277 lines
21 KiB
XML
277 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.36.7225" ID-PMC="PMC4023340" ID-Pensoft-Pub="1314-2003-36-27" ID-Pensoft-UUID="FFBEFA574912FF83FFB42C16FF81FF96" ID-PubMed="24843291" ID-Zenodo-Dep="576213" ModsDocID="1314-2003-36-27" checkinTime="1451251756806" checkinUser="pensoft" docAuthor="Droissart, Vincent, Cribb, Phillip J., Simo-Droissart, Murielle & Stevart, Tariq" docDate="2014" docId="C595858CB97B5102A217BEF8E077FA59" docLanguage="en" docName="PhytoKeys 36: 27-34" docOrigin="PhytoKeys 36" docPubDate="2014-04-09" docSource="http://dx.doi.org/10.3897/phytokeys.36.7225" docTitle="Distylodon sonkeanum Droissart, Stevart & P. J. Cribb 2014, sp. nov." docType="treatment" docVersion="3" id="FFBEFA574912FF83FFB42C16FF81FF96" lastPageNumber="30" masterDocId="FFBEFA574912FF83FFB42C16FF81FF96" masterDocTitle="Taxonomy of Atlantic Central African orchids 2. A second species of the rare genus Distylodon (Orchidaceae, Angraecinae) collected in Cameroon" masterLastPageNumber="34" masterPageNumber="27" pageNumber="27" updateTime="1668141088527" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomy of Atlantic Central African orchids 2. A second species of the rare genus Distylodon (Orchidaceae, Angraecinae) collected in Cameroon</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Droissart, Vincent</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9798-5616</mods:nameIdentifier>
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<mods:affiliation>Institut de Recherche pour le Developpement (IRD), Unite Mixte de Recherche AMAP (Botanique et Bioinformatique de l'Architecture des Plantes), Boulevard de la Lironde, TA A- 51 / PS 2, F- 34398 Montpellier Cedex 5, France & Missouri Botanical Garden, Africa & Madagascar Department, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. & Plant Systematic and Ecology Laboratory, Higher Teacher's Training College, University of Yaounde I, Yaounde, Cameroon & Herbarium et Bibliotheque de Botanique africaine, CP 265, Universite Libre de Bruxelles, Boulevard du Triomphe, B- 1050, Brussels, Belgium</mods:affiliation>
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<mods:nameIdentifier type="email">vincent.droissart@ird.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cribb, Phillip J.</mods:namePart>
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<mods:affiliation>Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AB, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Simo-Droissart, Murielle</mods:namePart>
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<mods:affiliation>Plant Systematic and Ecology Laboratory, Higher Teacher's Training College, University of Yaounde I, Yaounde, Cameroon</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stevart, Tariq</mods:namePart>
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<mods:affiliation>Missouri Botanical Garden, Africa & Madagascar Department, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. & Herbarium et Bibliotheque de Botanique africaine, CP 265, Universite Libre de Bruxelles, Boulevard du Triomphe, B- 1050, Brussels, Belgium & Botanic Garden, Meise, Domein van Bouchout, Nieuwelaan 38, B- 1860 Meise, Belgium</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-04-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>36</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>27</mods:start>
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<mods:end>34</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.36.7225</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.36.7225</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-36-27</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFBEFA574912FF83FFB42C16FF81FF96</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576213</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182228746" LSID="urn:lsid:plazi:treatment:C595858CB97B5102A217BEF8E077FA59" httpUri="http://treatment.plazi.org/id/C595858CB97B5102A217BEF8E077FA59" lastPageId="3" lastPageNumber="30" pageId="0" pageNumber="27">
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<subSubSection pageId="0" pageNumber="27" type="nomenclature">
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<paragraph pageId="0" pageNumber="27">
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<taxonomicName LSID="C595858C-B97B-5102-A217-BEF8E077FA59" authority="Droissart, Stévart & P. J. Cribb" authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Distylodon sonkeanum" order="Asparagales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="sonkeanum" status="sp. nov.">
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Distylodon sonkeanum Droissart,
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<normalizedToken originalValue="Stévart">Stevart</normalizedToken>
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& P.J.Cribb
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="27">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Illustration of Distylodon sonkeanum (Droissart, Stevart & Simo M. 585): A habitus B flower, diagonal view C flower, side view D dorsal sepal E lateral sepal F petal G lip H column with pollinium, sideview I column with anthercap, frontal J pollinium with stipe K stipe." httpUri="https://binary.pensoft.net/fig/10629" pageId="0" pageNumber="27">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of living specimen of Distylodon sonkeanum: A front view of the flower B side view of the flower C inflorescence." httpUri="https://binary.pensoft.net/fig/10630" pageId="0" pageNumber="27">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="27" type="diagnosis">
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<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="27">
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="0" pageNumber="27">Distylodon sonkeanum</emphasis>
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</taxonomicName>
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Droissart,
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<normalizedToken originalValue="Stévart">Stevart</normalizedToken>
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& P.J.Cribb, is close to
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<taxonomicName authorityName="Summerh" authorityYear="1966" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="comptum">
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<emphasis italics="true" pageId="0" pageNumber="27">Distylodon comptum</emphasis>
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</taxonomicName>
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Summerh. but differs from it in having several-flowered inflorescences, longer leaves and spur, and shorter pedicel and ovary.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="1" lastPageNumber="28" pageId="0" pageNumber="27" type="type">
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<paragraph pageId="0" pageNumber="27">Type.</paragraph>
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<paragraph lastPageId="1" lastPageNumber="28" pageId="0" pageNumber="27">
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Cameroon. Bifa (piste sur la route Kribi-Ebolowa),
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<normalizedToken originalValue="à">a</normalizedToken>
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5 km au SE de Zingui, le long
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<normalizedToken originalValue="d’une">d'une</normalizedToken>
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piste de chasseurs entre les
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<normalizedToken originalValue="rivières">rivieres</normalizedToken>
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<normalizedToken originalValue="Nyété">Nyete</normalizedToken>
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et Nyamefoo,
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<geoCoordinate degrees="02" direction="north" minutes="41.308" orientation="latitude" precision="1" value="2.6884665">
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<pageBreakToken pageId="1" pageNumber="28" start="start">02</pageBreakToken>
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°41.308'N
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</geoCoordinate>
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,
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<geoCoordinate degrees="010" direction="east" minutes="16.406" orientation="longitude" precision="1" value="10.273434">010°16.406'E</geoCoordinate>
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, 5 July 2007,
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<emphasis italics="true" pageId="1" pageNumber="28">
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Droissart,
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<normalizedToken originalValue="Stévart">Stevart</normalizedToken>
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& Simo M. 585
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</emphasis>
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(holotype: BRLU!; isotype: YA!).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28" type="description">
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<paragraph pageId="1" pageNumber="28">Description.</paragraph>
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<paragraph lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
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Dwarf epiphytic herb. Stem short, up to 8 mm long, 2-2.5 mm in diameter, leafy, unbranched. Roots more than 120 mm long, 1-1.8 mm in diameter, numerous, distributed at the base of the plant, unbranched. Leaves few (up to 5), 25-37
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<normalizedToken originalValue="×">x</normalizedToken>
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6-11 mm, obliquely narrowly elliptic, slightly coriaceous, with entire margins, with an indistinct and irregular reticulate venation; leaf apex obliquely subacute, unequally bilobed, with distance between the two lobes less than 1 mm. Inflorescences up to 41 mm long, 2- to 5-flowered, almost horizontal to pendant, unbranched, emerging at the base of the stem, with peduncle 13 mm long; floral internode about 6-7 mm, one flower per node; rachis terete. Bracts up to 1.2 mm long, tubular. Flowers 17-25
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<normalizedToken originalValue="×">x</normalizedToken>
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6-8 mm, not opening widely, green, the larger one situated at the apex of the inflorescence. Ovary and pedicel not twisted, 5-7 mm long. Dorsal sepal 4-7
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<normalizedToken originalValue="×">x</normalizedToken>
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1.6-1.9 mm, linear lanceolate, acute, slightly to markedly reflexed, with entire margins. Lateral sepals 5.5-9
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<normalizedToken originalValue="×">x</normalizedToken>
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1.8-2.0 mm, obliquely linear-lanceolate, acute, slightly to markedly reflexed, with entire margins. Petals 3.0-4.8
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<normalizedToken originalValue="×">x</normalizedToken>
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1.1-1.3 mm, linear lanceolate, acuminate, slightly curved forward, with entire margins. Lip 4.5-6.0
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<normalizedToken originalValue="×">x</normalizedToken>
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2.0-3.5 mm, slightly to markedly trilobed in the basal half; side lobe obliquely triangular, subacute to acute, 0.8-1.8 mm long; mid lobe much longer, 3-4 mm long, linear-subulate, acuminate, fleshy, somewhat curved forward; spur 13.5-18.0 mm long, 0.8-1.1 mm in diameter,
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<pageBreakToken pageId="2" pageNumber="29" start="start">cylindrical</pageBreakToken>
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, straight, slightly inflated in the apical part in larger flowers, apex rounded. Column 1-1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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1.0 mm, almost cylindrical with truncate apex. Rostellum 0.3 mm long, consists of two erect, subulate teeth or fangs. Anthercap 1.0
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<normalizedToken originalValue="×">x</normalizedToken>
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1.0 mm, deltoid. Pollinia two, spherical. Viscidia two, with two stipites 0.7-0.8 mm long, independent of each other, flattened, bifurcate.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/10629" pageId="2" pageNumber="29" start="Figure 1" startId="F1">
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<paragraph pageId="2" pageNumber="29">
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<emphasis bold="true" pageId="2" pageNumber="29">Figure 1.</emphasis>
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Illustration of
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="2" pageNumber="29">Distylodon sonkeanum</emphasis>
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</taxonomicName>
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(
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<emphasis italics="true" pageId="2" pageNumber="29">
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Droissart,
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<normalizedToken originalValue="Stévart">Stevart</normalizedToken>
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& Simo M. 585
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</emphasis>
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):
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<emphasis bold="true" pageId="2" pageNumber="29">A</emphasis>
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<emphasis italics="true" pageId="2" pageNumber="29">habitus</emphasis>
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<emphasis bold="true" pageId="2" pageNumber="29">B</emphasis>
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flower, diagonal view
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<emphasis bold="true" pageId="2" pageNumber="29">C</emphasis>
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flower, side view
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<emphasis bold="true" pageId="2" pageNumber="29">D</emphasis>
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dorsal sepal
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<emphasis bold="true" pageId="2" pageNumber="29">E</emphasis>
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lateral sepal
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<emphasis bold="true" pageId="2" pageNumber="29">F</emphasis>
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petal
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<emphasis bold="true" pageId="2" pageNumber="29">G</emphasis>
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lip
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<emphasis bold="true" pageId="2" pageNumber="29">H</emphasis>
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column with pollinium, sideview
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<emphasis bold="true" pageId="2" pageNumber="29">I</emphasis>
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column with anthercap, frontal
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<emphasis bold="true" pageId="2" pageNumber="29">J</emphasis>
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pollinium with stipe
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<emphasis bold="true" pageId="2" pageNumber="29">K</emphasis>
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stipe.
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</paragraph>
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</caption>
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<caption httpUri="https://binary.pensoft.net/fig/10630" pageId="2" pageNumber="29" start="Figure 2" startId="F2">
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<paragraph pageId="2" pageNumber="29">
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<emphasis bold="true" pageId="2" pageNumber="29">Figure 2.</emphasis>
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Photographs of living specimen of
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="2" pageNumber="29">Distylodon sonkeanum</emphasis>
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</taxonomicName>
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:
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<emphasis bold="true" pageId="2" pageNumber="29">A</emphasis>
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front view of the flower
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<emphasis bold="true" pageId="2" pageNumber="29">B</emphasis>
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side view of the flower
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<emphasis bold="true" pageId="2" pageNumber="29">C</emphasis>
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inflorescence.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="29" type="distribution and habitat">
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<paragraph pageId="2" pageNumber="29">Distribution and habitat.</paragraph>
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<paragraph pageId="2" pageNumber="29">
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="2" pageNumber="29">Distylodon sonkeanum</emphasis>
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</taxonomicName>
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is endemic to the coastal part of the Lower Guinea Domain (
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<bibRefCitation author="White, F" journalOrPublisher="Bulletin du Jardin botanique national de Belgique" pageId="5" pageNumber="32" pagination="11 - 55" refId="B11" refString="White, F, 1979. The Guineo-Congolian Region and its relationships to other phytochoria. Bulletin du Jardin botanique national de Belgique 49: 11 - 55" title="The Guineo-Congolian Region and its relationships to other phytochoria." volume="49" year="1979">White 1979</bibRefCitation>
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) of the Guineo-Congolian Region. It is known from a single locality in South Region of Cameroon (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of Distylodon sonkeanum and Distylodon comptum in tropical Africa." httpUri="https://binary.pensoft.net/fig/10631" pageId="2" pageNumber="29">Fig. 3</figureCitation>
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). The only population known so far was found in the lowland evergreen forest at 100 m elevation, growing epiphytically on a fallen branch.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/10631" pageId="2" pageNumber="29" start="Figure 3" startId="F3">
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<paragraph pageId="2" pageNumber="29">
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<emphasis bold="true" pageId="2" pageNumber="29">Figure 3.</emphasis>
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Distribution of
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="2" pageNumber="29">Distylodon sonkeanum</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Summerh" authorityYear="1966" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="comptum">
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<emphasis italics="true" pageId="2" pageNumber="29">Distylodon comptum</emphasis>
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</taxonomicName>
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in tropical Africa.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="conservation">
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<paragraph pageId="2" pageNumber="29">Conservation.</paragraph>
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<paragraph pageId="2" pageNumber="29">
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IUCN red list category: Critically Endangered, [CR B2ab(iii)].
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="2" pageNumber="29">Distylodon sonkeanum</emphasis>
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</taxonomicName>
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is only known from one very restricted subpopulation in Cameroon which represents one
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<normalizedToken originalValue="‘location’">'location'</normalizedToken>
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. The main threats known to the species in the southern Cameroon are deforestation for logging and oil palm (
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<taxonomicName class="Monocotyledoneae" family="Arecaceae" genus="Elaeis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Principes" pageId="2" pageNumber="29" phylum="Angiospermae" rank="species" species="guineensis">
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<emphasis italics="true" pageId="2" pageNumber="29">Elaeis guineensis</emphasis>
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</taxonomicName>
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Jacq.) and rubber (
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<taxonomicName class="Dicotyledoneae" family="Euphorbiaceae" genus="Hevea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Geraniales" pageId="2" pageNumber="29" phylum="Angiospermae" rank="species" species="brasiliensis">
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<emphasis italics="true" pageId="2" pageNumber="29">Hevea brasiliensis</emphasis>
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</taxonomicName>
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(Willd. Ex A. Juss.)
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<normalizedToken originalValue="Müll">Muell</normalizedToken>
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.Arg.) plantations, resulting from a strong international demand. The ongoing loss of the forest leads us to expect a continuous decline of its habitat in the only known locality, and therefore of its extent of occurrence and area of occupancy. Moreover, this location is accessible to local residents and through their practices of shifting agriculture and small-scale timber exploitation; they are gradually transforming this area into secondary forest.
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</paragraph>
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<paragraph pageId="3" pageNumber="30">
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<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="species" species="sonkeanum">
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<emphasis italics="true" pageId="3" pageNumber="30">
|
||
<pageBreakToken pageId="3" pageNumber="30" start="start">Distylodon</pageBreakToken>
|
||
sonkeanum
|
||
</emphasis>
|
||
</taxonomicName>
|
||
appears to be rare but, due to its inconspicuous habit, further fieldwork is required to ascertain its conservation status more objectively. Considering the general habitat of
|
||
<taxonomicName authorityName="Droissart, Stevart & P. J. Cribb" authorityYear="2014" class="Liliopsida" family="Orchidaceae" genus="Distylodon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="species" species="sonkeanum">
|
||
<emphasis italics="true" pageId="3" pageNumber="30">Distylodon sonkeanum</emphasis>
|
||
</taxonomicName>
|
||
, it is to be hoped that more specimens and additional sites will be found in the low-elevation parts of the adjacent Campo
|
||
<normalizedToken originalValue="Ma’an">Ma'an</normalizedToken>
|
||
National Park. Based on our current knowledge of the ecology and the distribution of the species (one location and AOO less than 10 km2), and using the Criteria B of the IUCN Red List Categories and Criteria, the species was assessed as CR B2ab(iii).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="30" type="etymology">
|
||
<paragraph pageId="3" pageNumber="30">Etymology.</paragraph>
|
||
<paragraph pageId="3" pageNumber="30">
|
||
The species epithet '
|
||
<emphasis italics="true" pageId="3" pageNumber="30">sonkeanum</emphasis>
|
||
' refers to the Professor Bonaventure
|
||
<normalizedToken originalValue="Sonké">Sonke</normalizedToken>
|
||
of the University of
|
||
<normalizedToken originalValue="Yaoundé">Yaounde</normalizedToken>
|
||
I, who is an internationally recognized taxonomist, specialized in the
|
||
<taxonomicName genus="Rubiaceae" lsidName="" pageId="3" pageNumber="30" rank="genus">Rubiaceae</taxonomicName>
|
||
's family. He has collected extensively in the Bipindi-Akom II area, and adjacent Ngovayang massif and
|
||
<normalizedToken originalValue="Campo-Ma’an">Campo-Ma'an</normalizedToken>
|
||
National Park. He was one of the first to highlight the high biodiversity of this area and he greatly facilitated our work in Cameroon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |