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<subSubSection id="E1F56531174C9D50FF8349F9A01FF9DF" box="[98,638,1509,1537]" pageId="1" pageNumber="72" type="nomenclature">
<paragraph id="A95036BA174C9D50FF8349F9A01FF9DF" blockId="1.[98,638,1509,1537]" box="[98,638,1509,1537]" pageId="1" pageNumber="72">
<heading id="F21881D6174C9D50FF8349F9A01FF9DF" box="[98,638,1509,1537]" fontSize="12" level="2" pageId="1" pageNumber="72" reason="2">
Family
<taxonomicName id="6EEF4D39174C9D50FF2349F9A01FF9DF" ID-CoL="6HF" authority="Handlirsch, 1922" authorityName="Handlirsch" authorityYear="1922" box="[194,638,1509,1537]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Anthracoptilidae
<bibRefCitation id="CD7E4B4B174C9D50FE4349F9A01FF9DF" author="Handlirsch, A." box="[418,638,1509,1537]" pageId="1" pageNumber="72" refId="ref13153" refString="Handlirsch, A. 1922. Animalia Insecta Palaeozoica. Fossilium Catalogus. 230 pp. W. Junk, Berlin." type="book" year="1922">Handlirsch, 1922</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="E1F56531174C9D50FF834A0DA0BAF99C" pageId="1" pageNumber="72" type="reference_group">
<paragraph id="A95036BA174C9D50FF834A0DA0C8F9F8" blockId="1.[98,731,1553,1602]" box="[98,681,1553,1574]" pageId="1" pageNumber="72">
2002
<taxonomicName id="6EEF4D39174C9D50FF794A0DA3B5F9F8" authority="Martynov, 1938" authorityName="Martynov" authorityYear="1938" box="[152,468,1553,1574]" class="Insecta" family="Strephocladidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Strephocladidae
<bibRefCitation id="CD7E4B4B174C9D50FED94A0DA3B5F9F8" author="Martynov, A. V." box="[312,468,1553,1574]" pageId="1" pageNumber="72" pagination="1 - 149" refId="ref13588" refString="Martynov, A. V. 1938. Etude sur l'histoire geologique et de phylogenie des ordres des insectes (Pterygota). Premiere partie. Palaeoptera et Neoptera - Polyneoptera [in Russian, with French summary]. Trudy Paleontologiceskogo Instituta Akademii nauk SSSR, Moskva 7 (4): 1 - 149." type="journal article" year="1938">Martynov, 1938</bibRefCitation>
</taxonomicName>
; Rasnitsyn 2002: 112.
</paragraph>
<paragraph id="A95036BA174C9D50FF834A31A0BAF99C" blockId="1.[98,731,1553,1602]" box="[98,731,1581,1602]" pageId="1" pageNumber="72">
2002
<taxonomicName id="6EEF4D39174C9D50FF794A31A3B6F99C" authority="Martynov, 1940" authorityName="Martynov" authorityYear="1940" box="[152,471,1581,1602]" class="Insecta" family="Strephoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Strephoneuridae
<bibRefCitation id="CD7E4B4B174C9D50FEDA4A31A3B6F99C" author="Martynov, A. V." box="[315,471,1581,1602]" pageId="1" pageNumber="72" pagination="1 - 62" refId="ref13645" refString="Martynov, A. V. 1940. Permian fossil insects from Tshekarda [in Russian, with English summary]. Trudy Paleontologiceskogo Instituta Akademii nauk SSSR 11: 1 - 62." type="journal article" year="1940">Martynov, 1940</bibRefCitation>
</taxonomicName>
; Rasnitsyn 2002: 112, 114.
</paragraph>
</subSubSection>
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<paragraph id="A95036BA174C9D50FF834A52A0EBF945" blockId="1.[97,771,1614,1691]" pageId="1" pageNumber="72">
<emphasis id="9B9BEAA8174C9D50FF834A52A2ADF9BD" box="[98,204,1614,1635]" italics="true" pageId="1" pageNumber="72">
<typeStatus id="76548818174C9D50FF834A52A2EEF9BD" box="[98,143,1614,1635]" pageId="1" pageNumber="72">Type</typeStatus>
genus
</emphasis>
:
<taxonomicName id="6EEF4D39174C9D50FF394A52A39DF9BD" authority="Lameere, 1917" authorityName="Lameere" authorityYear="1917" box="[216,508,1614,1635]" class="Insecta" genus="Anthracoptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FF394A52A306F9BD" box="[216,359,1614,1635]" italics="true" pageId="1" pageNumber="72">Anthracoptilus</emphasis>
<bibRefCitation id="CD7E4B4B174C9D50FE8C4A52A39DF9BD" author="Lameere, A." box="[365,508,1614,1635]" pageId="1" pageNumber="72" pagination="141 - 200" refId="ref13452" refString="Lameere, A. 1917. Revision sommaire des Insectes fossiles du Stephanien de Commentry. Bulletin du Museum National d'Histoire Naturelle, Paris 23: 141 - 200." type="journal article" year="1917">Lameere, 1917</bibRefCitation>
</taxonomicName>
for
<taxonomicName id="6EEF4D39174C9D50FDC24A52A298F9A1" authority="Meunier, 1909" authorityName="Meunier" authorityYear="1909" class="Insecta" genus="Anthracoptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="9B9BEAA8174C9D50FDC24A52A162F9BD" box="[547,771,1614,1635]" italics="true" pageId="1" pageNumber="72">Anthracoptilus perrieri</emphasis>
<bibRefCitation id="CD7E4B4B174C9D50FF894A76A298F9A1" author="Meunier, F." box="[104,249,1642,1663]" pageId="1" pageNumber="72" pagination="37 - 49" refId="ref13679" refString="Meunier, F. 1909. Nouveaux insectes du Stephanien de Commentry. Bulletin du Museum National d'Histoire Naturelle 15: 37 - 49." type="journal article" year="1909">Meunier, 1909</bibRefCitation>
</taxonomicName>
); Stephanian, Carboniferous (= Gzhelian, Upper Pennsylvanian), Commentry Basin, Allier department,
<collectingCountry id="D1F8762A174C9D50FDA04A9AA0E4F945" box="[577,645,1670,1691]" name="France" pageId="1" pageNumber="72">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E1F56531174C9D50FF834AB3A1EEFE67" pageId="1" pageNumber="72" type="diagnosis">
<paragraph id="A95036BA174C9D50FF834AB3A1EEFE67" blockId="1.[98,773,1711,1960]" lastBlockId="1.[808,1482,160,441]" pageId="1" pageNumber="72">
<emphasis id="9B9BEAA8174C9D50FF834AB3A35DF916" box="[98,316,1711,1736]" italics="true" pageId="1" pageNumber="72">Emended diagnosis</emphasis>
.—Area between C and
<collectionCode id="CFFEAE7F174C9D50FDA64AB3A013F916" box="[583,626,1711,1736]" pageId="1" pageNumber="72">ScP</collectionCode>
with a series of veinlets and crossveins (but probably not preserved in
<taxonomicName id="6EEF4D39174C9D50FF834AF3A3E6F8D6" authorityName="Guan &amp; Prokop &amp; Roques &amp; Lapeyrie &amp; Nel" authorityYear="2016" box="[98,391,1775,1800]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="permianus">
<emphasis id="9B9BEAA8174C9D50FF834AF3A3E6F8D6" box="[98,391,1775,1800]" italics="true" pageId="1" pageNumber="72">Strephocladus permianus</emphasis>
</taxonomicName>
); RA with one or two posterior branches (an apomorphic character, but not shared by all taxa in
<taxonomicName id="6EEF4D39174C9D50FF5C4B33A31BF896" authorityName="Handlirsch" authorityYear="1922" box="[189,378,1839,1864]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
) or simple; median vein strongly approximate or shortly fused with
<collectionCode id="CFFEAE7F174C9D50FE064B53A06BF8B6" box="[487,522,1871,1896]" pageId="1" pageNumber="72">RP</collectionCode>
near base of
<collectionCode id="CFFEAE7F174C9D50FD7E4B53A0A3F8B6" box="[671,706,1871,1896]" pageId="1" pageNumber="72">RP</collectionCode>
(synapomorphy); branches of
<collectionCode id="CFFEAE7F174C9D50FE9F4B73A3C0F856" box="[382,417,1903,1928]" pageId="1" pageNumber="72">RP</collectionCode>
, median vein and CuA strongly parallel at least in their basal halves(the presence of branches all simple is not shared by all taxa in
<taxonomicName id="6EEF4D39174C9D50FB1B4CBCA7DFFF67" authorityName="Handlirsch" authorityYear="1922" box="[1274,1470,160,185]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
); median vein braced to CuA by a short convex cross-vein mp-cua (a character present in Holometabola,
<taxonomicName id="6EEF4D39174C9D50FADA4CFCA7F6FF27" authorityName="Handlirsch" authorityYear="1906" box="[1339,1431,224,249]" class="Insecta" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="order">Paoliida</taxonomicName>
and Dictyoptera, but very useful to separate the
<taxonomicName id="6EEF4D39174C9D50FAE94D1CA7A9FEC7" authorityName="Handlirsch" authorityYear="1922" box="[1288,1480,256,281]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
from the archaeorthopteran taxa); CuA and
<collectionCode id="CFFEAE7F174C9D50FAC54D3CA734FEE7" box="[1316,1365,288,313]" pageId="1" pageNumber="72">CuP</collectionCode>
emerging from a long common stem; main stem of CuA strongly convex, but anteriorly pectinate with at least a series of parallel branches, all less convex than main stem of CuA (synapomorphy).
</paragraph>
</subSubSection>
<subSubSection id="E1F56531174C9D53FCC94DD7A052F876" lastPageId="2" lastPageNumber="73" pageId="1" pageNumber="72" type="discussion">
<paragraph id="A95036BA174C9D50FCC94DD7A776FC5B" blockId="1.[808,1482,459,1960]" pageId="1" pageNumber="72">
<emphasis id="9B9BEAA8174C9D50FCC94DD7A1EAFE3A" box="[808,907,459,484]" italics="true" pageId="1" pageNumber="72">Remarks</emphasis>
.—The family
<taxonomicName id="6EEF4D39174C9D50FBA44DD7A7A8FE3A" authority="Martynov, 1938" authorityName="Martynov" authorityYear="1938" box="[1093,1481,459,484]" class="Insecta" family="Strephocladidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Strephocladidae
<bibRefCitation id="CD7E4B4B174C9D50FAED4DD7A7A8FE3A" author="Martynov, A. V." box="[1292,1481,459,484]" pageId="1" pageNumber="72" pagination="1 - 149" refId="ref13588" refString="Martynov, A. V. 1938. Etude sur l'histoire geologique et de phylogenie des ordres des insectes (Pterygota). Premiere partie. Palaeoptera et Neoptera - Polyneoptera [in Russian, with French summary]. Trudy Paleontologiceskogo Instituta Akademii nauk SSSR, Moskva 7 (4): 1 - 149." type="journal article" year="1938">Martynov, 1938</bibRefCitation>
</taxonomicName>
(
<typeStatus id="76548818174C9D50FCD14DF7A103FDDA" box="[816,866,491,516]" pageId="1" pageNumber="72">type</typeStatus>
genus
<taxonomicName id="6EEF4D39174C9D50FC2C4DF7A746FDDA" authority="Scudder, 1885" authorityName="Scudder" authorityYear="1885" box="[973,1319,491,516]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FC2C4DF7A611FDDA" box="[973,1136,491,516]" italics="true" pageId="1" pageNumber="72">Strephocladus</emphasis>
<bibRefCitation id="CD7E4B4B174C9D50FB654DF7A746FDDA" author="Scudder, S. H." box="[1156,1319,491,516]" pageId="1" pageNumber="72" pagination="721 - 831" refId="ref14607" refString="Scudder, S. H. 1885. Systematische Ubersicht der fossilen Myriapoden, Arachnoiden und Insekten. In: K. A. Zittel (ed.), Handbuch der Palaontologie, 1 Abtheilung, Palaozoologie 2, 721 - 831. Oldenbourg, Munchen." type="journal article" year="1885">Scudder, 1885</bibRefCitation>
</taxonomicName>
) is currently considered as a junior synonym of the family
<taxonomicName id="6EEF4D39174C9D50FA914E17A612FD9A" authority="Handlirsch, 1922" authorityName="Handlirsch" authorityYear="1922" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Anthracoptilidae
<bibRefCitation id="CD7E4B4B174C9D50FC474E37A612FD9A" author="Handlirsch, A." box="[934,1139,555,580]" pageId="1" pageNumber="72" refId="ref13153" refString="Handlirsch, A. 1922. Animalia Insecta Palaeozoica. Fossilium Catalogus. 230 pp. W. Junk, Berlin." type="book" year="1922">Handlirsch, 1922</bibRefCitation>
</taxonomicName>
, also synonymized with the
<taxonomicName id="6EEF4D39174C9D50FCC94E57A6D6FDBA" authority="Martynov, 1931" authorityName="Martynov" authorityYear="1931" box="[808,1207,587,612]" class="Insecta" family="Permarrhaphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Permarrhaphidae
<bibRefCitation id="CD7E4B4B174C9D50FC1D4E57A6D6FDBA" author="Martynov, A. V." box="[1020,1207,587,612]" pageId="1" pageNumber="72" pagination="149 - 212" refId="ref13520" refString="Martynov, A. V. 1931. New fossil insects from Tikhie Gory. 2. Neoptera (excluding Miomoptera) [in Russian]. Trudy Geologiceskogo Muzea, Akademii nauk SSSR 8: 149 - 212." type="journal article" year="1931">Martynov, 1931</bibRefCitation>
</taxonomicName>
(late Permian, monotypic genus
<taxonomicName id="6EEF4D39174C9D50FC284E70A731FD5B" authority="Martynov, 1931" authorityName="Martynov" authorityYear="1931" box="[969,1360,620,645]" class="Insecta" family="Permarrhaphidae" genus="Permarrhaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FC284E70A61FFD5B" box="[969,1150,620,645]" italics="true" pageId="1" pageNumber="72">Permarrhaphus</emphasis>
<bibRefCitation id="CD7E4B4B174C9D50FB704E70A731FD5B" author="Martynov, A. V." box="[1169,1360,620,645]" pageId="1" pageNumber="72" pagination="149 - 212" refId="ref13520" refString="Martynov, A. V. 1931. New fossil insects from Tikhie Gory. 2. Neoptera (excluding Miomoptera) [in Russian]. Trudy Geologiceskogo Muzea, Akademii nauk SSSR 8: 149 - 212." type="journal article" year="1931">Martynov, 1931</bibRefCitation>
</taxonomicName>
) and the
<taxonomicName id="6EEF4D39174C9D50FCC94E90A6FBFD7B" authority="Martynov, 1940" authorityName="Martynov" authorityYear="1940" box="[808,1178,652,677]" class="Insecta" family="Strephoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Strephoneuridae
<bibRefCitation id="CD7E4B4B174C9D50FC064E90A6FBFD7B" author="Martynov, A. V." box="[999,1178,652,677]" pageId="1" pageNumber="72" pagination="1 - 62" refId="ref13645" refString="Martynov, A. V. 1940. Permian fossil insects from Tshekarda [in Russian, with English summary]. Trudy Paleontologiceskogo Instituta Akademii nauk SSSR 11: 1 - 62." type="journal article" year="1940">Martynov, 1940</bibRefCitation>
</taxonomicName>
(early Permian, monotypic genus
<taxonomicName id="6EEF4D39174C9D50FC9B4EB0A684FD1B" authority="Martynov, 1940" authorityName="Martynov" authorityYear="1940" box="[890,1253,684,709]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FC9B4EB0A677FD1B" box="[890,1046,684,709]" italics="true" pageId="1" pageNumber="72">Strephoneura</emphasis>
<bibRefCitation id="CD7E4B4B174C9D50FBC74EB0A684FD1B" author="Martynov, A. V." box="[1062,1253,684,709]" pageId="1" pageNumber="72" pagination="1 - 62" refId="ref13645" refString="Martynov, A. V. 1940. Permian fossil insects from Tshekarda [in Russian, with English summary]. Trudy Paleontologiceskogo Instituta Akademii nauk SSSR 11: 1 - 62." type="journal article" year="1940">Martynov, 1940</bibRefCitation>
</taxonomicName>
) (Rasnitsyn 2002;
<bibRefCitation id="CD7E4B4B174C9D50FCC94ED0A6E6FD3B" author="Novokshonov, V. G. &amp; Aristov, D. S." box="[808,1159,716,741]" pageId="1" pageNumber="72" pagination="60 - 66" refId="ref13844" refString="Novokshonov, V. G. and Aristov, D. S. 2004. New taxa of hypoperlids (Insecta: Hypoperlida) from the Upper Permian of the Arkhangelsk Region. Paleontological Journal 38: 60 - 66." type="journal article" year="2004">Novokshonov and Aristov 2004</bibRefCitation>
; Rasnitsyn and Aristov 2004; Rasnitsyn et al. 2004). Lastly
<bibRefCitation id="CD7E4B4B174C9D50FB9F4EF0A7A9FCDB" author="Rasnitsyn, A. P. &amp; Aristov, D. S." box="[1150,1480,748,773]" pageId="1" pageNumber="72" pagination="678 - 703" refId="ref14281" refString="Rasnitsyn, A. P. and Aristov, D. S. 2013. New fossil insects (Insecta: Caloneurida, Hypoperlida, Palaeomanteida, Jurinida) from the Middle and Upper Permian of European Russia. In: D. S. Aristov, A. S. Bashkuev, V. K. Golubev, A. V. Gorochov, E. V. Karasev, D. S. Kopylov, A. G. Ponomarenko, A. P. Rasnitsyn, D. A. Rasnitsyn, N. D. Sinitshenkova, I. D. Sukatsheva, and D. V. Vassilenko (eds.), Fossil Insects of the Middle and Upper Permian of European Russia. Paleontological Journal 47: 678 - 703." type="journal article" year="2013">Rasnitsyn and Aristov (2013)</bibRefCitation>
proposed again a synonymy of the
<taxonomicName id="6EEF4D39174C9D50FB2B4F10A7EAFCFB" authorityName="Martynov" authorityYear="1931" box="[1226,1419,780,805]" class="Insecta" family="Permarrhaphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Permarrhaphidae</taxonomicName>
with the
<taxonomicName id="6EEF4D39174C9D50FCBD4F30A641FC9B" authorityName="Handlirsch" authorityYear="1922" box="[860,1056,812,837]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
,
<taxonomicName id="6EEF4D39174C9D50FBD74F30A695FC9B" authorityName="Martynov" authorityYear="1940" box="[1078,1268,812,837]" class="Insecta" family="Strephoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Strephoneuridae</taxonomicName>
,
<taxonomicName id="6EEF4D39174C9D50FAEB4F30A7A5FC9B" authorityName="Martynov" authorityYear="1938" box="[1290,1476,812,837]" class="Insecta" family="Strephocladidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Strephocladidae</taxonomicName>
, but also with the
<taxonomicName id="6EEF4D39174C9D50FC124F50A70AFCBB" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[1011,1387,844,869]" class="Insecta" family="Ischnoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Ischnoneuridae
<bibRefCitation id="CD7E4B4B174C9D50FB494F50A70AFCBB" author="Handlirsch, A." box="[1192,1387,844,869]" pageId="1" pageNumber="72" pagination="661 - 820" refId="ref13126" refString="Handlirsch, A. 1906. Revision of American Paleozoic insects. Proceedings of the United States National Museum, Washington 29: 661 - 820." type="journal article" year="1906">Handlirsch, 1906</bibRefCitation>
</taxonomicName>
and the
<taxonomicName id="6EEF4D39174C9D50FCC94F70A6ECFC5B" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[808,1165,876,901]" class="Insecta" family="Adiphlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">
Adiphlebiidae
<bibRefCitation id="CD7E4B4B174C9D50FC2D4F70A6ECFC5B" author="Handlirsch, A." box="[972,1165,876,901]" pageId="1" pageNumber="72" pagination="661 - 820" refId="ref13126" refString="Handlirsch, A. 1906. Revision of American Paleozoic insects. Proceedings of the United States National Museum, Washington 29: 661 - 820." type="journal article" year="1906">Handlirsch, 1906</bibRefCitation>
</taxonomicName>
(see below).
</paragraph>
<paragraph id="A95036BA174C9D50FCAF4F90A19AF9B9" blockId="1.[808,1482,459,1960]" pageId="1" pageNumber="72">
Comparisonoftypegenera
<taxonomicName id="6EEF4D39174C9D50FB9B4F90A774FC7B" authorityName="Martynov" authorityYear="1940" box="[1146,1301,908,933]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FB9B4F90A774FC7B" box="[1146,1301,908,933]" italics="true" pageId="1" pageNumber="72">Strephoneura</emphasis>
</taxonomicName>
,
<taxonomicName id="6EEF4D39174C9D50FAC14F90A7A3FC7B" authorityName="Scudder" authorityYear="1885" box="[1312,1474,908,933]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FAC14F90A7A3FC7B" box="[1312,1474,908,933]" italics="true" pageId="1" pageNumber="72">Strephocladus</emphasis>
</taxonomicName>
,
<taxonomicName id="6EEF4D39174C9D50FCC94FB0A1BAFC1B" authorityName="Martynov" authorityYear="1931" box="[808,987,940,965]" class="Insecta" family="Permarrhaphidae" genus="Permarrhaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FCC94FB0A1BAFC1B" box="[808,987,940,965]" italics="true" pageId="1" pageNumber="72">Permarrhaphus</emphasis>
</taxonomicName>
, and
<taxonomicName id="6EEF4D39174C9D50FBFC4FB0A6A4FC1B" authorityName="Lameere" authorityYear="1917" box="[1053,1221,940,965]" class="Insecta" genus="Anthracoptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FBFC4FB0A6A4FC1B" box="[1053,1221,940,965]" italics="true" pageId="1" pageNumber="72">Anthracoptilus</emphasis>
</taxonomicName>
leads to the following conclusions.
<taxonomicName id="6EEF4D39174C9D50FC5E4FD0A63BFC3B" authorityName="Martynov" authorityYear="1940" box="[959,1114,972,997]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FC5E4FD0A63BFC3B" box="[959,1114,972,997]" italics="true" pageId="1" pageNumber="72">Strephoneura</emphasis>
</taxonomicName>
shares with
<taxonomicName id="6EEF4D39174C9D50FB164FD0A7FBFC3B" authorityName="Scudder" authorityYear="1885" box="[1271,1434,972,997]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FB164FD0A7FBFC3B" box="[1271,1434,972,997]" italics="true" pageId="1" pageNumber="72">Strephocladus</emphasis>
</taxonomicName>
the following potential apomorphies:
<collectionCode id="CFFEAE7F174C9D50FB534FF1A6BCFBD8" box="[1202,1245,1005,1030]" pageId="1" pageNumber="72">ScP</collectionCode>
and radius strongly approximate; RA with two apical posterior branches;
<collectionCode id="CFFEAE7F174C9D50FA964811A7FBFBF8" box="[1399,1434,1037,1062]" pageId="1" pageNumber="72">RP</collectionCode>
and median vein strongly approximate or touching near base of
<collectionCode id="CFFEAE7F174C9D50FCC94851A12FFBB8" box="[808,846,1101,1126]" pageId="1" pageNumber="72">RP</collectionCode>
; CuA with several pectinate anterior branches; branches of
<collectionCode id="CFFEAE7F174C9D50FCA44871A109FB58" box="[837,872,1133,1158]" pageId="1" pageNumber="72">RP</collectionCode>
, median vein, and CuA all simple and closely parallel;
<collectionCode id="CFFEAE7F174C9D50FCC94891A138FB78" box="[808,857,1165,1190]" pageId="1" pageNumber="72">CuP</collectionCode>
and first anal vein simple.
<taxonomicName id="6EEF4D39174C9D50FB4F4891A1B8FB18" authority="Martynov, 1931" authorityName="Martynov" authorityYear="1931" class="Insecta" family="Permarrhaphidae" genus="Permarrhaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="venosus">
<emphasis id="9B9BEAA8174C9D50FB4F4891A7A8FB78" box="[1198,1481,1165,1190]" italics="true" pageId="1" pageNumber="72">Permarrhaphus venosus</emphasis>
<bibRefCitation id="CD7E4B4B174C9D50FCC948B1A1B8FB18" author="Martynov, A. V." box="[808,985,1197,1222]" pageId="1" pageNumber="72" pagination="149 - 212" refId="ref13520" refString="Martynov, A. V. 1931. New fossil insects from Tikhie Gory. 2. Neoptera (excluding Miomoptera) [in Russian]. Trudy Geologiceskogo Muzea, Akademii nauk SSSR 8: 149 - 212." type="journal article" year="1931">Martynov, 1931</bibRefCitation>
</taxonomicName>
is based on a wing fragment that was originally interpreted as a beetle elytron. It cannot correspond to any part of a strephocladid wing, except, maybe the anal area, as supposed by Rasnitsyn and Aristov (2004: 160). Nothing looking like a radius, median or CuA vein can be seen on the original photograph (
<bibRefCitation id="CD7E4B4B174C9D50FBDE4952A691FAB9" author="Martynov, A. V." box="[1087,1264,1358,1383]" pageId="1" pageNumber="72" pagination="149 - 212" refId="ref13520" refString="Martynov, A. V. 1931. New fossil insects from Tikhie Gory. 2. Neoptera (excluding Miomoptera) [in Russian]. Trudy Geologiceskogo Muzea, Akademii nauk SSSR 8: 149 - 212." type="journal article" year="1931">Martynov 1931</bibRefCitation>
: fig. 28). It remains that nothing supports the attribution of this wing to the same clade with
<taxonomicName id="6EEF4D39174C9D50FC4D4992A626FA79" authorityName="Martynov" authorityYear="1940" box="[940,1095,1422,1447]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FC4D4992A626FA79" box="[940,1095,1422,1447]" italics="true" pageId="1" pageNumber="72">Strephoneura</emphasis>
</taxonomicName>
or
<taxonomicName id="6EEF4D39174C9D50FB954992A777FA79" authorityName="Scudder" authorityYear="1885" box="[1140,1302,1422,1447]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FB954992A777FA79" box="[1140,1302,1422,1447]" italics="true" pageId="1" pageNumber="72">Strephocladus</emphasis>
</taxonomicName>
. Therefore, we remove the first synonymy of the
<taxonomicName id="6EEF4D39174C9D50FB4B49B2A704FA19" authorityName="Martynov" authorityYear="1940" box="[1194,1381,1454,1479]" class="Insecta" family="Strephoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Strephoneuridae</taxonomicName>
with the
<taxonomicName id="6EEF4D39174C9D50FCC949D2A18AFA39" authorityName="Martynov" authorityYear="1931" box="[808,1003,1486,1511]" class="Insecta" family="Permarrhaphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Permarrhaphidae</taxonomicName>
proposed by Rasnitsyn (2002: 112114), but we confirm the synonymies of the
<taxonomicName id="6EEF4D39174C9D50FB3F49F2A7F9F9D9" authorityName="Martynov" authorityYear="1940" box="[1246,1432,1518,1543]" class="Insecta" family="Strephoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Strephoneuridae</taxonomicName>
and
<taxonomicName id="6EEF4D39174C9D50FCC94A12A1BFF9F9" authorityName="Martynov" authorityYear="1938" box="[808,990,1550,1575]" class="Insecta" family="Strephocladidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Strephocladidae</taxonomicName>
with the
<taxonomicName id="6EEF4D39174C9D50FBA64A12A76BF9F9" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[1095,1290,1550,1575]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
. We consider the
<taxonomicName id="6EEF4D39174C9D50FCC94A32A18AF999" authorityName="Martynov" authorityYear="1931" box="[808,1003,1582,1607]" class="Insecta" family="Permarrhaphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Permarrhaphidae</taxonomicName>
(and
<taxonomicName id="6EEF4D39174C9D50FBCF4A32A681F999" authorityName="Martynov" authorityYear="1931" box="[1070,1248,1582,1607]" class="Insecta" family="Permarrhaphidae" genus="Permarrhaphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FBCF4A32A681F999" box="[1070,1248,1582,1607]" italics="true" pageId="1" pageNumber="72">Permarrhaphus</emphasis>
</taxonomicName>
) as
<taxonomicName id="6EEF4D39174C9D50FAF24A32A705F999" ID-CoL="H6" box="[1299,1380,1582,1607]" class="Insecta" kingdom="Animalia" pageId="1" pageNumber="72" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
incertae sedis status novus.
</paragraph>
<paragraph id="A95036BA174C9D53FCAF4A72A325FE43" blockId="1.[808,1482,459,1960]" lastBlockId="2.[151,825,160,1851]" lastPageId="2" lastPageNumber="73" pageId="1" pageNumber="72">
<taxonomicName id="6EEF4D39174C9D50FCAF4A72A725F959" authority="(Meunier, 1909)" baseAuthorityName="Meunier" baseAuthorityYear="1909" box="[846,1348,1646,1671]" class="Insecta" genus="Anthracoptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="9B9BEAA8174C9D50FCAF4A72A60DF959" box="[846,1132,1646,1671]" italics="true" pageId="1" pageNumber="72">Anthracoptilus perrieri</emphasis>
(
<bibRefCitation id="CD7E4B4B174C9D50FB644A72A75DF959" author="Meunier, F." box="[1157,1340,1646,1671]" pageId="1" pageNumber="72" pagination="37 - 49" refId="ref13679" refString="Meunier, F. 1909. Nouveaux insectes du Stephanien de Commentry. Bulletin du Museum National d'Histoire Naturelle 15: 37 - 49." type="journal article" year="1909">Meunier, 1909</bibRefCitation>
)
</taxonomicName>
(Gzhelian of Commentry, see redescription below),
<typeStatus id="76548818174C9D50FAD34A92A707F979" box="[1330,1382,1678,1703]" pageId="1" pageNumber="72">type</typeStatus>
species of the
<taxonomicName id="6EEF4D39174C9D50FC9F4AB3A626F916" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[894,1095,1711,1736]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
, shares with
<taxonomicName id="6EEF4D39174C9D50FB134AB3A7F3F916" authorityName="Martynov" authorityYear="1940" box="[1266,1426,1711,1736]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FB134AB3A7F3F916" box="[1266,1426,1711,1736]" italics="true" pageId="1" pageNumber="72">Strephoneura</emphasis>
</taxonomicName>
and
<taxonomicName id="6EEF4D39174C9D50FCC94AD3A1AFF936" authorityName="Scudder" authorityYear="1885" box="[808,974,1743,1768]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FCC94AD3A1AFF936" box="[808,974,1743,1768]" italics="true" pageId="1" pageNumber="72">Strephocladus</emphasis>
</taxonomicName>
: the CuA and
<collectionCode id="CFFEAE7F174C9D50FB634AD3A6D5F936" box="[1154,1204,1743,1768]" pageId="1" pageNumber="72">CuP</collectionCode>
emerging from a common stem; a distinct brace between CuA and median vein; median vein and
<collectionCode id="CFFEAE7F174C9D50FBCE4B13A633F8F6" box="[1071,1106,1807,1832]" pageId="1" pageNumber="72">RP</collectionCode>
approximating near base of
<collectionCode id="CFFEAE7F174C9D50FA7F4B13A7A5F8F6" box="[1438,1476,1807,1832]" pageId="1" pageNumber="72">RP</collectionCode>
; an anteriorly pectinate CuA with branches less convex than main stem of CuA;
<collectionCode id="CFFEAE7F174C9D50FBB34B53A61AF8B6" box="[1106,1147,1871,1896]" pageId="1" pageNumber="72">ScP</collectionCode>
ending on RA; branches of main veins parallel between them but less clearly than in
<taxonomicName id="6EEF4D39174C9D50FCC94B93A1A7F876" authorityName="Martynov" authorityYear="1940" box="[808,966,1935,1960]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FCC94B93A1A7F876" box="[808,966,1935,1960]" italics="true" pageId="1" pageNumber="72">Strephoneura</emphasis>
</taxonomicName>
and
<taxonomicName id="6EEF4D39174C9D50FBEB4B93A6D1F876" authorityName="Scudder" authorityYear="1885" box="[1034,1200,1935,1960]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FBEB4B93A6D1F876" box="[1034,1200,1935,1960]" italics="true" pageId="1" pageNumber="72">Strephocladus</emphasis>
</taxonomicName>
.
<taxonomicName id="6EEF4D39174C9D50FB254B93A711F876" authorityName="Lameere" authorityYear="1917" box="[1220,1392,1935,1960]" class="Insecta" genus="Anthracoptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorthoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174C9D50FB254B93A711F876" box="[1220,1392,1935,1960]" italics="true" pageId="1" pageNumber="72">Anthracoptilus</emphasis>
</taxonomicName>
differs from them in the presence of several branches of
<collectionCode id="CFFEAE7F174F9D53FD3A4CBCA09FFF67" box="[731,766,160,185]" pageId="2" pageNumber="73">RP</collectionCode>
, median vein and CuA forked distally; presence of a net of very irregular sigmoidal crossveins, instead of being straight and defining rectangular cells between main veins; RA ending on anterior wing margin in a distinctly more basal position; RA without any posterior branches in its apical part (difference with
<taxonomicName id="6EEF4D39174F9D53FE684D78A04EFEA3" authorityName="Scudder" authorityYear="1885" box="[393,559,356,381]" class="Insecta" genus="Strephocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174F9D53FE684D78A04EFEA3" box="[393,559,356,381]" italics="true" pageId="2" pageNumber="73">Strephocladus</emphasis>
</taxonomicName>
, character unknown in
<taxonomicName id="6EEF4D39174F9D53FF764D98A355FE43" authorityName="Martynov" authorityYear="1940" box="[151,308,388,413]" class="Insecta" family="Anthracoptilidae" genus="Strephoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174F9D53FF764D98A355FE43" box="[151,308,388,413]" italics="true" pageId="2" pageNumber="73">Strephoneura</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="A95036BA174F9D53FF5C4DB9A0DFFAAC" blockId="2.[151,825,160,1851]" pageId="2" pageNumber="73">
An anteriorly pectinate CuA is convergently present in the Archaeorthoptera of the subclade
<taxonomicName id="6EEF4D39174F9D53FD944DDAA33DFE21" ID-CoL="YT" authority="Handlirsch, 1937" authorityName="Handlirsch" authorityYear="1937" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Cnemidolestodea Handlirsch, 1937</taxonomicName>
(sensu
<bibRefCitation id="CD7E4B4B174F9D53FE594DFAA008FE21" author="Bethoux, O." box="[440,617,486,511]" pageId="2" pageNumber="73" pagination="403 - 408" refId="ref12559" refString="Bethoux, O. 2005 a. Cnemidolestodea (Insecta): an ancient order reinstated. Journal of Systematic Palaeontology 3: 403 - 408." type="journal article" year="2005">Béthoux 2005a</bibRefCitation>
), but the patterns of venation of these orthopteroid insects are completely different from those of the
<taxonomicName id="6EEF4D39174F9D53FE3B4E34A0FCFD9F" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[474,669,552,577]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
, due to presence of basal fusion of CuA with M in the former and to a common stem Cu from which CuA and
<collectionCode id="CFFEAE7F174F9D53FD4B4E75A0BAFD5C" box="[682,731,617,642]" pageId="2" pageNumber="73">CuP</collectionCode>
emerge in the latter. Furthermore, the anterior branches of CuA (+CuPa) are clearly concave in the
<taxonomicName id="6EEF4D39174F9D53FD8B4EB6A155FD1D" ID-CoL="YT" authorityName="Handlirsch" authorityYear="1937" box="[618,820,682,707]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Cnemidolestodea</taxonomicName>
, while the same branches are relatively more convex in the
<taxonomicName id="6EEF4D39174F9D53FF764EF0A33DFCDB" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[151,348,748,773]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
.
<bibRefCitation id="CD7E4B4B174F9D53FE8D4EF0A0ABFCDB" author="Rasnitsyn, A. P. &amp; Aristov, D. S." box="[364,714,748,773]" pageId="2" pageNumber="73" pagination="678 - 703" refId="ref14281" refString="Rasnitsyn, A. P. and Aristov, D. S. 2013. New fossil insects (Insecta: Caloneurida, Hypoperlida, Palaeomanteida, Jurinida) from the Middle and Upper Permian of European Russia. In: D. S. Aristov, A. S. Bashkuev, V. K. Golubev, A. V. Gorochov, E. V. Karasev, D. S. Kopylov, A. G. Ponomarenko, A. P. Rasnitsyn, D. A. Rasnitsyn, N. D. Sinitshenkova, I. D. Sukatsheva, and D. V. Vassilenko (eds.), Fossil Insects of the Middle and Upper Permian of European Russia. Paleontological Journal 47: 678 - 703." type="journal article" year="2013">Rasnitsyn and Aristov (2013)</bibRefCitation>
, refusing the argument of convexity of the different veins, proposed to synonymize the
<taxonomicName id="6EEF4D39174F9D53FEB64F31A076FC98" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[343,535,813,838]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
with the
<taxonomicName id="6EEF4D39174F9D53FD6B4F31A156FC98" authorityName="Handlirsch" authorityYear="1906" box="[650,823,813,838]" class="Insecta" family="Ischnoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Ischnoneuridae</taxonomicName>
(
<typeStatus id="76548818174F9D53FF7E4F52A2B3FCB9" box="[159,210,846,871]" pageId="2" pageNumber="73">type</typeStatus>
genus
<taxonomicName id="6EEF4D39174F9D53FED14F52A0F7FCB9" authority="Brongniart, 1893" authorityName="Brongniart" authorityYear="1893" box="[304,662,846,871]" class="Insecta" family="Ischnoneuridae" genus="Ischnoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174F9D53FED14F52A3DEFCB9" box="[304,447,846,871]" italics="true" pageId="2" pageNumber="73">Ischnoneura</emphasis>
<bibRefCitation id="CD7E4B4B174F9D53FE2C4F52A0F7FCB9" author="Brongniart, C." box="[461,662,846,871]" pageId="2" pageNumber="73" pagination="124 - 615" refId="ref12823" refString="Brongniart, C. 1893. Recherches pour servir a l'histoire des insectes fossiles des temps primaires precedees d'une etude sur la nervation des ailes des insectes. Bulletin de la Societe d'Industrie Minerale de Saint-Etienne (3) 7: 124 - 615." type="journal article" year="1893">Brongniart, 1893</bibRefCitation>
</taxonomicName>
(Gzhelian of Commentry), without giving any argument against the attribution of
<taxonomicName id="6EEF4D39174F9D53FEC74F93A3D2FC76" authorityName="Brongniart" authorityYear="1893" box="[294,435,911,936]" class="Insecta" family="Ischnoneuridae" genus="Ischnoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174F9D53FEC74F93A3D2FC76" box="[294,435,911,936]" italics="true" pageId="2" pageNumber="73">Ischnoneura</emphasis>
</taxonomicName>
to the archaeorthopteran lineage (and more precisely to the
<taxonomicName id="6EEF4D39174F9D53FE354FACA0FAFC17" ID-CoL="YT" authorityName="Handlirsch" authorityYear="1937" box="[468,667,944,969]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Cnemidolestodea</taxonomicName>
; see
<bibRefCitation id="CD7E4B4B174F9D53FD394FACA2BCFC34" author="Bethoux, O." pageId="2" pageNumber="73" pagination="403 - 408" refId="ref12559" refString="Bethoux, O. 2005 a. Cnemidolestodea (Insecta): an ancient order reinstated. Journal of Systematic Palaeontology 3: 403 - 408." type="journal article" year="2005">Béthoux 2005a</bibRefCitation>
), as proposed by
<bibRefCitation id="CD7E4B4B174F9D53FE554FCDA0ADFC34" author="Bethoux, O. &amp; Nel, A." box="[436,716,977,1002]" pageId="2" pageNumber="73" pagination="1 - 25" refId="ref12655" refString="Bethoux, O. and Nel, A. 2005. Some Palaeozoic &quot; Protorthoptera &quot; are &quot; ancestral &quot; orthopteroids: major wing braces as clues to a new split among the &quot; Protorthoptera &quot;. Journal of Systematic Palaeontology 2: 1 - 25." type="journal article" year="2005">Béthoux and Nel (2005</bibRefCitation>
: fig. 19), on the basis of the pattern of forewing venation typical of those of the Archaeorthoptera, especially in the presence of a concave anterior branch of
<collectionCode id="CFFEAE7F174F9D53FDF3482FA022FB92" box="[530,579,1075,1100]" pageId="2" pageNumber="73">CuP</collectionCode>
reaching the convex CuA, itself emerging from a common stem with M. We reject this synonymy and restore the
<taxonomicName id="6EEF4D39174F9D53FDB54868A175FB53" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[596,788,1140,1165]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
as a separate family, in a different clade that does not belong to the Archaeorthoptera, unlike the
<taxonomicName id="6EEF4D39174F9D53FDDC48A9A08FFB10" authorityName="Handlirsch" authorityYear="1906" box="[573,750,1205,1230]" class="Insecta" family="Ischnoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Ischnoneuridae</taxonomicName>
. More recently,
<bibRefCitation id="CD7E4B4B174F9D53FF1C48CAA3FCFB31" author="Aristov, D. S." box="[253,413,1238,1263]" pageId="2" pageNumber="73" pagination="1 - 46" refId="ref12449" refString="Aristov, D. S. 2014. Classification of the order Cnemidolestida (Insecta: Perlidea) with descriptions of new taxa. Far Eastern Entomologist 277: 1 - 46." type="journal article" year="2014">Aristov (2014)</bibRefCitation>
synonymized the
<taxonomicName id="6EEF4D39174F9D53FD8848CAA176FB31" authorityName="Handlirsch" authorityYear="1906" box="[617,791,1238,1263]" class="Insecta" family="Ischnoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Ischnoneuridae</taxonomicName>
(at least its
<typeStatus id="76548818174F9D53FF1448EBA346FACE" box="[245,295,1271,1296]" pageId="2" pageNumber="73">type</typeStatus>
genus
<taxonomicName id="6EEF4D39174F9D53FE9848EBA064FACE" authorityName="Brongniart" authorityYear="1893" box="[377,517,1271,1296]" class="Insecta" family="Ischnoneuridae" genus="Ischnoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174F9D53FE9848EBA064FACE" box="[377,517,1271,1296]" italics="true" pageId="2" pageNumber="73">Ischnoneura</emphasis>
</taxonomicName>
) with the
<taxonomicName id="6EEF4D39174F9D53FD9A48EBA33FFAEE" ID-CoL="624CQ" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" class="Insecta" family="Cnemidolestidae" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">
Cnemidolestidae
<bibRefCitation id="CD7E4B4B174F9D53FF76490BA33FFAEE" author="Handlirsch, A." box="[151,350,1303,1328]" pageId="2" pageNumber="73" pagination="661 - 820" refId="ref13126" refString="Handlirsch, A. 1906. Revision of American Paleozoic insects. Proceedings of the United States National Museum, Washington 29: 661 - 820." type="journal article" year="1906">Handlirsch, 1906</bibRefCitation>
</taxonomicName>
,
<typeStatus id="76548818174F9D53FE8B490BA3FDFAEE" box="[362,412,1303,1328]" pageId="2" pageNumber="73">type</typeStatus>
family of the
<taxonomicName id="6EEF4D39174F9D53FDA3490BA169FAEE" ID-CoL="YT" authorityName="Handlirsch" authorityYear="1937" box="[578,776,1303,1328]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Cnemidolestodea</taxonomicName>
, but without speaking again his previous proposal of synonymy of the
<taxonomicName id="6EEF4D39174F9D53FF3F4945A3FFFAAC" ID-CoL="6HF" authorityName="Handlirsch" authorityYear="1922" box="[222,414,1369,1394]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
with the
<taxonomicName id="6EEF4D39174F9D53FDE84945A0DBFAAC" authorityName="Handlirsch" authorityYear="1906" box="[521,698,1369,1394]" class="Insecta" family="Ischnoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Ischnoneuridae</taxonomicName>
.
</paragraph>
<paragraph id="A95036BA174F9D53FF5C4965A0B2F8E5" blockId="2.[151,825,160,1851]" pageId="2" pageNumber="73">
Note that some holometabolous
<taxonomicName id="6EEF4D39174F9D53FDCC4965A093FA4C" box="[557,754,1401,1426]" class="Insecta" family="Permochoristidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Permochoristidae</taxonomicName>
(
<taxonomicName id="6EEF4D39174F9D53FCE54965A323FA6D" class="Insecta" family="Permochoristidae" genus="Protopanorpa" kingdom="Animalia" order="Mecoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9B9BEAA8174F9D53FCE54965A36CFA6D" italics="true" pageId="2" pageNumber="73">Protopanorpa</emphasis>
spp.
</taxonomicName>
) also have a similar pattern of a curved vein in mid part of wing with a series of anteriorly pectinate and parallel branches, separating basally from a simple vein, but the concerned veins are completely different, the anteriorly pectinate vein being
<collectionCode id="CFFEAE7F174F9D53FE694A01A3D0F9E8" box="[392,433,1565,1590]" country="0" name="Mohonk Preserve, Inc." pageId="2" pageNumber="73">MP</collectionCode>
while the simple posterior vein is CuA (see
<bibRefCitation id="CD7E4B4B174F9D53FEEA4A21A3DCF988" author="Bashkuev, A. S." box="[267,445,1597,1622]" pageId="2" pageNumber="73" pagination="291 - 296" refId="ref12528" refString="Bashkuev, A. S. 2010. New scorpionflies (Insecta: Mecoptera: Permochoristidae) from the Ufimian of Cisuralia. Paleontological Journal 44: 291 - 296." type="journal article" year="2010">Bashkuev 2010</bibRefCitation>
: fig. 1). Also the
<taxonomicName id="6EEF4D39174F9D53FD674A21A09CF988" authorityName="Burmeister" authorityYear="1839" box="[646,765,1597,1622]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
penturoperlid
<taxonomicName id="6EEF4D39174F9D53FEF34A42A3C9F9A9" authorityName="Illies" authorityYear="1960" box="[274,424,1630,1655]" class="Insecta" family="Austroperlidae" genus="Penturoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="9B9BEAA8174F9D53FEF34A42A3C9F9A9" box="[274,424,1630,1655]" italics="true" pageId="2" pageNumber="73">Penturoperla</emphasis>
</taxonomicName>
have a median vein with a series of parallel anterior branches (
<bibRefCitation id="CD7E4B4B174F9D53FE0D4A63A008F946" author="Illies, J." box="[492,617,1663,1688]" pageId="2" pageNumber="73" pagination="26 - 41" refId="ref13239" refString="Illies, J. 1960. Penturoperlidae, eine neue Plecopteren Familie. Zoologischer Anzeiger 164: 26 - 41." type="journal article" year="1960">Illies 1960</bibRefCitation>
), even in some of them could be in fact interpreted as branches of CuA captured by median vein following
<bibRefCitation id="CD7E4B4B174F9D53FDFF4ADCA085F907" author="Bethoux, O." box="[542,740,1728,1753]" pageId="2" pageNumber="73" pagination="52 - 81" refId="ref12586" refString="Bethoux, O. 2005 b. Wing venation pattern of Plecoptera (Insecta: Neoptera). Illiesia 1: 52 - 81." type="journal article" year="2005">Béthoux (2005b)</bibRefCitation>
. These similarities support the hypothesis of a convergent evolution between the Archaeorthoptera
<taxonomicName id="6EEF4D39174F9D53FDDE4B1DA163F8C4" authorityName="Handlirsch" authorityYear="1937" box="[575,770,1793,1818]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Cnemidolestodea</taxonomicName>
and the
<taxonomicName id="6EEF4D39174F9D53FF204B3EA37CF8E5" authorityName="Handlirsch" authorityYear="1906" box="[193,285,1826,1851]" class="Insecta" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="order">Paoliida</taxonomicName>
<taxonomicName id="6EEF4D39174F9D53FEC54B3EA386F8E5" authorityName="Handlirsch" authorityYear="1922" box="[292,487,1826,1851]" class="Insecta" family="Anthracoptilidae" kingdom="Animalia" order="Paoliida" pageId="2" pageNumber="73" phylum="Arthropoda" rank="family">Anthracoptilidae</taxonomicName>
, due to flight ability.
</paragraph>
<paragraph id="A95036BA174F9D53FF764B51A052F876" blockId="2.[151,824,1869,1960]" pageId="2" pageNumber="73">
<emphasis id="9B9BEAA8174F9D53FF764B51A05AF8B8" box="[151,571,1869,1894]" italics="true" pageId="2" pageNumber="73">Stratigraphic and geographic range</emphasis>
.—Bashkirian (Lower Pennsylvanian)Lopingian (upper Permian),
<collectingCountry id="D1F8762A174F9D53FD274B72A152F859" box="[710,819,1902,1927]" name="Australia" pageId="2" pageNumber="73">Australia</collectingCountry>
, Europe, North America, and
<collectingCountry id="D1F8762A174F9D53FE004B93A051F876" box="[481,560,1935,1960]" name="Russia" pageId="2" pageNumber="73">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>