205 lines
21 KiB
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205 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.592.8118" ID-GBIF-Dataset="8aa9b0f8-eb42-4d38-8e74-cd16596e22cf" ID-PMC="PMC4926634" ID-Pensoft-Pub="1313-2970-592-1" ID-PubMed="27408542" ID-ZBK="08A7D4E2B2C84D88822930ECE9793CB2" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-592-1" ModsDocOrigin="ZooKeys 592" ModsDocTitle="The family Plectopylidae (Gastropoda, Pulmonata) in Laos with the description of two new genera and a new species" checkinTime="1464247648205" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Muratov, Igor V. & Asami, Takahiro" docDate="2016" docId="21B4EA7327949CD2D6164EA47F8A5991" docLanguage="en" docName="ZooKeys 592: 1-26" docOrigin="ZooKeys 592" docSource="http://dx.doi.org/10.3897/zookeys.592.8118" docTitle="Naggsia Pall-Gergely & Muratov, gen. n." docType="treatment" docUuid="D8CDB123-6382-462E-A66E-ABFD7C838CA3" docUuidSource="ZooBank" docVersion="6" lastPageNumber="12" masterDocId="FFCC670A3014C95E8216FFF4FF9AFFD5" masterDocTitle="The family Plectopylidae (Gastropoda, Pulmonata) in Laos with the description of two new genera and a new species" masterLastPageNumber="26" masterPageNumber="1" pageNumber="11" updateTime="1668163127593" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The family Plectopylidae (Gastropoda, Pulmonata) in Laos with the description of two new genera and a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Muratov, Igor V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Asami, Takahiro</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>592</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>26</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.592.8118</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.592.8118</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-592-1</mods:identifier>
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<mods:identifier type="ZBK">08A7D4E2B2C84D88822930ECE9793CB2</mods:identifier>
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<mods:identifier type="ZooBank">08A7D4E2B2C84D88822930ECE9793CB2</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127881998" LSID="urn:lsid:zoobank.org:act:D8CDB123-6382-462E-A66E-ABFD7C838CA3" httpUri="http://treatment.plazi.org/id/21B4EA7327949CD2D6164EA47F8A5991" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<subSubSection pageId="10" pageNumber="11" type="multiple">
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<paragraph pageId="10" pageNumber="11">Taxon classification Animalia Pulmonata Plectopylidae</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="nomenclature">
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName LSID="http://zoobank.org/D8CDB123-6382-462E-A66E-ABFD7C838CA3" authority="Pall-Gergely & Muratov" class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
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Naggsia
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Muratov
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</taxonomicName>
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<taxonomicNameLabel pageId="10" pageNumber="11">gen. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="type species">
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<paragraph pageId="10" pageNumber="11">Type species.</paragraph>
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix laomontana" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="laomontana">Helix laomontana</taxonomicName>
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L. Pfeiffer, 1862.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="content">
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<paragraph pageId="10" pageNumber="11">Content.</paragraph>
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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(L. Pfeiffer, 1862).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph pageId="10" pageNumber="11">Diagnosis.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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Shell flat, widely umbilicated, body whorl rounded; protoconch with dense, regular, slightly waved ribs with extremely fine spiral striation. Epiphallus ab
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<pageBreakToken pageId="11" pageNumber="12" start="start">sent</pageBreakToken>
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, diverticulum and gametolytic sac are both very short, but diverticulum is still shorter than the gametolytic sac. Cusp of central tooth missing, only basal plate of central tooth present. Marginals bicuspid.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="diagnosis">
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<paragraph pageId="11" pageNumber="12">Differential diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Naggsia</taxonomicName>
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gen. n. differs from the genera having ribbed embryonic whorls (
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Endothyrella</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Gudeodiscus</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Halongella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halongella" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Halongella</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sicradiscus</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Sinicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinicola" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sinicola</taxonomicName>
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) by the absence of an epiphallus and the presence of a very short diverticulum. Moreover, although the protoconch of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Naggsia</taxonomicName>
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gen. n. is ribbed, the ribs (radial lines) are not straight, as in the other genera, but are somewhat wavy.
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Naggsia</taxonomicName>
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gen. n. differs from the genera without a ribbed protoconch (
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chersaecia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Chersaecia</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endoplon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Endoplon" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Endoplon</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Plectopylis</taxonomicName>
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) by the presence of regular, slightly waved ribs on the embryonic whorls. The latter genera are insufficiently known anatomically.
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis bensoni" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="bensoni">Plectopylis bensoni</taxonomicName>
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Gude, 1914 (mentioned as
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis achatina" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="achatina">Plectopylis achatina</taxonomicName>
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Pfeiffer, 1845) and
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis cyclaspis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="cyclaspis">Plectopylis cyclaspis</taxonomicName>
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Benson, 1859 have a well-developed epiphallus (
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<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="23" pageNumber="24" pagination="217 - 259" title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species." url="http://biodiversitylibrary.org/page/35631081" volume="40" year="1871">Stoliczka 1871</bibRefCitation>
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). Interestingly, these
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Plectopylis</taxonomicName>
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species have a relatively short and thickened diverticulum, which is somewhat similar to that of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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. Similarly to
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chersaecia simplex" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="simplex">Chersaecia simplex</taxonomicName>
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Solem, 1966 lacks the epiphallus (see
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<bibRefCitation author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="23" pageNumber="24" pagination="1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
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), but it at has no diverticulum, which is well developed in
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Naggsia</taxonomicName>
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. See differential diagnosis under
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Hunyadiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hunyadiscus" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Hunyadiscus</taxonomicName>
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and Table 1.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="etymology">
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<paragraph pageId="11" pageNumber="12">Etymology.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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The new genus is dedicated to Fred Naggs (NHM) in acknowledgement of his help with our studies on the
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<taxonomicName family="Plectopylidae" lsidName="" pageId="11" pageNumber="12" rank="family">Plectopylidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="distribution">
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<paragraph pageId="11" pageNumber="12">Distribution.</paragraph>
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<paragraph pageId="11" pageNumber="12">Northern Laos (Figure 15).</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="remarks">
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<paragraph pageId="11" pageNumber="12">Remarks.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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The anatomy of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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is rather similar to that of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Plectopylis</taxonomicName>
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in the short diverticulum and to
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Chersaecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chersaecia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Chersaecia</taxonomicName>
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in the absence of epiphallus. The shell characters, namely the ribbed protoconch and the reduced parietal plication is similar to the genera
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Endothyrella</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Gudeodiscus</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Halongella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halongella" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Halongella</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sicradiscus</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Sinicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinicola" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sinicola</taxonomicName>
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. The radula of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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shows similarities with those of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus (Veludiscus)" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="subGenus" subGenus="Veludiscus">Gudeodiscus (Veludiscus)</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Halongella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halongella" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Halongella</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Plectopylis</taxonomicName>
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(see
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<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="23" pageNumber="24" pagination="217 - 259" title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species." url="http://biodiversitylibrary.org/page/35631081" volume="40" year="1871">Stoliczka 1871</bibRefCitation>
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,
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="22" pageNumber="23" pagination="1 - 118" title="Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)." url="10.3897/zookeys.473.8659" volume="473" year="2015 a">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2015a
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</bibRefCitation>
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) in terms of the small central tooth and the simple marginals. In contrast, the central tooth of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Endothyrella</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus (Gudeodiscus)" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="subGenus" subGenus="Gudeodiscus">Gudeodiscus (Gudeodiscus)</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sicradiscus</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Sinicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinicola" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sinicola</taxonomicName>
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species is large (as large as or larger than the ectocones of the first laterals), (
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="22" pageNumber="23" pagination="1 - 118" title="Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)." url="10.3897/zookeys.473.8659" volume="473" year="2015 a">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2015a
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</bibRefCitation>
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,
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="22" pageNumber="23" pagination="1 - 70" title="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)." url="10.3897/zookeys.529.6139" volume="529" year="2015 b">2015b</bibRefCitation>
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), and the marginals are tricuspid or even quadricuspid. Before examining ethanol-preserved
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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specimens, we considered grouping the plectopylid genera into two tribes, namely one with a ribbed, and another with a smooth or granulated protoconch. The examination of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia laomontana" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laomontana">Naggsia laomontana</taxonomicName>
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revealed that the character states considered to be primarily important do not allow the placement of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Naggsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naggsia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Naggsia</taxonomicName>
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gen. n. in any of the two groups, and that the character states rather show a mosaic structure across genera.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |