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<mods:title id="03F048B0ABFF43D38C4C5BC6F65F9F35">Taxonomy of the ant genera Leptothorax Mayr, 1855 and Temnothorax Mayr, 1861 (Hymenoptera: Formicidae) of China with descriptions of twenty-eight new species and a key to the known Chinese species</mods:title>
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<mods:namePart id="03AB1A9970525A75A59D348DD62288C4">Qian, Yu-Han</mods:namePart>
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<mods:affiliation id="4E9627CA505A6D149098F5C7F9E39A33">Key Laboratory of Forest Disaster Warning and Control in Yunnan Province, College of Biodiversity Conservation, Southwest Forestry University, Kunming, Yunnan Province 650224, China. &amp; Key Laboratory for Forest Resources Conservation and Utilization in the Southwest Mountains of China, Ministry of Education, Southwest Forestry University, Kunming, Yunnan 650224, China.</mods:affiliation>
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<mods:namePart id="B2F28ECA50F0A701A1529D5D7BAC77AE">Xu, Zheng-Hui</mods:namePart>
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<mods:affiliation id="5D8EF452F4DA189671549F3BDE18CDEC">Key Laboratory of Forest Disaster Warning and Control in Yunnan Province, College of Biodiversity Conservation, Southwest Forestry University, Kunming, Yunnan Province 650224, China.</mods:affiliation>
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<heading id="6C6E81C9FFAF2A7EFD96A1C66AE4189A" box="[585,1004,1749,1776]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="F0994D26FFAF2A7EFD96A1C66A82189A" authority="Qian &amp; Xu, 2024" authorityName="Qian &amp; Xu" authorityYear="2024" box="[585,906,1749,1776]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="shuangjiang" status="sp. nov.">
<emphasis id="05EDEAB7FFAF2A7EFD96A1C66A82189A" bold="true" box="[585,906,1749,1776]" italics="true" pageId="51" pageNumber="52">Temnothorax shuangjiang</emphasis>
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<taxonomicNameLabel id="1EDE57CCFFAF2A7EFC4EA1C56AE4189A" box="[913,1004,1750,1776]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="372636A5FFAF2A7EFD2EA00F6A4B195C" blockId="51.[372,1216,1749,1846]" box="[753,835,1820,1846]" pageId="51" pageNumber="52">
<figureCitation id="AFA22A20FFAF2A7EFD2EA00F6A4B195C" box="[753,835,1820,1846]" captionStart="Fig" captionStartId="52.[187,230,1963,1989]" captionTargetBox="[187,1398,1291,1921]" captionTargetId="figure-292@52.[187,1398,1291,1921]" captionTargetPageId="52" captionText="Fig. 34. Temnothorax shuangjiang sp. nov., holotype worker (SWFU A13-198). A. Head in full-face view. B. Body in lateral view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11639321" httpUri="https://zenodo.org/record/11639321/files/figure.png" pageId="51" pageNumber="52">Fig. 34</figureCitation>
</paragraph>
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<subSubSection id="7F83652EFFAF2A79FF62A04E6D581CE9" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="372636A5FFAF2A7EFF62A04E683E191D" blockId="51.[189,310,1885,1911]" box="[189,310,1885,1911]" pageId="51" pageNumber="52">
<heading id="6C6E81C9FFAF2A7EFF62A04E683E191D" bold="true" box="[189,310,1885,1911]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="05EDEAB7FFAF2A7EFF62A04E683E191D" bold="true" box="[189,310,1885,1911]" pageId="51" pageNumber="52">Diagnosis</emphasis>
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<paragraph id="372636A5FFAF2A79FF62A09F6D581CE9" blockId="51.[189,1399,1932,1994]" lastBlockId="52.[187,1400,265,643]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
This new species is similar to
<taxonomicName id="F0994D26FFAF2A7EFDEBA09E6B7219CC" authority="Qian &amp; Xu, 2024" authorityName="Qian &amp; Xu" authorityYear="2024" box="[564,634,1933,1958]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="qiu" status="sp. nov.">
<emphasis id="05EDEAB7FFAF2A7EFDEBA09E6B7219CC" box="[564,634,1933,1958]" italics="true" pageId="51" pageNumber="52">T. qiu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EDE57CCFFAF2A7EFD5AA09F6BEB19CC" box="[645,739,1932,1958]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="AFA22A20FFAF2A7EFD28A09F6A5A19CC" box="[759,850,1932,1958]" captionStart="Fig" captionStartId="48.[189,232,1963,1989]" captionTargetBox="[189,1398,1307,1921]" captionTargetId="figure-379@48.[189,1398,1307,1921]" captionTargetPageId="48" captionText="Fig. 32. Temnothorax qiu sp. nov., holotype worker (SWFU C18-561). A. Head in full-face view. B. Body in lateral view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11639313" httpUri="https://zenodo.org/record/11639313/files/figure.png" pageId="51" pageNumber="52">Fig. 32</figureCitation>
), but differs in the new species head dorsum irregularly longitudinally rugose on the central area, apices of antennal scapes falled to reach posterior head margin, and mesosoma coarsely reticulate; in
<taxonomicName id="F0994D26FFA82A79FCC2A6196A681F49" authorityName="Qian &amp; Xu" authorityYear="2024" box="[797,864,266,291]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="qiu">
<emphasis id="05EDEAB7FFA82A79FCC2A6196A681F49" box="[797,864,266,291]" italics="true" pageId="52" pageNumber="53">T. qiu</emphasis>
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head dorsum parallelly longitudinally rugose on the central area, apices of antennal scapes surpassed posterior head margin, and mesosoma densely reticulate. The new species is also similar to
<taxonomicName id="F0994D26FFA82A79FD05A6436A251F03" authority="Qian &amp; Xu, 2024" authorityName="Qian &amp; Xu" authorityYear="2024" box="[730,813,335,361]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="lixia" status="sp. nov.">
<emphasis id="05EDEAB7FFA82A79FD05A6436A251F03" box="[730,813,335,361]" italics="true" pageId="52" pageNumber="53">T. lixia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EDE57CCFFA82A79FCE9A6436A9B1F00" box="[822,915,336,362]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="AFA22A20FFA82A79FC7AA65C6AF61F03" box="[933,1022,335,361]" captionStart="Fig" captionStartId="42.[188,229,1963,1989]" captionTargetBox="[189,1398,1436,1921]" captionTargetId="figure-357@42.[189,1398,1436,1921]" captionTargetPageId="42" captionText="Fig. 28. Temnothorax lixia sp. nov., holotype worker (SWFU A05-869). A. Head in full-face view. B. Body in lateral view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11639299" httpUri="https://zenodo.org/record/11639299/files/figure.png" pageId="52" pageNumber="53">Fig. 28</figureCitation>
), but differs in the new species propodeal spines longer than half length of declivity in lateral view, head dorsum longitudinally wavely rugose; in
<taxonomicName id="F0994D26FFA82A79FEE1A684689A1FDA" authorityName="Qian &amp; Xu" authorityYear="2024" box="[318,402,406,432]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="lixia">
<emphasis id="05EDEAB7FFA82A79FEE1A684689A1FDA" box="[318,402,406,432]" italics="true" pageId="52" pageNumber="53">T. lixia</emphasis>
</taxonomicName>
propodeal spines shorter than half length of declivity in lateral view, head dorsum longitudinally straightly rugose. This new species is also similar to
<taxonomicName id="F0994D26FFA82A79FC1CA6A96D201FB9" authority="Qian &amp; Xu, 2024" authorityName="Qian &amp; Xu" authorityYear="2024" box="[963,1064,441,467]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="qiufen" status="sp. nov.">
<emphasis id="05EDEAB7FFA82A79FC1CA6A96D201FB9" box="[963,1064,441,467]" italics="true" pageId="52" pageNumber="53">T. qiufen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EDE57CCFFA82A79FBF2A6AA6D8E1FB9" box="[1069,1158,441,467]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="AFA22A20FFA82A79FB4AA6AA6DE21FB9" box="[1173,1258,441,467]" captionStart="Fig" captionStartId="50.[189,232,1963,1989]" captionTargetBox="[189,1400,1326,1915]" captionTargetId="figure-275@50.[189,1400,1324,1915]" captionTargetPageId="50" captionText="Fig. 33. Temnothorax qiufen sp. nov., holotype worker (SWFU A18-972). A. Head in full-face view. B. Body in lateral view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11639315" httpUri="https://zenodo.org/record/11639315/files/figure.png" pageId="52" pageNumber="53">Fig. 33</figureCitation>
), but differs in the new species dorsum of mesonotum and propodeum weakly convex in lateral view, propodeal spines longer than half length of declivity, petiolar node with narrowly rounded dorsum, head dorsum wholly wavely rugose; in
<taxonomicName id="F0994D26FFA82A79FE44A5306B0C1C57" authorityName="Qian &amp; Xu" authorityYear="2024" box="[411,516,547,573]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="qiufen">
<emphasis id="05EDEAB7FFA82A79FE44A5306B0C1C57" box="[411,516,547,573]" italics="true" pageId="52" pageNumber="53">T. qiufen</emphasis>
</taxonomicName>
dorsum of mesonotum and propodeum straight in lateral view, propodeal spines about half length of declivity, petiolar node with rightly angled antertodorsal corner, head dorsum smooth on the central longitudinal narrow strip, longitudinally rugose laterally.
</paragraph>
</subSubSection>
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<paragraph id="372636A5FFA82A79FF62A5B8684C1CAF" blockId="52.[189,324,683,709]" box="[189,324,683,709]" pageId="52" pageNumber="53">
<heading id="6C6E81C9FFA82A79FF62A5B8684C1CAF" bold="true" box="[189,324,683,709]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="05EDEAB7FFA82A79FF62A5B8684C1CAF" bold="true" box="[189,324,683,709]" pageId="52" pageNumber="53">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="372636A5FFA82A79FF62A5CA6D6E1C99" blockId="52.[189,1126,729,755]" box="[189,1126,729,755]" pageId="52" pageNumber="53">
The specific epithet refers to shuangjiang, one of the 24 Solar Terms of
<collectingCountry id="4F8E7635FFA82A79FBC7A5CA6D6A1C99" box="[1048,1122,729,755]" name="China" pageId="52" pageNumber="53">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7F83652EFFA82A79FF62A40968DE1DA7" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="372636A5FFA82A79FF62A40968A21D5E" blockId="52.[189,426,794,820]" box="[189,426,794,820]" pageId="52" pageNumber="53">
<heading id="6C6E81C9FFA82A79FF62A40968A21D5E" bold="true" box="[189,426,794,820]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="05EDEAB7FFA82A79FF62A40968A21D5E" bold="true" box="[189,426,794,820]" pageId="52" pageNumber="53">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="372636A5FFA82A79FF35A45A68DE1DA7" blockId="52.[189,1399,841,973]" pageId="52" pageNumber="53">
<materialsCitation id="87F13CF8FFA82A79FF35A45A68DE1DA7" collectedFrom="soil sample from Pinus yunnanensis forest" collectingDate="2013-03-15" collectionCode="SWFU" collectorName="Shun-Rong Pu" country="China" county="Huaping County" elevation="2030" latitude="26.63645" location="Tianxing Forest Farm" longLatPrecision="1" longitude="101.14902" municipality="Xinzhuang Town" pageId="52" pageNumber="53" specimenCode="A13-198" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="05EDEAB7FFA82A79FF35A45A68B71D09" bold="true" box="[234,447,841,867]" pageId="52" pageNumber="53">
<typeStatus id="E8228807FFA82A79FF35A45A68541D09" box="[234,348,841,867]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
worker
</emphasis>
<collectingCountry id="4F8E7635FFA82A79FF62A47F68101DEC" box="[189,280,876,902]" name="China" pageId="52" pageNumber="53">CHINA</collectingCountry>
<collectingRegion id="F55DF847FFA82A79FEEDA47E68F71DEC" box="[306,511,876,903]" country="China" name="Yunnan" pageId="52" pageNumber="53">Yunnan Province</collectingRegion>
,
<collectingCounty id="DE474E29FFA82A79FDD3A47F6BD91DEC" box="[524,721,876,902]" pageId="52" pageNumber="53">Huaping County</collectingCounty>
,
<collectingMunicipality id="D742ACDFFFA82A79FD01A47F6AA31DED" box="[734,939,876,903]" pageId="52" pageNumber="53">Xinzhuang Town</collectingMunicipality>
,
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;
<geoCoordinate id="52AD5062FFA82A79FB16A47F6C7A1DED" box="[1225,1394,876,903]" degrees="26.636450" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="26.63645">26.636450° N</geoCoordinate>
,
<geoCoordinate id="52AD5062FFA82A79FF62A49C687A1DC0" box="[189,370,911,938]" degrees="101.149017" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="101.14902">101.149017° E</geoCoordinate>
;
<elevation id="BCB4D196FFA82A79FEA0A49C6B121DC3" box="[383,538,911,938]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="52" pageNumber="53" unit="m" value="2030.0">
<quantity id="F0619B40FFA82A79FEA0A49C68D11DC0" box="[383,473,911,938]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="52" pageNumber="53" unit="m" value="2030.0">2030 m</quantity>
a.s.l.
</elevation>
;
<date id="43271065FFA82A79FDF6A49C6BC41DC3" box="[553,716,911,938]" pageId="52" pageNumber="53" value="2013-03-15">
<collectingDate id="5363E98DFFA82A79FDF6A49C6BC41DC3" box="[553,716,911,938]" pageId="52" pageNumber="53" value="2013-03-15">15 Mar. 2013</collectingDate>
</date>
;
<collectorName id="9A6C5373FFA82A79FD06A49C6A8E1DC0" box="[729,902,911,938]" pageId="52" pageNumber="53">Shun-Rong Pu</collectorName>
leg.;
<collectedFrom id="84857558FFA82A79FC15A49C680C1DA6" pageId="52" pageNumber="53">
soil sample from
<taxonomicName id="F0994D26FFA82A79FB45A4836C7E1DC3" box="[1178,1398,912,937]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="05EDEAB7FFA82A79FB45A4836C7E1DC3" box="[1178,1398,912,937]" italics="true" pageId="52" pageNumber="53">Pinus yunnanensis</emphasis>
</taxonomicName>
forest
</collectedFrom>
;
<collectionCode id="5188AE60FFA82A79FED0A4A068681DA7" box="[271,352,947,973]" pageId="52" pageNumber="53">SWFU</collectionCode>
<specimenCode id="673F9EDEFFA82A79FEB9A4A068DA1DA7" box="[358,466,947,973]" collectionCode="SWFU" pageId="52" pageNumber="53">A13-198</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7F83652EFFA82A78FF62A4E06BD01B7D" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="372636A5FFA82A79FF62A4E068461A67" blockId="52.[189,334,1011,1037]" box="[189,334,1011,1037]" pageId="52" pageNumber="53">
<heading id="6C6E81C9FFA82A79FF62A4E068461A67" bold="true" box="[189,334,1011,1037]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="05EDEAB7FFA82A79FF62A4E068461A67" bold="true" box="[189,334,1011,1037]" pageId="52" pageNumber="53">Description</emphasis>
</heading>
</paragraph>
<paragraph id="372636A5FFA82A79FF62A3326B871A37" blockId="52.[188,1398,1056,1117]" pageId="52" pageNumber="53">
Measurements of
<typeStatus id="E8228807FFA82A79FE52A33368FC1A50" box="[397,500,1056,1082]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
worker: TL 3.5, HL 0.80, HW 0.72, CI 91, SL 0.59, SI 82, ED 0.18, PW 0.49, WL 1.05, PL 0.38,
<collectingCountry id="4F8E7635FFA82A79FE43A35068C91A37" box="[412,449,1091,1117]" name="Philippines" pageId="52" pageNumber="53">PH</collectingCountry>
0.28, DPW 0.20.
</paragraph>
<paragraph id="372636A5FFA82A78FF62A3906B2F1FC1" blockId="52.[188,1398,1155,1216]" lastBlockId="53.[189,1401,265,428]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">In full-face view head roughly rectangular, longer than broad, posterior margin almost straight, posterior corner narrowly rounded, lateral margins weakly convex. Mandibles triangular, masticatory margin with 5 teeth. Clypeal dorsum weakly convex, with waved median carina, anterior margin moderately convex. Frontal lobes broad, concealing antennal sockets. Frontal carinae short, reaching to the level of anterior eye margins. Antennae 12-segmented, scapes failing to reach posterior head margin by 1/7 of its length, antennal clubs 3-segmented. Eyes located slightly before midpoint of lateral head margin, occupying ¼ of lateral margin.</paragraph>
<caption id="63E6662DFFA82A79FF64A0B86A3D1982" ID-DOI="http://doi.org/10.5281/zenodo.11639321" ID-Zenodo-Dep="11639321" httpUri="https://zenodo.org/record/11639321/files/figure.png" pageId="52" pageNumber="53" startId="52.[187,230,1963,1989]" targetBox="[187,1398,1291,1921]" targetPageId="52">
<paragraph id="372636A5FFA82A79FF64A0B86A3D1982" blockId="52.[187,1396,1963,2025]" pageId="52" pageNumber="53">
<emphasis id="05EDEAB7FFA82A79FF64A0B8681319AF" bold="true" box="[187,283,1963,1989]" pageId="52" pageNumber="53">Fig. 34.</emphasis>
<taxonomicName id="F0994D26FFA82A79FEFBA0B86B5019AF" authority="Qian &amp; Xu, 2024" authorityName="Qian &amp; Xu" authorityYear="2024" box="[292,600,1963,1989]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="shuangjiang" status="sp. nov.">
<emphasis id="05EDEAB7FFA82A79FEFBA0B86B5019AF" box="[292,600,1963,1989]" italics="true" pageId="52" pageNumber="53">Temnothorax shuangjiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EDE57CCFFA82A79FDBBA0BF6BC819AF" box="[612,704,1964,1990]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
, holotype worker (SWFU A13-198).
<emphasis id="05EDEAB7FFA82A79FB59A0B86D9119AF" bold="true" box="[1158,1177,1963,1989]" pageId="52" pageNumber="53">A</emphasis>
. Head in full-face view.
<emphasis id="05EDEAB7FFA82A79FEDEA0DD681D1982" bold="true" box="[257,277,1998,2024]" pageId="52" pageNumber="53">B</emphasis>
. Body in lateral view.
<emphasis id="05EDEAB7FFA82A79FDC0A0DD6B3C1982" bold="true" box="[543,564,1998,2024]" pageId="52" pageNumber="53">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<paragraph id="372636A5FFA92A78FF62A6C16A101D4A" blockId="53.[189,1401,466,800]" pageId="53" pageNumber="54">In lateral view promesonotum weakly convex and weakly arched, antetrodorsal corner of pronotum bluntly angled, promesonotal suture and metanotal groove absent. Propodeal dorsum almost straight and gently sloping down posterriorly; propodeal spines long and slender, as long as declivity, pointed posterodorsally, its dorsal margin weakly convex, ventral margin straight; declivity straight; propodeal lobes relatively large and triangular. Petiole with anterior peduncle, about ⅔ length of petiolar node; petiolar node roughly trapezoidal and narrowing dorsally, anterior margin almost straight, dorsal and posterior margins weakly convex, anterodorsal corner narrowly rounded, posterodorsal corner broadly rounded; ventral margin straight anteriorly, weakly concave posteriorly, anteroventral corner bluntly angled. Postpetiole about as high as petiolar node, anterodorsal margin rounded, posterior margin weakly concave, ventral margin angularly concave.</paragraph>
<paragraph id="372636A5FFA92A78FF62A4556BD21A46" blockId="53.[188,1398,837,1069]" pageId="53" pageNumber="54">In dorsal view pronotum broadest, anterior and lateral margin moderately convex and rounded, humeral corners rounded. Promesonotal suture and metanotal groove absent. Mesothorax weakly constricted, lateral margins weakly concave. Propodeum roughly rectangular, lateral margins slightly concave, spines moderately long and straight, pointed posterolaterally. Petiole roughly trapezoidal and widening posteriorly, longer than broad, lateral margins weakly convex. Postpetiole roughly trapezoidal and narrowing posteriorly, about 1.4 times as broad as petiole, anterolateral corners broadly rounded, lateral margins weakly convex. Gaster elongate oval.</paragraph>
<paragraph id="372636A5FFA92A78FF62A3406BD01B7D" blockId="53.[189,1399,1107,1304]" pageId="53" pageNumber="54">Mandible longitudinally striate. Head dorsum longitudinally rugose and reticulate posteriorly and laterally, sides below eyes densely finely reticulate. Clypeus loosely irregularly rugose. Mososoma coarsely reticulate. Petiole and postpetiole densely finely reticulate, anterior face of postpetiole smooth. Gaster smooth and shiny. Body dorsum with sparse erect to suberect apically blunt short hairs and sparse decumbent pubescence. Scapes and tibiae with dense decumbent pubescence. Body color black, mandibles, antennae and legs blackish brown.</paragraph>
</subSubSection>
<subSubSection id="7F83652EFFA92A78FF62A22E69E71BCD" pageId="53" pageNumber="54" type="biology_ecology">
<paragraph id="372636A5FFA92A78FF62A22E688E1B32" blockId="53.[189,390,1341,1368]" box="[189,390,1341,1368]" pageId="53" pageNumber="54">
<heading id="6C6E81C9FFA92A78FF62A22E688E1B32" bold="true" box="[189,390,1341,1368]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="05EDEAB7FFA92A78FF62A22E688E1B32" bold="true" box="[189,390,1341,1368]" pageId="53" pageNumber="54">Ecological notes</emphasis>
</heading>
</paragraph>
<paragraph id="372636A5FFA92A78FF62A27869E71BCD" blockId="53.[189,1399,1387,1447]" pageId="53" pageNumber="54">
This new species inhabits
<taxonomicName id="F0994D26FFA92A78FE2EA27F6BC51BEF" box="[497,717,1388,1413]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="05EDEAB7FFA92A78FE2EA27F6BC51BEF" box="[497,717,1388,1413]" italics="true" pageId="53" pageNumber="54">Pinus yunnanensis</emphasis>
</taxonomicName>
forest (
<figureCitation id="AFA22A20FFA92A78FCF8A2786A8E1BEF" box="[807,902,1387,1413]" captionStart="Fig" captionStartId="86.[189,232,1227,1253]" captionTargetBox="[189,1398,265,1169]" captionTargetId="figure-17@86.[189,1398,265,1169]" captionTargetPageId="86" captionText="Fig. 69. Middle elevational vegetations. A. Pinus densata Masters forest (2750 m, Bomi, Tibet). B. Pinus armandi Franch. forest (2557 m, Qiaojia, Yunnan). C. Subalpine moist evergreen broadleaf forest (2500 m, Lushui, Yunnan). D. Conifer-broadleaf mixed forest (2500 m, Huaping, Yunnan). E. Moss evergreen broadleaf forest (2450 m. Cangyuan, Yunnan). F. Warm conifer forest (2230 m, Huili, Sichuan). G. Dry-warm valley shrubland (2230 m, Muli, Sichuan). H. Semi-evergreen broadleaf forest (2050 m, Danba, Sichuan). I. Pinus yunnanensis forest (2030 m, Huaping, Yunnan). Photos by Zheng-Hui Xu." figureDoi="http://doi.org/10.5281/zenodo.11639434" httpUri="https://zenodo.org/record/11639434/files/figure.png" pageId="53" pageNumber="54">Fig. 69I</figureCitation>
) at an elevation of
<quantity id="F0619B40FFA92A78FBB6A2786DCC1BEC" box="[1129,1220,1387,1414]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="53" pageNumber="54" unit="m" value="2030.0">2030 m</quantity>
, and forages in soil.
</paragraph>
</subSubSection>
<subSubSection id="7F83652EFFA92A78FF62A2DE687B187C" box="[189,371,1485,1558]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="372636A5FFA92A78FF62A2DE685E1B8D" blockId="53.[189,342,1485,1511]" box="[189,342,1485,1511]" pageId="53" pageNumber="54">
<heading id="6C6E81C9FFA92A78FF62A2DE685E1B8D" bold="true" box="[189,342,1485,1511]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="05EDEAB7FFA92A78FF62A2DE685E1B8D" bold="true" box="[189,342,1485,1511]" pageId="53" pageNumber="54">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="372636A5FFA92A78FF62A2E8687B187C" blockId="53.[189,371,1531,1558]" box="[189,371,1531,1558]" pageId="53" pageNumber="54">
<collectingCountry id="4F8E7635FFA92A78FF62A2E86800187F" box="[189,264,1531,1557]" name="China" pageId="53" pageNumber="54">China</collectingCountry>
:
<collectingRegion id="F55DF847FFA92A78FECEA2EF6867187C" box="[273,367,1532,1558]" country="China" name="Yunnan" pageId="53" pageNumber="54">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>