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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.91.83091" ID-Pensoft-Pub="1314-4049-91-113" ID-Pensoft-UUID="CF7C54854F6A5FF9A1E51BB07889ED33" ModsDocID="1314-4049-91-113" checkinTime="1658871291642" checkinUser="pensoft" docAuthor="Mongkolsamrit, Suchada, Noisripoom, Wasana, Tasanathai, Kanoksri, Kobmoo, Noppol, Thanakitpipattana, Donnaya, Khonsanit, Artit, Petcharad, Booppa, Sakolrak, Baramee &amp; Himaman, Winanda" docDate="2022" docId="664FD2BB96905008B84D3AC9DC881EE1" docLanguage="en" docName="MycoKeys 91: 113-149" docOrigin="MycoKeys 91" docPubDate="2022-07-26" docSource="http://dx.doi.org/10.3897/mycokeys.91.83091" docTitle="Polystromomyces araneae Mongkolsamrit, Noisripoom, Sakolrak &amp; Himaman 2022, sp. nov." docType="treatment" docVersion="1" id="CF7C54854F6A5FF9A1E51BB07889ED33" lastPageNumber="113" masterDocId="CF7C54854F6A5FF9A1E51BB07889ED33" masterDocTitle="Comprehensive treatise of Hevansia and three new genera Jenniferia, Parahevansia and Polystromomyces on spiders in Cordycipitaceae from Thailand" masterLastPageNumber="149" masterPageNumber="113" pageNumber="113" updateTime="1658871593950" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Comprehensive treatise of Hevansia and three new genera Jenniferia, Parahevansia and Polystromomyces on spiders in Cordycipitaceae from Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mongkolsamrit, Suchada</mods:namePart>
<mods:affiliation>Plant Microbe Interaction Research Team, National Center for Genetic Engineering and Biotechnology (BIOTEC), 113 Thailand Science Park, Phahonyothin Road, Khlong Nueng, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">suchada@biotec.or.th</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Noisripoom, Wasana</mods:namePart>
<mods:affiliation>Plant Microbe Interaction Research Team, National Center for Genetic Engineering and Biotechnology (BIOTEC), 113 Thailand Science Park, Phahonyothin Road, Khlong Nueng, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tasanathai, Kanoksri</mods:namePart>
<mods:affiliation>Plant Microbe Interaction Research Team, National Center for Genetic Engineering and Biotechnology (BIOTEC), 113 Thailand Science Park, Phahonyothin Road, Khlong Nueng, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kobmoo, Noppol</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3261-9970</mods:nameIdentifier>
<mods:affiliation>Plant Microbe Interaction Research Team, National Center for Genetic Engineering and Biotechnology (BIOTEC), 113 Thailand Science Park, Phahonyothin Road, Khlong Nueng, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thanakitpipattana, Donnaya</mods:namePart>
<mods:affiliation>Plant Microbe Interaction Research Team, National Center for Genetic Engineering and Biotechnology (BIOTEC), 113 Thailand Science Park, Phahonyothin Road, Khlong Nueng, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khonsanit, Artit</mods:namePart>
<mods:affiliation>Plant Microbe Interaction Research Team, National Center for Genetic Engineering and Biotechnology (BIOTEC), 113 Thailand Science Park, Phahonyothin Road, Khlong Nueng, Khlong Luang, Pathum Thani, 12120, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petcharad, Booppa</mods:namePart>
<mods:affiliation>Department of Biotechnology, Faculty of Science and Technology, Thammasat University, Pathum Thani, 12120, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sakolrak, Baramee</mods:namePart>
<mods:affiliation>Forest Entomology and Microbiology Research Group, Forest and Plant Conservation Research Office, 61 Department of National Parks, Wildlife and Plant Conservation, Phahonyothin Road, Chatuchak, Bangkok, 10900, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Himaman, Winanda</mods:namePart>
<mods:affiliation>Forest Entomology and Microbiology Research Group, Forest and Plant Conservation Research Office, 61 Department of National Parks, Wildlife and Plant Conservation, Phahonyothin Road, Chatuchak, Bangkok, 10900, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>MycoKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>91</mods:number>
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<mods:extent unit="page">
<mods:start>113</mods:start>
<mods:end>149</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.91.83091</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.91.83091</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-91-113</mods:identifier>
<mods:identifier type="Pensoft-UUID">CF7C54854F6A5FF9A1E51BB07889ED33</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="113" type="results">
<treatment LSID="urn:lsid:plazi:treatment:664FD2BB96905008B84D3AC9DC881EE1" httpUri="http://treatment.plazi.org/id/664FD2BB96905008B84D3AC9DC881EE1" lastPageNumber="113" pageId="0" pageNumber="113">
<subSubSection pageId="0" pageNumber="113" type="nomenclature">
<paragraph pageId="0" pageNumber="113">
<taxonomicName LSID="664FD2BB-9690-5008-B84D-3AC9DC881EE1" authority="Mongkolsamrit, Noisripoom, Sakolrak &amp; Himaman" authorityName="Mongkolsamrit, Noisripoom, Sakolrak &amp; Himaman" authorityYear="2022" class="Arachnida" family="Cordycipitaceae" genus="Polystromomyces" higherTaxonomySource="treatment-meta" kingdom="Fungi" lsidName="Polystromomyces araneae" order="Hypocreales" pageId="0" pageNumber="113" rank="species" species="araneae" status="sp. nov.">Polystromomyces araneae Mongkolsamrit, Noisripoom, Sakolrak &amp; Himaman</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="113">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="description">
<paragraph pageId="0" pageNumber="113">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Polystromomyces araneae a fungus on a spider egg sac (BBH 49054) b fertile heads c perithecia d asci e ascus tip f part-spores g, h colonies on OA at 21 days (g obverse, h reverse) i conidium formation from a hypha on OA j microcycle conidiation on OA k, l colonies on PDA at 21 days (k obverse, l reverse) m conidia formation from a hypha on PDA n microcycle conidiation on PDA. Scale bars: 10 mm (a); 3 mm (b); 200 µm (c, d); 10 µm (e, f, i, j, m, n)." figureDoi="10.3897/mycokeys.91.83091.figure8" httpUri="https://binary.pensoft.net/fig/719481" pageId="0" pageNumber="113">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="typification">
<paragraph pageId="0" pageNumber="113">Typification.</paragraph>
<paragraph pageId="0" pageNumber="113">
Thailand, Tak Province, Umphang Wildlife Sanctuary,
<geoCoordinate degrees="15" direction="north" minutes="55" orientation="latitude" precision="1" seconds="36.33" value="15.926759">15°55'36.33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="45" orientation="longitude" precision="1" seconds="12.15" value="98.75337">98°45'12.15&quot;E</geoCoordinate>
, on spider egg sac (
<taxonomicName class="Arachnida" family="Araneidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="family">Araneidae</taxonomicName>
<emphasis italics="true" pageId="0" pageNumber="113">sensu lato</emphasis>
) attached to the underside of a dicot leaf, 6 December 2020, B. Sakolrak, MY12684 (BBH 49054, holotype), ex-type culture BCC 93301 isolated from ascospores.
</paragraph>
<caption doi="10.3897/mycokeys.91.83091.figure8" httpUri="https://binary.pensoft.net/fig/719481" pageId="0" pageNumber="113" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="113">
<emphasis bold="true" pageId="0" pageNumber="113">Figure 8.</emphasis>
<taxonomicName authorityName="Mongkolsamrit, Noisripoom, Sakolrak &amp; Himaman" authorityYear="2022" class="Arachnida" family="Cordycipitaceae" genus="Polystromomyces" kingdom="Fungi" lsidName="Polystromomyces araneae" order="Hypocreales" pageId="0" pageNumber="113" rank="species" species="araneae">
<emphasis italics="true" pageId="0" pageNumber="113">Polystromomyces araneae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="113">a</emphasis>
fungus on a spider egg sac (BBH 49054)
<emphasis bold="true" pageId="0" pageNumber="113">b</emphasis>
fertile heads
<emphasis bold="true" pageId="0" pageNumber="113">c</emphasis>
perithecia
<emphasis bold="true" pageId="0" pageNumber="113">d</emphasis>
asci
<emphasis bold="true" pageId="0" pageNumber="113">e</emphasis>
ascus tip
<emphasis bold="true" pageId="0" pageNumber="113">f</emphasis>
part-spores
<emphasis bold="true" pageId="0" pageNumber="113">g, h</emphasis>
colonies on OA at 21 days (
<emphasis bold="true" pageId="0" pageNumber="113">g</emphasis>
obverse,
<emphasis bold="true" pageId="0" pageNumber="113">h</emphasis>
reverse)
<emphasis bold="true" pageId="0" pageNumber="113">i</emphasis>
conidium formation from a hypha on OA
<emphasis bold="true" pageId="0" pageNumber="113">j</emphasis>
microcycle conidiation on OA
<emphasis bold="true" pageId="0" pageNumber="113">k, l</emphasis>
colonies on PDA at 21 days (
<emphasis bold="true" pageId="0" pageNumber="113">k</emphasis>
obverse,
<emphasis bold="true" pageId="0" pageNumber="113">l</emphasis>
reverse)
<emphasis bold="true" pageId="0" pageNumber="113">m</emphasis>
conidia formation from a hypha on PDA
<emphasis bold="true" pageId="0" pageNumber="113">n</emphasis>
microcycle conidiation on PDA. Scale bars: 10 mm (
<emphasis bold="true" pageId="0" pageNumber="113">a</emphasis>
); 3 mm (
<emphasis bold="true" pageId="0" pageNumber="113">b</emphasis>
); 200
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="113">c, d</emphasis>
); 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="113">e, f, i, j, m, n</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="etymology">
<paragraph pageId="0" pageNumber="113">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="113">
From Latin,
<normalizedToken originalValue="“aranea”">&quot;aranea&quot;</normalizedToken>
refers to a spider host.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="description">
<paragraph pageId="0" pageNumber="113">Description.</paragraph>
<paragraph pageId="0" pageNumber="113">
Hosts covered by dense pale yellow mycelium (162D). Sexual morph:
<emphasis italics="true" pageId="0" pageNumber="113">Stromata</emphasis>
stipitate, arising from the host, multiple, cylindrical at the base, slightly enlarged midway to the terminal stipe below the fertile head, moderate yellow (162A-B), 8-12 mm long, 1-3 mm broad.
<emphasis italics="true" pageId="0" pageNumber="113">Fertile head</emphasis>
disc-shaped, upper surface slightly convex, 3-4
<normalizedToken originalValue="×">x</normalizedToken>
2-3.5 mm.
<emphasis italics="true" pageId="0" pageNumber="113">Perithecia</emphasis>
completely immersed, narrowly ovoid, 1000-1400
<normalizedToken originalValue="×">x</normalizedToken>
200-350
<normalizedToken originalValue="µm">µm</normalizedToken>
, ostiole pale brownish-orange (165B).
<emphasis italics="true" pageId="0" pageNumber="113">Asci</emphasis>
cylindrical, 8-spored, 400-1000
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 3.5-6
<normalizedToken originalValue="µm">µm</normalizedToken>
broad, with cap 2-5
<normalizedToken originalValue="µm">µm</normalizedToken>
thick.
<emphasis italics="true" pageId="0" pageNumber="113">Ascospores</emphasis>
hyaline, dissociating into 128 part-spores, cylindrical, 2-6
<normalizedToken originalValue="×">x</normalizedToken>
1-3
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="description">
<paragraph pageId="0" pageNumber="113">Culture characteristics.</paragraph>
<paragraph pageId="0" pageNumber="113">
Colonies on OA attaining a diam. of 8-10 mm in 21 d, mycelium sparse, white, reverse pale yellow (161C).
<emphasis italics="true" pageId="0" pageNumber="113">Conidia</emphasis>
forming on vegetative hyphae or by microcyclic conidiation, hyaline, clavate to cylindrical, 2-10
<normalizedToken originalValue="×">x</normalizedToken>
1-5
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="113">
Colonies on PDA attaining a diam. of 8-10 mm in 20 d, mycelium sparse, white, reverse pale yellow (161C).
<emphasis italics="true" pageId="0" pageNumber="113">Conidia</emphasis>
forming on vegetative hyphae or by microcyclic conidiation, hyaline, clavate to cylindrical, 2-12
<normalizedToken originalValue="×">x</normalizedToken>
1-5
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="host">
<paragraph pageId="0" pageNumber="113">Host.</paragraph>
<paragraph pageId="0" pageNumber="113">Spider egg sac.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="habitat">
<paragraph pageId="0" pageNumber="113">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="113">Specimen was found on the underside of a dicot leaf of a forest plant.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="113" type="notes">
<paragraph pageId="0" pageNumber="113">Notes.</paragraph>
<paragraph pageId="0" pageNumber="113">
Based on natural specimens,
<taxonomicName class="Sordariomycetes" family="Cordycipitaceae" genus="Po." kingdom="Fungi" lsidName="Po. araneae" order="Hypocreales" pageId="0" pageNumber="113" phylum="Ascomycota" rank="species" species="araneae">
<emphasis italics="true" pageId="0" pageNumber="113">Po. araneae</emphasis>
</taxonomicName>
closely resembles
<taxonomicName class="Arachnida" genus="H." kingdom="Animalia" lsidName="H. nelumboides" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="nelumboides">
<emphasis italics="true" pageId="0" pageNumber="113">H. nelumboides</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" genus="H." kingdom="Animalia" lsidName="H. novoguineensis" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="0" pageNumber="113">H. novoguineensis</emphasis>
</taxonomicName>
by producing fertile heads at the end of the stipes. The perithecia of these species are completely immersed. The ascospores of
<taxonomicName class="Sordariomycetes" family="Cordycipitaceae" genus="Po." kingdom="Fungi" lsidName="Po. araneae" order="Hypocreales" pageId="0" pageNumber="113" phylum="Ascomycota" rank="species" species="araneae">
<emphasis italics="true" pageId="0" pageNumber="113">Po. araneae</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" genus="H." kingdom="Animalia" lsidName="H. nelumboides" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="nelumboides">
<emphasis italics="true" pageId="0" pageNumber="113">H. nelumboides</emphasis>
</taxonomicName>
are filamentous, multiseptate ascospores disarticulating into part-spores, whereas
<taxonomicName class="Arachnida" genus="H." kingdom="Animalia" lsidName="H. novoguineensis" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="novoguineensis">
<emphasis italics="true" pageId="0" pageNumber="113">H. novoguineensis</emphasis>
</taxonomicName>
produces filiform, whole ascospores. However,
<taxonomicName class="Sordariomycetes" family="Cordycipitaceae" genus="Po." kingdom="Fungi" lsidName="Po. araneae" order="Hypocreales" pageId="0" pageNumber="113" phylum="Ascomycota" rank="species" species="araneae">
<emphasis italics="true" pageId="0" pageNumber="113">Po. araneae</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Arachnida" genus="H." kingdom="Animalia" lsidName="H. nelumboides" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="nelumboides">
<emphasis italics="true" pageId="0" pageNumber="113">H. nelumboides</emphasis>
</taxonomicName>
in the size of the perithecia. In
<taxonomicName class="Sordariomycetes" family="Cordycipitaceae" genus="Po." kingdom="Fungi" lsidName="Po. araneae" order="Hypocreales" pageId="0" pageNumber="113" phylum="Ascomycota" rank="species" species="araneae">
<emphasis italics="true" pageId="0" pageNumber="113">Po. araneae</emphasis>
</taxonomicName>
, perithecia are larger than those reported for
<taxonomicName class="Arachnida" genus="H." kingdom="Animalia" lsidName="H. nelumboides" order="Araneae" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="nelumboides">
<emphasis italics="true" pageId="0" pageNumber="113">H. nelumboides</emphasis>
</taxonomicName>
(1000-1400
<normalizedToken originalValue="×">x</normalizedToken>
200-350
<normalizedToken originalValue="µm">µm</normalizedToken>
vs. 535-545
<normalizedToken originalValue="×">x</normalizedToken>
180-190
<normalizedToken originalValue="µm">µm</normalizedToken>
) (Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological comparisons of sexual morphs in Hevansia, Jenniferia and Polystromomyces." httpUri="http://table.plazi.org/id/649844D1D32BE234FC567534FADE3ABC" pageId="0" pageNumber="113" tableUuid="649844D1D32BE234FC567534FADE3ABC">2</tableCitation>
).
<taxonomicName authorityName="Mongkolsamrit, Noisripoom, Sakolrak &amp; Himaman" authorityYear="2022" class="Arachnida" family="Cordycipitaceae" genus="Polystromomyces" kingdom="Fungi" lsidName="Polystromomyces araneae" order="Hypocreales" pageId="0" pageNumber="113" rank="species" species="araneae">
<emphasis italics="true" pageId="0" pageNumber="113">Polystromomyces araneae</emphasis>
</taxonomicName>
produces microcycle conidiation from conidia on culture, while the microcyclic sporulation is often seen in discharged ascospores in
<taxonomicName authorityName="Luangsa-ard, Mongkolsamrit, Himaman, Thanakitpipattana &amp; Samson" authorityYear="2020" class="Sordariomycetes" family="Clavicipitaceae" genus="Metarhizium" kingdom="Fungi" lsidName="Metarhizium phuwiangense" order="Hypocreales" pageId="0" pageNumber="113" phylum="Ascomycota" rank="species" species="phuwiangense">
<emphasis italics="true" pageId="0" pageNumber="113">Metarhizium phuwiangense</emphasis>
</taxonomicName>
Luangsa-ard, Mongkols., Himaman, Thanakitp. &amp; Samson and
<taxonomicName authorityName="Luangsa-ard, Samson &amp; Thanakitpipattana" authorityYear="2020" baseAuthorityName="Hywel-Jones" class="Sordariomycetes" family="Clavicipitaceae" genus="Purpureomyces" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="Purpureomyces khaoyaiensis" order="Hypocreales" pageId="0" pageNumber="113" phylum="Ascomycota" rank="species" species="khaoyaiensis">
<emphasis italics="true" pageId="0" pageNumber="113">Purpureomyces khaoyaiensis</emphasis>
</taxonomicName>
(Hywel-Jones) Luangsa-ard, Samson &amp; Thanakitp (
<bibRefCitation DOI="https://doi.org/10.1016/j.simyco.2020.04.001" author="Mongkolsamrit, S" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="113" pagination="171 - 251" refId="B32" refString="Mongkolsamrit, S, Khonsanit, A, Thanakitpipattana, D, Tasanathai, K, Noisripoom, W, Lamlertthon, S, Himaman, W, Houbraken, J, Samson, RA, Luangsa-ard, J, 2020a. Revisiting Metarhizium and the description of new species from Thailand. Studies in Mycology 95: 171 - 251, DOI: https://doi.org/10.1016/j.simyco.2020.04.001" title="Revisiting Metarhizium and the description of new species from Thailand." url="https://doi.org/10.1016/j.simyco.2020.04.001" volume="95" year="2020 a">Mongkolsamrit et al. 2020a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>