrow="11" pageId="2" pageNumber="380">13.6</td>
<td id="924D1678FFA1007CDA55FD7E5ADE1DAE" box="[1105,1232,698,720]" gridcol="3" gridrow="11" pageId="2" pageNumber="380">13.0–14.4</td>
<td id="924D1678FFA1007CDB27FD7E5B4D1DAE" box="[1315,1347,698,720]" gridcol="4" gridrow="11" pageId="2" pageNumber="380">0.4</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FD205B4D1D84" box="[199,1347,740,762]" gridrow="12" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FD205C321D84" box="[199,572,740,762]" gridcol="0" gridrow="12" pageId="2" pageNumber="380">Body depth</th>
<td id="924D1678FFA1007CDCC3FD205D2E1D84" box="[711,800,740,762]" gridcol="1" gridrow="12" pageId="2" pageNumber="380">16.9</td>
<td id="924D1678FFA1007CDDCBFD205A061D84" box="[975,1032,740,762]" gridcol="2" gridrow="12" pageId="2" pageNumber="380">15.9</td>
<td id="924D1678FFA1007CDA55FD205ADE1D84" box="[1105,1232,740,762]" gridcol="3" gridrow="12" pageId="2" pageNumber="380">15.0–17.1</td>
<td id="924D1678FFA1007CDB27FD205B4D1D84" box="[1315,1347,740,762]" gridcol="4" gridrow="12" pageId="2" pageNumber="380">0.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FCC95B4D1C5D" box="[199,1347,781,803]" gridrow="13" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FCC95C321C5D" box="[199,572,781,803]" gridcol="0" gridrow="13" pageId="2" pageNumber="380">Caudal-peduncle depth</th>
<td id="924D1678FFA1007CDCC3FCC95D2E1C5D" box="[711,800,781,803]" gridcol="1" gridrow="13" pageId="2" pageNumber="380">13.8</td>
<td id="924D1678FFA1007CDDCBFCC95A061C5D" box="[975,1032,781,803]" gridcol="2" gridrow="13" pageId="2" pageNumber="380">13.3</td>
<td id="924D1678FFA1007CDA55FCC95ADE1C5D" box="[1105,1232,781,803]" gridcol="3" gridrow="13" pageId="2" pageNumber="380">12.5–13.8</td>
<td id="924D1678FFA1007CDB27FCC95B4D1C5D" box="[1315,1347,781,803]" gridcol="4" gridrow="13" pageId="2" pageNumber="380">0.5</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FCF25B4D1C32" box="[199,1347,822,844]" gridrow="14" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FCF25C321C32" box="[199,572,822,844]" gridcol="0" gridrow="14" pageId="2" pageNumber="380">Caudal-peduncle length</th>
<td id="924D1678FFA1007CDCC3FCF25D2E1C32" box="[711,800,822,844]" gridcol="1" gridrow="14" pageId="2" pageNumber="380">14.4</td>
<td id="924D1678FFA1007CDDCBFCF25A061C32" box="[975,1032,822,844]" gridcol="2" gridrow="14" pageId="2" pageNumber="380">14.1</td>
<td id="924D1678FFA1007CDA55FCF25ADE1C32" box="[1105,1232,822,844]" gridcol="3" gridrow="14" pageId="2" pageNumber="380">12.9–15.4</td>
<td id="924D1678FFA1007CDB27FCF25B4D1C32" box="[1315,1347,822,844]" gridcol="4" gridrow="14" pageId="2" pageNumber="380">0.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FCA45B4D1C08" box="[199,1347,864,886]" gridrow="15" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FCA45C321C08" box="[199,572,864,886]" gridcol="0" gridrow="15" pageId="2" pageNumber="380">Snout length</th>
<td id="924D1678FFA1007CDCC3FCA45D2E1C08" box="[711,800,864,886]" gridcol="1" gridrow="15" pageId="2" pageNumber="380">11.3</td>
<td id="924D1678FFA1007CDDCBFCA45A061C08" box="[975,1032,864,886]" gridcol="2" gridrow="15" pageId="2" pageNumber="380">10.6</td>
<td id="924D1678FFA1007CDA55FCA45ADE1C08" box="[1105,1232,864,886]" gridcol="3" gridrow="15" pageId="2" pageNumber="380">9.2–12.1</td>
<td id="924D1678FFA1007CDB27FCA45B4D1C08" box="[1315,1347,864,886]" gridcol="4" gridrow="15" pageId="2" pageNumber="380">0.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FC4D5B4D1CE1" box="[199,1347,905,927]" gridrow="16" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FC4D5C321CE1" box="[199,572,905,927]" gridcol="0" gridrow="16" pageId="2" pageNumber="380">Head width (at nares)</th>
<td id="924D1678FFA1007CDCC3FC4D5D2E1CE1" box="[711,800,905,927]" gridcol="1" gridrow="16" pageId="2" pageNumber="380">9.7</td>
<td id="924D1678FFA1007CDDCBFC4D5A061CE1" box="[975,1032,905,927]" gridcol="2" gridrow="16" pageId="2" pageNumber="380">10.6</td>
<td id="924D1678FFA1007CDA55FC4D5ADE1CE1" box="[1105,1232,905,927]" gridcol="3" gridrow="16" pageId="2" pageNumber="380">9.7–11.3</td>
<td id="924D1678FFA1007CDB27FC4D5B4D1CE1" box="[1315,1347,905,927]" gridcol="4" gridrow="16" pageId="2" pageNumber="380">0.6</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FC765B4D1CB6" box="[199,1347,946,968]" gridrow="17" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FC765C321CB6" box="[199,572,946,968]" gridcol="0" gridrow="17" pageId="2" pageNumber="380">Maximum head width</th>
<td id="924D1678FFA1007CDCC3FC765D2E1CB6" box="[711,800,946,968]" gridcol="1" gridrow="17" pageId="2" pageNumber="380">17.2</td>
<td id="924D1678FFA1007CDDCBFC765A061CB6" box="[975,1032,946,968]" gridcol="2" gridrow="17" pageId="2" pageNumber="380">16.7</td>
<td id="924D1678FFA1007CDA55FC765ADE1CB6" box="[1105,1232,946,968]" gridcol="3" gridrow="17" pageId="2" pageNumber="380">15.4–18.6</td>
<td id="924D1678FFA1007CDB27FC765B4D1CB6" box="[1315,1347,946,968]" gridcol="4" gridrow="17" pageId="2" pageNumber="380">0.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FC185B4D1C8C" box="[199,1347,988,1010]" gridrow="18" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FC185C321C8C" box="[199,572,988,1010]" gridcol="0" gridrow="18" pageId="2" pageNumber="380">Body width (at dorsal-fin origin)</th>
<td id="924D1678FFA1007CDCC3FC185D2E1C8C" box="[711,800,988,1010]" gridcol="1" gridrow="18" pageId="2" pageNumber="380">12.1</td>
<td id="924D1678FFA1007CDDCBFC185A061C8C" box="[975,1032,988,1010]" gridcol="2" gridrow="18" pageId="2" pageNumber="380">13.0</td>
<td id="924D1678FFA1007CDA55FC185ADE1C8C" box="[1105,1232,988,1010]" gridcol="3" gridrow="18" pageId="2" pageNumber="380">12.1–14.6</td>
<td id="924D1678FFA1007CDB27FC185B4D1C8C" box="[1315,1347,988,1010]" gridcol="4" gridrow="18" pageId="2" pageNumber="380">0.8</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FBC15B4D1B65" box="[199,1347,1029,1051]" gridrow="19" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FBC15C321B65" box="[199,572,1029,1051]" gridcol="0" gridrow="19" pageId="2" pageNumber="380">Body width (at anal-fin origin)</th>
<td id="924D1678FFA1007CDCC3FBC15D2E1B65" box="[711,800,1029,1051]" gridcol="1" gridrow="19" pageId="2" pageNumber="380">7.4</td>
<td id="924D1678FFA1007CDDCBFBC15A061B65" box="[975,1032,1029,1051]" gridcol="2" gridrow="19" pageId="2" pageNumber="380">7.2</td>
<td id="924D1678FFA1007CDA55FBC15ADE1B65" box="[1105,1232,1029,1051]" gridcol="3" gridrow="19" pageId="2" pageNumber="380">6.3–7.6</td>
<td id="924D1678FFA1007CDB27FBC15B4D1B65" box="[1315,1347,1029,1051]" gridcol="4" gridrow="19" pageId="2" pageNumber="380">0.5</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FBEA5B4D1B3A" box="[199,1347,1070,1092]" gridrow="20" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FBEA5C321B3A" box="[199,572,1070,1092]" gridcol="0" gridrow="20" pageId="2" pageNumber="380">Eye diameter</th>
<td id="924D1678FFA1007CDCC3FBEA5D2E1B3A" box="[711,800,1070,1092]" gridcol="1" gridrow="20" pageId="2" pageNumber="380">5.9</td>
<td id="924D1678FFA1007CDDCBFBEA5A061B3A" box="[975,1032,1070,1092]" gridcol="2" gridrow="20" pageId="2" pageNumber="380">6.3</td>
<td id="924D1678FFA1007CDA55FBEA5ADE1B3A" box="[1105,1232,1070,1092]" gridcol="3" gridrow="20" pageId="2" pageNumber="380">5.1–7.3</td>
<td id="924D1678FFA1007CDB27FBEA5B4D1B3A" box="[1315,1347,1070,1092]" gridcol="4" gridrow="20" pageId="2" pageNumber="380">0.7</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FB9C5B4D1B10" box="[199,1347,1112,1134]" gridrow="21" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FB9C5C321B10" box="[199,572,1112,1134]" gridcol="0" gridrow="21" pageId="2" pageNumber="380">Interorbital width</th>
<td id="924D1678FFA1007CDCC3FB9C5D2E1B10" box="[711,800,1112,1134]" gridcol="1" gridrow="21" pageId="2" pageNumber="380">7.4</td>
<td id="924D1678FFA1007CDDCBFB9C5A061B10" box="[975,1032,1112,1134]" gridcol="2" gridrow="21" pageId="2" pageNumber="380">7.8</td>
<td id="924D1678FFA1007CDA55FB9C5ADE1B10" box="[1105,1232,1112,1134]" gridcol="3" gridrow="21" pageId="2" pageNumber="380">6.7–8.9</td>
<td id="924D1678FFA1007CDB27FB9C5B4D1B10" box="[1315,1347,1112,1134]" gridcol="4" gridrow="21" pageId="2" pageNumber="380">0.8</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FB455B4D1BE9" box="[199,1347,1153,1175]" gridrow="22" pageId="2" pageNumber="380" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="924D1678FFA1007CDEC3FB455C321BE9" box="[199,572,1153,1175]" gridcol="0" gridrow="22" pageId="2" pageNumber="380">In percent of HL</th>
</tr>
<tr id="D19C7F04FFA1007CDEC3FB6E5B4D1BBE" box="[199,1347,1194,1216]" gridrow="23" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FB6E5C321BBE" box="[199,572,1194,1216]" gridcol="0" gridrow="23" pageId="2" pageNumber="380">Head depth (at eye)</th>
<td id="924D1678FFA1007CDCC3FB6E5D2E1BBE" box="[711,800,1194,1216]" gridcol="1" gridrow="23" pageId="2" pageNumber="380">50.0</td>
<td id="924D1678FFA1007CDDCBFB6E5A061BBE" box="[975,1032,1194,1216]" gridcol="2" gridrow="23" pageId="2" pageNumber="380">52.2</td>
<td id="924D1678FFA1007CDA55FB6E5ADE1BBE" box="[1105,1232,1194,1216]" gridcol="3" gridrow="23" pageId="2" pageNumber="380">50.0–54.7</td>
<td id="924D1678FFA1007CDB27FB6E5B4D1BBE" box="[1315,1347,1194,1216]" gridcol="4" gridrow="23" pageId="2" pageNumber="380">1.7</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FB105B4D1B94" box="[199,1347,1236,1258]" gridrow="24" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FB105C321B94" box="[199,572,1236,1258]" gridcol="0" gridrow="24" pageId="2" pageNumber="380">Head depth (at nape)</th>
<td id="924D1678FFA1007CDCC3FB105D2E1B94" box="[711,800,1236,1258]" gridcol="1" gridrow="24" pageId="2" pageNumber="380">58.3</td>
<td id="924D1678FFA1007CDDCBFB105A061B94" box="[975,1032,1236,1258]" gridcol="2" gridrow="24" pageId="2" pageNumber="380">58.1</td>
<td id="924D1678FFA1007CDA55FB105ADE1B94" box="[1105,1232,1236,1258]" gridcol="3" gridrow="24" pageId="2" pageNumber="380">55.7–61.0</td>
<td id="924D1678FFA1007CDB27FB105B4D1B94" box="[1315,1347,1236,1258]" gridcol="4" gridrow="24" pageId="2" pageNumber="380">2.0</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FB395B4D1A6D" box="[199,1347,1277,1299]" gridrow="25" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FB395C321A6D" box="[199,572,1277,1299]" gridcol="0" gridrow="25" pageId="2" pageNumber="380">Body depth</th>
<td id="924D1678FFA1007CDCC3FB395D2E1A6D" box="[711,800,1277,1299]" gridcol="1" gridrow="25" pageId="2" pageNumber="380">68.8</td>
<td id="924D1678FFA1007CDDCBFB395A061A6D" box="[975,1032,1277,1299]" gridcol="2" gridrow="25" pageId="2" pageNumber="380">68.2</td>
<td id="924D1678FFA1007CDA55FB395ADE1A6D" box="[1105,1232,1277,1299]" gridcol="3" gridrow="25" pageId="2" pageNumber="380">64.8–74.7</td>
<td id="924D1678FFA1007CDB27FB395B4D1A6D" box="[1315,1347,1277,1299]" gridcol="4" gridrow="25" pageId="2" pageNumber="380">3.3</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FAE25B4D1A42" box="[199,1347,1318,1340]" gridrow="26" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FAE25C321A42" box="[199,572,1318,1340]" gridcol="0" gridrow="26" pageId="2" pageNumber="380">Caudal-peduncle depth</th>
<td id="924D1678FFA1007CDCC3FAE25D2E1A42" box="[711,800,1318,1340]" gridcol="1" gridrow="26" pageId="2" pageNumber="380">56.3</td>
<td id="924D1678FFA1007CDDCBFAE25A061A42" box="[975,1032,1318,1340]" gridcol="2" gridrow="26" pageId="2" pageNumber="380">56.8</td>
<td id="924D1678FFA1007CDA55FAE25ADE1A42" box="[1105,1232,1318,1340]" gridcol="3" gridrow="26" pageId="2" pageNumber="380">53.4–60.0</td>
<td id="924D1678FFA1007CDB27FAE25B4D1A42" box="[1315,1347,1318,1340]" gridcol="4" gridrow="26" pageId="2" pageNumber="380">2.2</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FA945B4D1A18" box="[199,1347,1360,1382]" gridrow="27" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FA945C321A18" box="[199,572,1360,1382]" gridcol="0" gridrow="27" pageId="2" pageNumber="380">Caudal-peduncle length</th>
<td id="924D1678FFA1007CDCC3FA945D2E1A18" box="[711,800,1360,1382]" gridcol="1" gridrow="27" pageId="2" pageNumber="380">58.3</td>
<td id="924D1678FFA1007CDDCBFA945A061A18" box="[975,1032,1360,1382]" gridcol="2" gridrow="27" pageId="2" pageNumber="380">60.4</td>
<td id="924D1678FFA1007CDA55FA945ADE1A18" box="[1105,1232,1360,1382]" gridcol="3" gridrow="27" pageId="2" pageNumber="380">54.4–67.7</td>
<td id="924D1678FFA1007CDB27FA945B4D1A18" box="[1315,1347,1360,1382]" gridcol="4" gridrow="27" pageId="2" pageNumber="380">4.6</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FABD5B4D1AF1" box="[199,1347,1401,1423]" gridrow="28" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FABD5C321AF1" box="[199,572,1401,1423]" gridcol="0" gridrow="28" pageId="2" pageNumber="380">Snout length</th>
<td id="924D1678FFA1007CDCC3FABD5D2E1AF1" box="[711,800,1401,1423]" gridcol="1" gridrow="28" pageId="2" pageNumber="380">45.8</td>
<td id="924D1678FFA1007CDDCBFABD5A061AF1" box="[975,1032,1401,1423]" gridcol="2" gridrow="28" pageId="2" pageNumber="380">45.5</td>
<td id="924D1678FFA1007CDA55FABD5ADE1AF1" box="[1105,1232,1401,1423]" gridcol="3" gridrow="28" pageId="2" pageNumber="380">38.0–52.6</td>
<td id="924D1678FFA1007CDB27FABD5B4D1AF1" box="[1315,1347,1401,1423]" gridcol="4" gridrow="28" pageId="2" pageNumber="380">4.2</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FA665B4D1AC6" box="[199,1347,1442,1464]" gridrow="29" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FA665C321AC6" box="[199,572,1442,1464]" gridcol="0" gridrow="29" pageId="2" pageNumber="380">Head width (at nares)</th>
<td id="924D1678FFA1007CDCC3FA665D2E1AC6" box="[711,800,1442,1464]" gridcol="1" gridrow="29" pageId="2" pageNumber="380">39.6</td>
<td id="924D1678FFA1007CDDCBFA665A061AC6" box="[975,1032,1442,1464]" gridcol="2" gridrow="29" pageId="2" pageNumber="380">45.6</td>
<td id="924D1678FFA1007CDA55FA665ADE1AC6" box="[1105,1232,1442,1464]" gridcol="3" gridrow="29" pageId="2" pageNumber="380">39.6–49.3</td>
<td id="924D1678FFA1007CDB27FA665B4D1AC6" box="[1315,1347,1442,1464]" gridcol="4" gridrow="29" pageId="2" pageNumber="380">3.5</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FA085B4D1A9C" box="[199,1347,1484,1506]" gridrow="30" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FA085C321A9C" box="[199,572,1484,1506]" gridcol="0" gridrow="30" pageId="2" pageNumber="380">Maximum head width</th>
<td id="924D1678FFA1007CDCC3FA085D2E1A9C" box="[711,800,1484,1506]" gridcol="1" gridrow="30" pageId="2" pageNumber="380">69.8</td>
<td id="924D1678FFA1007CDDCBFA085A061A9C" box="[975,1032,1484,1506]" gridcol="2" gridrow="30" pageId="2" pageNumber="380">71.7</td>
<td id="924D1678FFA1007CDA55FA085ADE1A9C" box="[1105,1232,1484,1506]" gridcol="3" gridrow="30" pageId="2" pageNumber="380">65.9–81.1</td>
<td id="924D1678FFA1007CDB27FA085B4D1A9C" box="[1315,1347,1484,1506]" gridcol="4" gridrow="30" pageId="2" pageNumber="380">4.4</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3FA315B4D1975" box="[199,1347,1525,1547]" gridrow="31" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3FA315C321975" box="[199,572,1525,1547]" gridcol="0" gridrow="31" pageId="2" pageNumber="380">Body width (at dorsal-fin origin)</th>
<td id="924D1678FFA1007CDCC3FA315D2E1975" box="[711,800,1525,1547]" gridcol="1" gridrow="31" pageId="2" pageNumber="380">49.0</td>
<td id="924D1678FFA1007CDDCBFA315A061975" box="[975,1032,1525,1547]" gridcol="2" gridrow="31" pageId="2" pageNumber="380">55.9</td>
<td id="924D1678FFA1007CDA55FA315ADE1975" box="[1105,1232,1525,1547]" gridcol="3" gridrow="31" pageId="2" pageNumber="380">49.0–64.0</td>
<td id="924D1678FFA1007CDB27FA315B4D1975" box="[1315,1347,1525,1547]" gridcol="4" gridrow="31" pageId="2" pageNumber="380">4.4</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3F9DA5B4D194A" box="[199,1347,1566,1588]" gridrow="32" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3F9DA5C32194A" box="[199,572,1566,1588]" gridcol="0" gridrow="32" pageId="2" pageNumber="380">Body width (at anal-fin origin)</th>
<td id="924D1678FFA1007CDCC3F9DA5D2E194A" box="[711,800,1566,1588]" gridcol="1" gridrow="32" pageId="2" pageNumber="380">30.2</td>
<td id="924D1678FFA1007CDDCBF9DA5A06194A" box="[975,1032,1566,1588]" gridcol="2" gridrow="32" pageId="2" pageNumber="380">30.7</td>
<td id="924D1678FFA1007CDA55F9DA5ADE194A" box="[1105,1232,1566,1588]" gridcol="3" gridrow="32" pageId="2" pageNumber="380">27.4–32.9</td>
<td id="924D1678FFA1007CDB27F9DA5B4D194A" box="[1315,1347,1566,1588]" gridcol="4" gridrow="32" pageId="2" pageNumber="380">1.9</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3F98C5B4D1920" box="[199,1347,1608,1630]" gridrow="33" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3F98C5C321920" box="[199,572,1608,1630]" gridcol="0" gridrow="33" pageId="2" pageNumber="380">Eye diameter</th>
<td id="924D1678FFA1007CDCC3F98C5D2E1920" box="[711,800,1608,1630]" gridcol="1" gridrow="33" pageId="2" pageNumber="380">24.0</td>
<td id="924D1678FFA1007CDDCBF98C5A061920" box="[975,1032,1608,1630]" gridcol="2" gridrow="33" pageId="2" pageNumber="380">26.9</td>
<td id="924D1678FFA1007CDA55F98C5ADE1920" box="[1105,1232,1608,1630]" gridcol="3" gridrow="33" pageId="2" pageNumber="380">22.2–32.0</td>
<td id="924D1678FFA1007CDB27F98C5B4D1920" box="[1315,1347,1608,1630]" gridcol="4" gridrow="33" pageId="2" pageNumber="380">3.1</td>
</tr>
<tr id="D19C7F04FFA1007CDEC3F9B55B4D19F9" box="[199,1347,1649,1671]" gridrow="34" pageId="2" pageNumber="380">
<th id="924D1678FFA1007CDEC3F9B55C3219F9" box="[199,572,1649,1671]" gridcol="0" gridrow="34" pageId="2" pageNumber="380">Interorbital width</th>
<td id="924D1678FFA1007CDCC3F9B55D2E19F9" box="[711,800,1649,1671]" gridcol="1" gridrow="34" pageId="2" pageNumber="380">30.2</td>
<td id="924D1678FFA1007CDDCBF9B55A0619F9" box="[975,1032,1649,1671]" gridcol="2" gridrow="34" pageId="2" pageNumber="380">33.5</td>
<td id="924D1678FFA1007CDA55F9B55ADE19F9" box="[1105,1232,1649,1671]" gridcol="3" gridrow="34" pageId="2" pageNumber="380">28.8–39.0</td>
<td id="924D1678FFA1007CDB27F9B55B4D19F9" box="[1315,1347,1649,1671]" gridcol="4" gridrow="34" pageId="2" pageNumber="380">3.9</td>
</tr>
</table>
</paragraph>
<paragraph id="6F137D46FFA1FF81DEC3F9085F3A180B" blockId="2.[151,1437,1740,2016]" pageId="2" pageNumber="380">
Vertebrae 36–38 [37]; 18–19 [19] abdominal and 18–20 [18] caudal (n=8). Gas bladder in ossified capsule; two halves spatially separated, connected via a massive, flattened duct; secondary chamber small, conical, not ossified, in immediate contact with capsule (n=2) (
<figureCitation id="F79761C3FFA1FF81DC74F8D05CBB1852" box="[624,693,1812,1837]" captionStart="FIGURE 3" captionStartId="5.[151,250,935,957]" captionTargetBox="[155,1426,202,904]" captionTargetId="figure@5.[150,1436,193,913]" captionTargetPageId="5" captionText="FIGURE 3. Physoschistura absumbra, ZFMK-ICH 103627, paratype, 37.6 mm SL; gas bladder." httpUri="https://zenodo.org/record/573197/files/figure.png" pageId="2" pageNumber="380">Fig. 3</figureCitation>
). Gastrointestinal tract with simple, U-shaped stomach; intestines with single bend immediately after stomach. Largest recorded length 
<quantity id="A854D0A3FFA1FF81DDC5F8F35A271831" box="[961,1065,1847,1872]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" pageId="2" pageNumber="380" unit="mm" value="43.5">43.5 mm</quantity>
SL, 
<quantity id="A854D0A3FFA1FF81DA61F8F35AC31831" box="[1125,1229,1847,1872]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="2" pageNumber="380" unit="mm" value="53.6">53.6 mm</quantity>
total length (KIZ 2014006022).
</paragraph>
<paragraph id="6F137D46FFA1FF81DEC3F8BB5A8718A1" blockId="2.[151,1437,1740,2016]" pageId="2" pageNumber="380">Body covered by small, cycloid scales; body in front of dorsal fin and abdomen in front of pelvic fin naked. Lateral line complete, reaching caudal-fin base, with 79–91 pores. Cephalic lateralis system with 7–10 supraorbital, 3+12 infraorbital, 10–11 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="6F137D46FFA0FF80DEC3FF535FE61E1A" blockId="3.[151,1437,151,357]" pageId="3" pageNumber="381">Dorsal fin with 4 simple and 9½–10½ [9½] branched rays; reaching vertical through anal-fin origin; distal margin straight. Anal fin with 3 simple and 5½ branched rays; distal margin convex; not reaching caudal-fin base. Caudal fin deeply emarginate, with 9+8 branched rays; lobes rounded, lower lobe slightly longer. Pelvic fin with 8 rays, reaching anus, inserted about opposite to second branched dorsal-fin ray; fifth ray longest; distal margin convex. Pectoral fin with 11–12 [11] rays, reaching or nearly reaching vertical through dorsal-fin origin; fifth ray longest; distal margin convex.</paragraph>
</subSubSection>
<caption id="3BD32DCEFFA0FF80DE93FB715B5D1BB4" box="[151,1363,1204,1227]" httpUri="https://zenodo.org/record/573194/files/figure.png" pageId="3" pageNumber="381" targetBox="[150,1436,409,1183]" targetPageId="3">
<paragraph id="6F137D46FFA0FF80DE93FB715B5D1BB4" blockId="3.[151,1363,1204,1227]" box="[151,1363,1204,1227]" pageId="3" pageNumber="381">
<emphasis id="5DD8A154FFA0FF80DE93FB715F1D1BB4" bold="true" box="[151,275,1205,1227]" pageId="3" pageNumber="381">FIGURE 1.</emphasis>
<taxonomicName id="A8AC06C5FFA0FF80DF1DFB705C111BB4" box="[281,543,1204,1226]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="381" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA0FF80DF1DFB705C111BB4" box="[281,543,1204,1226]" italics="true" pageId="3" pageNumber="381">Physoschistura absumbra</emphasis>
</taxonomicName>
, KIZ 2014006018, holotype, 39.0 mm SL; Yunnan: Cangyuan; Nangunhe River.
</paragraph>
</caption>
<subSubSection id="27B62ECDFFA0FF80DEC3FB365AAC1930" pageId="3" pageNumber="381" type="materials_examined">
<paragraph id="6F137D46FFA0FF80DEC3FB365AAC1930" blockId="3.[151,1437,1266,2011]" pageId="3" pageNumber="381">
<emphasis id="5DD8A154FFA0FF80DEC3FB365C601A75" bold="true" box="[199,622,1266,1291]" pageId="3" pageNumber="381">Coloration of preserved specimens.</emphasis>
Body beige-gray, dorsum darker, abdomen lighter, with 9–12 dark gray bars, wider as interspaces; intensity of bars decreasing towards head. Bars straight, rather regular; interconnected over dorsum behind dorsal fin and over ventral midline behind anus; interrupted on caudal peduncle in some specimens. A conspicuous, black humeral spot; a black chevron-shaped stripe along midlateral from caudal-fin base reaching forward to about mid of dorsal fin, absent in some specimens. Bar at caudal-fin base conspicuous, black, straight, complete, not reaching dorsal and ventral extremities. Head gray, top above infraorbital cephalic lateralis canal and upper part of operculum dark gray; lower side with numerous densely set melanophores. Snout with a beige hue. Dorsal fin with a conspicuous, basal, black spot anterior to first branched ray and with a broad, transverse, median black band on light gray ground. Caudal fin with a broad transverse, median, dark gray band on light gray ground. Other fins with a narrow, transverse, dark gray band on light gray ground.
</paragraph>
</subSubSection>
<subSubSection id="27B62ECDFFA0FF80DEC3F99E5A47187D" pageId="3" pageNumber="381" type="distribution">
<paragraph id="6F137D46FFA0FF80DEC3F99E5F5719E8" blockId="3.[151,1437,1266,2011]" pageId="3" pageNumber="381">
<emphasis id="5DD8A154FFA0FF80DEC3F99E5F6E190D" bold="true" box="[199,352,1626,1651]" pageId="3" pageNumber="381">Distribution.</emphasis>
Known only from the 
<typeStatus id="B017C3E4FFA0FF80DC61F99F5C9B190D" box="[613,661,1627,1651]" pageId="3" pageNumber="381">type</typeStatus>
locality, a small nameless tributary to the Nangunhe River, Salween drainage (
<figureCitation id="F79761C3FFA0FF80DF02F9B95F4219E8" box="[262,332,1661,1686]" captionStart="FIGURE 4" captionStartId="6.[151,250,1748,1770]" captionTargetBox="[170,1416,354,1708]" captionTargetId="figure@6.[151,1436,337,1726]" captionTargetPageId="6" captionText="FIGURE 4. Map of Sino-Burmese border region, taken from Endruweit (2017). Symbol indicates the type locality of Physoschistura absumbra." httpUri="https://zenodo.org/record/573199/files/figure.png" pageId="3" pageNumber="381">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="6F137D46FFA0FF80DEC3F9665A47187D" blockId="3.[151,1437,1266,2011]" pageId="3" pageNumber="381">
<emphasis id="5DD8A154FFA0FF80DEC3F9665F2519C5" bold="true" box="[199,299,1698,1723]" pageId="3" pageNumber="381">Ecology.</emphasis>
Obtained from a small tributary, about 
<quantity id="A854D0A3FFA0FF80DCEAF9665D1519C4" box="[750,795,1698,1722]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="381" unit="m" value="2.0">2 m</quantity>
wide and 
<quantity id="A854D0A3FFA0FF80DD96F9665DD619C4" box="[914,984,1698,1722]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="381" unit="cm" value="30.0">30 cm</quantity>
deep at the collection site, running into the Nangunhe River; could not be obtained from the mainstem of the Nangunhe River. Co-occurring nemacheilids were 
<taxonomicName id="A8AC06C5FFA0FF80DED0F92E5FEB187C" box="[212,485,1770,1794]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="381" phylum="Chordata" rank="species" species="cryptofasciata">
<emphasis id="5DD8A154FFA0FF80DED0F92E5FEB187C" box="[212,485,1770,1794]" italics="true" pageId="3" pageNumber="381">Schistura cryptofasciata</emphasis>
</taxonomicName>
and 
<taxonomicName id="A8AC06C5FFA0FF80DC18F92E5CB2187C" box="[540,700,1770,1794]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="381" phylum="Chordata" rank="species" species="disparizona">
<emphasis id="5DD8A154FFA0FF80DC18F92E5CB2187C" box="[540,700,1770,1794]" italics="true" pageId="3" pageNumber="381">S. disparizona</emphasis>
</taxonomicName>
, both also present in the mainstem.
</paragraph>
</subSubSection>
<subSubSection id="27B62ECDFFA0FF80DEC3F8C95B281810" pageId="3" pageNumber="381" type="etymology">
<paragraph id="6F137D46FFA0FF80DEC3F8C95B281810" blockId="3.[151,1437,1266,2011]" pageId="3" pageNumber="381">
<emphasis id="5DD8A154FFA0FF80DEC3F8C95F401858" bold="true" box="[199,334,1805,1830]" pageId="3" pageNumber="381">Etymology.</emphasis>
From the mythology of the Wa people, the predominant ethnic minority along the Sinoburmese border in this region. A composite of the Latin words absum, to be absent; and umbra, shadow; an allusion to the Wa’s mythic ancestors, from whom it is said to cast no shadows (
<bibRefCitation id="0B3D00B7FFA0FF80DD74F8915A111810" author="Obayashi" box="[880,1055,1877,1902]" pageId="3" pageNumber="381" refString="Obayashi, T. (1966) Authropogonic Myths of the Wa in Northern Indo-China. Hitotsubashi Journal of Social Studies, 3, 43 - 66." type="journal article" year="1966">Obayashi 1966</bibRefCitation>
). A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="27B62ECDFFA0FF84DEC3F8BE5F961D42" lastPageId="7" lastPageNumber="385" pageId="3" pageNumber="381" type="discussion">
<paragraph id="6F137D46FFA0FF87DEC3F8BE5D111ED3" blockId="3.[151,1437,1266,2011]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="382" pageId="3" pageNumber="381">
<emphasis id="5DD8A154FFA0FF80DEC3F8BE5F4218ED" bold="true" box="[199,332,1914,1939]" pageId="3" pageNumber="381">Discussion.</emphasis>
Considering the quite high heterogeneity of species currently allocated in 
<taxonomicName id="A8AC06C5FFA0FF80DAABF8BE5B5518EC" box="[1199,1371,1914,1938]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="381" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA0FF80DAABF8BE5B5518EC" box="[1199,1371,1914,1938]" italics="true" pageId="3" pageNumber="381">Physoschistura</emphasis>
</taxonomicName>
there seem to be rather different opinions about its generic definition. Banarescu and Nalbant’s (1982) original generic concept encompasses species with a scaled body, at least posteriorly; an incomplete lateral line, not reaching behind the dorsal fin; a forked caudal fin; 8½–9½ branched dorsal-fin rays; a weak to moderate processus dentiformis; the two halves of the ossified air-bladder capsule merged and coalescent on the inner surfaces lacking a connective duct; a posterior non-ossified air-bladder chamber free in abdominal cavity; and a brownish barred color pattern. By the inclusion of 
<taxonomicName id="A8AC06C5FFA7FF87DC13FEC15C981E62" box="[535,662,260,284]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="382" phylum="Chordata" rank="species" species="elongata">
<emphasis id="5DD8A154FFA7FF87DC13FEC15C981E62" box="[535,662,260,284]" italics="true" pageId="4" pageNumber="382">P. elongata</emphasis>
</taxonomicName>
in the genus the authors stretched the concept to also accommodate species whose two halves of the air-bladder capsule are not merged but connected via a massive somewhat flattened duct. The peculiar structure of the gas-bladder capsule as present in the genus’ 
<typeStatus id="B017C3E4FFA7FF87DA68FE895A921E1B" box="[1132,1180,333,357]" pageId="4" pageNumber="382">type</typeStatus>
species, 
<emphasis id="5DD8A154FFA7FF87DB05FE895B921E1A" box="[1281,1436,332,356]" italics="true" pageId="4" pageNumber="382">
<taxonomicName id="A8AC06C5FFA7FF87DB05FE895B961E1A" box="[1281,1432,332,356]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="382" phylum="Chordata" rank="species" species="brunneana">P. brunneana</taxonomicName>
,
</emphasis>
of two merged halves with their inner surfaces coalescent and thereby making a connective duct-like structure redundant, has not been described from another congener.
</paragraph>
<caption id="3BD32DCEFFA7FF87DE93F8BC5AC918F0" box="[151,1223,1912,1934]" httpUri="https://zenodo.org/record/573195/files/figure.png" pageId="4" pageNumber="382" targetBox="[300,1287,481,1890]" targetPageId="4">
<paragraph id="6F137D46FFA7FF87DE93F8BC5AC918F0" blockId="4.[151,1223,1912,1934]" box="[151,1223,1912,1934]" pageId="4" pageNumber="382">
<emphasis id="5DD8A154FFA7FF87DE93F8BC5F1C18F3" bold="true" box="[151,274,1912,1934]" pageId="4" pageNumber="382">FIGURE 2.</emphasis>
<taxonomicName id="A8AC06C5FFA7FF87DF1DF8BC5C1118F0" box="[281,543,1912,1934]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="382" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA7FF87DF1DF8BC5C1118F0" box="[281,543,1912,1934]" italics="true" pageId="4" pageNumber="382">Physoschistura absumbra</emphasis>
</taxonomicName>
, ZFMK-ICH 103627, paratype, 37.6 mm SL; head in ventral view.
</paragraph>
</caption>
<caption id="3BD32DCEFFA6FF86DE93FC635A601CC3" box="[151,1134,935,957]" httpUri="https://zenodo.org/record/573197/files/figure.png" pageId="5" pageNumber="383" targetBox="[155,1426,202,904]" targetPageId="5">
<paragraph id="6F137D46FFA6FF86DE93FC635A601CC3" blockId="5.[151,1134,935,957]" box="[151,1134,935,957]" pageId="5" pageNumber="383">
<emphasis id="5DD8A154FFA6FF86DE93FC635F1C1CC2" bold="true" box="[151,274,935,957]" pageId="5" pageNumber="383">FIGURE 3.</emphasis>
<taxonomicName id="A8AC06C5FFA6FF86DF1DFC635C111CC3" box="[281,543,935,957]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA6FF86DF1DFC635C111CC3" box="[281,543,935,957]" italics="true" pageId="5" pageNumber="383">Physoschistura absumbra</emphasis>
</taxonomicName>
, ZFMK-ICH 103627, paratype, 37.6 mm SL; gas bladder.
</paragraph>
</caption>
<paragraph id="6F137D46FFA6FF86DEC3FC205DBD1A1B" blockId="5.[151,1437,996,2029]" pageId="5" pageNumber="383">
<bibRefCitation id="0B3D00B7FFA6FF86DEC3FC205F7A1C83" author="Kottelat" box="[199,372,996,1021]" pageId="5" pageNumber="383" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Fiet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat (1990)</bibRefCitation>
argued that the sole presence of a secondary gas-bladder chamber is insufficient to diagnose 
<taxonomicName id="A8AC06C5FFA6FF86DE93FBCC5F4C1B5E" box="[151,322,1032,1056]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DE93FBCC5F4C1B5E" box="[151,322,1032,1056]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
since some species of other genera also have a secondary chamber, as evidently present in 
<taxonomicName id="A8AC06C5FFA6FF86DB33FBCC5F3B1B3B" authority="Kottelat 1990" authorityName="Kottelat" authorityYear="1990" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="similis">
<emphasis id="5DD8A154FFA6FF86DB33FBCC5B921B5E" box="[1335,1436,1032,1056]" italics="true" pageId="5" pageNumber="383">S. similis</emphasis>
<bibRefCitation id="0B3D00B7FFA6FF86DE93FBE85F3B1B3B" author="Kottelat" box="[151,309,1068,1093]" pageId="5" pageNumber="383" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Fiet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat 1990</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="A8AC06C5FFA6FF86DF71FBE85DE61B3B" authority="(Boulenger 1893) (Kottelat 1990)" authorityName="(Boulenger 1893) (Kottelat" authorityYear="1990" box="[373,1000,1068,1093]" class="Actinopterygii" family="Nemacheilidae" genus="Petruichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="brevis">
<emphasis id="5DD8A154FFA6FF86DF71FBE85C5F1B3A" box="[373,593,1068,1092]" italics="true" pageId="5" pageNumber="383">Petruichthys brevis</emphasis>
(
<bibRefCitation id="0B3D00B7FFA6FF86DC61FBE85D2E1B3B" author="Boulenger" box="[613,800,1068,1093]" pageId="5" pageNumber="383" refString="Boulenger, G. A. (1893) List of the fishes collected by Mr. E. W. Oates in the southern Shan States, and presented by him to the British Museum. Annals and Magazine of Natural History, 12, 198 - 203." type="journal article" year="1893">Boulenger 1893</bibRefCitation>
) (
<bibRefCitation id="0B3D00B7FFA6FF86DD3FFBE85DD11B3B" author="Kottelat" box="[827,991,1068,1093]" pageId="5" pageNumber="383" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Fiet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat 1990</bibRefCitation>
)
</taxonomicName>
. At least two more 
<taxonomicName id="A8AC06C5FFA6FF86DADDFBE85B4D1B3A" box="[1241,1347,1068,1092]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DADDFBE85B4D1B3A" box="[1241,1347,1068,1092]" italics="true" pageId="5" pageNumber="383">Schistura</emphasis>
</taxonomicName>
have a secondary chamber; namely 
<taxonomicName id="A8AC06C5FFA6FF86DFE6FB945A051B16" authority="Lokeshwor &amp; Vishwanath 2012" authorityName="Lokeshwor &amp; Vishwanath" authorityYear="2012" box="[482,1035,1104,1129]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="koladynensis">
<emphasis id="5DD8A154FFA6FF86DFE6FB945C9C1B16" box="[482,658,1104,1128]" italics="true" pageId="5" pageNumber="383">S. koladynensis</emphasis>
Lokeshwor &amp; Vishwanath 2012
</taxonomicName>
and 
<taxonomicName id="A8AC06C5FFA6FF86DA4CFB945F521BF3" authority="Lokeshwor &amp; Vishwanath 2013" authorityName="Lokeshwor &amp; Vishwanath" authorityYear="2013" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="porocephala">
<emphasis id="5DD8A154FFA6FF86DA4CFB945AFA1B16" box="[1096,1268,1104,1128]" italics="true" pageId="5" pageNumber="383">S. porocephala</emphasis>
<bibRefCitation id="0B3D00B7FFA6FF86DAFAFB945F521BF3" author="Lokeshwor" pageId="5" pageNumber="383" refString="Lokeshwor, Y. &amp; Vishwanath, W. (2013) Schistura porocephala, a new nemacheilid loach from Koladyne basin, Mizoram, India (Teleostei: Nemacheilidae). Ichthyological Research, 60, 159 - 162." type="journal article" year="2013">Lokeshwor &amp; Vishwanath 2013</bibRefCitation>
</taxonomicName>
, both from the Kaladan River. 
<bibRefCitation id="0B3D00B7FFA6FF86DCB7FBB05D561BF3" author="Kottelat" box="[691,856,1140,1165]" pageId="5" pageNumber="383" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Fiet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat (1990</bibRefCitation>
, 
<bibRefCitation id="0B3D00B7FFA6FF86DD67FBB15D911BF3" author="Kottelat" box="[867,927,1141,1165]" pageId="5" pageNumber="383" refString="Kottelat, M. (2001) Fishes of Laos. WHT Publications, Colombo, 198 pp." type="book" year="2001">2001</bibRefCitation>
), instead, considered a strongly arched mouth (1.5–2.0 times wider than long) and a conspicuously notched lower lip that forms triangular furrowed lateral pads chief characters to rediagnose the genus. Contradictory, 
<bibRefCitation id="0B3D00B7FFA6FF86DD0FFB785DF81BAB" box="[779,1014,1212,1237]" pageId="5" pageNumber="383" refString="Kottelat, M., Harries, D. R. &amp; Proudlove, G. S. (2007) Schistura papulifera, a new species of cave loach from Meghalaya, India (Teleostei: Balitoridae). Zootaxa, 1393, 34 - 44." type="journal article">
Kottelat 
<emphasis id="5DD8A154FFA6FF86DD68FB795D931BAA" box="[876,925,1212,1236]" italics="true" pageId="5" pageNumber="383">et al</emphasis>
. (2007)
</bibRefCitation>
placed a subterranean nemacheilid in 
<taxonomicName id="A8AC06C5FFA6FF86DE93FB245F0F1B86" box="[151,257,1248,1272]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DE93FB245F0F1B86" box="[151,257,1248,1272]" italics="true" pageId="5" pageNumber="383">Schistura</emphasis>
</taxonomicName>
although it possesses the diagnostic peculiar mouth structure. 
<bibRefCitation id="0B3D00B7FFA6FF86DDC1FB245A7C1B86" author="Kottelat" box="[965,1138,1248,1273]" pageId="5" pageNumber="383" refString="Kottelat, M. (2001) Fishes of Laos. WHT Publications, Colombo, 198 pp." type="book" year="2001">Kottelat (2001)</bibRefCitation>
and 
<bibRefCitation id="0B3D00B7FFA6FF86DAAEFB245B761B86" box="[1194,1400,1248,1273]" pageId="5" pageNumber="383" refString="Chen, X. Y., Kottelat, M. &amp; Neely, D. A. (2011) Physoschistura yunnaniloides, a new species of loach from Myanmar (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 22, 179 - 183." type="journal article">
Chen 
<emphasis id="5DD8A154FFA6FF86DAE9FB255B2E1B86" box="[1261,1312,1248,1272]" italics="true" pageId="5" pageNumber="383">et al</emphasis>
. (2011)
</bibRefCitation>
do not even list the secondary chamber in their definition of 
<taxonomicName id="A8AC06C5FFA6FF86DD18FAC05DC91A62" box="[796,967,1284,1308]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DD18FAC05DC91A62" box="[796,967,1284,1308]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
. Following this generic definition several other 
<taxonomicName id="A8AC06C5FFA6FF86DEDDFAEC5F4C1A3E" box="[217,322,1320,1344]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DEDDFAEC5F4C1A3E" box="[217,322,1320,1344]" italics="true" pageId="5" pageNumber="383">Schistura</emphasis>
</taxonomicName>
would have to be relocated to 
<taxonomicName id="A8AC06C5FFA6FF86DCA7FAEC5D401A3E" box="[675,846,1320,1344]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DCA7FAEC5D401A3E" box="[675,846,1320,1344]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
; for example 
<taxonomicName id="A8AC06C5FFA6FF86DDEAFAEC5AFD1A3F" authority="Nguyen 2005" authorityName="Nguyen" authorityYear="2005" box="[1006,1267,1320,1345]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="hoai">
<emphasis id="5DD8A154FFA6FF86DDEAFAEC5A301A3E" box="[1006,1086,1320,1344]" italics="true" pageId="5" pageNumber="383">S. hoai</emphasis>
(
<bibRefCitation id="0B3D00B7FFA6FF86DA49FAEC5AE51A3F" author="Nguyen" box="[1101,1259,1320,1345]" pageId="5" pageNumber="383" refString="Nguyen, V. H. (2005) Freshwater Fishes of Fietnam. Volume 2. Agriculture Publishing House, Hanoi, 760 pp. [in Vietnamese]" type="book" year="2005">Nguyen 2005</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DB04FAEC5ED41A1B" authority="Zhu 1982" authorityName="Zhu" authorityYear="1982" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="longa">
<emphasis id="5DD8A154FFA6FF86DB04FAEC5B531A3E" box="[1280,1373,1320,1344]" italics="true" pageId="5" pageNumber="383">S. longa</emphasis>
(
<bibRefCitation id="0B3D00B7FFA6FF86DB69FAEC5EDD1A1B" author="Zhu" pageId="5" pageNumber="383" refString="Zhu, S. Q. (1982) Five new species of fishes of the genus Nemachilus from Yunnan Province, China. Acta Zootaxonomica Sinica, 7, 104 - 111. [in Chinese]" type="journal article" year="1982">Zhu 1982</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DEE2FA895FE81A1B" authority="Smith 1945" authorityName="Smith" authorityYear="1945" box="[230,486,1356,1381]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="poculi">
<emphasis id="5DD8A154FFA6FF86DEE2FA895F461A1A" box="[230,328,1356,1380]" italics="true" pageId="5" pageNumber="383">S. poculi</emphasis>
(
<bibRefCitation id="0B3D00B7FFA6FF86DF53FA885FD31A1B" author="Smith" box="[343,477,1356,1381]" pageId="5" pageNumber="383" refString="Smith, H. M. (1945) The fresh-water fishes of Siam, or Thailand. Bulletin of the United States National Museum, 188, 1 - 622. https: // doi. org / 10.5479 / si. 03629236.188.1" type="journal article" year="1945">Smith 1945</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="A8AC06C5FFA6FF86DC25FA895DBE1A1B" authority="Zheng et al. 2012" authorityName="Zheng et al." authorityYear="2012" box="[545,944,1356,1381]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="prolixifasciata">
<emphasis id="5DD8A154FFA6FF86DC25FA895CD11A1A" box="[545,735,1356,1380]" italics="true" pageId="5" pageNumber="383">S. prolixifasciata</emphasis>
<bibRefCitation id="0B3D00B7FFA6FF86DCE2FA885DBE1A1B" author="Zheng" box="[742,944,1356,1381]" pageId="5" pageNumber="383" refString="Zheng, L. P., Yang, J. X. &amp; Chen, X. Y. (2012) Schistura prolixifasciata, a new species of loach (Teleostei: Nemacheilidae) from the Salween basin in Yunnan, China. Ichthyological Exploration of Freshwaters, 23, 63 - 68." type="journal article" year="2012">
Zheng 
<emphasis id="5DD8A154FFA6FF86DD31FA895D681A1A" box="[821,870,1356,1380]" italics="true" pageId="5" pageNumber="383">et al</emphasis>
. 2012
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="6F137D46FFA6FF86DEC3FAB45A9C191E" blockId="5.[151,1437,996,2029]" pageId="5" pageNumber="383">
The definition of 
<taxonomicName id="A8AC06C5FFA6FF86DF8EFAB45C3B1AF6" box="[394,565,1392,1416]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DF8EFAB45C3B1AF6" box="[394,565,1392,1416]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
, as applied by 
<bibRefCitation id="0B3D00B7FFA6FF86DCDFFAB45D881AF6" author="Kottelat" box="[731,902,1392,1417]" pageId="5" pageNumber="383" refString="Kottelat, M. (2001) Fishes of Laos. WHT Publications, Colombo, 198 pp." type="book" year="2001">Kottelat (2001)</bibRefCitation>
and 
<bibRefCitation id="0B3D00B7FFA6FF86DDBFFAB45A881AF6" box="[955,1158,1392,1417]" pageId="5" pageNumber="383" refString="Chen, X. Y., Kottelat, M. &amp; Neely, D. A. (2011) Physoschistura yunnaniloides, a new species of loach from Myanmar (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 22, 179 - 183." type="journal article">
Chen 
<emphasis id="5DD8A154FFA6FF86DDF8FAB55A231AF6" box="[1020,1069,1392,1416]" italics="true" pageId="5" pageNumber="383">et al</emphasis>
. (2011)
</bibRefCitation>
, is evidently stretched. It is far distant from Banarescu and Nalbant’s (1982) original generic concept and is herein rejected. ‘ 
<emphasis id="5DD8A154FFA6FF86DAE3FA505F2C1AAE" italics="true" pageId="5" pageNumber="383">Physoschistura’ walongensis</emphasis>
<bibRefCitation id="0B3D00B7FFA6FF86DF2CFA7D5C2D1AAF" author="Tamang" box="[296,547,1464,1489]" pageId="5" pageNumber="383" refString="Tamang, L. &amp; Sinha, B. (2016) Physoschistura walongensis, a new species of loach (Teleostei: Nemacheilidae) from Arunachal Pradesh, northeastern India. Zootaxa, 4173, 280 - 288." type="journal article" year="2016">Tamang &amp; Sinha 2016</bibRefCitation>
and ‘ 
<emphasis id="5DD8A154FFA6FF86DC66FA7D5C8E1AAE" box="[610,640,1465,1488]" italics="true" pageId="5" pageNumber="383">P.’</emphasis>
<taxonomicName id="A8AC06C5FFA6FF86DC82FA7C5DEF1AAE" authority="Chen et al. 2011" authorityName="Chen et al." authorityYear="2011" box="[646,993,1464,1489]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="yunnaniloides">
<emphasis id="5DD8A154FFA6FF86DC82FA7C5D2D1AAE" box="[646,803,1464,1488]" italics="true" pageId="5" pageNumber="383">yunnaniloides</emphasis>
<bibRefCitation id="0B3D00B7FFA6FF86DD2EFA7C5DEF1AAE" author="Chen" box="[810,993,1464,1489]" pageId="5" pageNumber="383" refString="Chen, X. Y., Kottelat, M. &amp; Neely, D. A. (2011) Physoschistura yunnaniloides, a new species of loach from Myanmar (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 22, 179 - 183." type="journal article" year="2011">
Chen 
<emphasis id="5DD8A154FFA6FF86DD68FA7D5D931AAE" box="[876,925,1464,1488]" italics="true" pageId="5" pageNumber="383">et al</emphasis>
. 2011
</bibRefCitation>
</taxonomicName>
cannot be considered congeneric, since they lack the secondary chamber and are, consequently, relocated to 
<taxonomicName id="A8AC06C5FFA6FF86DD9EFA185A0A1A8A" box="[922,1028,1500,1524]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DD9EFA185A0A1A8A" box="[922,1028,1500,1524]" italics="true" pageId="5" pageNumber="383">Schistura</emphasis>
</taxonomicName>
. Apparently, these two species have been described in 
<taxonomicName id="A8AC06C5FFA6FF86DF69F9C45C161966" box="[365,536,1536,1560]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DF69F9C45C161966" box="[365,536,1536,1560]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
simply based on the mouth structure. Six more species with a complete (vs. incomplete in the original generic concept) lateral line have been placed in 
<taxonomicName id="A8AC06C5FFA6FF86DA64F9E05B051942" box="[1120,1291,1572,1596]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DA64F9E05B051942" box="[1120,1291,1572,1596]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
, namely 
<taxonomicName id="A8AC06C5FFA6FF86DB83F9E15F38191E" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="chindwinensis">
<emphasis id="5DD8A154FFA6FF86DB83F9E15F38191E" italics="true" pageId="5" pageNumber="383">P. chindwinensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DF40F98D5FCC191E" box="[324,450,1608,1632]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="prashadi">
<emphasis id="5DD8A154FFA6FF86DF40F98D5FCC191E" box="[324,450,1608,1632]" italics="true" pageId="5" pageNumber="383">P. prashadi</emphasis>
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DFD4F98D5C56191E" box="[464,600,1608,1632]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="shanensis">
<emphasis id="5DD8A154FFA6FF86DFD4F98D5C56191E" box="[464,600,1608,1632]" italics="true" pageId="5" pageNumber="383">P. shanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DC63F98D5D4B191E" box="[615,837,1608,1632]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="shuangjiangensis">
<emphasis id="5DD8A154FFA6FF86DC63F98D5D4B191E" box="[615,837,1608,1632]" italics="true" pageId="5" pageNumber="383">P. shuangjiangensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DD57F98D5DB4191E" box="[851,954,1609,1632]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5DD8A154FFA6FF86DD57F98D5DB4191E" box="[851,954,1609,1632]" italics="true" pageId="5" pageNumber="383">P. tigrina</emphasis>
</taxonomicName>
, and 
<taxonomicName id="A8AC06C5FFA6FF86DDFCF98D5A85191E" box="[1016,1163,1609,1632]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="tuivaiensis">
<emphasis id="5DD8A154FFA6FF86DDFCF98D5A85191E" box="[1016,1163,1609,1632]" italics="true" pageId="5" pageNumber="383">P. tuivaiensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="6F137D46FFA6FF86DEC3F9A85A0A19B3" blockId="5.[151,1437,996,2029]" pageId="5" pageNumber="383">
The new species, 
<taxonomicName id="A8AC06C5FFA6FF86DF90F9A85CB719FA" box="[404,697,1644,1668]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA6FF86DF90F9A85CB719FA" box="[404,697,1644,1668]" italics="true" pageId="5" pageNumber="383">Physoschistura absumbra</emphasis>
</taxonomicName>
, is a member of 
<taxonomicName id="A8AC06C5FFA6FF86DD85F9A85A2219FA" box="[897,1068,1644,1668]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="genus">
<emphasis id="5DD8A154FFA6FF86DD85F9A85A2219FA" box="[897,1068,1644,1668]" italics="true" pageId="5" pageNumber="383">Physoschistura</emphasis>
</taxonomicName>
as diagnosed by 
<bibRefCitation id="0B3D00B7FFA6FF86DAF1F9A95F4D19D6" author="Banarescu" pageId="5" pageNumber="383" refString="Banarescu, P. M. &amp; Nalbant, T. T. (1982) In: Singh, A., Sen, N., Banarescu, P. M. &amp; Nalbant, T. T. New noemacheiline loaches from India (Pisces, Cobitidae). Travaux du Museum d'Histoire Naturelle &quot; Grigore Antipa &quot;, 23, 201 - 212." type="journal article" year="1982">Banarescu and Nalbant (1982)</bibRefCitation>
. It also possesses an arched mouth (1.5–1.8 times wider than long) and a notched lower lip, laterally slightly triangular, characters used by 
<bibRefCitation id="0B3D00B7FFA6FF86DC45F9705CE319B3" author="Kottelat" box="[577,749,1716,1741]" pageId="5" pageNumber="383" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Fiet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat (1990)</bibRefCitation>
to rediagnose the genus.
</paragraph>
<paragraph id="6F137D46FFA6FF86DED6F91C5A3A1847" blockId="5.[151,1437,996,2029]" pageId="5" pageNumber="383">
<taxonomicName id="A8AC06C5FFA6FF86DED6F91C5FF6198E" box="[210,504,1752,1776]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA6FF86DED6F91C5FF6198E" box="[210,504,1752,1776]" italics="true" pageId="5" pageNumber="383">Physoschistura absumbra</emphasis>
</taxonomicName>
is readily distinguished from all congeners in having a complete lateral line, a continuous lower jaw, and in lacking a suborbital lobe. Its occurrence in the Nangunhe River extends the distributional range of the genus further northward (upstream) within the Salween.
</paragraph>
<paragraph id="6F137D46FFA6FF86DEC3F8805B921893" blockId="5.[151,1437,996,2029]" pageId="5" pageNumber="383">
The new species somewhat resembles the other Salween species 
<taxonomicName id="A8AC06C5FFA6FF86DDBCF8815A411822" box="[952,1103,1860,1884]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="brunneana">
<emphasis id="5DD8A154FFA6FF86DDBCF8815A411822" box="[952,1103,1860,1884]" italics="true" pageId="5" pageNumber="383">P. brunneana</emphasis>
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA6FF86DA65F8815AA41822" box="[1121,1194,1861,1884]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="raoi">
<emphasis id="5DD8A154FFA6FF86DA65F8815AA41822" box="[1121,1194,1861,1884]" italics="true" pageId="5" pageNumber="383">P. raoi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="A8AC06C5FFA6FF86DAF5F8815B721822" box="[1265,1404,1860,1884]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="rivulicola">
<emphasis id="5DD8A154FFA6FF86DAF5F8815B721822" box="[1265,1404,1860,1884]" italics="true" pageId="5" pageNumber="383">P. rivulicola</emphasis>
</taxonomicName>
in lacking a suborbital lobe and in having a small reported adult size of less than 
<quantity id="A854D0A3FFA6FF86DDFBF8AC5A4318FE" box="[1023,1101,1896,1920]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="383" unit="mm" value="44.0">44 mm</quantity>
and a barred color pattern but differs from them in having a lower jaw that is not notched (vs. notched in all three species); a complete (vs. incomplete) lateral line; basicaudal bar complete (vs. dissociated); head depth at nape 13–14% SL (vs. 
<quantity id="A854D0A3FFA6FF86DB19F8755B8E18B7" box="[1309,1408,1968,1993]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.318" metricValueMin="3.556" pageId="5" pageNumber="383" unit="in" value="15.5" valueMax="17.0" valueMin="14.0">14–17 in</quantity>
<taxonomicName id="A8AC06C5FFA6FF86DB83F8755F1E1892" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="brunneana">
<emphasis id="5DD8A154FFA6FF86DB83F8755F1E1892" italics="true" pageId="5" pageNumber="383">P. brunneana</emphasis>
</taxonomicName>
; 
<quantity id="A854D0A3FFA6FF86DF25F8105F521893" box="[289,348,2004,2029]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="5" pageNumber="383" unit="in" value="15.0">15 in</quantity>
<taxonomicName id="A8AC06C5FFA6FF86DF61F8115FBF1892" box="[357,433,2005,2028]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="raoi">
<emphasis id="5DD8A154FFA6FF86DF61F8115FBF1892" box="[357,433,2005,2028]" italics="true" pageId="5" pageNumber="383">P. raoi</emphasis>
</taxonomicName>
; 
<quantity id="A854D0A3FFA6FF86DFC6F8105C291893" box="[450,551,2004,2029]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="5" pageNumber="383" unit="in" value="15.5" valueMax="16.0" valueMin="15.0">15–16 in</quantity>
<taxonomicName id="A8AC06C5FFA6FF86DC34F8115CB41892" box="[560,698,2004,2028]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="383" phylum="Chordata" rank="species" species="rivulicola">
<emphasis id="5DD8A154FFA6FF86DC34F8115CB41892" box="[560,698,2004,2028]" italics="true" pageId="5" pageNumber="383">P. rivulicola</emphasis>
</taxonomicName>
); and caudal-peduncle depth 13–14% SL (vs. 9–12; 13; 12–13).
</paragraph>
<paragraph id="6F137D46FFA5FF85DE93FF535B101E62" blockId="6.[151,1437,151,285]" pageId="6" pageNumber="384">
Likewise, 
<taxonomicName id="A8AC06C5FFA5FF85DF08FF5D5F971FCE" box="[268,409,152,176]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA5FF85DF08FF5D5F971FCE" box="[268,409,152,176]" italics="true" pageId="6" pageNumber="384">P. absumbra</emphasis>
</taxonomicName>
shares with the remaining Salween species 
<taxonomicName id="A8AC06C5FFA5FF85DD8AFF5D5A161FCE" box="[910,1048,152,176]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="shanensis">
<emphasis id="5DD8A154FFA5FF85DD8AFF5D5A161FCE" box="[910,1048,152,176]" italics="true" pageId="6" pageNumber="384">P. shanensis</emphasis>
</taxonomicName>
a small reported adult size of less than 
<quantity id="A854D0A3FFA5FF85DED5FF785F2D1FAA" box="[209,291,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="384" unit="mm" value="44.0">44 mm</quantity>
and, in addition, a complete lateral line. 
<taxonomicName id="A8AC06C5FFA5FF85DD00FF785A271FAA" box="[772,1065,188,212]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA5FF85DD00FF785A271FAA" box="[772,1065,188,212]" italics="true" pageId="6" pageNumber="384">Physoschistura absumbra</emphasis>
</taxonomicName>
is readily distinguished from 
<taxonomicName id="A8AC06C5FFA5FF85DB83FF795F0A1F86" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="shanensis">
<emphasis id="5DD8A154FFA5FF85DB83FF795F0A1F86" italics="true" pageId="6" pageNumber="384">P. shanensis</emphasis>
</taxonomicName>
in having a lower that is not notched (vs. notched); a barred (vs. mottled) color pattern; basicaudal bar complete (vs. dissociated); and head length dorsally 23–25% SL (vs. 21), laterally 26–28% SL (vs. 22).
</paragraph>
<caption id="3BD32DCEFFA5FF85DE93F9105FAB1876" httpUri="https://zenodo.org/record/573199/files/figure.png" pageId="6" pageNumber="384" targetBox="[170,1416,354,1708]" targetPageId="6">
<paragraph id="6F137D46FFA5FF85DE93F9105FAB1876" blockId="6.[151,1436,1748,1800]" pageId="6" pageNumber="384">
<emphasis id="5DD8A154FFA5FF85DE93F9105F161997" bold="true" box="[151,280,1748,1770]" pageId="6" pageNumber="384">FIGURE 4.</emphasis>
Map of Sino-Burmese border region, taken from Endruweit (2017). Symbol indicates the type locality of 
<taxonomicName id="A8AC06C5FFA5FF85DE93F9365FAF1876" box="[151,417,1778,1800]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="absumbra">Physoschistura absumbra</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="6F137D46FFA5FF84DEC3F8F15CB11E62" blockId="6.[151,1436,1845,2014]" lastBlockId="7.[151,1437,151,573]" lastPageId="7" lastPageNumber="385" pageId="6" pageNumber="384">
Three valid species of 
<emphasis id="5DD8A154FFA5FF85DFC7F8F15C7B1833" box="[451,629,1845,1869]" italics="true" pageId="6" pageNumber="384">
<taxonomicName id="A8AC06C5FFA5FF85DFC7F8F15C7F1833" box="[451,625,1845,1869]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="genus">Physoschistura</taxonomicName>
,
</emphasis>
namely 
<taxonomicName id="A8AC06C5FFA5FF85DCD1F8F25D9E1833" box="[725,912,1845,1869]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="chindwinensis">
<emphasis id="5DD8A154FFA5FF85DCD1F8F25D9E1833" box="[725,912,1845,1869]" italics="true" pageId="6" pageNumber="384">P. chindwinensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A8AC06C5FFA5FF85DD99F8F25A121833" box="[925,1052,1845,1869]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="prashadi">
<emphasis id="5DD8A154FFA5FF85DD99F8F25A121833" box="[925,1052,1845,1869]" italics="true" pageId="6" pageNumber="384">P. prashadi</emphasis>
</taxonomicName>
and 
<taxonomicName id="A8AC06C5FFA5FF85DA56F8F25AB71833" box="[1106,1209,1846,1869]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5DD8A154FFA5FF85DA56F8F25AB71833" box="[1106,1209,1846,1869]" italics="true" pageId="6" pageNumber="384">P. tigrina</emphasis>
</taxonomicName>
, have been reported from the drainage of the Chindwin River, a right bank tributary to the 
<collectingRegion id="AD68B3A4FFA5FF85DDBEF89D5A3F180C" box="[954,1073,1881,1906]" country="Myanmar" name="Ayeyawady" pageId="6" pageNumber="384">Irrawaddy</collectingRegion>
; none from other tributaries nor the mainstem of the 
<collectingRegion id="AD68B3A4FFA5FF85DF7AF8B95FFF18E8" box="[382,497,1917,1942]" country="Myanmar" name="Ayeyawady" pageId="6" pageNumber="384">Irrawaddy</collectingRegion>
hitherto. The Chindwin River is formed by a network of rivers in the Indoburmese border region and is geographically far distant from the Salween. 
<taxonomicName id="A8AC06C5FFA5FF85DD81F8655AA618C7" box="[901,1192,1953,1977]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="384" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA5FF85DD81F8655AA618C7" box="[901,1192,1953,1977]" italics="true" pageId="6" pageNumber="384">Physoschistura absumbra</emphasis>
</taxonomicName>
shares with the three species occurring in the Chindwin drainage a continuous lower jaw and a complete lateral line. It is easily distinguished from them in lacking (vs. having) a suborbital, having a barred (vs. irregularly barred and blotched) color pattern; HL 23–25% SL (vs. 
<quantity id="A854D0A3FFA4FF84DC27FF785C891FAB" box="[547,647,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.588" metricValueMin="4.826" pageId="7" pageNumber="385" unit="in" value="20.5" valueMax="22.0" valueMin="19.0">19–22 in</quantity>
<taxonomicName id="A8AC06C5FFA4FF84DC8AFF795D441FAA" box="[654,842,188,212]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="chindwinensis">
<emphasis id="5DD8A154FFA4FF84DC8AFF795D441FAA" box="[654,842,188,212]" italics="true" pageId="7" pageNumber="385">P. chindwinensis</emphasis>
</taxonomicName>
; 
<quantity id="A854D0A3FFA4FF84DD5EFF785D9D1FAB" box="[858,915,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="7" pageNumber="385" unit="in" value="22.0">22 in</quantity>
<taxonomicName id="A8AC06C5FFA4FF84DD9FFF795A151FAA" box="[923,1051,188,212]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="prashadi">
<emphasis id="5DD8A154FFA4FF84DD9FFF795A151FAA" box="[923,1051,188,212]" italics="true" pageId="7" pageNumber="385">P. prashadi</emphasis>
</taxonomicName>
; 
<quantity id="A854D0A3FFA4FF84DA2FFF785A811FAB" box="[1067,1167,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="7" pageNumber="385" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
<taxonomicName id="A8AC06C5FFA4FF84DA93FF795AF31FAA" box="[1175,1277,189,212]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5DD8A154FFA4FF84DA93FF795AF31FAA" box="[1175,1277,189,212]" italics="true" pageId="7" pageNumber="385">P. tigrina</emphasis>
</taxonomicName>
); and caudalpeduncle depth 13–14% SL (vs. 9–10; 12; 10–11). Besides, 
<taxonomicName id="A8AC06C5FFA4FF84DD43FF255DA31F89" box="[839,941,224,248]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="tigrina">
<emphasis id="5DD8A154FFA4FF84DD43FF255DA31F89" box="[839,941,224,248]" italics="true" pageId="7" pageNumber="385">P. tigrina</emphasis>
</taxonomicName>
is a large species reaching an adult size of about 
<quantity id="A854D0A3FFA4FF84DED8FEC05F251E62" box="[220,299,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="385" unit="mm" value="74.0">74 mm</quantity>
SL (vs. 
<quantity id="A854D0A3FFA4FF84DF8DFEC05FD71E62" box="[393,473,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="385" unit="mm" value="44.0">44 mm</quantity>
SL in 
<taxonomicName id="A8AC06C5FFA4FF84DC20FEC15CA11E62" box="[548,687,260,284]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA4FF84DC20FEC15CA11E62" box="[548,687,260,284]" italics="true" pageId="7" pageNumber="385">P. absumbra</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="6F137D46FFA4FF84DEC3FEEC5F961D42" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="385">
<taxonomicName id="A8AC06C5FFA4FF84DEC3FEEC5FFD1E3E" box="[199,499,296,320]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA4FF84DEC3FEEC5FFD1E3E" box="[199,499,296,320]" italics="true" pageId="7" pageNumber="385">Physoschistura absumbra</emphasis>
</taxonomicName>
shares with 
<taxonomicName id="A8AC06C5FFA4FF84DC9BFEED5D6F1E3E" box="[671,865,296,320]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="chulabhornae">
<emphasis id="5DD8A154FFA4FF84DC9BFEED5D6F1E3E" box="[671,865,296,320]" italics="true" pageId="7" pageNumber="385">P. chulabhornae</emphasis>
</taxonomicName>
from the Chao Phraya drainage and with 
<taxonomicName id="A8AC06C5FFA4FF84DB83FEED5F6E1E1A" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="pseudobrunneana">
<emphasis id="5DD8A154FFA4FF84DB83FEED5F6E1E1A" italics="true" pageId="7" pageNumber="385">P. pseudobrunneana</emphasis>
</taxonomicName>
and 
<taxonomicName id="A8AC06C5FFA4FF84DF91FE895C7D1E1A" box="[405,627,332,356]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="shuangjiangensis">
<emphasis id="5DD8A154FFA4FF84DF91FE895C7D1E1A" box="[405,627,332,356]" italics="true" pageId="7" pageNumber="385">P. shuangjiangensis</emphasis>
</taxonomicName>
both from the Mekong a continuous lower jaw, but is distinct from them in lacking (vs. having) a suborbital lobe and having a complete (vs. incomplete in 
<taxonomicName id="A8AC06C5FFA4FF84DA83FEB55B4B1EF6" box="[1159,1349,368,392]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="chulabhornae">
<emphasis id="5DD8A154FFA4FF84DA83FEB55B4B1EF6" box="[1159,1349,368,392]" italics="true" pageId="7" pageNumber="385">P. chulabhornae</emphasis>
</taxonomicName>
and 
<taxonomicName id="A8AC06C5FFA4FF84DB83FEB55F6E1ED2" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="pseudobrunneana">
<emphasis id="5DD8A154FFA4FF84DB83FEB55F6E1ED2" italics="true" pageId="7" pageNumber="385">P. pseudobrunneana</emphasis>
</taxonomicName>
; complete in 
<taxonomicName id="A8AC06C5FFA4FF84DFF9FE515CD21ED2" box="[509,732,404,428]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="shuangjiangensis">
<emphasis id="5DD8A154FFA4FF84DFF9FE515CD21ED2" box="[509,732,404,428]" italics="true" pageId="7" pageNumber="385">P. shuangjiangensis</emphasis>
</taxonomicName>
) lateral line; HL 23–25% SL (18–23; 21–23; 20–23); caudalpeduncle depth 13–14% SL (vs. 8–12; 
<quantity id="A854D0A3FFA4FF84DC4DFE7C5CA51EAE" box="[585,683,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="7" pageNumber="385" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11–13 in</quantity>
the latter two); and body depth 15–17% SL (vs. 16–23; 20–22; 18– 21). Besides, the four species differ notably in color pattern: 
<taxonomicName id="A8AC06C5FFA4FF84DD68FE195DF21E8A" box="[876,1020,476,500]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="absumbra">
<emphasis id="5DD8A154FFA4FF84DD68FE195DF21E8A" box="[876,1020,476,500]" italics="true" pageId="7" pageNumber="385">P. absumbra</emphasis>
</taxonomicName>
possesses a rather regularly barred pattern whereas it is spotted in 
<taxonomicName id="A8AC06C5FFA4FF84DFF7FDC55CA71D66" box="[499,681,512,536]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="chulabhornae">
<emphasis id="5DD8A154FFA4FF84DFF7FDC55CA71D66" box="[499,681,512,536]" italics="true" pageId="7" pageNumber="385">P. chulabhornae</emphasis>
</taxonomicName>
, irregularly barred or blotched in 
<taxonomicName id="A8AC06C5FFA4FF84DA21FDC55B071D66" box="[1061,1289,512,536]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="pseudobrunneana">
<emphasis id="5DD8A154FFA4FF84DA21FDC55B071D66" box="[1061,1289,512,536]" italics="true" pageId="7" pageNumber="385">P. pseudobrunneana</emphasis>
</taxonomicName>
, and mottled in 
<taxonomicName id="A8AC06C5FFA4FF84DEB0FDE15F9F1D42" box="[180,401,548,572]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="385" phylum="Chordata" rank="species" species="shuangjiangensis">
<emphasis id="5DD8A154FFA4FF84DEB0FDE15F9F1D42" box="[180,401,548,572]" italics="true" pageId="7" pageNumber="385">P. shuangjiangensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>