treatments-xml/data/88/40/A2/8840A2035857FFC035EEFB91FE87B722.xml

<document id="6E0672892E6CF9E8FDDA17D7914A4EB9" ID-DOI="10.11646/zootaxa.3957.5.4" ID-GBIF-Dataset="c53fe374-dbf8-4f3e-869b-b2a0c7cdee19" ID-ISSN="1175-5326" ID-Zenodo-Dep="234560" ID-ZooBank="471D1EA1-9BAF-4396-90DB-540ED37318A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461066273047" checkinUser="plazi" docAuthor="Kotov, Alexey A. &amp; Fuentes-Reinés, Juan M." docDate="2015" docId="8840A2035857FFC035EEFB91FE87B722" docLanguage="en" docName="zt03957p566.pdf" docOrigin="Zootaxa 3957 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leberis colombiensis Kotov &amp; Fuentes-Reinés, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="564" masterDocId="7479DA7B5856FFCB3579FF9BFFBFB337" masterDocTitle="A new species of Leberis Smirnov, 1989 (Cladocera: Chydoridae) from Colombia" masterLastPageNumber="566" masterPageNumber="553" pageNumber="554" updateTime="1698583394311" updateUser="plazi">
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<mods:title id="3818585319D530F1C794F17B5EA1DB55">A new species of Leberis Smirnov, 1989 (Cladocera: Chydoridae) from Colombia</mods:title>
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<mods:namePart id="9C45B9F99174C110572BABE1680FA8AF">Kotov, Alexey A.</mods:namePart>
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<mods:namePart id="4DF255F3463128B8700E0170D463109A">Fuentes-Reinés, Juan M.</mods:namePart>
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<mods:date id="CC07F471372019FBA7BF6EE13061E96E">2015</mods:date>
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<treatment id="8840A2035857FFC035EEFB91FE87B722" ID-DOI="http://doi.org/10.5281/zenodo.5667126" ID-GBIF-Taxon="119614950" ID-Zenodo-Dep="5667126" LSID="urn:lsid:plazi:treatment:8840A2035857FFC035EEFB91FE87B722" httpUri="http://treatment.plazi.org/id/8840A2035857FFC035EEFB91FE87B722" lastPageId="11" lastPageNumber="564" pageId="1" pageNumber="554">
<subSubSection id="48F3409E5857FFCA35EEFB91FEADB771" pageId="1" pageNumber="554" type="nomenclature">
<paragraph id="005613155857FFCA35EEFB91FE46B713" blockId="1.[151,505,1034,1094]" box="[151,505,1034,1060]" pageId="1" pageNumber="554">
<heading id="5B1EA4795857FFCA35EEFB91FE46B713" bold="true" box="[151,505,1034,1060]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
<emphasis id="329DCF075857FFCA35EEFB91FE46B713" bold="true" box="[151,505,1034,1060]" pageId="1" pageNumber="554">
<taxonomicName id="C7E968965857FFCA35EEFB91FE2BB713" box="[151,404,1034,1060]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF075857FFCA35EEFB91FE2BB713" bold="true" box="[151,404,1034,1060]" italics="true" pageId="1" pageNumber="554">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C5857FFCA34E2FB91FE46B713" box="[411,505,1034,1060]" pageId="1" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="005613155857FFCA35EEFBB6FEADB771" blockId="1.[151,505,1034,1094]" box="[151,274,1069,1094]" pageId="1" pageNumber="554">
(
<figureCitation id="98D20F905857FFCA35E6FBB6FEB7B771" box="[159,264,1069,1094]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1922,1944]" captionStartId-1="4.[151,250,1953,1975]" captionStartId-2="5.[151,250,1915,1937]" captionStartId-3="6.[151,250,1888,1910]" captionStartId-4="7.[151,250,1917,1939]" captionTargetBox-0="[176,1400,198,1888]" captionTargetBox-1="[201,1386,200,1926]" captionTargetBox-2="[180,1407,195,1875]" captionTargetBox-3="[154,1418,204,1859]" captionTargetBox-4="[244,1334,196,1884]" captionTargetId-0="figure@3.[164,1414,193,1895]" captionTargetId-1="figure@4.[195,1391,193,1932]" captionTargetId-2="figure@5.[176,1410,193,1879]" captionTargetId-3="figure@6.[151,1436,193,1867]" captionTargetId-4="figure@7.[240,1337,193,1886]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." captionText-1="FIGURE 2. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A – B, general view. C, head. D, head shield. E, major head pores. F, labrum. G, armature of postero-ventral portion of valve. H, antenna I. Scale bars 0.1 mm for A – D; 0.01 mm for E – H." captionText-2="FIGURE 3. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E – F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D – H." captionText-3="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." captionText-4="FIGURE 5. Leberis colombiensis sp. nov., thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C – D, corm of limb I. E, limb II. F, Limb III. G – H, distal portion of limb III. Scale bars 0.1 mm." httpUri-0="https://zenodo.org/record/234561/files/figure.png" httpUri-1="https://zenodo.org/record/234562/files/figure.png" httpUri-2="https://zenodo.org/record/234563/files/figure.png" httpUri-3="https://zenodo.org/record/234564/files/figure.png" httpUri-4="https://zenodo.org/record/234565/files/figure.png" pageId="1" pageNumber="554">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="48F3409E5857FFCA35EEFBEEFB1DB7B9" box="[151,1186,1141,1166]" pageId="1" pageNumber="554" type="etymology">
<paragraph id="005613155857FFCA35EEFBEEFB1DB7B9" blockId="1.[151,1436,1141,1994]" box="[151,1186,1141,1166]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35EEFBEEFEA9B7B9" bold="true" box="[151,278,1141,1166]" pageId="1" pageNumber="554">Etymology</emphasis>
. The new species is named after 
<collectingCountry id="78FE53855857FFCA37F3FBEEFD44B7B9" box="[650,763,1141,1166]" name="Colombia" pageId="1" pageNumber="554">Colombia</collectingCountry>
, the country where it was discovered.
</paragraph>
</subSubSection>
<subSubSection id="48F3409E5857FFCA35BEFB02FC83B6E5" pageId="1" pageNumber="554" type="materials_examined">
<paragraph id="005613155857FFCA35BEFB02FDD3B7CD" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEFB02FED9B785" bold="true" box="[199,358,1177,1202]" pageId="1" pageNumber="554">
<typeStatus id="DF52ADB75857FFCA35BEFB02FEBDB785" box="[199,258,1177,1202]" pageId="1" pageNumber="554">Type</typeStatus>
locality.
</emphasis>
Ciénaga El Covento, Atlántico Department, 
<collectingCountry id="78FE53855857FFCA3619FB02FC71B785" box="[864,974,1177,1202]" name="Colombia" pageId="1" pageNumber="554">Colombia</collectingCountry>
(
<geoCoordinate id="65DD75D25857FFCA36A7FB01FBFCB785" box="[990,1091,1178,1202]" direction="north" orientation="latitude" pageId="1" pageNumber="554" precision="925" value="10.633333">10º 38'N</geoCoordinate>
; 
<geoCoordinate id="65DD75D25857FFCA3129FB01FB03B785" box="[1104,1212,1177,1202]" direction="west" orientation="longitude" pageId="1" pageNumber="554" precision="925" value="-74.916664">74º 55'W</geoCoordinate>
). From this locality, 70 parthenogenetic females, four ephippial females, ten juvenile and two adult males were collected in June and October of 2014 by J. M. Fuentes-Reinés.
</paragraph>
<paragraph id="005613155857FFCA35BEFA9EFD64B675" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEFA9EFE50B629" bold="true" box="[199,495,1285,1310]" pageId="1" pageNumber="554">
<typeStatus id="DF52ADB75857FFCA35BEFA9EFEBDB629" box="[199,258,1285,1310]" pageId="1" pageNumber="554">Type</typeStatus>
material. 
<typeStatus id="DF52ADB75857FFCA34FBFA9EFE50B629" box="[386,495,1285,1310]" pageId="1" pageNumber="554" type="holotype">Holotype</typeStatus>
</emphasis>
. Parthenogenetic female deposited at the Museo de Colecciones Biológicas de la Universidad del Atlántico, 
<collectingCountry id="78FE53855857FFCA34BFFAB2FD88B675" box="[454,567,1321,1346]" name="Colombia" pageId="1" pageNumber="554">Colombia</collectingCountry>
, 
<collectionCode id="66F88BD05857FFCA373AFAB1FD2AB675" box="[579,661,1322,1346]" pageId="1" pageNumber="554">UARC</collectionCode>
<specimenCount id="16EFD89C5857FFCA37ECFAB1FD64B675" box="[661,731,1322,1346]" pageId="1" pageNumber="554" type="generic">221M.</specimenCount>
</paragraph>
<paragraph id="005613155857FFCA35BEFAD6FC13B651" blockId="1.[151,1436,1141,1994]" box="[199,940,1357,1382]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEFAD6FE8CB651" bold="true" box="[199,307,1357,1382]" pageId="1" pageNumber="554">Allotype.</emphasis>
A dissected adult male from 
<typeStatus id="DF52ADB75857FFCA3704FAD5FD12B651" box="[637,685,1358,1382]" pageId="1" pageNumber="554">type</typeStatus>
locality, 
<collectionCode id="66F88BD05857FFCA366DFAD5FCDAB651" box="[788,869,1358,1382]" pageId="1" pageNumber="554">UARC</collectionCode>
<specimenCount id="16EFD89C5857FFCA361CFAD5FC13B651" box="[869,940,1358,1382]" pageId="1" pageNumber="554" type="generic">222M.</specimenCount>
</paragraph>
<paragraph id="005613155857FFCA35BEFAEAFC83B6E5" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEFAEAFEF9B6BD" bold="true" box="[199,326,1393,1418]" pageId="1" pageNumber="554">
<typeStatus id="DF52ADB75857FFCA35BEFAEAFEFEB6BD" box="[199,321,1393,1418]" pageId="1" pageNumber="554" type="paratype">Paratypes</typeStatus>
.
</emphasis>
20 undisected females in alcohol, 
<collectionCode id="66F88BD05857FFCA37AAFAE9FC9BB6BD" box="[723,804,1394,1418]" pageId="1" pageNumber="554">UARC</collectionCode>
223M; 11 undissected females in alcohol, 
<collectionCode id="66F88BD05857FFCA307DFAE9FAEAB6BD" box="[1284,1365,1394,1418]" pageId="1" pageNumber="554">UARC</collectionCode>
224M; adult male, 
<collectionCode id="66F88BD05857FFCA3458FA0EFECDB699" box="[289,370,1429,1454]" pageId="1" pageNumber="554">UARC</collectionCode>
226M-230M; 
<specimenCount id="16EFD89C5857FFCA3769FA0DFDC0B699" box="[528,639,1429,1454]" pageId="1" pageNumber="554" type="female">3 females</specimenCount>
from Laguna Navio Quebrado, 
<specimenCount id="16EFD89C5857FFCA368EFA0DFBCBB699" box="[1015,1140,1429,1454]" pageId="1" pageNumber="554" type="female">10 females</specimenCount>
from Ciénaga Grande de Santa 
<collectingCountry id="78FE53855857FFCA35A5FA21FE9DB6E6" box="[220,290,1466,1489]" name="Malta" pageId="1" pageNumber="554">Marta</collectingCountry>
, personal collection of Juan M. Fuentes-Reinés.
</paragraph>
</subSubSection>
<subSubSection id="48F3409E5857FFC035BEFA46FB1DB00F" lastPageId="11" lastPageNumber="564" pageId="1" pageNumber="554" type="description">
<paragraph id="005613155857FFCA35BEFA46FDA2B5B1" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEFA46FD3EB6C1" bold="true" box="[199,641,1501,1526]" pageId="1" pageNumber="554">Description. Parthenogenetic female</emphasis>
. 
<emphasis id="329DCF075857FFCA37EDFA46FD4BB6C1" box="[660,756,1501,1526]" italics="true" pageId="1" pageNumber="554">Habitus.</emphasis>
In lateral view, body ovoid (
<figureCitation id="98D20F905857FFCA312BFA46FB1FB6C1" box="[1106,1184,1501,1526]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="1" pageNumber="554">Figs. 1</figureCitation>
A, 2A–B), maximum height in middle, height/length about 
<quantity id="C711BEF05857FFCA373AF999FD3BB52D" box="[579,644,1537,1562]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="1" pageNumber="554" unit="in" value="0.8">0.8 in</quantity>
both adults and juveniles. Dorsal margin generally uniformly curved, postero-dorsal angle smoothed, postero-ventral angle broadly rounded. Head small, not keeled, rostrum short and blunt, compound eye and ocellus subequal in size, distance from tip of rostrum to ocellus somewhat larger than between ocellus and eye (
<figureCitation id="98D20F905857FFCA34CEF9F6FE45B5B1" box="[439,506,1645,1670]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[201,1386,200,1926]" captionTargetId="figure@4.[195,1391,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A – B, general view. C, head. D, head shield. E, major head pores. F, labrum. G, armature of postero-ventral portion of valve. H, antenna I. Scale bars 0.1 mm for A – D; 0.01 mm for E – H." httpUri="https://zenodo.org/record/234562/files/figure.png" pageId="1" pageNumber="554">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="005613155857FFCA35BEF90AFC2BB422" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEF90AFEF1B59D" box="[199,334,1681,1706]" italics="true" pageId="1" pageNumber="554">Head shield</emphasis>
elongated, length/width ratio about 1.6–1.8; anterior extremity truncated, posterior portion widely rounded, with three closely located major head pores of equal size, with connection between them (
<figureCitation id="98D20F905857FFCA306BF92EFAE1B5F9" box="[1298,1374,1717,1742]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="1" pageNumber="554">Figs. 1</figureCitation>
B–C, 2D–E), post-pore distance about 1.0–1.2 inter-pore distance. Lateral head pores about 1.2–1.4 inter-pore distances from midline, located at level of central or anterior major head pore.
</paragraph>
<paragraph id="005613155857FFCA35BEF8BAFAC4B469" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEF8BAFE9FB40D" box="[199,288,1825,1850]" italics="true" pageId="1" pageNumber="554">Labrum</emphasis>
with a broad, regularly convex labral keel, apex more or less blunt (
<figureCitation id="98D20F905857FFCA315AF8BAFBD4B40D" box="[1059,1131,1825,1850]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="1" pageNumber="554">Figs. 1</figureCitation>
D–E, 2F); in single female a shallow depression presents at anterior margin near apex (
<figureCitation id="98D20F905857FFCA364DF8DEFCCBB469" box="[820,884,1861,1886]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="1" pageNumber="554">Fig. 1</figureCitation>
F). No setulation, teeth etc. on margin of keel.
</paragraph>
<paragraph id="005613155857FFCA35BEF8F2FAD3B4FD" blockId="1.[151,1436,1141,1994]" pageId="1" pageNumber="554">
<emphasis id="329DCF075857FFCA35BEF8F2FEA9B4B5" box="[199,278,1897,1922]" italics="true" pageId="1" pageNumber="554">Valves.</emphasis>
Ventral margin with 67–75 setae of different size in different regions of the margin, between them series of small setules. The posteriormost 6–7 setae with sparsely located, pointed, spine-like short setules, followed by a row of spinules, starting marginally, continued along inner margin to the dorsum (
<figureCitation id="98D20F905857FFCA31B4F82AFAA9B4FD" box="[1229,1302,1969,1994]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="1" pageNumber="554">Figs. 1</figureCitation>
G, 2G).
</paragraph>
<paragraph id="005613155854FFC935BEFF0CFE4DB29A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFF0CFEDFB387" box="[199,352,151,176]" italics="true" pageId="2" pageNumber="555">Postabdomen</emphasis>
(
<figureCitation id="98D20F905854FFC93409FF0CFE03B387" box="[368,444,151,176]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="2" pageNumber="555">Figs. 1</figureCitation>
H, 3C) elongated, somewhat tapering distally, length/width ratio about 2.4–2.6. Ventral margin almost straight, with no ridges or setules. Preanal and postanal angle well-expressed, distal angle broadly rounded. Preanal margin short, anal margin 1.3 times longer than the former, postanal margin two times longer than preanal margin. Postanal margin provided with 9–11 clusters of denticles, continued on anal margin (
<figureCitation id="98D20F905854FFC93075FE9FFAE6B22B" box="[1292,1369,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[180,1407,195,1875]" captionTargetId="figure@5.[176,1410,193,1879]" captionTargetPageId="5" captionText="FIGURE 3. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E – F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D – H." httpUri="https://zenodo.org/record/234563/files/figure.png" pageId="2" pageNumber="555">Figs. 3</figureCitation>
D–F). On each side of postabdomen, 9–11 lateral fascicles of relatively long setules (
<figureCitation id="98D20F905854FFC9317CFEBCFBF8B277" box="[1029,1095,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[180,1407,195,1875]" captionTargetId="figure@5.[176,1410,193,1879]" captionTargetPageId="5" captionText="FIGURE 3. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E – F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D – H." httpUri="https://zenodo.org/record/234563/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
G). Postabdominal claw (
<figureCitation id="98D20F905854FFC9301DFEBCFF1DB253" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="2" pageNumber="555">Figs. 1</figureCitation>
H, 3H) regularly curved, somewhat longer than preanal portion of postabdomen, with fine setulation along ventral margin. Basal spine length about 0.3 of claw length, with a row of setules along its dorsal margin, also there are three long setules at claw base.
</paragraph>
<paragraph id="005613155854FFC935BEFE23FA99B120" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFE23FE8AB2E7" box="[199,309,440,464]" italics="true" pageId="2" pageNumber="555">Antenna I</emphasis>
(
<figureCitation id="98D20F905854FFC9343CFE2CFE2EB2E7" box="[325,401,439,464]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="2" pageNumber="555">
Figs. 
<date id="745735D55854FFC934FCFE23FE2EB2E7" box="[389,401,440,464]" pageId="2" pageNumber="555">1</date>
</figureCitation>
I, 2H) reaching tip of rostrum, with a smooth surface lacking any ridges, or transverse rows of setules. Antenular sensory seta slender, inserted at 1/3 of antennular length from tip, projecting beyond tip. Nine asthetascs, two of them markedly longer than the others, all aesthetascs projected beyond tip of rostrum.
</paragraph>
<paragraph id="005613155854FFC935BEFDBFFDDBB19F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFDBFFE80B10B" box="[199,319,548,572]" italics="true" pageId="2" pageNumber="555">Antenna II</emphasis>
short (
<figureCitation id="98D20F905854FFC934E9FDBFFE62B10B" box="[400,477,548,573]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="2" pageNumber="555">Figs. 1</figureCitation>
J, 3A), antennular formula: setae 0-0-3/1-1-3; spines 1-0-1/0-0-1. Setae on proximal and next segment of endopod thinner and shorter than apical setae. Spine on first exopod segment robust, shorter than 0.75 of second segment length. Spines on both apical segment of exopod and endopod relatively short, with a hillock near base of each spine (
<figureCitation id="98D20F905854FFC93486FD14FDFEB19F" box="[511,577,655,680]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[180,1407,195,1875]" captionTargetId="figure@5.[176,1410,193,1879]" captionTargetPageId="5" captionText="FIGURE 3. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E – F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D – H." httpUri="https://zenodo.org/record/234563/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="005613155854FFC935BEFD2FFB40B0DA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFD2FFEAEB1FB" box="[199,273,692,717]" italics="true" pageId="2" pageNumber="555">Limb I</emphasis>
(
<figureCitation id="98D20F905854FFC93459FD2FFED0B1FB" box="[288,367,692,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
A, 5A–D) with small, ovoid epipodite (removed in 
<figureCitation id="98D20F905854FFC936D6FD2FFC42B1FB" box="[943,1021,692,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
A, 5A). Accessory seta absent. Outer distal lobe (
<figureCitation id="98D20F905854FFC93458FD4CFECDB1C7" box="[289,370,727,752]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
A, 5B, ODL) large, bearing only a single bisegmented seta with unilaterally setulated distal segment. Inner distal lobe (
<figureCitation id="98D20F905854FFC934B1FD67FDAAB022" box="[456,533,764,789]" captionStart="FIGURE 5" captionStartId="7.[151,250,1917,1939]" captionTargetBox="[244,1334,196,1884]" captionTargetId="figure@7.[240,1337,193,1886]" captionTargetPageId="7" captionText="FIGURE 5. Leberis colombiensis sp. nov., thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C – D, corm of limb I. E, limb II. F, Limb III. G – H, distal portion of limb III. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234565/files/figure.png" pageId="2" pageNumber="555">Figs. 5</figureCitation>
B, IDL) with several clusters of short setules, two bisegmented setae of different length, armed by short and thick setules distally, and a small sensillum (
<figureCitation id="98D20F905854FFC93686FC84FBF8B00F" box="[1023,1095,799,824]" captionStart="FIGURE 5" captionStartId="7.[151,250,1917,1939]" captionTargetBox="[244,1334,196,1884]" captionTargetId="figure@7.[240,1337,193,1886]" captionTargetPageId="7" captionText="FIGURE 5. Leberis colombiensis sp. nov., thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C – D, corm of limb I. E, limb II. F, Limb III. G – H, distal portion of limb III. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234565/files/figure.png" pageId="2" pageNumber="555">Fig. 5</figureCitation>
B, incut). Limb corm almost rectangular in lateral view. Endite 3 with three posterior setae subequal in length (
<figureCitation id="98D20F905854FFC93143FCDFFB37B06B" box="[1082,1160,836,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
F, 5C, a–c), and a single anterior seta (1). On endite 2 three bi-segmented setae (d–f), two longer ones (e–f) unequal in length, unilaterally armed by strong setules distally, third seta (d) short. Endite 1 with three setae (g–i). Fascicles of thin setules on inner face of limb, plus bunches of longer and robuster setules at ventral margin of limb (
<figureCitation id="98D20F905854FFC931E4FC34FB5EB0FF" box="[1181,1249,943,968]" captionStart="FIGURE 5" captionStartId="7.[151,250,1917,1939]" captionTargetBox="[244,1334,196,1884]" captionTargetId="figure@7.[240,1337,193,1886]" captionTargetPageId="7" captionText="FIGURE 5. Leberis colombiensis sp. nov., thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C – D, corm of limb I. E, limb II. F, Limb III. G – H, distal portion of limb III. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234565/files/figure.png" pageId="2" pageNumber="555">Fig. 5</figureCitation>
D, arrows). Two serrate ejector hooks of similar size. A small projection with a single seta on limb basalmost portion.
</paragraph>
<paragraph id="005613155854FFC935BEFC6CFA97B76F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFC6CFEA5B727" box="[199,282,1015,1040]" italics="true" pageId="2" pageNumber="555">Limb II</emphasis>
(
<figureCitation id="98D20F905854FFC93450FC6CFEC8B727" box="[297,375,1015,1040]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
C, 5E) with exopodite small, lacking setae; inner portion of limb with eight scrapers increasing in size distally (
<figureCitation id="98D20F905854FFC93433FB87FE27B702" box="[330,408,1052,1077]" captionStart="FIGURE 5" captionStartId="7.[151,250,1917,1939]" captionTargetBox="[244,1334,196,1884]" captionTargetId="figure@7.[240,1337,193,1886]" captionTargetPageId="7" captionText="FIGURE 5. Leberis colombiensis sp. nov., thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C – D, corm of limb I. E, limb II. F, Limb III. G – H, distal portion of limb III. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234565/files/figure.png" pageId="2" pageNumber="555">Figs. 5</figureCitation>
E, 1–8). Distal armature of gnathobase with four elements, one of them a minute sensillum (
<figureCitation id="98D20F905854FFC935E6FBA4FF5EB76F" box="[159,225,1087,1112]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[180,1407,195,1875]" captionTargetId="figure@5.[176,1410,193,1879]" captionTargetPageId="5" captionText="FIGURE 3. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E – F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D – H." httpUri="https://zenodo.org/record/234563/files/figure.png" pageId="2" pageNumber="555">Fig. 3</figureCitation>
E, arrow). Filter plate with seven setae, two distalmost setae shorter than others (
<figureCitation id="98D20F905854FFC93113FBA4FB13B76F" box="[1130,1196,1087,1112]" captionStart="FIGURE 5" captionStartId="7.[151,250,1917,1939]" captionTargetBox="[244,1334,196,1884]" captionTargetId="figure@7.[240,1337,193,1886]" captionTargetPageId="7" captionText="FIGURE 5. Leberis colombiensis sp. nov., thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C – D, corm of limb I. E, limb II. F, Limb III. G – H, distal portion of limb III. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234565/files/figure.png" pageId="2" pageNumber="555">Fig. 5</figureCitation>
E, arrows).
</paragraph>
<paragraph id="005613155854FFC935BEFBFFFE7CB607" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFBFFFE9CB74B" box="[199,291,1124,1149]" italics="true" pageId="2" pageNumber="555">Limb III</emphasis>
(
<figureCitation id="98D20F905854FFC9344AFBFFFE3EB74B" box="[307,385,1124,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
D, 5F–H) with ovoid epipodite. Exopodite rectangular, with two lateral setae (
<figureCitation id="98D20F905854FFC93193FBFFFA87B74B" box="[1258,1336,1124,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
D, 5F–G, 5–6) and four distal setae (1–4), seta 4 longest. Distal endite (sensu 
<bibRefCitation id="64786EE45854FFC936B1FB12FBEDB797" author="Kotov" box="[968,1106,1160,1184]" pageId="2" pageNumber="555" refString="Kotov, A. A. (1999) Redescription of Macrothrix tripectinata Weisig, 1934 (Anomopoda, Branchiopoda), with a discussion of some features rarely used in the systematics of the genus. Hydrobiologia, 403, 63 - 80. http: // dx. doi. org / 10.1023 / A: 1003714909471" type="journal article" year="1999">Kotov 1999</bibRefCitation>
) sub-rectangular, with three anterior setae (
<figureCitation id="98D20F905854FFC93442FB37FE36B7F3" box="[315,393,1196,1221]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
D, 5H, 1–3); setae 1 and 2 stout, basalmost seta 3 with long setules distally. A small sensillum between setae 1 and 2, and between setae 2 and 3. On proximal endite, four anterior setae and four soft posterior setae (
<figureCitation id="98D20F905854FFC935A6FB6FFE9CB63A" box="[223,291,1268,1293]" captionStart="FIGURE 6" captionStartId="8.[151,250,1954,1976]" captionTargetBox="[233,1353,200,1928]" captionTargetId="figure@8.[228,1360,193,1933]" captionTargetPageId="8" captionText="FIGURE 6. Leberis colombiensis sp. nov., female from Ciénaga El Convento, Colombia (type locality). A, limb III, distal portion. B, gnathobase of limb III. C, limb IV. D, inner portion of limb IV. E, limb V. F, inner portion of limb V. G – H, ephippial female. Scale bars 0.1 mm for G – H; 0.01 mm for A – F." httpUri="https://zenodo.org/record/234566/files/figure.png" pageId="2" pageNumber="555">Fig. 6</figureCitation>
A, a–d), all armed with sparse, fine setules. Distal armature of gnathobase as in 
<taxonomicName id="C7E968965854FFC931CAFB6EFAA7B63A" box="[1203,1304,1268,1293]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="329DCF075854FFC931CAFB6EFAA7B63A" box="[1203,1304,1268,1293]" italics="true" pageId="2" pageNumber="555">L. davidi</emphasis>
</taxonomicName>
, filter plate with seven setae (
<figureCitation id="98D20F905854FFC93427FA8CFE1EB607" box="[350,417,1303,1328]" captionStart="FIGURE 6" captionStartId="8.[151,250,1954,1976]" captionTargetBox="[233,1353,200,1928]" captionTargetId="figure@8.[228,1360,193,1933]" captionTargetPageId="8" captionText="FIGURE 6. Leberis colombiensis sp. nov., female from Ciénaga El Convento, Colombia (type locality). A, limb III, distal portion. B, gnathobase of limb III. C, limb IV. D, inner portion of limb IV. E, limb V. F, inner portion of limb V. G – H, ephippial female. Scale bars 0.1 mm for G – H; 0.01 mm for A – F." httpUri="https://zenodo.org/record/234566/files/figure.png" pageId="2" pageNumber="555">Fig. 6</figureCitation>
B).
</paragraph>
<paragraph id="005613155854FFC935BEFAA7FB3AB6D3" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFAA7FE9EB663" box="[199,289,1340,1365]" italics="true" pageId="2" pageNumber="555">Limb IV</emphasis>
(
<figureCitation id="98D20F905854FFC93448FAA7FEC1B663" box="[305,382,1340,1365]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
E, 6C–D) with ovoid epipodite. Exopodite wide, sub-quadrangular, with six setae (1–6), seta 3 longest. Inner limb portion of endopodite with four marginal setae (
<figureCitation id="98D20F905854FFC936E2FAC4FC54B64F" box="[923,1003,1375,1400]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
E, 6D, 1–4). Distalmost seta (1) stout, naked; setae 2–4 with inflated proximal segments and slender, unilaterally setulated distal segments, seta 2 largest among the latter. On posterior limb face, three soft setae (a–c). Distal armature of gnathobase with a large sensillum (1), a peculiar brush-shaped seta (2) and two short setae (3–4). Filter plate with five setae.
</paragraph>
<paragraph id="005613155854FFC935BEFA74FDC2B567" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEFA74FEA6B53F" box="[199,281,1519,1544]" italics="true" pageId="2" pageNumber="555">Limb V</emphasis>
(
<figureCitation id="98D20F905854FFC93450FA74FEC8B53F" box="[297,375,1519,1544]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
F, 6E–F). Epipodite ovoid. Exopodite subovoid, with a single distal (
<figureCitation id="98D20F905854FFC93107FA74FB73B53F" box="[1150,1228,1519,1544]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="2" pageNumber="555">Figs. 4</figureCitation>
F, 6E, 1) and three lateral (2–4) setae. Inner limb portion elongated, with setulated inner margin, with two setulated setae of different length (
<figureCitation id="98D20F905854FFC93595F9ACFE90B567" box="[236,303,1591,1616]" captionStart="FIGURE 6" captionStartId="8.[151,250,1954,1976]" captionTargetBox="[233,1353,200,1928]" captionTargetId="figure@8.[228,1360,193,1933]" captionTargetPageId="8" captionText="FIGURE 6. Leberis colombiensis sp. nov., female from Ciénaga El Convento, Colombia (type locality). A, limb III, distal portion. B, gnathobase of limb III. C, limb IV. D, inner portion of limb IV. E, limb V. F, inner portion of limb V. G – H, ephippial female. Scale bars 0.1 mm for G – H; 0.01 mm for A – F." httpUri="https://zenodo.org/record/234566/files/figure.png" pageId="2" pageNumber="555">Fig. 6</figureCitation>
F). Filter plate was not found.
</paragraph>
<paragraph id="005613155854FFC935BEF9C7FA92B5AF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEF9C7FE2BB542" bold="true" box="[199,404,1628,1653]" pageId="2" pageNumber="555">Ephippial female</emphasis>
. Habitus similar to that in parthenogenetic female, with a bit elongated body (body height/ body length = 0.7), ephippium slightly additionally pigmented in brown, with a single egg (
<figureCitation id="98D20F905854FFC931E1F9E4FB5AB5AF" box="[1176,1253,1663,1688]" captionStart="FIGURE 6" captionStartId="8.[151,250,1954,1976]" captionTargetBox="[233,1353,200,1928]" captionTargetId="figure@8.[228,1360,193,1933]" captionTargetPageId="8" captionText="FIGURE 6. Leberis colombiensis sp. nov., female from Ciénaga El Convento, Colombia (type locality). A, limb III, distal portion. B, gnathobase of limb III. C, limb IV. D, inner portion of limb IV. E, limb V. F, inner portion of limb V. G – H, ephippial female. Scale bars 0.1 mm for G – H; 0.01 mm for A – F." httpUri="https://zenodo.org/record/234566/files/figure.png" pageId="2" pageNumber="555">Figs. 6</figureCitation>
G–H).
</paragraph>
<paragraph id="005613155854FFC935BEF93FFDEBB41E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEF93FFEBBB58A" bold="true" box="[199,260,1700,1725]" pageId="2" pageNumber="555">Male</emphasis>
. 
<emphasis id="329DCF075854FFC9346AF93FFED3B58A" box="[275,364,1700,1725]" italics="true" pageId="2" pageNumber="555">Habitus</emphasis>
in general similar to female, but maximum height smaller (height/length = 0.7–0.75); posterodorsal angle expressed; postero-ventral angle much more developed as compared with female (
<figureCitation id="98D20F905854FFC931AFF95CFA9EB5D7" box="[1238,1313,1735,1760]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="2" pageNumber="555">Figs. 1</figureCitation>
K, 7A–B). Head larger than in female, rostrum truncated, ocellus in middle of distance between compound eye and tip of rostrum, or more closely located to eye.
</paragraph>
<paragraph id="005613155854FFC935BEF8AFFE88B447" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEF8AFFEB0B47A" box="[199,271,1844,1869]" italics="true" pageId="2" pageNumber="555">Valves</emphasis>
with naked setae at postero-ventral angle, followed by a submarginal row of strong spinules of similar size (
<figureCitation id="98D20F905854FFC935ABF8CCFEAAB447" box="[210,277,1879,1904]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[226,1353,202,1873]" captionTargetId="figure@9.[214,1361,193,1882]" captionTargetPageId="9" captionText="FIGURE 7. Leberis colombiensis sp. nov., adult male from Ciénaga El Convento, Colombia (type locality). A – B, general view. C, postero-ventral portion of valve. D, postabdomen. E, armature of its postanal margin. F, lateral fascicles of setules. G, postabdominal claw. H, antenna I. I, limb I. Scale bars 0.1 mm for A – D; 0.01 mm for E – I." httpUri="https://zenodo.org/record/234567/files/figure.png" pageId="2" pageNumber="555">Fig. 7</figureCitation>
C).
</paragraph>
<paragraph id="005613155854FFC135BEF8E7FF07B6E3" blockId="2.[151,1437,151,2013]" lastBlockId="10.[119,1732,1438,1641]" lastPageId="10" lastPageNumber="563" pageId="2" pageNumber="555">
<emphasis id="329DCF075854FFC935BEF8E7FEDFB4A2" box="[199,352,1916,1941]" italics="true" pageId="2" pageNumber="555">Postabdomen</emphasis>
elongated (length/width ratio about 2.55), distal portion of postabdomen more narrow than in female, gonopore opens near base of postabdominal claw (
<figureCitation id="98D20F905854FFC9364CF83BFC3EB48F" box="[821,897,1952,1977]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="2" pageNumber="555">Figs. 1</figureCitation>
L, 7D). Preanal angle projected, postanal angle smoothed, dorso-distal angle widely rounded. Anal margin somewhat longer than preanal margin, postanal margin Postabdomen postanal/ preanal margin About 2.8 About 2.5 About 3.0 About 1.5 1.7–2.0 length
</paragraph>
<caption id="5496439D5855FFC835EEF819FF79B4C3" httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="3" pageNumber="556" targetBox="[176,1400,198,1888]" targetPageId="3">
<paragraph id="005613155855FFC835EEF819FF79B4C3" blockId="3.[151,1436,1922,2036]" pageId="3" pageNumber="556">
<emphasis id="329DCF075855FFC835EEF819FEAFB4A0" bold="true" box="[151,272,1922,1944]" pageId="3" pageNumber="556">FIGURE 1</emphasis>
. 
<taxonomicName id="C7E968965855FFC83459F819FE46B4AF" box="[288,505,1922,1944]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF075855FFC83459F819FE46B4AF" box="[288,505,1922,1944]" italics="true" pageId="3" pageNumber="556">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C5855FFC8377AF818FDE8B4AF" box="[515,599,1923,1944]" pageId="3" pageNumber="556" rank="species">
<emphasis id="329DCF075855FFC8377AF818FDE8B4AF" bold="true" box="[515,599,1923,1944]" pageId="3" pageNumber="556">sp. nov.</emphasis>
</taxonomicNameLabel>
from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D–F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="5496439D5852FFCF35EEF83AFD61B4C3" httpUri="https://zenodo.org/record/234562/files/figure.png" pageId="4" pageNumber="557" targetBox="[201,1386,200,1926]" targetPageId="4">
<paragraph id="005613155852FFCF35EEF83AFD61B4C3" blockId="4.[151,1436,1953,2036]" pageId="4" pageNumber="557">
<emphasis id="329DCF075852FFCF35EEF83AFEADB481" bold="true" box="[151,274,1953,1975]" pageId="4" pageNumber="557">FIGURE 2.</emphasis>
<taxonomicName id="C7E968965852FFCF3461F83AFE50B480" box="[280,495,1953,1975]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF075852FFCF3461F83AFE50B480" box="[280,495,1953,1975]" italics="true" pageId="4" pageNumber="557">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C5852FFCF348CF83AFDFAB481" box="[501,581,1953,1974]" pageId="4" pageNumber="557" rank="species">
<emphasis id="329DCF075852FFCF348CF83AFDFAB481" bold="true" box="[501,581,1953,1974]" pageId="4" pageNumber="557">sp. nov.</emphasis>
</taxonomicNameLabel>
, parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A–B, general view. C, head. D, head shield. E, major head pores. F, labrum. G, armature of postero-ventral portion of valve. H, antenna I. Scale bars 0.1 mm for A–D; 0.01 mm for E–H.
</paragraph>
</caption>
<caption id="5496439D5853FFCE35EEF8E0FF52B4DA" httpUri="https://zenodo.org/record/234563/files/figure.png" pageId="5" pageNumber="558" targetBox="[180,1407,195,1875]" targetPageId="5">
<paragraph id="005613155853FFCE35EEF8E0FF52B4DA" blockId="5.[151,1435,1915,2029]" pageId="5" pageNumber="558">
<emphasis id="329DCF075853FFCE35EEF8E0FEABB4A7" bold="true" box="[151,276,1915,1937]" pageId="5" pageNumber="558">FIGURE 3.</emphasis>
<taxonomicName id="C7E968965853FFCE3465F8E0FE4AB4A6" box="[284,501,1915,1937]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF075853FFCE3465F8E0FE4AB4A6" box="[284,501,1915,1937]" italics="true" pageId="5" pageNumber="558">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C5853FFCE3484F8E0FDF0B4A7" box="[509,591,1915,1936]" pageId="5" pageNumber="558" rank="species">
<emphasis id="329DCF075853FFCE3484F8E0FDF0B4A7" bold="true" box="[509,591,1915,1936]" pageId="5" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
, parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E–F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D–H.
</paragraph>
</caption>
<caption id="5496439D5850FFCD35EEF8FBFF54B484" httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="6" pageNumber="559" targetBox="[154,1418,204,1859]" targetPageId="6">
<paragraph id="005613155850FFCD35EEF8FBFF54B484" blockId="6.[151,1436,1888,1971]" pageId="6" pageNumber="559">
<emphasis id="329DCF075850FFCD35EEF8FBFEADB442" bold="true" box="[151,274,1888,1910]" pageId="6" pageNumber="559">FIGURE 4.</emphasis>
<taxonomicName id="C7E968965850FFCD3461F8FBFE51B441" box="[280,494,1888,1910]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF075850FFCD3461F8FBFE51B441" box="[280,494,1888,1910]" italics="true" pageId="6" pageNumber="559">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C5850FFCD348DF8FBFDFCB442" box="[500,579,1888,1909]" pageId="6" pageNumber="559" rank="species">
<emphasis id="329DCF075850FFCD348DF8FBFDFCB442" bold="true" box="[500,579,1888,1909]" pageId="6" pageNumber="559">sp. nov.</emphasis>
</taxonomicNameLabel>
, limbs of parthenogenetic female (A, C–F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="5496439D5851FFCC35EEF8E6FEE6B4E7" httpUri="https://zenodo.org/record/234565/files/figure.png" pageId="7" pageNumber="560" targetBox="[244,1334,196,1884]" targetPageId="7">
<paragraph id="005613155851FFCC35EEF8E6FEE6B4E7" blockId="7.[151,1436,1917,2000]" pageId="7" pageNumber="560">
<emphasis id="329DCF075851FFCC35EEF8E6FEB0B4A5" bold="true" box="[151,271,1917,1939]" pageId="7" pageNumber="560">FIGURE 5</emphasis>
. 
<taxonomicName id="C7E968965851FFCC3464F8E6FE49B4A4" box="[285,502,1917,1939]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF075851FFCC3464F8E6FE49B4A4" box="[285,502,1917,1939]" italics="true" pageId="7" pageNumber="560">Leberis colombiensis</emphasis>
</taxonomicName>
<emphasis id="329DCF075851FFCC3486F8E6FDE6B4A5" bold="true" box="[511,601,1917,1938]" pageId="7" pageNumber="560">
<taxonomicNameLabel id="29AE727C5851FFCC3486F8E6FDEBB4A5" box="[511,596,1917,1938]" pageId="7" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
thoracic limbs of parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I. C–D, corm of limb I. E, limb II. F, Limb III. G–H, distal portion of limb III. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="5496439D585EFFC335EEF839FD7DB4C2" httpUri="https://zenodo.org/record/234566/files/figure.png" pageId="8" pageNumber="561" targetBox="[233,1353,200,1928]" targetPageId="8">
<paragraph id="00561315585EFFC335EEF839FD7DB4C2" blockId="8.[151,1436,1954,2038]" pageId="8" pageNumber="561">
<emphasis id="329DCF07585EFFC335EEF839FEB0B480" bold="true" box="[151,271,1954,1976]" pageId="8" pageNumber="561">FIGURE 6</emphasis>
. 
<taxonomicName id="C7E96896585EFFC33467F839FE47B48F" box="[286,504,1954,1976]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="8" pageNumber="561" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF07585EFFC33467F839FE47B48F" box="[286,504,1954,1976]" italics="true" pageId="8" pageNumber="561">Leberis colombiensis</emphasis>
</taxonomicName>
<emphasis id="329DCF07585EFFC33779F838FDE5B48F" bold="true" box="[512,602,1955,1976]" pageId="8" pageNumber="561">
<taxonomicNameLabel id="29AE727C585EFFC33779F838FDEAB48F" box="[512,597,1955,1976]" pageId="8" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female from Ciénaga El Convento, Colombia (type locality). A, limb III, distal portion. B, gnathobase of limb III. C, limb IV. D, inner portion of limb IV. E, limb V. F, inner portion of limb V. G–H, ephippial female. Scale bars 0.1 mm for G–H; 0.01 mm for A–F.
</paragraph>
</caption>
<caption id="5496439D585FFFC235EEF8E7FB97B4E0" httpUri="https://zenodo.org/record/234567/files/figure.png" pageId="9" pageNumber="562" targetBox="[226,1353,202,1873]" targetPageId="9">
<paragraph id="00561315585FFFC235EEF8E7FB97B4E0" blockId="9.[151,1436,1916,2007]" pageId="9" pageNumber="562">
<emphasis id="329DCF07585FFFC235EEF8E7FEAFB4A5" bold="true" box="[151,272,1916,1938]" pageId="9" pageNumber="562">FIGURE 7</emphasis>
. 
<taxonomicName id="C7E96896585FFFC23466F8E7FE46B4A5" box="[287,505,1916,1938]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF07585FFFC23466F8E7FE46B4A5" box="[287,505,1916,1938]" italics="true" pageId="9" pageNumber="562">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C585FFFC23778F8E6FDEAB4A5" box="[513,597,1917,1938]" pageId="9" pageNumber="562" rank="species">
<emphasis id="329DCF07585FFFC23778F8E6FDEAB4A5" bold="true" box="[513,597,1917,1938]" pageId="9" pageNumber="562">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult male from Ciénaga El Convento, Colombia (type locality). A–B, general view. C, postero-ventral portion of valve. D, postabdomen. E, armature of its postanal margin. F, lateral fascicles of setules. G, postabdominal claw. H, antenna I. I, limb I. Scale bars 0.1 mm for A–D; 0.01 mm for E–I.
</paragraph>
</caption>
<caption id="5496439D585CFFC1350EFDA2FDDDB159" ID-Table-UUID="5496439D585CFFC1350EFDA2FDDDB159" httpUri="http://table.plazi.org/id/5496439D585CFFC1350EFDA2FDDDB159" pageId="10" pageNumber="563" targetBox="[119,1833,660,1402]" targetIsTable="true" targetPageId="10">
<paragraph id="00561315585CFFC1350EFDA2FDDDB159" blockId="10.[119,1875,568,622]" pageId="10" pageNumber="563">
<emphasis id="329DCF07585CFFC1350EFDA2FF55B167" bold="true" box="[119,234,569,592]" pageId="10" pageNumber="563">TABLE 1.</emphasis>
Main morphological differences of 
<taxonomicName id="C7E96896585CFFC13718FDA3FC81B167" box="[609,830,568,592]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF07585CFFC13718FDA3FC81B167" box="[609,830,568,592]" italics="true" pageId="10" pageNumber="563">Leberis colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C585CFFC1363DFDA2FC2DB167" box="[836,914,568,592]" pageId="10" pageNumber="563" rank="species">
<emphasis id="329DCF07585CFFC1363DFDA2FC34B167" bold="true" box="[836,907,569,592]" pageId="10" pageNumber="563">sp.nov</emphasis>
.
</taxonomicNameLabel>
from 
<taxonomicName id="C7E96896585CFFC136A8FDA3FB92B167" box="[977,1069,568,592]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="329DCF07585CFFC136A8FDA3FB92B167" box="[977,1069,568,592]" italics="true" pageId="10" pageNumber="563">L. davidi</emphasis>
</taxonomicName>
, 
<taxonomicName id="C7E96896585CFFC13142FDA3FB52B167" box="[1083,1261,568,592]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="chihuahuensis">
<emphasis id="329DCF07585CFFC13142FDA3FBF6B167" box="[1083,1097,568,592]" italics="true" pageId="10" pageNumber="563">L</emphasis>
. 
<emphasis id="329DCF07585CFFC1312FFDA3FB52B167" box="[1110,1261,568,592]" italics="true" pageId="10" pageNumber="563">chihuahuensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C7E96896585CFFC13181FDA3FAC0B167" box="[1272,1407,568,592]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="diaphanus">L. diaphanus</taxonomicName>
and 
<taxonomicName id="C7E96896585CFFC130CBFDA3F9E3B167" box="[1458,1628,568,592]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="aenigmatosus">
<emphasis id="329DCF07585CFFC130CBFDA3F9E3B167" box="[1458,1628,568,592]" italics="true" pageId="10" pageNumber="563">L. aenigmatosus</emphasis>
</taxonomicName>
(after Frey 1991, 1998; Sinev 
<emphasis id="329DCF07585CFFC135C0FDCDFF59B159" box="[185,230,598,622]" italics="true" pageId="10" pageNumber="563">et al</emphasis>
. 2005; Sinev &amp; Sanoamuang 2011).
</paragraph>
</caption>
<paragraph id="00561315585CFFC1350EFD0FF9C3B66B" pageId="10" pageNumber="563">
<table id="72E9E1B5585C0034350EFD0FF896B64D" box="[119,1833,660,1402]" gridcols="6" gridrows="4" pageId="10" pageNumber="563">
<tr id="BED91157585C0034350EFD0FF896B19C" box="[119,1833,660,683]" gridrow="0" pageId="10" pageNumber="563">
<th id="FD08782B585C0034350EFD0FFD8EB19C" box="[119,561,660,683]" gridcol="0" gridrow="0" pageId="10" pageNumber="563">
<emphasis id="329DCF07585CFFC1350EFD0FFF56B19C" bold="true" box="[119,233,660,683]" pageId="10" pageNumber="563">Character</emphasis>
</th>
<th id="FD08782B585C00343728FD0FFC82B19C" box="[593,829,660,683]" gridcol="1" gridrow="0" pageId="10" pageNumber="563">
<taxonomicName id="C7E96896585CFFC13728FD0FFD46B19C" box="[593,761,660,683]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="colombiensis">
<emphasis id="329DCF07585CFFC13728FD0FFD46B19C" bold="true" box="[593,761,660,683]" italics="true" pageId="10" pageNumber="563">L. colombiensis</emphasis>
</taxonomicName>
</th>
<th id="FD08782B585C00343620FD0FFB98B19C" box="[857,1063,660,683]" gridcol="2" gridrow="0" pageId="10" pageNumber="563">
<taxonomicName id="C7E96896585CFFC13620FD0FFC08B19C" box="[857,951,660,683]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="329DCF07585CFFC13620FD0FFC08B19C" bold="true" box="[857,951,660,683]" italics="true" pageId="10" pageNumber="563">L. davidi</emphasis>
</taxonomicName>
</th>
<th id="FD08782B585C0034313AFD0FFA85B19C" box="[1091,1338,660,683]" gridcol="3" gridrow="0" pageId="10" pageNumber="563">
<taxonomicName id="C7E96896585CFFC1313AFD0FFB41B19C" box="[1091,1278,660,683]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="chihuahuensis">
<emphasis id="329DCF07585CFFC1313AFD0FFB41B19C" bold="true" box="[1091,1278,660,683]" italics="true" pageId="10" pageNumber="563">L. chihuahuensis</emphasis>
</taxonomicName>
</th>
<th id="FD08782B585C00343022FD0FF997B19C" box="[1371,1576,660,683]" gridcol="4" gridrow="0" pageId="10" pageNumber="563">
<taxonomicName id="C7E96896585CFFC13022FD0FFA49B19C" box="[1371,1526,660,683]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="diaphanous">
<emphasis id="329DCF07585CFFC13022FD0FFA49B19C" bold="true" box="[1371,1526,660,683]" italics="true" pageId="10" pageNumber="563">L. diaphanous</emphasis>
</taxonomicName>
</th>
<th id="FD08782B585C00343325FD0FF896B19C" box="[1628,1833,660,683]" gridcol="5" gridrow="0" pageId="10" pageNumber="563">
<taxonomicName id="C7E96896585CFFC13324FD0FF8B3B19C" box="[1629,1804,660,683]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="aenigmatosus">
<emphasis id="329DCF07585CFFC13324FD0FF8B3B19C" bold="true" box="[1629,1804,660,683]" italics="true" pageId="10" pageNumber="563">L. aenigmatosus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="BED91157585C0034350EFD28F896B075" box="[119,1833,691,834]" gridrow="1" pageId="10" pageNumber="563">
<th id="FD08782B585C0034350EFD28FD8EB075" box="[119,561,691,834]" gridcol="0" gridrow="1" pageId="10" pageNumber="563">
<emphasis id="329DCF07585CFFC1350EFD28FEB8B1FD" bold="true" box="[119,263,691,714]" pageId="10" pageNumber="563">Adult female</emphasis>
Maximum size of female Number of setae on ventral margin of valve Post-pore/inter-pore distance
</th>
<td id="FD08782B585C00343728FD28FC82B075" box="[593,829,691,834]" gridcol="1" gridrow="1" pageId="10" pageNumber="563">0.50 mm 67–75 1.0–1.2</td>
<td id="FD08782B585C00343620FD28FB98B075" box="[857,1063,691,834]" gridcol="2" gridrow="1" pageId="10" pageNumber="563">0.69 mm 84–95 1.5–2.0</td>
<td id="FD08782B585C0034313AFD28FA85B075" box="[1091,1338,691,834]" gridcol="3" gridrow="1" pageId="10" pageNumber="563">0.53 mm 51–55 2.3–2.6</td>
<td id="FD08782B585C00343022FD28F997B075" box="[1371,1576,691,834]" gridcol="4" gridrow="1" pageId="10" pageNumber="563">0.56 mm 58–67 &lt;1</td>
<td id="FD08782B585C00343325FD28F896B075" box="[1628,1833,691,834]" gridcol="5" gridrow="1" pageId="10" pageNumber="563">1.1 mm 101–129 3–4</td>
</tr>
<tr id="BED91157585C0034350EFCD2F896B7BC" box="[119,1833,841,1163]" gridrow="2" pageId="10" pageNumber="563">
<th id="FD08782B585C0034350EFCD2FD8EB7BC" box="[119,561,841,1163]" gridcol="0" gridrow="2" pageId="10" pageNumber="563">Anterior margin of labrum Spine on proximal segment of exopod Outer distal lobe of limb I Exopodite III Exopodite IV Finger-like projections on epipodite III-IV</th>
<td id="FD08782B585C00343728FCD2FC82B7BC" box="[593,829,841,1163]" gridcol="1" gridrow="2" pageId="10" pageNumber="563">Convex Shorter than second segment With sensillum, without accessory seta Seta 2 and 1 of the same length Seta 2 longer than seta 1 Absent</td>
<td id="FD08782B585C00343620FCD2FB98B7BC" box="[857,1063,841,1163]" gridcol="2" gridrow="2" pageId="10" pageNumber="563">Wavy Shorter than second segment With sensillum and with accessory seta Setae 1 and 2 of the same length Seta 1 and 2 of same length Present</td>
<td id="FD08782B585C0034313AFCD2FA85B7BC" box="[1091,1338,841,1163]" gridcol="3" gridrow="2" pageId="10" pageNumber="563">Convex Longer than second segment Without sensillum, whithout accessory seta Seta 2 longer than seta 1 Seta 2 longer than seta 1 Absent</td>
<td id="FD08782B585C00343022FCD2F997B7BC" box="[1371,1576,841,1163]" gridcol="4" gridrow="2" pageId="10" pageNumber="563">Convex Longer than second segment Without sensillum, with accessory seta Seta 2 longer than seta 1 Seta 2 longer than seta 1 Absent</td>
<td id="FD08782B585C00343325FCD2F896B7BC" box="[1628,1833,841,1163]" gridcol="5" gridrow="2" pageId="10" pageNumber="563">Convex Longer than second segment Without sensillum, without accessory seta Seta 2 longer than seta 1 No information Absent</td>
</tr>
<tr id="BED91157585C0034350EFB08F896B64D" box="[119,1833,1171,1402]" gridrow="3" pageId="10" pageNumber="563">
<th id="FD08782B585C0034350EFB08FD8EB64D" box="[119,561,1171,1402]" gridcol="0" gridrow="3" pageId="10" pageNumber="563">
<emphasis id="329DCF07585CFFC1350EFB08FF10B79D" bold="true" box="[119,175,1171,1194]" pageId="10" pageNumber="563">Male</emphasis>
Setules in middle of posterior margin longer than in region of postero-venral angle Postabdomen distal portion remarkably asymmetrical relative to longitudinal axis Gonopore region on postabdomen inflated, overhangs base of postabdominal
</th>
<td id="FD08782B585C00343728FB08FC82B64D" box="[593,829,1171,1402]" gridcol="1" gridrow="3" pageId="10" pageNumber="563">No No No</td>
<td id="FD08782B585C00343620FB08FB98B64D" box="[857,1063,1171,1402]" gridcol="2" gridrow="3" pageId="10" pageNumber="563">Yes Yes Yes</td>
<td id="FD08782B585C0034313AFB08FA85B64D" box="[1091,1338,1171,1402]" gridcol="3" gridrow="3" pageId="10" pageNumber="563">No Yes Yes</td>
<td id="FD08782B585C00343022FB08F997B64D" box="[1371,1576,1171,1402]" gridcol="4" gridrow="3" pageId="10" pageNumber="563">Yes Yes No</td>
<td id="FD08782B585C00343325FB08F896B64D" box="[1628,1833,1171,1402]" gridcol="5" gridrow="3" pageId="10" pageNumber="563">No No No</td>
</tr>
</table>
</paragraph>
<paragraph id="00561315585CFFC135EDFA41F9C3B6C5" blockId="10.[119,1732,1438,1641]" box="[148,1660,1498,1522]" pageId="10" pageNumber="563">each lateral fascicle on postanal portion Yes No Yes No No</paragraph>
<paragraph id="00561315585CFFC135EDFA6CFDA4B538" blockId="10.[119,1732,1438,1641]" box="[148,539,1527,1551]" pageId="10" pageNumber="563">postabdomen distal 1-2 setae stronger</paragraph>
<paragraph id="00561315585CFFC135D2F98EFF55B51A" blockId="10.[119,1732,1438,1641]" box="[171,234,1557,1581]" pageId="10" pageNumber="563">others</paragraph>
<paragraph id="00561315585CFFC1350EF9A8FE84B55E" blockId="10.[119,1732,1438,1641]" pageId="10" pageNumber="563">Length of basal spine/ length of About 0.4 About 0.2–0.3 About 0.4 About 0.5 About 0.4 postabominal claw</paragraph>
<paragraph id="00561315585DFFC035EEFF03FB48B22B" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
2.8 times longer than preanal margin. Successive series of fine setules on preanal margin (
<figureCitation id="98D20F90585DFFC031C6FF0CFABAB387" box="[1215,1285,151,176]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[226,1353,202,1873]" captionTargetId="figure@9.[214,1361,193,1882]" captionTargetPageId="9" captionText="FIGURE 7. Leberis colombiensis sp. nov., adult male from Ciénaga El Convento, Colombia (type locality). A – B, general view. C, postero-ventral portion of valve. D, postabdomen. E, armature of its postanal margin. F, lateral fascicles of setules. G, postabdominal claw. H, antenna I. I, limb I. Scale bars 0.1 mm for A – D; 0.01 mm for E – I." httpUri="https://zenodo.org/record/234567/files/figure.png" pageId="11" pageNumber="564">Fig. 7</figureCitation>
E) instead of denticles in female (
<figureCitation id="98D20F90585DFFC03403FF27FE79B3E3" box="[378,454,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[180,1407,195,1875]" captionTargetId="figure@5.[176,1410,193,1879]" captionTargetPageId="5" captionText="FIGURE 3. Leberis colombiensis sp. nov., parthenogenetic female from Ciénaga El Convento, Colombia (type locality). A, antenna II. B, distal portions of its branches. C, postabdomen. D, postabdominal anal margin. E – F, armature of postabdominal distal portion; G, lateral fascicles of setules on postabdomen. H, postabdominal claw. Scale bars 0.1 mm for A, C; 0.01 mm for B, D – H." httpUri="https://zenodo.org/record/234563/files/figure.png" pageId="11" pageNumber="564">Figs. 3</figureCitation>
E–F); fascicles of setules on each side of postabdomen, 1–2 distal setules in each group longer and robuster than others (
<figureCitation id="98D20F90585DFFC03768FF44FDE8B3CF" box="[529,599,223,248]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[226,1353,202,1873]" captionTargetId="figure@9.[214,1361,193,1882]" captionTargetPageId="9" captionText="FIGURE 7. Leberis colombiensis sp. nov., adult male from Ciénaga El Convento, Colombia (type locality). A – B, general view. C, postero-ventral portion of valve. D, postabdomen. E, armature of its postanal margin. F, lateral fascicles of setules. G, postabdominal claw. H, antenna I. I, limb I. Scale bars 0.1 mm for A – D; 0.01 mm for E – I." httpUri="https://zenodo.org/record/234567/files/figure.png" pageId="11" pageNumber="564">Fig. 7</figureCitation>
F). Postabdominal claws shorter and robuster as compared with females. Basal spine long (length about 0.4 claw length), with large setules along its dorsal margin (
<figureCitation id="98D20F90585DFFC031E8FE9FFB6BB22B" box="[1169,1236,260,285]" captionStart="FIGURE 7" captionStartId="9.[151,250,1916,1938]" captionTargetBox="[226,1353,202,1873]" captionTargetId="figure@9.[214,1361,193,1882]" captionTargetPageId="9" captionText="FIGURE 7. Leberis colombiensis sp. nov., adult male from Ciénaga El Convento, Colombia (type locality). A – B, general view. C, postero-ventral portion of valve. D, postabdomen. E, armature of its postanal margin. F, lateral fascicles of setules. G, postabdominal claw. H, antenna I. I, limb I. Scale bars 0.1 mm for A – D; 0.01 mm for E – I." httpUri="https://zenodo.org/record/234567/files/figure.png" pageId="11" pageNumber="564">Fig. 7</figureCitation>
G).
</paragraph>
<paragraph id="00561315585DFFC035BEFEB3FDDAB253" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
<emphasis id="329DCF07585DFFC035BEFEB3FE8BB277" box="[199,308,296,320]" italics="true" pageId="11" pageNumber="564">Antenna I</emphasis>
relatively thick, with seven terminal aesthetascs and two lateral aesthetascs, male seta near them, and with a small sensory seta (
<figureCitation id="98D20F90585DFFC034C7FED7FDB8B253" box="[446,519,332,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1922,1944]" captionTargetBox="[176,1400,198,1888]" captionTargetId="figure@3.[164,1414,193,1895]" captionTargetPageId="3" captionText="FIGURE 1. Leberis colombiensis sp. nov. from Ciénaga El Convento, Colombia (type locality). A, adult parthenogenetic female, general view. B, head shield. C, dorsal head pores. D – F, labrum and labral keel. G, armature of postero-ventral portion of valve. H, postabdomen. I, antenna I. J, antenna II. K, adult male, lateral view. L, postabdomen. M, antenna I. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234561/files/figure.png" pageId="11" pageNumber="564">Figs. 1</figureCitation>
M, 7H).
</paragraph>
<paragraph id="00561315585DFFC035BEFEF4FE3EB2F8" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
<emphasis id="329DCF07585DFFC035BEFEF4FEEFB2BF" box="[199,336,367,392]" italics="true" pageId="11" pageNumber="564">Trunk limb I</emphasis>
with outer distal lobe as in female; inner distal lobe with a strong, U-shaped copulatory hook; two setae as in female but armed by somewhat finer setules, and additional male seta (
<figureCitation id="98D20F90585DFFC0314DFE0FFB3CB29B" box="[1076,1155,404,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[154,1418,204,1859]" captionTargetId="figure@6.[151,1436,193,1867]" captionTargetPageId="6" captionText="FIGURE 4. Leberis colombiensis sp. nov., limbs of parthenogenetic female (A, C – F) and male (B) from Ciénaga El Convento, Colombia (type locality). A, limb I. B, distal portion of limb I in male. C, limb II. D, limb III. E, limb IV. F, limb V. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/234564/files/figure.png" pageId="11" pageNumber="564">Figs. 4</figureCitation>
D, 
<date id="745735D5585DFFC031DCFE0EFB01B29B" box="[1189,1214,405,428]" pageId="11" pageNumber="564">7I</date>
, arrow) longer than half of a former seta.
</paragraph>
<paragraph id="00561315585DFFC035BEFE47FD84B120" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
<emphasis id="329DCF07585DFFC035BEFE47FF41B2C2" bold="true" box="[199,254,476,501]" pageId="11" pageNumber="564">Size.</emphasis>
Juvenile females 
<quantity id="C711BEF0585DFFC034B2FE47FDCDB2C3" box="[459,626,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="3.35" metricValueMax="4.0" metricValueMin="2.7" pageId="11" pageNumber="564" unit="mm" value="0.335" valueMax="0.4" valueMin="0.27">0.27–0.40 mm</quantity>
; adult parthenogenetic females 
<quantity id="C711BEF0585DFFC036A3FE47FB3EB2C3" box="[986,1153,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="5.0" metricValueMin="4.2" pageId="11" pageNumber="564" unit="mm" value="0.45999999999999996" valueMax="0.5" valueMin="0.42">0.42–0.50 mm</quantity>
; ephippial females 
<quantity id="C711BEF0585DFFC03024FE47FF41B120" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="5.0" metricValueMin="3.9" pageId="11" pageNumber="564" unit="mm" value="0.445" valueMax="0.5" valueMin="0.39">0.39– 0.50 mm</quantity>
; adult males 
<quantity id="C711BEF0585DFFC034E9FE64FD89B120" box="[400,566,511,536]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.5999999999999996" pageId="11" pageNumber="564" unit="mm" value="0.39" valueMax="0.42" valueMin="0.36">0.36–0.42 mm</quantity>
.
</paragraph>
<paragraph id="00561315585DFFC035BEFDBFFB53B157" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
<emphasis id="329DCF07585DFFC035BEFDBFFEFAB10A" bold="true" box="[199,325,548,573]" pageId="11" pageNumber="564">Variability</emphasis>
. In a single female, an incomplete scraper 4 of limb II was found. In a single female, the spine on the second exopod segment of antenna II was very short, not reaching the half of the next segment.
</paragraph>
<paragraph id="00561315585DFFC035BEFDF7FB1DB00F" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
<emphasis id="329DCF07585DFFC035BEFDF7FE70B1B2" bold="true" box="[199,463,620,645]" pageId="11" pageNumber="564">Differential diagnosis.</emphasis>
In the Americas, 
<taxonomicName id="C7E96896585DFFC037E2FDF6FCF3B1B2" box="[667,844,620,645]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF07585DFFC037E2FDF6FCF3B1B2" box="[667,844,620,645]" italics="true" pageId="11" pageNumber="564">L. colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C585DFFC0362DFDF7FC11B1B2" box="[852,942,620,645]" pageId="11" pageNumber="564" rank="species">
<emphasis id="329DCF07585DFFC0362DFDF7FC11B1B2" bold="true" box="[852,942,620,645]" pageId="11" pageNumber="564">sp. nov.</emphasis>
</taxonomicNameLabel>
could be confused with 
<taxonomicName id="C7E96896585DFFC031BFFDF6FF67B19F" authority="Richard, 1985" authorityName="Richard" authorityYear="1985" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="329DCF07585DFFC031BFFDF6FA94B1B2" box="[1222,1323,620,645]" italics="true" pageId="11" pageNumber="564">L. davidi</emphasis>
(Richard, 1985)
</taxonomicName>
and 
<taxonomicName id="C7E96896585DFFC0346BFD0AFC28B19F" authority="Elias-Gutierrez &amp; Velez-Moreno, 2008" authorityName="Elias-Gutierrez &amp; Velez-Moreno" authorityYear="2008" box="[274,919,655,680]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="chihuahuensis">
<emphasis id="329DCF07585DFFC0346BFD0AFE6EB19F" box="[274,465,655,680]" italics="true" pageId="11" pageNumber="564">L. chihuahuensis</emphasis>
<bibRefCitation id="64786EE4585DFFC034A0FD14FC28B19F" author="Elias-Gutierrez" box="[473,919,655,680]" pageId="11" pageNumber="564" refString="Elias-Gutierrez, M. &amp; Valdez-Moreno, M. (2008) A new cryptic species of Leberis Smirnov, 1989 (Crustacea, Cladocera, Chydoridae) from the Mexican semi-desert region, highlighted by DNA barcoding. Hidrobiologica, 18, 63 - 74." type="journal article" year="2008">Elías-Gutiérrez &amp; Velez-Moreno, 2008</bibRefCitation>
</taxonomicName>
. But the former could be easily distinguished from the latter by: (1) smaller post-pore distance in both sexes; (2) in male, postabdomen distal portion symmetrical relative to longitudinal axis, with gonopore region un-inflated, not overhangs base of postabdominal claw; (3) shorter postanal portion of male postabdomen. In addition, 
<taxonomicName id="C7E96896585DFFC0368EFD66FB10B022" box="[1015,1199,764,789]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="329DCF07585DFFC0368EFD66FB10B022" box="[1015,1199,764,789]" italics="true" pageId="11" pageNumber="564">L. colombiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29AE727C585DFFC031B9FD67FA9CB022" box="[1216,1315,764,789]" pageId="11" pageNumber="564" rank="species">
<emphasis id="329DCF07585DFFC031B9FD67FA9CB022" bold="true" box="[1216,1315,764,789]" pageId="11" pageNumber="564">sp. nov.</emphasis>
</taxonomicNameLabel>
could be differentiated from 
<taxonomicName id="C7E96896585DFFC0340BFCBAFE6AB00F" box="[370,469,799,824]" class="Branchiopoda" genus="Leberis" kingdom="Animalia" order="Diplostraca" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="329DCF07585DFFC0340BFCBAFE6AB00F" box="[370,469,799,824]" italics="true" pageId="11" pageNumber="564">L. davidi</emphasis>
</taxonomicName>
by its smaller maximum size. For other differences see 
<tableCitation id="4D6B26AE585DFFC03131FC84FB23B00F" box="[1096,1180,799,824]" captionStart="TABLE 1" captionStartId="10.[119,207,569,592]" captionTargetBox="[119,1833,660,1402]" captionTargetPageId="10" captionText="TABLE 1. Main morphological differences of Leberis colombiensis sp. nov. from L. davidi, L. chihuahuensis, L. diaphanus and L. aenigmatosus (after Frey 1991, 1998; Sinev et al. 2005; Sinev &amp; Sanoamuang 2011)." httpUri="http://table.plazi.org/id/5496439D585CFFC1350EFDA2FDDDB159" pageId="11" pageNumber="564" tableUuid="5496439D585CFFC1350EFDA2FDDDB159">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="48F3409E585DFFC035BEFCDFFE87B722" pageId="11" pageNumber="564" type="distribution">
<paragraph id="00561315585DFFC035BEFCDFFE87B722" blockId="11.[151,1437,151,1046]" pageId="11" pageNumber="564">
<emphasis id="329DCF07585DFFC035BEFCDFFE4AB06A" bold="true" box="[199,501,836,861]" pageId="11" pageNumber="564">Distribution and ecology.</emphasis>
The species is known to date from Atlántico Department, Magdalena Department and La Guajira Department in 
<collectingCountry id="78FE5385585DFFC03492FCFCFDE2B0B7" box="[491,605,871,896]" name="Colombia" pageId="11" pageNumber="564">Colombia</collectingCountry>
. The 
<typeStatus id="DF52ADB7585DFFC037E0FCF3FD76B0B7" box="[665,713,872,896]" pageId="11" pageNumber="564">type</typeStatus>
locality, La Ciénaga El Convento, is a small (surface area of 250 ha) lagoon system, with shallow water bodies (depth 
<quantity id="C711BEF0585DFFC0366BFC14FC36B090" box="[786,905,911,935]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="11" pageNumber="564" unit="m" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 m</quantity>
) covered by submerged and floating aquatic vegetation, represented by patches of 
<taxonomicName id="C7E96896585DFFC03713FC29FBA9B0FC" authority="(Mart.) Solms" authorityName="(Mart.) Solms" box="[618,1046,946,971]" class="Liliopsida" family="Pontederiaceae" genus="Eichhornia" kingdom="Plantae" order="Commelinales" pageId="11" pageNumber="564" phylum="Tracheophyta" rank="species" species="crassipes">
<emphasis id="329DCF07585DFFC03713FC29FCDFB0FC" box="[618,864,946,971]" italics="true" pageId="11" pageNumber="564">Eichhornia crassipes</emphasis>
(Mart.) Solms
</taxonomicName>
and 
<taxonomicName id="C7E96896585DFFC03126FC29FA27B0FC" authority="Gaertn." authorityName="Gaertn." box="[1119,1432,946,971]" class="Magnoliopsida" family="Nelumbonaceae" genus="Nelumbo" kingdom="Plantae" order="Proteales" pageId="11" pageNumber="564" phylum="Tracheophyta" rank="species" species="nucifera">
<emphasis id="329DCF07585DFFC03126FC29FA8EB0FC" box="[1119,1329,946,971]" italics="true" pageId="11" pageNumber="564">Nelumbo nucifera</emphasis>
Gaertn.
</taxonomicName>
; temperature varies over the seasons in the range of 27.5–30 ºC; during the sampling pH was 8.6 and conductivity 2,828 µS.cm -1.
</paragraph>
</subSubSection>
</treatment>
</document>