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<document id="B0F39603B52A0533A626F0EEE93A236D" ID-DOI="10.1080/00222930802002535" ID-ISSN="1464-5262" ID-Zenodo-Dep="5219397" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629340601867" checkinUser="felipe" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Scholtz, Clarke H." docDate="2008" docId="1F77BC72FFCD306E23C9FB14FD7FF926" docLanguage="en" docName="JNatHist.42.21-24.1477-1508.pdf" docOrigin="Journal of Natural History 42 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222930802002535" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Coptorhina excavata Frolov &amp; Akhmetova &amp; Scholtz 2008, sp. n." docType="treatment" docVersion="3" lastPageNumber="1502" masterDocId="E34EC40AFFD430752211FF81FFF5FF8C" masterDocTitle="Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1508" masterPageNumber="1477" pageNumber="1500" updateTime="1699068592934" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="21FDAF000A8BB5F469EB2B8FD8797078" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9F7A5BDFD5AC72629710B3716C9696F6">
<mods:title id="5C848174A33506E852634219B2487FB8">Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="251F75913F5975105BBE67AA657BE953" type="personal">
<mods:role id="B37F6C8A67AFE456EE85933EEEBF564C">
<mods:roleTerm id="861AB3E63C5E09AE32CF378643AFDD2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="570E91142D2AAF3B9E5E4CB256894B3E">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:name id="1CABF11D584EE6099FD1EAA14E64AEA6" type="personal">
<mods:role id="EA1CF499722D13F111C638D1B6D44975">
<mods:roleTerm id="1E8AC95C563B7FDC3EBC7BAD32E622B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="103B31B952B9370AB1FC990F04ADE783">Akhmetova, Lilia A.</mods:namePart>
</mods:name>
<mods:name id="C8788F3C1A1B262DACE20831E7E991BD" type="personal">
<mods:role id="743E453406380964DB2CC34F537056FB">
<mods:roleTerm id="E514CD88028E666BCC527BB83261B5D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC7CE894044D1C93CAEB03208B50144B">Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E3DBE7176FEA3653E6B1975A58F2466A">text</mods:typeOfResource>
<mods:relatedItem id="21D02B772D69908168AB7FDC0CF28018" type="host">
<mods:titleInfo id="F55CA5CB75A69B91748236AED2F8A9B5">
<mods:title id="1E473C869443F3564D1D28A9AF17CB23">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="E1A7C9B4F313CA318DEAF32000174535">
<mods:date id="E884930AC5182CA7AD8EE762C360DDAC">2008</mods:date>
<mods:detail id="8DE7E95C906EF2C949DC86A8356F3640" type="pubDate">
<mods:number id="F0DCEA23A7C6CCE510349B610F26235B">2008-06-30</mods:number>
</mods:detail>
<mods:detail id="6638342F5D3D396D90C98EEBAD577B88" type="volume">
<mods:number id="D51FBEAC59B75A10EE6B9D2FE55E64B1">42</mods:number>
</mods:detail>
<mods:detail id="20A9E8100D0B8E6627B691B626B0D8B3" type="issue">
<mods:number id="001840B49CA572B5A9EECC17CE9E16F4">21 - 24</mods:number>
</mods:detail>
<mods:extent id="8E7B8B060F7061ADF8CB63EA40D2FCE4" unit="page">
<mods:start id="69C2E9AFB72BE0247A02669B6C29CBA9">1477</mods:start>
<mods:end id="708B3A1B00736A27DE5A53C0E102B55C">1508</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0D332837E432F0F5358E2ED38318A88F">
<mods:url id="83F99E532E38F9AC7552B7772E863510">http://dx.doi.org/10.1080/00222930802002535</mods:url>
</mods:location>
<mods:classification id="E935D27EAE28B4CA1F319BA75C911115">journal article</mods:classification>
<mods:identifier id="3BF958CF310996C7BC2DC168F247BA4A" type="DOI">10.1080/00222930802002535</mods:identifier>
<mods:identifier id="A0975062D318F25B184417F6E81AEA66" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="DDE2F711D2B4D549834FBA7667081B4D" type="Zenodo-Dep">5219397</mods:identifier>
</mods:mods>
<treatment id="1F77BC72FFCD306E23C9FB14FD7FF926" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926" httpUri="http://treatment.plazi.org/id/1F77BC72FFCD306E23C9FB14FD7FF926" lastPageId="27" lastPageNumber="1502" pageId="25" pageNumber="1500">
<subSubSection id="DFC45EEFFFCD306C23C9FB14FCFDFB21" box="[472,776,1173,1197]" pageId="25" pageNumber="1500" type="nomenclature">
<paragraph id="97610D64FFCD306C23C9FB14FCFDFB21" blockId="25.[472,776,1173,1229]" box="[472,776,1173,1197]" pageId="25" pageNumber="1500">
<heading id="CC29BA08FFCD306C23C9FB14FCFDFB21" box="[472,776,1173,1197]" centered="true" fontSize="36" level="2" pageId="25" pageNumber="1500" reason="7">
<taxonomicName id="50DE76E7FFCD306C23C9FB14FD35FB21" ID-CoL="Y87Z" authority="Frolov &amp; Akhmetova &amp; Scholtz, 2008" authorityName="Frolov &amp; Akhmetova &amp; Scholtz" authorityYear="2008" box="[472,704,1173,1197]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1500" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="A5AAD176FFCD306C23C9FB14FD35FB21" bold="true" box="[472,704,1173,1197]" italics="true" pageId="25" pageNumber="1500">Coptorhina excavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BE996C0DFFCD306C20DBFB14FCFDFB21" box="[714,776,1173,1197]" pageId="25" pageNumber="1500" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DFC45EEFFFCD306E23CBFB34FD7FF926" lastPageId="27" lastPageNumber="1502" pageId="25" pageNumber="1500" type="description">
<paragraph id="97610D64FFCD306C23CBFB34FCF3FB41" blockId="25.[472,776,1173,1229]" box="[474,774,1205,1229]" pageId="25" pageNumber="1500">
(
<figureCitation id="0FE511E1FFCD306C23F3FB34FDB8FB41" box="[482,589,1205,1229]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="25" pageNumber="1500">Figures 4</figureCitation>
,
<figureCitation id="0FE511E1FFCD306C204BFB34FD5EFB41" box="[602,683,1205,1229]" captionStart="Figures 9–14" captionStartId="7.[146,222,1424,1446]" captionTargetBox="[146,1104,165,1387]" captionTargetId="figure-1@7.[146,1104,165,1387]" captionTargetPageId="7" captionText="Figures 9–14. Coptorhina spp. 9, wing; 10–11, aedeagus in dorsal and lateral view; 12, internal sac of aedeagus; 13, internal sac sclerite arrowed in Figure 12; 14, abdomen in lateral view (14A, female, 14B, male). 9, 10, 12, 13C–E, 14, C. auspicata (13C, lectotype, Enkeldoorn, Zimbabwe, 13D, Cinergy Game farm, RSA, 13E, Shesheke, Zambia); 11, 13I–J, C. nitidipennis (13I, Cinergy Game farm, RSA, 13J, paratype, ‘‘Caffraria’’); 13A–B, C. excavata sp. n. (13A, holotype, 13B, paratype, Mamathes, Lesotho); 13F, C. klugii, Blouberg, RSA; 13G, C. davidi, paratype, Mpika, Congo; 13H, C. nitefacta, holotype, Bura, Kenya. 9, 13 and 14 are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219401" httpUri="https://zenodo.org/record/5219401/files/figure.png" pageId="25" pageNumber="1500">13A–B</figureCitation>
,
<figureCitation id="0FE511E1FFCD306C20A9FB34FD20FB41" box="[696,725,1205,1229]" captionStart="Figures 15–17" captionStartId="9.[146,222,1409,1431]" captionTargetBox="[152,1096,178,1370]" captionTargetId="figure-62@9.[146,1104,165,1372]" captionTargetPageId="9" captionText="Figures 15–17. Coptorhina spp., general view. 15, C. auspicata, Cinergy Game farm, RSA; 16, C. excavata sp. n., holotype; 17, C. nitidipennis, Cinergy Game farm, RSA." figureDoi="http://doi.org/10.5281/zenodo.5219403" httpUri="https://zenodo.org/record/5219403/files/figure.png" pageId="25" pageNumber="1500">16</figureCitation>
,
<figureCitation id="0FE511E1FFCD306C20F2FB34FD0BFB41" box="[739,766,1205,1229]" captionStart="Figures 20–22" captionStartId="26.[213,289,1398,1420]" captionTargetBox="[213,1171,165,1362]" captionTargetId="figure-56@26.[213,1171,165,1362]" captionTargetPageId="26" captionText="Figures 20–22. Coptorhina spp., distribution map. 20, C. auspicata; 21, C. excavata sp. n.; 22, C. nitefacta." figureDoi="http://doi.org/10.5281/zenodo.5219409" httpUri="https://zenodo.org/record/5219409/files/figure.png" pageId="25" pageNumber="1500">21</figureCitation>
)
</paragraph>
<paragraph id="97610D64FFCD306C2283FA87FE57FA92" blockId="25.[146,418,1286,1310]" box="[146,418,1286,1310]" pageId="25" pageNumber="1500">
<emphasis id="A5AAD176FFCD306C2283FA87FE57FA92" box="[146,418,1286,1310]" italics="true" pageId="25" pageNumber="1500">Type material examined</emphasis>
</paragraph>
<paragraph id="97610D64FFCD306F2283FAAEFC2AF926" blockId="25.[146,1103,1327,1706]" lastBlockId="26.[213,1169,1552,1706]" lastPageId="26" lastPageNumber="1501" pageId="25" pageNumber="1500">
<materialsCitation id="27B60739FFCD306C2283FAAEFF0EFAEB" collectingDate="1954-12-04" collectingDateMax="1954-12-14" collectingDateMin="1954-12-04" collectionCode="AMGS" collectorName="C. Jacot Guillarmod'" country="Lesotho" location="Mamathes" pageId="25" pageNumber="1500" specimenCount="1" typeStatus="holotype">
<typeStatus id="4865B3C6FFCD306C2283FAAEFF08FACB" box="[146,253,1327,1351]" pageId="25" pageNumber="1500" type="holotype">Holotype</typeStatus>
<emphasis id="A5AAD176FFCD306C2312FAB3FEE0FACA" box="[259,277,1330,1350]" italics="true" pageId="25" pageNumber="1500">„</emphasis>
with label ‘‘
<location id="92015BBFFFCD306C238BFAAEFDE3FACB" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCD306C238BFAAEFDE3FACB" box="[410,534,1327,1351]" country="Lesotho" name="Mamathes" pageId="25" pageNumber="1500">Mamathes</location>
<collectingCountry id="EFC94DF4FFCD306C200CFAAEFD6BFACB" box="[541,670,1327,1351]" name="Lesotho" pageId="25" pageNumber="1500">Basutoland</collectingCountry>
<date id="E3602BA4FFCD306C20B4FAAEFCA7FACB" box="[677,850,1327,1351]" pageId="25" pageNumber="1500" value="1954-12-04" valueMax="1954-12-14" valueMin="1954-12-04">
<collectingDate id="F324D24CFFCD306C20B4FAAEFCA7FACB" box="[677,850,1327,1351]" pageId="25" pageNumber="1500" value="1954-12-04" valueMax="1954-12-14" valueMin="1954-12-04">4 - 14-XII-1954</collectingDate>
</date>
<collectorName id="3A2B68B2FFCD306C2149FAAEFBBFFACB" box="[856,1098,1327,1351]" pageId="25" pageNumber="1500">C. Jacot Guillarmod’</collectorName>
’ (
<collectionCode id="F1CF95A1FFCD306C228BFACEFF07FAEB" box="[154,242,1359,1383]" country="South Africa" httpUri="http://grbio.org/cool/jk9j-qt42" name="Albany Museum" pageId="25" pageNumber="1500">AMGS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCD306C2314FACEFC10FAEB" box="[261,997,1359,1383]" collectionCode="AMGS, ZIN" country="Lesotho" pageId="25" pageNumber="1500" specimenCount="2" typeStatus="paratype">
<typeStatus id="4865B3C6FFCD306C2314FACEFE8DFAEB" box="[261,376,1359,1383]" pageId="25" pageNumber="1500" type="paratype">Paratypes</typeStatus>
. 7 spm. with same label as the
<typeStatus id="4865B3C6FFCD306C20C3FACEFCC3FAEB" box="[722,822,1359,1383]" pageId="25" pageNumber="1500" type="holotype">holotype</typeStatus>
(
<collectionCode id="F1CF95A1FFCD306C2154FACEFC69FAEB" box="[837,924,1359,1383]" country="South Africa" httpUri="http://grbio.org/cool/jk9j-qt42" name="Albany Museum" pageId="25" pageNumber="1500">AMGS</collectionCode>
,
<collectionCode id="F1CF95A1FFCD306C21B6FACEFC28FAEB" box="[935,989,1359,1383]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="25" pageNumber="1500" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCD306C21E1FACEFC34FA24" collectingDate="1930-01-05" collectingDateMax="1930-01-31" collectingDateMin="1930-01-05" collectionCode="AMGS, UPSA" collectorName="J. Hewitt' &amp; C. Jaco" country="Lesotho" location="Entomology University of Pretoria'" municipality="Mamathes" pageId="25" pageNumber="1500" specimenCount="1" typeStatus="paratype">
1
<emphasis id="A5AAD176FFCD306C2615FAD3FBE3FAEA" box="[1028,1046,1362,1382]" italics="true" pageId="25" pageNumber="1500">„</emphasis>
with label ‘‘
<collectingCountry id="EFC94DF4FFCD306C22C8FAEEFEABFA0B" box="[217,350,1391,1415]" name="Lesotho" pageId="25" pageNumber="1500">Basutoland</collectingCountry>
Masite,
<date id="E3602BA4FFCD306C23D0FAEEFDD9FA0B" box="[449,556,1391,1415]" pageId="25" pageNumber="1500" value="1930-01-31">
<collectingDate id="F324D24CFFCD306C23D0FAEEFDD9FA0B" box="[449,556,1391,1415]" pageId="25" pageNumber="1500" value="1930-01-31">31.I.1930</collectingDate>
</date>
.
<collectorName id="3A2B68B2FFCD306C2026FAEEFD5CFA0B" box="[567,681,1391,1415]" pageId="25" pageNumber="1500">J. Hewitt’</collectorName>
’ (
<collectionCode id="F1CF95A1FFCD306C20AAFAEEFCE1FA0B" box="[699,788,1391,1415]" country="South Africa" httpUri="http://grbio.org/cool/jk9j-qt42" name="Albany Museum" pageId="25" pageNumber="1500">AMGS</collectionCode>
). 1
<emphasis id="A5AAD176FFCD306C212BFAF2FCB9FA0B" box="[826,844,1395,1415]" italics="true" pageId="25" pageNumber="1500">„</emphasis>
with label ‘‘
<collectingMunicipality id="7705971EFFCD306C21C3FAEEFBBBFA0B" box="[978,1102,1391,1415]" pageId="25" pageNumber="1500">Mamathes</collectingMunicipality>
<date id="E3602BA4FFCD306C2283FA11FF04FA24" box="[146,241,1424,1448]" pageId="25" pageNumber="1500" value="1930-01-05">
<collectingDate id="F324D24CFFCD306C2283FA11FF04FA24" box="[146,241,1424,1448]" pageId="25" pageNumber="1500" value="1930-01-05">5-I-1930</collectingDate>
</date>
<collectorName id="3A2B68B2FFCD306C22E9FA11FEBAFA24" box="[248,335,1424,1448]" pageId="25" pageNumber="1500">C. Jaco</collectorName>
, Dept. of
<location id="92015BBFFFCD306C23D1FA11FCA3FA24" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCD306C23D1FA11FCA3FA24" box="[448,854,1424,1448]" country="Lesotho" municipality="Mamathes" name="Entomology University of Pretoria'" pageId="25" pageNumber="1500">Entomology University of Pretoria’</location>
’ (
<collectionCode id="F1CF95A1FFCD306C217BFA11FC4DFA24" box="[874,952,1424,1448]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="25" pageNumber="1500">UPSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCD306C21DAFA11FC23FA44" collectingDate="1933-12-02" collectionCode="UPSA" collectorName="C. Jacot Guillarmod'" country="Lesotho" location="Mamathes" pageId="25" pageNumber="1500" specimenCount="1" typeStatus="paratype">
1 spm. with labels ‘‘
<collectingCountry id="EFC94DF4FFCD306C22F5FA31FE9CFA44" box="[228,361,1456,1480]" name="Lesotho" pageId="25" pageNumber="1500">Basutoland</collectingCountry>
<location id="92015BBFFFCD306C2360FA31FE1CFA44" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCD306C2360FA31FE1CFA44" box="[369,489,1456,1480]" country="Lesotho" name="Mamathes" pageId="25" pageNumber="1500">Mamathes</location>
<date id="E3602BA4FFCD306C23E0FA31FD9AFA44" box="[497,623,1456,1480]" pageId="25" pageNumber="1500" value="1933-12-02">
<collectingDate id="F324D24CFFCD306C23E0FA31FD9AFA44" box="[497,623,1456,1480]" pageId="25" pageNumber="1500" value="1933-12-02">2-XII-1933</collectingDate>
</date>
<collectorName id="3A2B68B2FFCD306C2067FA31FC9EFA44" box="[630,875,1456,1480]" pageId="25" pageNumber="1500">C. Jacot Guillarmod’</collectorName>
’ (
<collectionCode id="F1CF95A1FFCD306C216EFA31FC3BFA44" box="[895,974,1456,1480]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="25" pageNumber="1500">UPSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCD306C21F0FA31FBBEF906" collectingDate="1983-10-19" collectingDateMax="1984-01" collectingDateMin="1983-10-19" collectionCode="BMSA" collectorName="S. Africa &amp; O. F. S. Kromrant &amp; S. Africa O. F. S. Schoongezicht &amp; O. F. S. Driekloof" country="Lesotho" location="Ca'" pageId="25" pageNumber="1500" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
1
<emphasis id="A5AAD176FFCD306C21E7FA32FBFDFA4B" box="[1014,1032,1459,1479]" italics="true" pageId="25" pageNumber="1500">„</emphasis>
and
<specimenCount id="81D8C6EDFFCD306C2650FA31FF55FA64" pageId="25" pageNumber="1500" type="female">1 ♀</specimenCount>
with labels ‘‘
<collectorName id="3A2B68B2FFCD306C2354FA51FE46FA64" box="[325,435,1488,1512]" pageId="25" pageNumber="1500">S. Africa</collectorName>
,
<collectorName id="3A2B68B2FFCD306C23D2FA51FD7BFA64" box="[451,654,1488,1512]" pageId="25" pageNumber="1500">O.F.S. Kromrant</collectorName>
123 BOSHOF SE2825
<location id="92015BBFFFCD306C21BFFA51FC2FFA64" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCD306C21BFFA51FC2FFA64" box="[942,986,1488,1512]" country="Lesotho" name="Ca'" pageId="25" pageNumber="1500">Ca’</location>
’, ‘‘BMSA 14055’’ and ‘‘
<date id="E3602BA4FFCD306C2320FA70FE1AF985" box="[305,495,1521,1545]" pageId="25" pageNumber="1500" value="1983-11">
<collectingDate id="F324D24CFFCD306C2320FA70FE1AF985" box="[305,495,1521,1545]" pageId="25" pageNumber="1500" value="1983-11">November 1983</collectingDate>
</date>
–
<date id="E3602BA4FFCD306C23EFFA70FD68F985" box="[510,669,1521,1545]" pageId="25" pageNumber="1500" value="1984-01">
<collectingDate id="F324D24CFFCD306C23EFFA70FD68F985" box="[510,669,1521,1545]" pageId="25" pageNumber="1500" value="1984-01">January 1984</collectingDate>
</date>
. Pres. traps.’’ (
<collectionCode id="F1CF95A1FFCD306C2145FA70FC5CF985" box="[852,937,1521,1545]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="25" pageNumber="1500">BMSA</collectionCode>
). 1 spm. with labels ‘‘
<collectorName id="3A2B68B2FFCD306C22E2F990FD8DF9A5" box="[243,632,1553,1577]" pageId="25" pageNumber="1500">S. Africa O.F.S. Schoongezicht</collectorName>
56. Bloemfontein 29
<emphasis id="A5AAD176FFCD306C2196F990FC64F9A5" bold="true" box="[903,913,1553,1577]" italics="true" pageId="25" pageNumber="1500">°</emphasis>
149 S 26
<emphasis id="A5AAD176FFCD306C21E4F990FC0AF9A5" bold="true" box="[1013,1023,1553,1577]" italics="true" pageId="25" pageNumber="1500">°</emphasis>
099 E’’, ‘‘
<date id="E3602BA4FFCD306C228CF9B0FEE6F9C5" box="[157,275,1585,1609]" pageId="25" pageNumber="1500" value="1983-10-19">
<collectingDate id="F324D24CFFCD306C228CF9B0FEE6F9C5" box="[157,275,1585,1609]" pageId="25" pageNumber="1500" value="1983-10-19">19.X.1983</collectingDate>
</date>
Van Rensburg, Earle´’’ (
<collectionCode id="F1CF95A1FFCD306C2051F9B0FD60F9C5" box="[576,661,1585,1609]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="25" pageNumber="1500">BMSA</collectionCode>
). 1
<emphasis id="A5AAD176FFCD306C20D8F9B5FD2EF9C4" box="[713,731,1588,1608]" italics="true" pageId="25" pageNumber="1500">„</emphasis>
with labels ‘‘
<collectorName id="3A2B68B2FFCD306C2192F9B0FC07F9C5" box="[899,1010,1585,1609]" pageId="25" pageNumber="1500">S. Africa</collectorName>
,
<collectorName id="3A2B68B2FFCD306C2615F9B0FEF4F9E5" pageId="25" pageNumber="1500">O.F.S. Driekloof</collectorName>
2836 Bloemfontein 29
<emphasis id="A5AAD176FFCD306C2014F9D0FDFAF9E5" bold="true" box="[517,527,1617,1641]" italics="true" pageId="25" pageNumber="1500">°</emphasis>
129 S 25
<emphasis id="A5AAD176FFCD306C207BF9D0FD81F9E5" bold="true" box="[618,628,1617,1641]" italics="true" pageId="25" pageNumber="1500">°</emphasis>
549 E’’, ‘‘National Museum Bloemfontein Dept. Entomology’’, ‘‘
<date id="E3602BA4FFCD306C2382F9F3FDFCF906" box="[403,521,1650,1674]" pageId="25" pageNumber="1500" value="1983-12">
<collectingDate id="F324D24CFFCD306C2382F9F3FDFCF906" box="[403,521,1650,1674]" pageId="25" pageNumber="1500" value="1983-12">Dec. 1983</collectingDate>
</date>
Van Rensburg, Earle´’’, ‘‘BMSA 14943’’ (
<collectionCode id="F1CF95A1FFCD306C21FFF9F3FBB6F906" box="[1006,1091,1650,1674]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="25" pageNumber="1500">BMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCD306C2283F913FD02F926" box="[146,759,1682,1706]" collectionCode="TMSA" collectorName="Lichtenburg Dr. Brauns'" country="Lesotho" pageId="25" pageNumber="1500" specimenCount="1" typeStatus="paratype">
spm. with label ‘‘
<collectorName id="3A2B68B2FFCD306C234AF913FD76F926" box="[347,643,1682,1706]" pageId="25" pageNumber="1500">Lichtenburg Dr. Brauns’</collectorName>
’ (
<collectionCode id="F1CF95A1FFCD306C208BF913FD1AF926" box="[666,751,1682,1706]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="25" pageNumber="1500">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCD306F2117F913FE1AF9C5" collectionCode="R, MNHN" collectorName="Museum Paris Basoutolanad Lessouto Vallee du Ht Orange &amp; E. Haug" country="Lesotho" elevation="1500" lastPageId="26" lastPageNumber="1501" location="Ellenberger" pageId="25" pageNumber="1500" specimenCount="1" typeStatus="paratype">
1 spm. with label ‘‘
<collectorName id="3A2B68B2FFCD306F21F6F913FCCCF9A4" lastPageId="26" lastPageNumber="1501" pageId="25" pageNumber="1500">Museum Paris Basoutolanad Lessouto Vallee du Ht Orange</collectorName>
<quantity id="5026A081FFCE306F2157F991FC63F9A4" box="[838,918,1552,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="1501" unit="m" value="1500.0">
<elevation id="1CF3EA57FFCE306F2157F991FC63F9A4" box="[838,918,1552,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="1501" unit="m" value="1500.0">1500 m</elevation>
</quantity>
(legit
<collectionCode id="F1CF95A1FFCE306F21FBF991FC08F9A4" box="[1002,1021,1552,1576]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="1501">R</collectionCode>
.
<location id="92015BBFFFCE306F2611F991FB7CF9A4" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCE306F2611F991FB7CF9A4" box="[1024,1161,1552,1576]" country="Lesotho" name="Ellenberger" pageId="26" pageNumber="1501">Ellenberger</location>
)
<collectorName id="3A2B68B2FFCE306F22C4F9B0FED8F9C5" box="[213,301,1585,1609]" pageId="26" pageNumber="1501">E.Haug</collectorName>
1906’’ (
<collectionCode id="F1CF95A1FFCE306F2397F9B0FE12F9C5" box="[390,487,1585,1609]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="1501">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCE306F23EBF9B0FD30F9E5" collectionCode="MNHN" collectorName="Museum Paris &amp; Ellenberger" country="South Africa" location="Pays des Bassoutos" pageId="26" pageNumber="1501" specimenCount="1" typeStatus="paratype">
1 spm. with label ‘‘
<collectorName id="3A2B68B2FFCE306F20DDF9B0FC86F9C5" box="[716,883,1585,1609]" pageId="26" pageNumber="1501">Museum Paris</collectorName>
<collectingCountry id="EFC94DF4FFCE306F216BF9B0FBDCF9C5" box="[890,1065,1585,1609]" name="South Africa" pageId="26" pageNumber="1501">Afrique du Sud</collectingCountry>
<location id="92015BBFFFCE306F2621F9B0FEBDF9E5" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCE306F2621F9B0FEBDF9E5" country="South Africa" name="Pays des Bassoutos" pageId="26" pageNumber="1501">Pays des Bassoutos</location>
V.
<collectorName id="3A2B68B2FFCE306F2369F9D0FE0FF9E5" box="[376,506,1617,1641]" pageId="26" pageNumber="1501">Ellenberger</collectorName>
1928’’ (
<collectionCode id="F1CF95A1FFCE306F204DF9D0FD48F9E5" box="[604,701,1617,1641]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="1501">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCE306F20C5F9D0FD62F906" collectingDate="1930-12-19" collectionCode="TMSA" collectorName="C. Jacot Guillarmod'" country="South Africa" location="Moletsane" pageId="26" pageNumber="1501" specimenCount="1" typeStatus="paratype">
1 spm. with label ‘‘
<location id="92015BBFFFCE306F21A7F9D0FBC4F9E5" LSID="urn:lsid:plazi:treatment:1F77BC72FFCD306E23C9FB14FD7FF926:92015BBFFFCE306F21A7F9D0FBC4F9E5" box="[950,1073,1617,1641]" country="South Africa" name="Moletsane" pageId="26" pageNumber="1501">Moletsane</location>
<date id="E3602BA4FFCE306F262DF9D0FEFFF906" pageId="26" pageNumber="1501" value="1930-12-19">
<collectingDate id="F324D24CFFCE306F262DF9D0FEFFF906" pageId="26" pageNumber="1501" value="1930-12-19">19-XII- 1930</collectingDate>
</date>
<collectorName id="3A2B68B2FFCE306F2308F9F3FDEBF906" box="[281,542,1650,1674]" pageId="26" pageNumber="1501">C. Jacot Guillarmod’</collectorName>
’ (
<collectionCode id="F1CF95A1FFCE306F202BF9F3FD7AF906" box="[570,655,1650,1674]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="26" pageNumber="1501">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="27B60739FFCE306F20BBF9F3FC2EF926" collectionCode="SAMC" country="South Africa" location="LIMPOPO" pageId="26" pageNumber="1501" specimenCount="1" stateProvince="Limpopo" typeStatus="paratype">
1
<emphasis id="A5AAD176FFCE306F20D7F9F4FD2DF905" box="[710,728,1653,1673]" italics="true" pageId="26" pageNumber="1501">„</emphasis>
with labels ‘‘BETW.
<collectingRegion id="551AC386FFCE306F21FCF9F3FB99F906" box="[1005,1132,1650,1674]" country="South Africa" name="Limpopo" pageId="26" pageNumber="1501">LIMPOPO</collectingRegion>
&amp; ZAMBEZI RS.’’, ‘‘C. optata’’, ‘‘SAMC-COL. AO32416’’ (
<collectionCode id="F1CF95A1FFCE306F216CF913FC26F926" box="[893,979,1682,1706]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="26" pageNumber="1501" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="C3A15DECFFCE306F22C4FAF7FEA4FA25" ID-DOI="http://doi.org/10.5281/zenodo.5219409" ID-Zenodo-Dep="5219409" httpUri="https://zenodo.org/record/5219409/files/figure.png" pageId="26" pageNumber="1501" startId="26.[213,289,1398,1420]" targetBox="[213,1171,165,1362]" targetPageId="26">
<paragraph id="97610D64FFCE306F22C4FAF7FEA4FA25" blockId="26.[213,1169,1398,1449]" pageId="26" pageNumber="1501">
Figures 20–22.
<taxonomicName id="50DE76E7FFCE306F2367FAF7FDE3FA00" box="[374,534,1398,1420]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1484" phylum="Arthropoda" rank="species" species="spp">
<emphasis id="A5AAD176FFCE306F2367FAF7FE12FA07" box="[374,487,1398,1419]" italics="true" pageId="26" pageNumber="1501">Coptorhina</emphasis>
spp
</taxonomicName>
., distribution map. 20,
<taxonomicName id="50DE76E7FFCE306F2113FAF7FC8AFA07" box="[770,895,1398,1419]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1501" phylum="Arthropoda" rank="species" species="auspicata">
<emphasis id="A5AAD176FFCE306F2113FAF7FC8AFA07" box="[770,895,1398,1419]" italics="true" pageId="26" pageNumber="1501">C. auspicata</emphasis>
</taxonomicName>
; 21,
<taxonomicName id="50DE76E7FFCE306F21A0FAF7FBD2FA07" authority="Frolov &amp; Akhmetova &amp; Scholtz, 2008" authorityName="Frolov &amp; Akhmetova &amp; Scholtz" authorityYear="2008" box="[945,1063,1398,1419]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1501" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="A5AAD176FFCE306F21A0FAF7FBD2FA07" box="[945,1063,1398,1419]" italics="true" pageId="26" pageNumber="1501">C. excavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BE996C0DFFCE306F263FFAF7FB92FA00" box="[1070,1127,1398,1420]" pageId="26" pageNumber="1501" rank="species">sp. n.</taxonomicNameLabel>
; 22,
<taxonomicName id="50DE76E7FFCE306F22C4FA15FEBEFA25" authorityName="Gillet" authorityYear="1932" box="[213,331,1428,1449]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1501" phylum="Arthropoda" rank="species" species="nitefacta">
<emphasis id="A5AAD176FFCE306F22C4FA15FEBEFA25" box="[213,331,1428,1449]" italics="true" pageId="26" pageNumber="1501">C. nitefacta</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="97610D64FFCF306E2283FF29FF0AFF4C" blockId="27.[146,255,168,192]" box="[146,255,168,192]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FF29FF0AFF4C" box="[146,255,168,192]" italics="true" pageId="27" pageNumber="1502">Diagnosis</emphasis>
</paragraph>
<paragraph id="97610D64FFCF306E2283FF51FE05FECB" blockId="27.[146,1103,208,327]" pageId="27" pageNumber="1502">
This species differs from other
<taxonomicName id="50DE76E7FFCF306E23E3FF51FD36FF64" box="[498,707,208,232]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1502" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A5AAD176FFCF306E23E3FF51FD9AFF64" box="[498,623,208,232]" italics="true" pageId="27" pageNumber="1502">Coptorhina</emphasis>
species
</taxonomicName>
by the shape of the pronotum and its lateral border (
<figureCitation id="0FE511E1FFCF306E2376FF71FE27FE84" box="[359,466,240,264]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="27" pageNumber="1502">Figures 4</figureCitation>
,
<figureCitation id="0FE511E1FFCF306E23F0FF71FE0BFE84" box="[481,510,240,264]" captionStart="Figures 15–17" captionStartId="9.[146,222,1409,1431]" captionTargetBox="[152,1096,178,1370]" captionTargetId="figure-62@9.[146,1104,165,1372]" captionTargetPageId="9" captionText="Figures 15–17. Coptorhina spp., general view. 15, C. auspicata, Cinergy Game farm, RSA; 16, C. excavata sp. n., holotype; 17, C. nitidipennis, Cinergy Game farm, RSA." figureDoi="http://doi.org/10.5281/zenodo.5219403" httpUri="https://zenodo.org/record/5219403/files/figure.png" pageId="27" pageNumber="1502">16</figureCitation>
), somewhat serrate transverse pronotal carina and mostly brown coloration of elytra. From
<taxonomicName id="50DE76E7FFCF306E2076FE91FD1CFEA4" authorityName="Gillet" authorityYear="1932" box="[615,745,272,296]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1502" phylum="Arthropoda" rank="species" species="nitefacta">
<emphasis id="A5AAD176FFCF306E2076FE91FD1CFEA4" box="[615,745,272,296]" italics="true" pageId="27" pageNumber="1502">C. nitefacta</emphasis>
</taxonomicName>
it also differs in having densely punctured upper side of body.
</paragraph>
<paragraph id="97610D64FFCF306E2283FEFBFEE4FE1E" blockId="27.[146,273,378,402]" box="[146,273,378,402]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FEFBFEE4FE1E" box="[146,273,378,402]" italics="true" pageId="27" pageNumber="1502">Description</emphasis>
</paragraph>
<paragraph id="97610D64FFCF306E2283FE20FBBAFE55" blockId="27.[146,1103,417,473]" pageId="27" pageNumber="1502">
<typeStatus id="4865B3C6FFCF306E2283FE20FF0EFE35" box="[146,251,417,441]" pageId="27" pageNumber="1502" type="holotype">Holotype</typeStatus>
<emphasis id="A5AAD176FFCF306E2314FE24FEE2FE35" box="[261,279,421,441]" italics="true" pageId="27" pageNumber="1502">„</emphasis>
. Strongly convex, dark-brown with slightly lighter elytra, opaque beetle (
<figureCitation id="0FE511E1FFCF306E228BFE40FEF3FE55" box="[154,262,449,473]" captionStart="Figures 15–17" captionStartId="9.[146,222,1409,1431]" captionTargetBox="[152,1096,178,1370]" captionTargetId="figure-62@9.[146,1104,165,1372]" captionTargetPageId="9" captionText="Figures 15–17. Coptorhina spp., general view. 15, C. auspicata, Cinergy Game farm, RSA; 16, C. excavata sp. n., holotype; 17, C. nitidipennis, Cinergy Game farm, RSA." figureDoi="http://doi.org/10.5281/zenodo.5219403" httpUri="https://zenodo.org/record/5219403/files/figure.png" pageId="27" pageNumber="1502">Figure 16</figureCitation>
). Body length 20.5 mm, width 14.0 mm. Dorsal surface without visible setae.
</paragraph>
<paragraph id="97610D64FFCF306E2283FD8DFBEDFD4F" blockId="27.[146,1103,524,707]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FD8DFF3BFDA8" box="[146,206,524,548]" italics="true" pageId="27" pageNumber="1502">Head</emphasis>
. Clypeus with protruding anterior angles separated by deep sinuation (
<figureCitation id="0FE511E1FFCF306E228BFDADFF0FFDC8" box="[154,250,556,580]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="27" pageNumber="1502">Figure 4</figureCitation>
). Sinuation right-angled to broadly rounded. Protruding angles acute to narrowly rounded apically, with short longitudinal carinae on the upper side. Genae obtusely rounded. Genal sutures feebly distinct. Frontal suture feebly visible, broadly interrupted in the middle. Head very densely and evenly punctuate, some punctures adjacent and sculpture appears rugose. Antennal clubs dark-brown.
</paragraph>
<paragraph id="97610D64FFCF306E2283FD77FD1FFC21" blockId="27.[146,1103,757,941]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FD77FEF4FC82" box="[146,257,758,782]" italics="true" pageId="27" pageNumber="1502">Pronotum</emphasis>
. Trapezoidal, two times wider than long. Anterior margin and base finely bordered. Sides somewhat undulate. Lateral border very wide near the middle and tapering basally and apically. Pronotum divided into larger anterior part and smaller posterior part by a transversal carina. The carina is almost as wide as pronotum, very narrowly interrupted in the middle, serrate laterally. Anterior part of pronotum rugose to granulate, very deeply excavated laterally.
</paragraph>
<paragraph id="97610D64FFCF306E2283FC64FC13FBD0" blockId="27.[146,1103,997,1116]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FC64FF22FC71" box="[146,215,997,1021]" italics="true" pageId="27" pageNumber="1502">Elytra</emphasis>
. Colour brown, somewhat paler than in pronotum and head. Striae fine but distinct, punctate (punctures separated by 3–4 puncture diameter). Intervals flat, very densely punctuate, punctures separated by about 1 puncture diameter, sometimes almost adjacent; their margins sometimes indistinct especially in apical part.
</paragraph>
<paragraph id="97610D64FFCF306E2283FB15FD9EFB60" blockId="27.[146,1103,1172,1260]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FB15FEF4FB20" box="[146,257,1172,1196]" italics="true" pageId="27" pageNumber="1502">Underside</emphasis>
. Pygidium with fine border, its disc densely punctate. Abdominal and thoracic sternites coarsely punctate except for disc of metasternum which is very sparsely to almost indistinctly punctuate.
</paragraph>
<paragraph id="97610D64FFCF306E2283FAA5FC8CFAB7" blockId="27.[146,889,1315,1340]" box="[146,889,1315,1340]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FAA5FF09FAB0" box="[146,252,1316,1340]" italics="true" pageId="27" pageNumber="1502">Aedeagus</emphasis>
. Parameres with widely rounded apices in lateral view.
</paragraph>
<paragraph id="97610D64FFCF306E2283FAF2FEFFFA07" blockId="27.[146,266,1395,1419]" box="[146,266,1395,1419]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283FAF2FEFFFA07" box="[146,266,1395,1419]" italics="true" pageId="27" pageNumber="1502">Variability</emphasis>
</paragraph>
<paragraph id="97610D64FFCF306E2283FA1AFDF3F99F" blockId="27.[146,1103,1435,1555]" pageId="27" pageNumber="1502">
Body size of
<typeStatus id="4865B3C6FFCF306E2322FA1AFE50FA3F" box="[307,421,1435,1459]" pageId="27" pageNumber="1502" type="paratype">paratypes</typeStatus>
: length 10.5–20.5 mm, width 7.5–14.0 mm. Except for the body size variability the beetles differ in shape and sculpture of pronotum. Smaller specimens have less developed transversal pronotal carina and lesser excavated anterolateral sides of pronotum.
</paragraph>
<paragraph id="97610D64FFCF306E2283F9CAFEEDF9EF" blockId="27.[146,280,1611,1635]" box="[146,280,1611,1635]" pageId="27" pageNumber="1502">
<emphasis id="A5AAD176FFCF306E2283F9CAFEEDF9EF" box="[146,280,1611,1635]" italics="true" pageId="27" pageNumber="1502">Distribution</emphasis>
</paragraph>
<paragraph id="97610D64FFCF306E2283F9F3FD7FF926" blockId="27.[146,1102,1650,1706]" pageId="27" pageNumber="1502">
The species is known from a few localities in
<collectingCountry id="EFC94DF4FFCF306E2089F9F3FCD9F906" box="[664,812,1650,1674]" name="South Africa" pageId="27" pageNumber="1502">South Africa</collectingCountry>
(mostly in the
<collectingRegion id="551AC386FFCF306E21CBF9F3FF02F926" country="South Africa" name="Free State" pageId="27" pageNumber="1502">Free State Province</collectingRegion>
) and Western
<collectingCountry id="EFC94DF4FFCF306E23B3F913FE0BF926" box="[418,510,1682,1706]" name="Lesotho" pageId="27" pageNumber="1502">Lesotho</collectingCountry>
(
<figureCitation id="0FE511E1FFCF306E201EF913FD8BF926" box="[527,638,1682,1706]" captionStart="Figures 20–22" captionStartId="26.[213,289,1398,1420]" captionTargetBox="[213,1171,165,1362]" captionTargetId="figure-56@26.[213,1171,165,1362]" captionTargetPageId="26" captionText="Figures 20–22. Coptorhina spp., distribution map. 20, C. auspicata; 21, C. excavata sp. n.; 22, C. nitefacta." figureDoi="http://doi.org/10.5281/zenodo.5219409" httpUri="https://zenodo.org/record/5219409/files/figure.png" pageId="27" pageNumber="1502">Figure 20</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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