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<mods:title id="319B6F3B59E41973C0ED4F88258E9B9C">Correcting sponge names: nomenclatural update of lower taxa level Porifera</mods:title>
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<heading id="6C103F05FFA331179786FCBEBFDB048B" bold="true" box="[151,697,870,897]" fontSize="11" level="2" pageId="68" pageNumber="69" reason="4">
<taxonomicName id="F0E7F3EAFFA331179786FCBEBFDB048B" ID-CoL="5PH59" authority="Hentschel, 1929" authorityName="Hentschel" authorityYear="1929" box="[151,697,870,897]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="variety" species="frigidus" variety="gracilis">
<emphasis id="0593547BFFA331179786FCBEBFDB048B" bold="true" box="[151,697,870,897]" pageId="68" pageNumber="69">
<emphasis id="0593547BFFA331179786FCBEBC34048A" bold="true" box="[151,342,870,897]" italics="true" pageId="68" pageNumber="69">Iophon frigidus</emphasis>
var.
<emphasis id="0593547BFFA331179682FCBEBC8F048A" bold="true" box="[403,493,870,897]" italics="true" pageId="68" pageNumber="69">gracilis</emphasis>
<bibRefCitation id="5376F598FFA3311796E5FCBFBFDB048B" author="Hentschel, E." box="[500,697,870,897]" pageId="68" pageNumber="69" pagination="857 - 1042" refId="ref97583" refString="Hentschel, E. (1929) Die Kiesel- und Hornschwamme des N ˆ rdlichen Eismeers. In: R ˆ mer, F., Schaudinn, F., Brauer, A. &amp; Arndt, W. (Eds.), Fauna Arctica. Eine Zusammenstellung der arktischen Tierformen mit besonderer Ber ¸ cksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das N ˆ rdliche Eismeer im Jahre 1898. 5 (4). G. Fischer, Jena, pp. 857 - 1042, pls. XII - XIV." type="book chapter" year="1929">Hentschel, 1929</bibRefCitation>
</emphasis>
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</paragraph>
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<treatmentCitation id="B646AE78FFA331179786FC73BFF004C8" author="Hentschel, E." box="[151,658,939,963]" page="884" pageId="68" pageNumber="69" year="1929">
<taxonomicName id="F0E7F3EAFFA331179786FC73BFF004C8" ID-CoL="5PH59" authority="Hentschel, 1929: 884" authorityName="Hentschel" authorityPageNumber="884" authorityYear="1929" box="[151,658,939,963]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="variety" species="frigidus" variety="gracilis">
<emphasis id="0593547BFFA331179786FC73BC5604C8" box="[151,308,939,963]" italics="true" pageId="68" pageNumber="69">Iophon frigidus</emphasis>
var.
<emphasis id="0593547BFFA331179677FC73BCD104C8" box="[358,435,939,963]" italics="true" pageId="68" pageNumber="69">gracilis</emphasis>
<bibRefCitation id="5376F598FFA3311796ABFC73BFF004C8" author="Hentschel, E." box="[442,658,939,963]" pageId="68" pageNumber="69" pagination="857 - 1042" refId="ref97583" refString="Hentschel, E. (1929) Die Kiesel- und Hornschwamme des N ˆ rdlichen Eismeers. In: R ˆ mer, F., Schaudinn, F., Brauer, A. &amp; Arndt, W. (Eds.), Fauna Arctica. Eine Zusammenstellung der arktischen Tierformen mit besonderer Ber ¸ cksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das N ˆ rdliche Eismeer im Jahre 1898. 5 (4). G. Fischer, Jena, pp. 857 - 1042, pls. XII - XIV." type="book chapter" year="1929">Hentschel, 1929: 884</bibRefCitation>
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(no illustration).
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<paragraph id="37588869FFA331179786FC29BD89020C" blockId="68.[151,1437,1009,1828]" pageId="68" pageNumber="69">
The variety described by Hentschel from North Spitsbergen,
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N of Ross Island,
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, depth
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(
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in ZMH), and the White Sea,
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, depth
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(collected by the Deutschen Expedition in das N̂rdliche Eismeer im Jahre 1898) was erected primarily on the basis of a single character: the possession of bipocillae, which were lacking in the typical variety described by
<bibRefCitation id="5376F598FFA331179492FB85B91D037C" author="Lundbeck, W." box="[899,1151,1117,1144]" pageId="68" pageNumber="69" pagination="1 - 219" refId="ref101887" refString="Lundbeck, W. (1905) Porifera. (Part II.) Desmacidonidae (Pars). The Danish Ingolf-Expedition, 6 (2), 1 - 219. [https: // biodiversitylibrary. org / page / 2177257]" type="journal article" year="1905">Lundbeck (1905: 183)</bibRefCitation>
from the East
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shelf,
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, depth
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,
<typeStatus id="E85C36CBFFA33117954DFB5ABFDC0397" box="[604,702,1154,1180]" pageId="68" pageNumber="69" type="syntype">syntypes</typeStatus>
ZMUC DEM 18 and 26), and by
<bibRefCitation id="5376F598FFA33117933DFB59B8720390" author="Levinsen, G. M. R." box="[1068,1296,1153,1179]" pageId="68" pageNumber="69" pagination="339 - 372" refId="ref101143" refString="Levinsen, G. M. R. (1887) Kara-Havets Svampe (Porifera). Dijmphna-Togtets zoologisk-botaniske Udbytte, 1, 339 - 372, pls. XXIX - XXXI. [https: // books. google. nl / books? id = PDzPAAAAMAAJ &amp; printsec = frontcover &amp; source = gbs _ atb &amp; redir _ esc = y # v = onepage &amp; q &amp; f = false]" type="journal article" year="1887">Levinsen (1887: 360</bibRefCitation>
as
<taxonomicName id="F0E7F3EAFFA331179223FB59BDB603B4" baseAuthorityName="Vosmaer" baseAuthorityYear="1882" class="Demospongiae" family="Hymedesmiidae" genus="Esperella" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="picea">
<emphasis id="0593547BFFA331179223FB59BDB603B4" italics="true" pageId="68" pageNumber="69">Esperella picea</emphasis>
</taxonomicName>
) from the Kara Sea, and by
<bibRefCitation id="5376F598FFA331179530FB7DBE7003B4" author="Hentschel, E." box="[545,786,1189,1215]" pageId="68" pageNumber="69" pagination="1 - 18" refId="ref97529" refString="Hentschel, E. (1916) 3 - Die Spongien des Eisfjords In: Zoologische Ergebnisse der Schwedischen Expedition nach Spitzbergen, 1908. Teil II. Kungliga Svenska vetenskapsakademiens Handlingar, 54 (3), 1 - 18. [https: // biodiversitylibrary. org / page / 41924362]" type="journal article" year="1916">Hentschel (1916: 10)</bibRefCitation>
from Spitsbergen. Further differences were more subtle: slightly smaller megascleres and anisochelae with a narrow upper half distinct from the broader shape in the typical variety.
</paragraph>
</subSubSection>
<subSubSection id="7FFDDBE2FFA3311797D6FACAB9220028" pageId="68" pageNumber="69" type="discussion">
<paragraph id="37588869FFA3311797D6FACABFC8029C" blockId="68.[151,1437,1009,1828]" pageId="68" pageNumber="69">
Burton (1932: 348) discussed the global diversity of species of
<taxonomicName id="F0E7F3EAFFA3311794B5FAC9BE910220" ID-CoL="84P9X" authorityName="Gray" authorityYear="1867" box="[932,1011,1297,1323]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="genus">
<emphasis id="0593547BFFA3311794B5FAC9BE910220" box="[932,1011,1297,1323]" italics="true" pageId="68" pageNumber="69">Iophon</emphasis>
</taxonomicName>
and in a footnote he synonymized
<taxonomicName id="F0E7F3EAFFA33117929DFACABD8C0244" ID-CoL="6N8WG" authorityName="Lundbeck" authorityYear="1905" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="frigidus">
<emphasis id="0593547BFFA33117929DFACABD8C0244" italics="true" pageId="68" pageNumber="69">I. frigidus</emphasis>
</taxonomicName>
, (ignoring
<taxonomicName id="F0E7F3EAFFA33117967AFAEDBF740244" ID-CoL="5PH59" authorityName="Hentschel" authorityYear="1929" box="[363,534,1333,1360]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="variety" species="frigidus" variety="gracilis">
<emphasis id="0593547BFFA33117967AFAEDBCE90244" box="[363,395,1333,1359]" italics="true" pageId="68" pageNumber="69">I.f.</emphasis>
var.
<emphasis id="0593547BFFA3311796D1FAEDBF740244" box="[448,534,1333,1359]" italics="true" pageId="68" pageNumber="69">gracilis</emphasis>
</taxonomicName>
), and
<taxonomicName id="F0E7F3EAFFA33117954AFAEEBE3F025B" ID-CoL="6N98H" authority="Hansen, 1885" authorityName="Hansen" authorityYear="1885" box="[603,861,1333,1360]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="dubius">
<emphasis id="0593547BFFA33117954AFAEEBFDF0244" box="[603,701,1333,1359]" italics="true" pageId="68" pageNumber="69">I. dubius</emphasis>
<bibRefCitation id="5376F598FFA3311795D2FAEEBE3F025B" author="Hansen, G. A." box="[707,861,1333,1360]" pageId="68" pageNumber="69" pagination="1 - 26" refId="ref97258" refString="Hansen, G. A. (1885) Spongiadae. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 13, 1 - 26, pls. I - VII. [https: // biodiversitylibrary. org / page / 38792336]" type="journal article" year="1885">Hansen, 1885</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="F0E7F3EAFFA33117948AFAEEB9A0025B" ID-CoL="7VJKJ" authority="(Vosmaer, 1885)" baseAuthorityName="Vosmaer" baseAuthorityYear="1885" box="[923,1218,1333,1360]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="piceum">
<emphasis id="0593547BFFA33117948AFAEEB9610244" box="[923,1027,1334,1359]" italics="true" pageId="68" pageNumber="69">I. piceum</emphasis>
(
<bibRefCitation id="5376F598FFA331179302FAEEB9D8025B" author="Vosmaer, G. C. J." box="[1043,1210,1333,1360]" pageId="68" pageNumber="69" refId="ref111455" refString="Vosmaer, G. C. J. (1885) Parts Vll - XI. Porifera. In: Bronn, H. G. (Ed.), Die Klassen und Ordnungen des Thier-Reichs. Vol. 1. Leipzig &amp; Heidelberg, pp. 177 - 368, pls. XIX - XXV." type="book" year="1885">Vosmaer, 1885</bibRefCitation>
)
</taxonomicName>
citing observations of the variability in spiculation in species from the southern oceans [
<taxonomicName id="F0E7F3EAFFA3311794A4FA81B9B40278" ID-CoL="6MX88" authority="Topsent, 1901" authorityName="Topsent" authorityYear="1901" box="[949,1238,1369,1396]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="radiatum">
<emphasis id="0593547BFFA3311794A4FA81B9490278" box="[949,1067,1369,1395]" italics="true" pageId="68" pageNumber="69">I.radiatum</emphasis>
<bibRefCitation id="5376F598FFA331179324FA82B9B40278" author="Topsent, E." box="[1077,1238,1369,1396]" pageId="68" pageNumber="69" pagination="1 - 54" refId="ref108552" refString="Topsent, E. (1901) Spongiaires. Resultats du voyage du S. Y. ' Belgica'en 1897 - 99 sous le commandement de A. de Gerlache de Gomery. Expedition antarctique belge, Zoologie, 4, 1 - 54, pls. I - VI." type="journal article" year="1901">Topsent, 1901</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F0E7F3EAFFA331179203FA82BC59029C" ID-CoL="6MX7W" authority="(Ridley, 1881)" baseAuthorityName="Ridley" baseAuthorityYear="1881" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="proximum">
<emphasis id="0593547BFFA331179203FA82B8FF0278" box="[1298,1437,1370,1395]" italics="true" pageId="68" pageNumber="69">I. proximum</emphasis>
(
<bibRefCitation id="5376F598FFA33117978EFAA5BC50029C" author="Ridley, S. O." box="[159,306,1405,1431]" pageId="68" pageNumber="69" pagination="107 - 141" refId="ref104189" refString="Ridley, S. O. (1881) XI. Spongida. Horny and Siliceous Sponges of Magellan Straits, S. W. Chili, and Atlantic off SW Brazil. In: Gunther, A. (Ed.), Account of the zoological collections made during the Survey of H. M. S. ' Alert' in the Straits of Magellan and on the Coast of Patagonia. Proceedings of the Zoological Society of London. Academic Press, London, pp. 107 - 141, pls. X - XI. [https: // www. biodiversitylibrary. org / page / 30759222]" type="book chapter" year="1881">Ridley, 1881</bibRefCitation>
)
</taxonomicName>
] as evidence for this conclusion.
</paragraph>
<paragraph id="37588869FFA3311797D6FA79BC750164" blockId="68.[151,1437,1009,1828]" pageId="68" pageNumber="69">
<bibRefCitation id="5376F598FFA3311797D6FA79BCC102B7" author="Koltun, V. M." box="[199,419,1441,1468]" pageId="68" pageNumber="69" pagination="1 - 236" refId="ref99720" refString="Koltun, V. M. (1959) [Siliceous horny sponges of the northern and fareastern seas of the U. S. S. R.]. Opredeliteli po faune SSR, izdavaemye Zoologicheskim muzeem Akademii nauk, 67, 1 - 236. [in Russian]" type="journal article" year="1959">Koltun (1959: 151)</bibRefCitation>
largely followed Burtons example but proposed a more differentiated widespread Arctic species
<taxonomicName id="F0E7F3EAFFA3311797FFFA1DBCF602D4" baseAuthorityName="Vosmaer" baseAuthorityYear="1885" box="[238,404,1477,1503]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="piceum">
<emphasis id="0593547BFFA3311797FFFA1DBCF602D4" box="[238,404,1477,1503]" italics="true" pageId="68" pageNumber="69">Iophon piceum</emphasis>
</taxonomicName>
five subspecies. Of these, he confirmed that
<taxonomicName id="F0E7F3EAFFA331179482FA1DB95D02D4" authorityName="Lundbeck" authorityYear="1905" box="[915,1087,1477,1503]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="frigidus">
<emphasis id="0593547BFFA331179482FA1DB95D02D4" box="[915,1087,1477,1503]" italics="true" pageId="68" pageNumber="69">Iophon frigidus</emphasis>
</taxonomicName>
and the variety
<taxonomicName id="F0E7F3EAFFA3311793FFFA1EB8FF02D4" authorityName="Hentschel" authorityYear="1929" box="[1262,1437,1477,1504]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="variety" species="frigidus" variety="gracilis">
<emphasis id="0593547BFFA3311793FFFA1EB87602D4" box="[1262,1300,1477,1503]" italics="true" pageId="68" pageNumber="69">I. f.</emphasis>
var.
<emphasis id="0593547BFFA331179256FA1DB8FF02D4" box="[1351,1437,1477,1503]" italics="true" pageId="68" pageNumber="69">gracilis</emphasis>
</taxonomicName>
were synonymous with
<taxonomicName id="F0E7F3EAFFA3311796B7FA32BE43010F" authority="(Hansen, 1885)" baseAuthorityName="Hansen" baseAuthorityYear="1885" box="[422,801,1513,1540]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="subSpecies" species="piceum" subSpecies="dubium">
<emphasis id="0593547BFFA3311796B7FA32BF090108" box="[422,619,1513,1539]" italics="true" pageId="68" pageNumber="69">I. piceum dubium</emphasis>
(
<bibRefCitation id="5376F598FFA33117956AFA31BE7B010F" author="Hansen, G. A." box="[635,793,1513,1540]" pageId="68" pageNumber="69" pagination="1 - 26" refId="ref97258" refString="Hansen, G. A. (1885) Spongiadae. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoology, 13, 1 - 26, pls. I - VII. [https: // biodiversitylibrary. org / page / 38792336]" type="journal article" year="1885">Hansen, 1885</bibRefCitation>
)
</taxonomicName>
rather than with the typical subspecies. Remarkably, he proposed a subspecies
<taxonomicName id="F0E7F3EAFFA33117968AF9D5BFB1012C" authorityName="Koltun" authorityYear="1959" box="[411,723,1549,1575]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="subSpecies" species="piceum" subSpecies="abipocillum">
<emphasis id="0593547BFFA33117968AF9D5BFB1012C" box="[411,723,1549,1575]" italics="true" pageId="68" pageNumber="69">Iophon piceum abipocillum</emphasis>
</taxonomicName>
, for a specimen which lacked bipocillae similar to
<taxonomicName id="F0E7F3EAFFA33117920DF9D6B8F7012C" authorityName="Lundbeck" authorityYear="1905" box="[1308,1429,1549,1575]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="frigidum">
<emphasis id="0593547BFFA33117920DF9D6B8F7012C" box="[1308,1429,1549,1575]" italics="true" pageId="68" pageNumber="69">I. frigidum</emphasis>
</taxonomicName>
. Koltuns proposed diversity of largely sympatric subspecies is not consistent with current hypotheses about subspecies.
</paragraph>
<paragraph id="37588869FFA3311797D6F9A2B96B01BC" blockId="68.[151,1437,1009,1828]" pageId="68" pageNumber="69">
I am not convinced that
<taxonomicName id="F0E7F3EAFFA3311796CFF9A1BFE50198" baseAuthorityName="Vosmaer" baseAuthorityYear="1885" box="[478,647,1657,1683]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="piceum">
<emphasis id="0593547BFFA3311796CFF9A1BFE50198" box="[478,647,1657,1683]" italics="true" pageId="68" pageNumber="69">Iophon piceum</emphasis>
</taxonomicName>
is the senior synonym of
<taxonomicName id="F0E7F3EAFFA3311794A0F9A2B9480198" authorityName="Lundbeck" authorityYear="1905" box="[945,1066,1657,1683]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="frigidum">
<emphasis id="0593547BFFA3311794A0F9A2B9480198" box="[945,1066,1657,1683]" italics="true" pageId="68" pageNumber="69">I. frigidum</emphasis>
</taxonomicName>
, as no comparative studies have been published. Burtons and Koltuns opinions are not sufficiently underbuilt.
</paragraph>
<paragraph id="37588869FFA3311797D6F919B9220028" blockId="68.[151,1437,1009,1828]" pageId="68" pageNumber="69">
If
<taxonomicName id="F0E7F3EAFFA3311797F5F91ABC3C01D0" ID-CoL="6N98G" authorityName="Lundbeck" authorityYear="1905" box="[228,350,1729,1755]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="frigidum">
<emphasis id="0593547BFFA3311797F5F91ABC3C01D0" box="[228,350,1729,1755]" italics="true" pageId="68" pageNumber="69">I. frigidum</emphasis>
</taxonomicName>
is a distinct species,
<taxonomicName id="F0E7F3EAFFA331179548F919BE6601D0" ID-CoL="5PH58" authorityName="Hentschel" authorityYear="1929" box="[601,772,1729,1756]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="variety" species="frigidum" variety="gracile">
<emphasis id="0593547BFFA331179548F919BF1B01D0" box="[601,633,1729,1755]" italics="true" pageId="68" pageNumber="69">I.f.</emphasis>
var.
<emphasis id="0593547BFFA3311795A4F919BE6601D0" box="[693,772,1729,1755]" italics="true" pageId="68" pageNumber="69">gracile</emphasis>
</taxonomicName>
cannot be synonymized and prudence dictates a possibly temporary elevation in rank to species level, since the difference (possession of bipocillae) precludes synonymy. Accordingly, I propose to name the present variety
<taxonomicName id="F0E7F3EAFFA3311795C7F8D1B9590028" authority="Hentschel, 1929" authorityName="Hentschel" authorityYear="1929" box="[726,1083,1801,1827]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="68" pageNumber="69" phylum="Porifera" rank="species" species="gracile">
<emphasis id="0593547BFFA3311795C7F8D1BE190028" box="[726,891,1801,1827]" italics="true" pageId="68" pageNumber="69">Iophon gracile</emphasis>
<bibRefCitation id="5376F598FFA331179493F8D1B9590028" author="Hentschel, E." box="[898,1083,1801,1827]" pageId="68" pageNumber="69" pagination="857 - 1042" refId="ref97583" refString="Hentschel, E. (1929) Die Kiesel- und Hornschwamme des N ˆ rdlichen Eismeers. In: R ˆ mer, F., Schaudinn, F., Brauer, A. &amp; Arndt, W. (Eds.), Fauna Arctica. Eine Zusammenstellung der arktischen Tierformen mit besonderer Ber ¸ cksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das N ˆ rdliche Eismeer im Jahre 1898. 5 (4). G. Fischer, Jena, pp. 857 - 1042, pls. XII - XIV." type="book chapter" year="1929">Hentschel, 1929</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>