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<document id="7D6CF3F841691D3A7D7F52577E97FF7B" ID-DOI="10.5281/zenodo.194870" ID-GBIF-Dataset="3e0d4d97-b5d0-472e-bc4d-87134c7565b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="194870" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460218486540" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2010" docId="1F53FD42D67BFF8EFF1971DBFBFEFF34" docLanguage="en" docName="zt02438p036.pdf" docOrigin="Zootaxa 2438" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptobasis Selys 1877" docType="treatment" docVersion="13" lastPageNumber="4" masterDocId="E36A853AD679FF8AFF8E703CFF92FFC8" masterDocTitle="Redefinition of Leptobasis Selys with the synonymy of Chrysobasis Rácenis and description of L. mauffrayi sp. nov. from Peru (Odonata: Coenagrionidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="3" updateTime="1698237345539" updateUser="plazi">
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<mods:title id="FC71D131E5ED614D89320318B2F53736">Redefinition of Leptobasis Selys with the synonymy of Chrysobasis Rácenis and description of L. mauffrayi sp. nov. from Peru (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="E2B105C32D38F93089FD2F3445A891A7">Garrison, Rosser W.</mods:namePart>
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<mods:namePart id="D08B6817DFDC86E53A7857E8BD11ACD2">Ellenrieder, Natalia Von</mods:namePart>
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<mods:date id="38ED047EF6D6CB3CBFC89A6A3FE714E7">2010</mods:date>
<mods:detail id="4B5F5214F23367C1C8E3E3A11DF54A8A" type="volume">
<mods:number id="5AB154DD1318C7771D4DE70529F93B5A">2438</mods:number>
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<treatment id="1F53FD42D67BFF8EFF1971DBFBFEFF34" ID-DOI="http://doi.org/10.5281/zenodo.6203220" ID-GBIF-Taxon="119384939" ID-Zenodo-Dep="6203220" LSID="urn:lsid:plazi:treatment:1F53FD42D67BFF8EFF1971DBFBFEFF34" httpUri="http://treatment.plazi.org/id/1F53FD42D67BFF8EFF1971DBFBFEFF34" lastPageId="4" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="DFE01FDFD67BFF88FF1971DBFE32FDC9" box="[151,416,487,513]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="97454C54D67BFF88FF1971DBFE32FDC9" blockId="2.[151,416,487,513]" box="[151,416,487,513]" pageId="2" pageNumber="3">
<heading id="CC0DFB38D67BFF88FF1971DBFE32FDC9" bold="true" box="[151,416,487,513]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="50FA37D7D67BFF88FF1971DBFE32FDC9" ID-CoL="62X9T" authority="Selys 1877" authorityName="Selys" authorityYear="1877" box="[151,416,487,513]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FF1971DBFE32FDC9" bold="true" box="[151,416,487,513]" pageId="2" pageNumber="3">
<emphasis id="A58E9046D67BFF88FF1971DBFE8AFDC9" bold="true" box="[151,280,487,513]" italics="true" pageId="2" pageNumber="3">Leptobasis</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FE9171DBFE32FDC9" author="Selys" box="[287,416,487,513]" pageId="2" pageNumber="16" refString="Selys, E. (1877) Synopsis des Agrionines, 5 me legion: Agrion (suite et fin). Les genres Telebasis, Argiocnemis et Hemiphlebia. Bulletin de l'Academie royale de Belgique, (2) 43, 97 - 159 (1 - 65 reprint)." type="journal article" year="1877">Selys 1877</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DFE01FDFD67BFF88FF19720CFE66FDAF" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="97454C54D67BFF88FF19720CFE66FDAF" blockId="2.[151,658,560,615]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="B7EA6B7AD67BFF88FF19720CFE66FDAF" pageId="2" pageNumber="3">
<taxonomicName id="50FA37D7D67BFF88FF19720CFEC4FD8F" ID-CoL="62X9T" authority="Hagen" authorityName="Hagen" box="[151,342,560,583]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FF19720CFE99FD8F" box="[151,267,560,583]" italics="true" pageId="2" pageNumber="3">Leptobasis</emphasis>
Hagen
</taxonomicName>
<emphasis id="A58E9046D67BFF88FED3720CFEE0FD8F" box="[349,370,560,583]" italics="true" pageId="2" pageNumber="3">in</emphasis>
<treatmentCitation id="165B6A45D67BFF88FEF7720CFD8AFD8F" author="Selys" box="[377,536,560,583]" page="99" pageId="2" pageNumber="3" year="1877">
<bibRefCitation id="F36B31A5D67BFF88FEF7720CFE60FD8F" author="Selys" box="[377,498,560,583]" pageId="2" pageNumber="16" refString="Selys, E. (1877) Synopsis des Agrionines, 5 me legion: Agrion (suite et fin). Les genres Telebasis, Argiocnemis et Hemiphlebia. Bulletin de l'Academie royale de Belgique, (2) 43, 97 - 159 (1 - 65 reprint)." type="journal article" year="1877">Selys 1877</bibRefCitation>
: 99
</treatmentCitation>
(5 reprint).
<emphasis id="A58E9046D67BFF88FF19726CFF3BFDAF" box="[151,169,592,615]" italics="true" pageId="2" pageNumber="3">=</emphasis>
<taxonomicName id="50FA37D7D67BFF88FF21726CFE62FDAF" ID-CoL="9WZ8S" authority="Forster 1906: 15" authorityName="Forster" authorityPageNumber="15" authorityYear="1906" box="[175,496,592,615]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FF21726CFEA4FDAF" box="[175,310,592,615]" italics="true" pageId="2" pageNumber="3">Hylaeagrion</emphasis>
<treatmentCitation id="165B6A45D67BFF88FEB3726CFE62FDAF" author="Forster" box="[317,496,592,615]" page="15" pageId="2" pageNumber="3" year="1906">
<bibRefCitation id="F36B31A5D67BFF88FEB3726CFE5AFDAF" author="Forster" box="[317,456,592,615]" pageId="2" pageNumber="14" refString="Forster, F. (1906) Neotropische Libellen. III. Entomologisches Wochenblatt (Insektenbosse), 23, 15 - 16." type="journal article" year="1906">Förster 1906</bibRefCitation>
: 15
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67BFF88FF1972ACFEA3FBCF" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="97454C54D67BFF88FF1972ACFACFFD6F" blockId="2.[151,1373,656,679]" box="[151,1373,656,679]" pageId="2" pageNumber="3">
<typeStatus id="4841F2F6D67BFF88FF1972ACFF5EFD6F" box="[151,204,656,679]" pageId="2" pageNumber="3">Type</typeStatus>
species:
<taxonomicName id="50FA37D7D67BFF88FEA372ACFDFCFD6F" authority="Forster 1906" authorityName="Forster" authorityYear="1906" box="[301,622,656,679]" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FEA372ACFE4BFD6F" box="[301,473,656,679]" italics="true" pageId="2" pageNumber="3">Agrion croceum</emphasis>
:
<bibRefCitation id="F36B31A5D67BFF88FE6972ACFDFCFD6F" author="Forster" box="[487,622,656,679]" pageId="2" pageNumber="14" refString="Forster, F. (1906) Neotropische Libellen. III. Entomologisches Wochenblatt (Insektenbosse), 23, 15 - 16." type="journal article" year="1906">Förster 1906</bibRefCitation>
</taxonomicName>
(
<emphasis id="A58E9046D67BFF88FDF072ACFD31FD6F" box="[638,675,656,679]" italics="true" pageId="2" pageNumber="3">nec</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FD2472ACFCF2FD6F" author="Burmeister" box="[682,864,656,679]" pageId="2" pageNumber="14" refString="Burmeister, H. (1839) Handbuch der Entomologie. Zweiter Band. Besondere Entomologie. Zweite Abtheilung. Kaukerfe. Gymnognatha. (Zweite Halfte; vulgo Neuroptera). T. C. F. Enslin, Berlin." type="book" year="1839">Burmeister 1839</bibRefCitation>
), subsequently designated by
<bibRefCitation id="F36B31A5D67BFF88FB1072ACFACBFD6F" author="Williamson" box="[1182,1369,656,679]" pageId="2" pageNumber="17" refString="Williamson, E. B. (1917) Some species of Leptagrion with descriptions of a new genus and a new species (Odonata). Entomological News, 28, 241 - 255." type="journal article" year="1917">Williamson 1917</bibRefCitation>
.
</paragraph>
<paragraph id="97454C54D67BFF88FF2F72ECFEA3FBCF" blockId="2.[161,1436,720,1031]" pageId="2" pageNumber="3">
(Note: All catalogs (
<bibRefCitation id="F36B31A5D67BFF88FE0B72ECFDE8FD2F" author="Davies" box="[389,634,720,743]" pageId="2" pageNumber="14" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The dragonflies of the world: a systematic list of the extant species of Odonata. Volume 1. Zygoptera, Anisozygoptera. Societas Internationalis Odonatologica Rapid Communications (Supplement), 3, 1 - 127." type="journal article" year="1984">Davies &amp; Tobin 1984</bibRefCitation>
: 78;
<bibRefCitation id="F36B31A5D67BFF88FD3A72ECFCD8FD2F" author="Bridges" box="[692,842,720,743]" pageId="2" pageNumber="13" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group, and species-group names of the Odonata of the world (third edition). Bridges, Urbana, Illinois, 905 pp." type="book" year="1994">Bridges 1994</bibRefCitation>
:
<date id="E3446A94D67BFF88FCD672ECFC04FD2F" box="[856,918,720,743]" pageId="2" pageNumber="3" value="1922-03">III.22</date>
,
<bibRefCitation id="F36B31A5D67BFF88FC2B72ECFBC8FD2F" author="Steinmann" box="[933,1114,720,743]" pageId="2" pageNumber="16" refString="Steinmann, H. (1997). World catalogue of Odonata. Volume I. Zygoptera. In: Wermuth, H. &amp; Fischer, M. (Eds.), Das Tierreich. The Animal Kingdom. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen, Volume 110. Walter de Gruyter, Berlin, 500 pp." type="book" year="1997">Steinmann 1997</bibRefCitation>
: 290;
<bibRefCitation id="F36B31A5D67BFF88FB2F72ECFAB1FD2F" author="Tsuda" box="[1185,1315,720,743]" pageId="2" pageNumber="16" refString="Tsuda, S. (2000) A distributional list of world Odonata 2000. Privately published, Osaka, 430 pp." type="book" year="2000">Tsuda 2000</bibRefCitation>
: 208) treat
<taxonomicName id="50FA37D7D67BFF88FF4972CCFEDCFCCF" box="[199,334,752,775]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FF4972CCFEDCFCCF" box="[199,334,752,775]" italics="true" pageId="2" pageNumber="3">Hylaeagrion</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="50FA37D7D67BFF88FD8072CCFCD7FCCF" authority="Williamson 1917" authorityName="Williamson" authorityYear="1917" box="[526,837,752,775]" class="Insecta" family="Coenagrionidae" genus="Aeolagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FD8072CCFD14FCCF" box="[526,646,752,775]" italics="true" pageId="2" pageNumber="3">Aeolagrion</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FD0372CCFCD7FCCF" author="Williamson" box="[653,837,752,775]" pageId="2" pageNumber="17" refString="Williamson, E. B. (1917) Some species of Leptagrion with descriptions of a new genus and a new species (Odonata). Entomological News, 28, 241 - 255." type="journal article" year="1917">Williamson 1917</bibRefCitation>
</taxonomicName>
with its
<typeStatus id="4841F2F6D67BFF88FC2B72CCFC41FCCF" box="[933,979,752,775]" pageId="2" pageNumber="3">type</typeStatus>
species
<taxonomicName id="50FA37D7D67BFF88FBA172CCFA0EFCCF" authority="Forster 1906" authorityName="Forster" authorityYear="1906" box="[1071,1436,752,775]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="argenteolineatum">
<emphasis id="A58E9046D67BFF88FBA172CCFA9EFCCF" box="[1071,1292,752,775]" italics="true" pageId="2" pageNumber="3">H. argenteolineatum</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FA9A72CCFA0EFCCF" author="Forster" box="[1300,1436,752,775]" pageId="2" pageNumber="14" refString="Forster, F. (1906) Neotropische Libellen. III. Entomologisches Wochenblatt (Insektenbosse), 23, 15 - 16." type="journal article" year="1906">Förster 1906</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="50FA37D7D67BFF88FF69732CFDD9FCEF" authority="Burmeister 1839" authorityName="Burmeister" authorityYear="1839" box="[231,587,784,807]" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dorsale">
<emphasis id="A58E9046D67BFF88FF69732CFE18FCEF" box="[231,394,784,807]" italics="true" pageId="2" pageNumber="3">Agrion dorsale</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FE1D732CFDD9FCEF" author="Burmeister" box="[403,587,784,807]" pageId="2" pageNumber="14" refString="Burmeister, H. (1839) Handbuch der Entomologie. Zweiter Band. Besondere Entomologie. Zweite Abtheilung. Kaukerfe. Gymnognatha. (Zweite Halfte; vulgo Neuroptera). T. C. F. Enslin, Berlin." type="book" year="1839">Burmeister 1839</bibRefCitation>
</taxonomicName>
). Förster described
<taxonomicName id="50FA37D7D67BFF88FCAC732CFC3BFCEF" box="[802,937,784,807]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FCAC732CFC3BFCEF" box="[802,937,784,807]" italics="true" pageId="2" pageNumber="3">Hylaeagrion</emphasis>
</taxonomicName>
including two species, &quot;
<taxonomicName id="50FA37D7D67BFF88FB38732CFABEFCEF" box="[1206,1324,784,807]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FB38732CFABEFCEF" box="[1206,1324,784,807]" italics="true" pageId="2" pageNumber="3">Leptagrion</emphasis>
</taxonomicName>
?
<taxonomicName id="50FA37D7D67BFF88FACF732CFE11FC8F" authority="Selys-Burmeister" authorityName="Selys-Burmeister" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FACF732CFA0EFCEF" box="[1345,1436,784,807]" italics="true" pageId="2" pageNumber="3">croceum</emphasis>
Selys-Burmeister
</taxonomicName>
&quot; and
<taxonomicName id="50FA37D7D67BFF88FE4B730CFC83FC8F" box="[453,785,816,839]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="argenteolineatum">
<emphasis id="A58E9046D67BFF88FE4B730CFC83FC8F" box="[453,785,816,839]" italics="true" pageId="2" pageNumber="3">Hylaeagrion argenteolineatum</emphasis>
</taxonomicName>
(sp. nov.) both from
<collectingCountry id="EFED0CC4D67BFF88FB8C730CFBF2FC8F" box="[1026,1120,816,839]" name="Suriname" pageId="2" pageNumber="3">Surinam</collectingCountry>
; he did not designate a
<typeStatus id="4841F2F6D67BFF88FAE0730CFA0EFC8F" box="[1390,1436,816,839]" pageId="2" pageNumber="3">type</typeStatus>
species.
<bibRefCitation id="F36B31A5D67BFF88FEA8736CFDBDFCAF" author="Williamson" box="[294,559,848,871]" pageId="2" pageNumber="17" refString="Williamson, E. B. (1917) Some species of Leptagrion with descriptions of a new genus and a new species (Odonata). Entomological News, 28, 241 - 255." type="journal article" year="1917">Williamson (1917: 241)</bibRefCitation>
considered
<taxonomicName id="50FA37D7D67BFF88FD37736CFC04FCAF" box="[697,918,848,871]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FD37736CFC04FCAF" box="[697,918,848,871]" italics="true" pageId="2" pageNumber="3">Leptagrion croceum</emphasis>
</taxonomicName>
(=
<taxonomicName id="50FA37D7D67BFF88FC4B736CFAABFCAF" authority="Burmeister 1839" authorityName="Burmeister" authorityYear="1839" box="[965,1337,848,871]" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FC4B736CFBE7FCAF" box="[965,1141,848,871]" italics="true" pageId="2" pageNumber="3">Agrion croceum</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FB0F736CFAABFCAF" author="Burmeister" box="[1153,1337,848,871]" pageId="2" pageNumber="14" refString="Burmeister, H. (1839) Handbuch der Entomologie. Zweiter Band. Besondere Entomologie. Zweite Abtheilung. Kaukerfe. Gymnognatha. (Zweite Halfte; vulgo Neuroptera). T. C. F. Enslin, Berlin." type="book" year="1839">Burmeister 1839</bibRefCitation>
</taxonomicName>
) as
<typeStatus id="4841F2F6D67BFF88FAE0736CFA0EFCAF" box="[1390,1436,848,871]" pageId="2" pageNumber="3">type</typeStatus>
species (&quot;The second new genus proposed by Foerster is
<taxonomicName id="50FA37D7D67BFF88FCD3734CFC76FC4F" box="[861,996,880,903]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FCD3734CFC76FC4F" box="[861,996,880,903]" italics="true" pageId="2" pageNumber="3">Hylaeagrion</emphasis>
</taxonomicName>
, of which
<taxonomicName id="50FA37D7D67BFF88FBE6734CFAD5FC4F" box="[1128,1351,880,903]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FBE6734CFAD5FC4F" box="[1128,1351,880,903]" italics="true" pageId="2" pageNumber="3">Leptagrion croceum</emphasis>
</taxonomicName>
, of his determination must be the
<typeStatus id="4841F2F6D67BFF88FE7D73ACFDB1FC6F" box="[499,547,912,935]" pageId="2" pageNumber="3">type</typeStatus>
, congeneric with which, according to Foerster, is his new
<taxonomicName id="50FA37D7D67BFF88FB3B73ACFA07FC6F" box="[1205,1429,912,935]" class="Insecta" family="Coenagrionidae" genus="Hylaeagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="argenteolineatum">
<emphasis id="A58E9046D67BFF88FB3B73ACFA07FC6F" box="[1205,1429,912,935]" italics="true" pageId="2" pageNumber="3">H. argenteolineatum</emphasis>
</taxonomicName>
, known from the male only…&quot;). From Förster's description, Williamson believed
<taxonomicName id="50FA37D7D67BFF88FBB6738CFAF4FC0F" authority="Forster" authorityName="Forster" box="[1080,1382,944,967]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FBB6738CFA80FC0F" box="[1080,1298,944,967]" italics="true" pageId="2" pageNumber="3">Leptagrion croceum</emphasis>
Förster
</taxonomicName>
(
<emphasis id="A58E9046D67BFF88FAF9738CFA0EFC0F" box="[1399,1436,944,967]" italics="true" pageId="2" pageNumber="3">nec</emphasis>
Burmeister) to be a
<taxonomicName id="50FA37D7D67BFF88FE3F73ECFDB7FC2F" box="[433,549,976,999]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FE3F73ECFDB7FC2F" box="[433,549,976,999]" italics="true" pageId="2" pageNumber="3">Leptobasis</emphasis>
</taxonomicName>
—probably
<taxonomicName id="50FA37D7D67BFF88FD2473ECFCBEFC2F" box="[682,812,976,999]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67BFF88FD2473ECFCBEFC2F" box="[682,812,976,999]" italics="true" pageId="2" pageNumber="3">L. vacillans</emphasis>
</taxonomicName>
—based on illustrations of Burmeister's
<typeStatus id="4841F2F6D67BFF88FB7B73ECFAB1FC2F" box="[1269,1315,976,999]" pageId="2" pageNumber="3">type</typeStatus>
of
<taxonomicName id="50FA37D7D67BFF88FADC73ECFEB0FBCF" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="croceum">
<emphasis id="A58E9046D67BFF88FADC73ECFEB0FBCF" italics="true" pageId="2" pageNumber="3">Agrion croceum</emphasis>
</taxonomicName>
.)
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67BFF88FF13740CFEB8FBAF" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="97454C54D67BFF88FF13740CFEB8FBAF" blockId="2.[157,1435,1072,1127]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="B7EA6B7AD67BFF88FF13740CFEB8FBAF" pageId="2" pageNumber="3">
=
<taxonomicName id="50FA37D7D67BFF88FF3D740CFE67FB8F" authority="Racenis 1959: 55" authorityName="Racenis" authorityPageNumber="55" authorityYear="1959" box="[179,501,1072,1095]" class="Insecta" family="Coenagrionidae" genus="Chrysobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FF3D740CFEA5FB8F" box="[179,311,1072,1095]" italics="true" pageId="2" pageNumber="3">Chrysobasis</emphasis>
<treatmentCitation id="165B6A45D67BFF88FEB3740CFE67FB8F" author="Racenis" box="[317,501,1072,1095]" page="55" pageId="2" pageNumber="3" year="1959">
<bibRefCitation id="F36B31A5D67BFF88FEB3740CFE42FB8F" author="Racenis" box="[317,464,1072,1095]" pageId="2" pageNumber="16" refString="Racenis, J. (1959) Zwei neue Gattungen und Arten der Familie Coenagrionidae (Odonata) aus Venezuela. Senckenberger Biologie, 40 (112), 55 - 61." type="journal article" year="1959">Rácenis 1959</bibRefCitation>
: 55
</treatmentCitation>
</taxonomicName>
(description,
<typeStatus id="4841F2F6D67BFF88FD06740CFD24FB8F" box="[648,694,1072,1095]" pageId="2" pageNumber="3">type</typeStatus>
species
<taxonomicName id="50FA37D7D67BFF88FC81740CFB05FB8F" ID-CoL="9WYBS" authority="Racenis 1959" authorityName="Racenis" authorityYear="1959" box="[783,1175,1072,1095]" class="Insecta" family="Coenagrionidae" genus="Chrysobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="buchholzi">
<emphasis id="A58E9046D67BFF88FC81740CFB93FB8F" box="[783,1025,1072,1095]" italics="true" pageId="2" pageNumber="3">Chrysobasis buchholzi</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FB89740CFB05FB8F" author="Racenis" box="[1031,1175,1072,1095]" pageId="2" pageNumber="16" refString="Racenis, J. (1959) Zwei neue Gattungen und Arten der Familie Coenagrionidae (Odonata) aus Venezuela. Senckenberger Biologie, 40 (112), 55 - 61." type="journal article" year="1959">Rácenis 1959</bibRefCitation>
</taxonomicName>
by original designation)
<emphasis id="A58E9046D67BFF88FF497473FEB8FBAF" bold="true" box="[199,298,1103,1127]" pageId="2" pageNumber="3">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67BFF88FF1974A9FC72F82C" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="97454C54D67BFF88FF1974A9FAB7FB67" blockId="2.[151,1437,1173,2020]" box="[151,1317,1173,1199]" pageId="2" pageNumber="3">
<emphasis id="A58E9046D67BFF88FF1974A9FEABFB67" bold="true" box="[151,313,1173,1199]" pageId="2" pageNumber="3">
<typeStatus id="4841F2F6D67BFF88FF1974A9FF46FB67" box="[151,212,1173,1199]" pageId="2" pageNumber="3">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="50FA37D7D67BFF88FECE74A9FD16FB67" authority="Hagen" authorityName="Hagen" box="[320,644,1173,1199]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67BFF88FECE74A9FDA3FB66" box="[320,561,1173,1198]" italics="true" pageId="2" pageNumber="3">Leptobasis vacillans</emphasis>
Hagen
</taxonomicName>
<emphasis id="A58E9046D67BFF88FD0274A9FD31FB66" box="[652,675,1173,1198]" italics="true" pageId="2" pageNumber="3">in</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FD2474A9FCBCFB67" author="Selys" box="[682,814,1173,1199]" pageId="2" pageNumber="16" refString="Selys, E. (1877) Synopsis des Agrionines, 5 me legion: Agrion (suite et fin). Les genres Telebasis, Argiocnemis et Hemiphlebia. Bulletin de l'Academie royale de Belgique, (2) 43, 97 - 159 (1 - 65 reprint)." type="journal article" year="1877">Selys 1877</bibRefCitation>
, subsequently designated by
<bibRefCitation id="F36B31A5D67BFF88FB0D74A9FAB3FB67" author="Kirby" box="[1155,1313,1173,1199]" pageId="2" pageNumber="15" refString="Kirby, W. F. (1890) A synonymic catalogue of Neuroptera Odonata, or dragonflies. With an appendix of fossil species. Gurney &amp; Jackson, London, 202 pp." type="book" year="1890">Kirby (1890)</bibRefCitation>
.
</paragraph>
<paragraph id="97454C54D67BFF88FF487486FB14FAD7" blockId="2.[151,1437,1173,2020]" pageId="2" pageNumber="3">
<emphasis id="A58E9046D67BFF88FF487486FE78FB1C" bold="true" box="[198,490,1210,1236]" pageId="2" pageNumber="3">Other species included.</emphasis>
<taxonomicName id="50FA37D7D67BFF88FE7C7487FC37FB1C" authority="Racenis 1959" authorityName="Racenis" authorityYear="1959" box="[498,933,1210,1236]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="buchholzi" status="comb. nov.">
<emphasis id="A58E9046D67BFF88FE7C7487FD79FB1C" box="[498,747,1211,1236]" italics="true" pageId="2" pageNumber="3">Leptobasis buchholzi</emphasis>
(
<bibRefCitation id="F36B31A5D67BFF88FD757486FC0CFB1C" author="Racenis" box="[763,926,1210,1236]" pageId="2" pageNumber="16" refString="Racenis, J. (1959) Zwei neue Gattungen und Arten der Familie Coenagrionidae (Odonata) aus Venezuela. Senckenberger Biologie, 40 (112), 55 - 61." type="journal article" year="1959">Rácenis 1959</bibRefCitation>
)
</taxonomicName>
<emphasis id="A58E9046D67BFF88FC207486FBA7FB1C" bold="true" box="[942,1077,1210,1236]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="BEBD2D3DD67BFF88FC207486FBA7FB1C" box="[942,1077,1210,1236]" pageId="2" pageNumber="3">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="50FA37D7D67BFF88FBCA7487FAE5FB1C" authority="Alayo 1968" authorityName="Alayo" authorityYear="1968" box="[1092,1399,1210,1236]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67BFF88FBCA7487FB71FB1C" box="[1092,1251,1211,1236]" italics="true" pageId="2" pageNumber="3">L. candelaria</emphasis>
Alayo 1968
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FA0A7487FE59FB32" authority="Paulson 2009" authorityName="Paulson" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67BFF88FA0A7487FE8DFB31" italics="true" pageId="2" pageNumber="3">L. guanacaste</emphasis>
Paulson 2009
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FE5674DCFC87FB32" authority="Donnelly 1967" authorityName="Donnelly" authorityYear="1967" box="[472,789,1248,1274]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lucifer" status="comb. nov.">
<emphasis id="A58E9046D67BFF88FE5674DCFDD5FB31" box="[472,583,1248,1273]" italics="true" pageId="2" pageNumber="3">L. lucifer</emphasis>
(
<bibRefCitation id="F36B31A5D67BFF88FDD974DCFC9FFB32" author="Donnelly" box="[599,781,1248,1274]" pageId="2" pageNumber="14" refString="Donnelly, T. W. (1967) The discovery of Chrysobasis in Central America, with the description of a new species (Odonata: Coenagrionidae). Florida Entomologist, 50 (1), 47 - 52." type="journal article" year="1967">Donnelly 1967</bibRefCitation>
)
</taxonomicName>
<emphasis id="A58E9046D67BFF88FC9374E3FC34FB31" bold="true" box="[797,934,1247,1273]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="BEBD2D3DD67BFF88FC9374E3FC34FB31" box="[797,934,1247,1273]" pageId="2" pageNumber="3">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="50FA37D7D67BFF88FC3874DCFB29FB32" authority="Garrison" authorityName="Garrison" box="[950,1211,1248,1274]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="A58E9046D67BFF88FC3874DCFBD8FB31" box="[950,1098,1248,1273]" italics="true" pageId="2" pageNumber="3">L. mauffrayi</emphasis>
Garrison
</taxonomicName>
&amp; von
<bibRefCitation id="F36B31A5D67BFF88FA9974DCFF40FAD7" author="Ellenrieder" pageId="2" pageNumber="15" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2009) Redefinition of Mesoleptobasis Sjostedt 1918 with the inclusion of Metaleptobasis [sic. Mesoleptobasis] cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa, 2145, 47 - 68." type="journal article" year="2009">Ellenrieder 2009</bibRefCitation>
<emphasis id="A58E9046D67BFF88FF577539FEA4FAD7" bold="true" box="[217,310,1285,1311]" pageId="2" pageNumber="3">sp. nov.</emphasis>
,
<taxonomicName id="50FA37D7D67BFF88FECB7539FC8FFAD7" authority="Gonzalez-Soriano 2002" authorityName="Gonzalez-Soriano" authorityYear="2002" box="[325,797,1285,1311]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="melinogaster">
<emphasis id="A58E9046D67BFF88FECB7539FE6FFAD6" box="[325,509,1285,1310]" italics="true" pageId="2" pageNumber="3">L. melinogaster</emphasis>
<bibRefCitation id="F36B31A5D67BFF88FD8A7539FC8FFAD7" author="Gonzalez-Soriano" box="[516,797,1285,1311]" pageId="2" pageNumber="15" refString="Gonzalez-Soriano, E. (2002) Leptobasis melinogaster spec. nov., a new species from Mexico (Zygoptera: Coenagrionidae). Odonatologica, 31 (2), 181 - 185." type="journal article" year="2002">González-Soriano 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FCA67539FB10FAD7" authority="Williamson 1915" authorityName="Williamson" authorityYear="1915" box="[808,1154,1285,1311]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="raineyi">
<emphasis id="A58E9046D67BFF88FCA67539FC08FAD6" box="[808,922,1285,1310]" italics="true" pageId="2" pageNumber="3">L. raineyi</emphasis>
(
<bibRefCitation id="F36B31A5D67BFF88FC277539FBEBFAD7" author="Williamson" box="[937,1145,1285,1311]" pageId="2" pageNumber="17" refString="Williamson, E. B. (1915) Notes on neotropical dragonflies, or Odonata. Proceedings of the United States National Museum, 48 (2089), 601 - 638." type="journal article" year="1915">Williamson 1915</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="97454C54D67BFF88FF487516FCB4FA12" blockId="2.[151,1437,1173,2020]" pageId="2" pageNumber="3">
<bibRefCitation id="F36B31A5D67BFF88FF487516FEE7FA8C" author="Calvert" box="[198,373,1322,1348]" pageId="2" pageNumber="14" refString="Calvert, P. P. (1909) Contributions to a knowledge of the Odonata of the Neotropical region, exclusive of Mexico and Central America. Annals of the Carnegie Museum, 6 (3), 73 - 280." type="journal article" year="1909">Calvert (1909)</bibRefCitation>
described
<taxonomicName id="50FA37D7D67BFF88FE757517FC82FA8C" box="[507,784,1323,1348]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mammilaris">
<emphasis id="A58E9046D67BFF88FE757517FC82FA8C" box="[507,784,1323,1348]" italics="true" pageId="2" pageNumber="3">Leptobasis mammilaris</emphasis>
</taxonomicName>
based on two males (one incomplete) from Chapada,
<collectingCountry id="EFED0CC4D67BFF88FF19756CFF7BFAA2" box="[151,233,1360,1386]" name="Brazil" pageId="2" pageNumber="3">Brazil</collectingCountry>
, and a female (Rio de Janeiro) which he doubtfully assigned to this species.
<bibRefCitation id="F36B31A5D67BFF88FB44756CFA0FFAA2" author="Machado" box="[1226,1437,1360,1386]" pageId="2" pageNumber="15" refString="Machado, A. B. M. (2009) Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa, 2108, 1 - 36." type="journal article" year="2009">Machado (2009)</bibRefCitation>
transferred
<taxonomicName id="50FA37D7D67BFF88FE927549FE55FA46" box="[284,455,1397,1422]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mammilaris">
<emphasis id="A58E9046D67BFF88FE927549FE55FA46" box="[284,455,1397,1422]" italics="true" pageId="2" pageNumber="3">L. mammilaris</emphasis>
</taxonomicName>
to his new genus
<taxonomicName id="50FA37D7D67BFF88FD187549FCDAFA46" box="[662,840,1397,1422]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FD187549FCDAFA46" box="[662,840,1397,1422]" italics="true" pageId="2" pageNumber="3">Tuberculobasis</emphasis>
</taxonomicName>
, and
<bibRefCitation id="F36B31A5D67BFF88FC097549FBF0FA47" author="Williamson" box="[903,1122,1397,1423]" pageId="2" pageNumber="17" refString="Williamson, E. B. (1917) Some species of Leptagrion with descriptions of a new genus and a new species (Odonata). Entomological News, 28, 241 - 255." type="journal article" year="1917">Williamson (1917)</bibRefCitation>
and Garrison (in
<bibRefCitation id="F36B31A5D67BFF88FAA17549FF41FA7C" author="Machado" pageId="2" pageNumber="15" refString="Machado, A. B. M. (2009) Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae). Zootaxa, 2108, 1 - 36." type="journal article" year="2009">Machado 2009</bibRefCitation>
) both expressed doubt as to the identity of Calvert's female. Our examination of the female from Rio de Janeiro shows it to be a female
<taxonomicName id="50FA37D7D67BFF88FD8975FCFCB4FA12" authority="Selys." authorityName="Selys." box="[519,806,1472,1498]" class="Insecta" family="Coenagrionidae" genus="Telagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="longum">
<emphasis id="A58E9046D67BFF88FD8975FCFD4AFA11" box="[519,728,1472,1497]" italics="true" pageId="2" pageNumber="3">Telagrion longum</emphasis>
Selys.
</taxonomicName>
</paragraph>
<paragraph id="97454C54D67BFF88FF4875D9FC72F82C" blockId="2.[151,1437,1173,2020]" pageId="2" pageNumber="3">
<emphasis id="A58E9046D67BFF88FF4875D9FD9EFA37" bold="true" box="[198,524,1509,1535]" pageId="2" pageNumber="3">
Synonymy of
<taxonomicName id="50FA37D7D67BFF88FEFD75D9FD94FA37" box="[371,518,1509,1535]" class="Insecta" family="Coenagrionidae" genus="Chrysobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FEFD75D9FD94FA37" bold="true" box="[371,518,1509,1535]" italics="true" pageId="2" pageNumber="3">Chrysobasis</emphasis>
</taxonomicName>
.
</emphasis>
Of the characters proposed for
<taxonomicName id="50FA37D7D67BFF88FC0A75D9FB41FA37" authority="Racenis 1959" authorityName="Racenis" authorityYear="1959" box="[900,1235,1509,1535]" class="Insecta" family="Coenagrionidae" genus="Chrysobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FC0A75D9FB87FA36" box="[900,1045,1509,1534]" italics="true" pageId="2" pageNumber="3">Chrysobasis</emphasis>
(
<bibRefCitation id="F36B31A5D67BFF88FBAB75D9FB58FA37" author="Racenis" box="[1061,1226,1509,1535]" pageId="2" pageNumber="16" refString="Racenis, J. (1959) Zwei neue Gattungen und Arten der Familie Coenagrionidae (Odonata) aus Venezuela. Senckenberger Biologie, 40 (112), 55 - 61." type="journal article" year="1959">Rácenis 1959</bibRefCitation>
)
</taxonomicName>
, an equally short CuA is found in
<taxonomicName id="50FA37D7D67BFF88FED97637FDCEF9EC" box="[343,604,1547,1572]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67BFF88FED97637FDCEF9EC" box="[343,604,1547,1572]" italics="true" pageId="2" pageNumber="3">Leptobasis candelaria</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FDE47637FC9FF9EC" box="[618,781,1547,1572]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67BFF88FDE47637FC9FF9EC" box="[618,781,1547,1572]" italics="true" pageId="2" pageNumber="3">L. guanacaste</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FC927637FC40F9EC" box="[796,978,1547,1572]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="melinogaster">
<emphasis id="A58E9046D67BFF88FC927637FC40F9EC" box="[796,978,1547,1572]" italics="true" pageId="2" pageNumber="3">L. melinogaster</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FC6E7637FBC0F9EC" box="[992,1106,1547,1572]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="raineyi">
<emphasis id="A58E9046D67BFF88FC6E7637FBC0F9EC" box="[992,1106,1547,1572]" italics="true" pageId="2" pageNumber="3">L. raineyi</emphasis>
</taxonomicName>
, and often in
<taxonomicName id="50FA37D7D67BFF88FB7F7637FAEAF9EC" box="[1265,1400,1547,1572]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67BFF88FB7F7637FAEAF9EC" box="[1265,1400,1547,1572]" italics="true" pageId="2" pageNumber="3">L. vacillans</emphasis>
</taxonomicName>
. A short ventro-basal process is present on the cercus of
<taxonomicName id="50FA37D7D67BFF88FC92760CFC21F981" box="[796,947,1584,1609]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="buchholzi">
<emphasis id="A58E9046D67BFF88FC92760CFC21F981" box="[796,947,1584,1609]" italics="true" pageId="2" pageNumber="3">C. buchholzi</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67BFF88FC4A760CFB8CF982" box="[964,1054,1584,1610]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="2" pageNumber="3">Fig. 20</figureCitation>
a), which we observed similarly developed in
<taxonomicName id="50FA37D7D67BFF88FEB87669FE36F9A6" box="[310,420,1621,1646]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lucifer">
<emphasis id="A58E9046D67BFF88FEB87669FE36F9A6" box="[310,420,1621,1646]" italics="true" pageId="2" pageNumber="3">C. lucifer</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FE3A7669FDC0F9A6" box="[436,594,1621,1646]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67BFF88FE3A7669FDC0F9A6" box="[436,594,1621,1646]" italics="true" pageId="2" pageNumber="3">L. candelaria</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FDEF7669FC94F9A6" box="[609,774,1621,1646]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67BFF88FDEF7669FC94F9A6" box="[609,774,1621,1646]" italics="true" pageId="2" pageNumber="3">L. guanacaste</emphasis>
</taxonomicName>
, and
<taxonomicName id="50FA37D7D67BFF88FCC67669FB92F9A6" box="[840,1024,1621,1646]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="melinogaster">
<emphasis id="A58E9046D67BFF88FCC67669FB92F9A6" box="[840,1024,1621,1646]" italics="true" pageId="2" pageNumber="3">L. melinogaster</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67BFF88FB9F7669FBE0F9A7" box="[1041,1138,1621,1647]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
b–d, f), reduced to a very low bump in
<taxonomicName id="50FA37D7D67BFF88FEB77647FE5CF95C" box="[313,462,1659,1684]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="A58E9046D67BFF88FEB77647FE5CF95C" box="[313,462,1659,1684]" italics="true" pageId="2" pageNumber="3">L. mauffrayi</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67BFF88FE507647FDC6F95C" box="[478,596,1659,1684]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="raineyi">
<emphasis id="A58E9046D67BFF88FE507647FDC6F95C" box="[478,596,1659,1684]" italics="true" pageId="2" pageNumber="3">L. raineyi</emphasis>
</taxonomicName>
, and
<taxonomicName id="50FA37D7D67BFF88FD167647FCB4F95C" box="[664,806,1659,1684]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67BFF88FD167647FCB4F95C" box="[664,806,1659,1684]" italics="true" pageId="2" pageNumber="3">L. vacillans</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67BFF88FCB97646FC0FF95C" box="[823,925,1658,1684]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
e, g–h). The pair of lobe-like processes on sides of medial cleft of postero-dorsal margin of male S10 described by
<bibRefCitation id="F36B31A5D67BFF88FC53769CFB1CF972" author="Racenis" box="[989,1166,1696,1722]" pageId="2" pageNumber="16" refString="Racenis, J. (1959) Zwei neue Gattungen und Arten der Familie Coenagrionidae (Odonata) aus Venezuela. Senckenberger Biologie, 40 (112), 55 - 61." type="journal article" year="1959">Rácenis (1959)</bibRefCitation>
for
<taxonomicName id="50FA37D7D67BFF88FB30769CFAC0F971" box="[1214,1362,1696,1721]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="buchholzi">
<emphasis id="A58E9046D67BFF88FB30769CFAC0F971" box="[1214,1362,1696,1721]" italics="true" pageId="2" pageNumber="3">C. buchholzi</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67BFF88FAEF769CFF25F917" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
a, 21a) is present also, although less developed, in
<taxonomicName id="50FA37D7D67BFF88FC8A76F9FCE0F916" box="[772,882,1733,1758]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lucifer">
<emphasis id="A58E9046D67BFF88FC8A76F9FCE0F916" box="[772,882,1733,1758]" italics="true" pageId="2" pageNumber="3">C. lucifer</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67BFF88FC0D76F9FC76F917" box="[899,996,1733,1759]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
d, 21d), and even less developed in
<taxonomicName id="50FA37D7D67BFF88FA0A76F9FEA0F8CC" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="melinogaster">
<emphasis id="A58E9046D67BFF88FA0A76F9FEA0F8CC" italics="true" pageId="2" pageNumber="3">L. melinogaster</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67BFF88FECD76D6FE34F8CC" box="[323,422,1770,1796]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
f, 21f). Thus, none of these three characters, and no other combination of characters, can be employed to confidently diagnose two genera within this complex of species. Moreover, these eight species share a unique genital ligula morphology within
<taxonomicName id="50FA37D7D67BFF88FCB57709FC68F887" box="[827,1018,1845,1871]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
, indicating their close relationship: the distal segment has a pair of chitinized, flap-like, movable processes directed posteriorly (shared only with the nearctic genus
<taxonomicName id="50FA37D7D67BFF88FEF077BCFD1FF852" authority="Calvert" authorityName="Calvert" box="[382,653,1920,1946]" class="Insecta" family="Coenagrionidae" genus="Hesperagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FEF077BCFDB5F851" box="[382,551,1920,1945]" italics="true" pageId="2" pageNumber="3">Hesperagrion</emphasis>
Calvert
</taxonomicName>
,
<figureCitation id="0FC150D1D67BFF88FD1577BCFCB3F852" box="[667,801,1920,1946]" captionStart-0="FIGURES 9 - 12" captionStart-1="FIGURES 13 - 16" captionStart-2="FIGURES 17 – 18" captionStartId-0="23.[151,269,1661,1685]" captionStartId-1="24.[151,274,1852,1876]" captionStartId-2="25.[151,270,940,964]" captionTargetBox-0="[236,1337,209,1625]" captionTargetBox-1="[155,1435,186,1820]" captionTargetBox-2="[151,1428,207,899]" captionTargetId-0="figure@23.[226,1360,194,1639]" captionTargetId-1="figure@24.[151,1436,176,1831]" captionTargetId-2="figure@25.[151,1436,194,919]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURES 9 - 12. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 9: Leptobasis buchholzi, Colombia, Puerto Colombia. 10: L. candelaria, Belize, Toledo, Monkey River. 11: L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; 12: L. lucifer, Mexico, Campeche, Laguna Zoh." captionText-1="FIGURES 13 - 16. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 13: Leptobasis mauffrayi, paratype, Peru, Huanuco, N of Cucharas; 14: L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; 15: L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; 16: L. vacillans, Dominican Republic, La Altagracia, Nisibon." captionText-2="FIGURES 17 – 18. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 17: Hesperagrion heterodoxum, USA, Arizona, Yarnell; 18: Apanisagrion lais, USA, Arizona, Hot Springs Canyon" httpUri-0="https://zenodo.org/record/194877/files/figure.png" httpUri-1="https://zenodo.org/record/194878/files/figure.png" httpUri-2="https://zenodo.org/record/194879/files/figure.png" pageId="2" pageNumber="3">Figs. 9–17</figureCitation>
), an inner fold proximal to ligula flexure (absent in
<taxonomicName id="50FA37D7D67BFF88FF197799FEA8F876" box="[151,314,1957,1982]" class="Insecta" family="Coenagrionidae" genus="Hesperagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FF197799FEA8F876" box="[151,314,1957,1982]" italics="true" pageId="2" pageNumber="3">Hesperagrion</emphasis>
</taxonomicName>
, which has a transverse fold distal to ligula flexure; see discussion), and no lateral lobes. For all these reasons we consider
<taxonomicName id="50FA37D7D67BFF88FE4377F7FDCCF82C" box="[461,606,1995,2020]" class="Insecta" family="Coenagrionidae" genus="Chrysobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FE4377F7FDCCF82C" box="[461,606,1995,2020]" italics="true" pageId="2" pageNumber="3">Chrysobasis</emphasis>
</taxonomicName>
a junior synonym of
<taxonomicName id="50FA37D7D67BFF88FCD477F7FC4BF82C" box="[858,985,1995,2020]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67BFF88FCD477F7FC4BF82C" box="[858,985,1995,2020]" italics="true" pageId="2" pageNumber="3">Leptobasis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67AFF89FF4870ABFEECFC54" pageId="3" pageNumber="4" type="description">
<paragraph id="97454C54D67AFF89FF4870ABFEECFC54" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="4">
<emphasis id="A58E9046D67AFF89FF4870ABFE31FF79" bold="true" box="[198,419,151,177]" pageId="3" pageNumber="4">Characterization.</emphasis>
Small to medium coenagrionids (
<quantity id="5002E1B1D67AFF89FCBE70A4FC26FF7A" box="[816,948,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="4.1" metricValueMin="2.6" pageId="3" pageNumber="4" unit="mm" value="33.5" valueMax="41.0" valueMin="26.0">26–41 mm</quantity>
), with a relatively long abdomen (ratio of 4–5.25 to length of head plus thorax); dorsum of head, thorax, and S3–8 black, brown, or orange; pterothoracic stripes blue or green, S7–10 to S9–10 blue, yellowish, orange, reddish, or black; pale areas on sides of thorax and abdomen yellow and green (
<figureCitation id="0FC150D1D67AFF89FD6F7134FCE5FEEA" box="[737,887,264,290]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStart-3="FIGURE 26" captionStartId-0="30.[151,254,1524,1548]" captionStartId-1="31.[151,254,1798,1822]" captionStartId-2="31.[151,257,1894,1918]" captionStartId-3="34.[151,256,151,175]" captionTargetBox-0="[151,1436,194,1501]" captionTargetBox-1="[151,1436,184,1775]" captionTargetId-0="figure@30.[151,1436,194,1503]" captionTargetId-1="figure@31.[151,1436,184,1777]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 23. Adults of Leptobasis. (a) L. guanacaste immature male, Costa Rica, Guanacaste, Río Cañas, photographed by William A. Haber; (b) L. guanacaste female, Costa Rica, Guanacaste, Río Cañas, photographed by William A. Haber." captionText-1="FIGURE 24. Adults of Leptobasis lucifer, USA, Florida, Collier, Corkscrew Swamp Sanctuary. (a) male, photographed by Dennis R. Paulson; (b) female, photographed by Netta Smith." captionText-2="FIGURE 25. Adults of Leptobasis. (a) L. mauffrayi female, Peru, Madre de Dios, Explorer’s Inn, photographed by Dennis R. Paulson; (b) L. melinogaster male, USA, Texas, Hidalgo, Santa Ana National Wildlife Refuge, photographed by J. C. Abbott; (c) L. melinogaster female, USA, Texas, Hidalgo, Santa Ana National Wildlife Refuge, photographed by J. C. Abbott." captionText-3="FIGURE 26. Adults of Leptobasis vacillans. (a) mature male, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber; (b) immature male, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber; (c) mature female, Colombia, Antioquía, Municipio de la Pintada, Farallón de la Paz, cañón del río Cauca, photographed by Cornelio A. Bota Sierra; (d) teneral female, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber." httpUri-0="https://zenodo.org/record/194884/files/figure.png" httpUri-1="https://zenodo.org/record/194885/files/figure.png" pageId="3" pageNumber="4">Figs. 23–26</figureCitation>
). Pale postocular spots and pale occipital bar present (
<figureCitation id="0FC150D1D67AFF89FF717111FEC2FE8F" box="[255,336,301,327]" captionStart="FIGURE 1" captionStartId="17.[151,255,1606,1630]" captionTargetBox="[229,1344,194,1567]" captionTargetId="figure@17.[227,1360,175,1582]" captionTargetPageId="17" captionText="FIGURE 1. Head, dorsal view of right half. (a) Leptobasis candelaria, male, Mexico, Veracruz, E of Córdoba; (b) L. guanacaste, male paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. mauffrayi, male holotype, Peru, Manu, Pakitza; (d) L. vacillans, male, Dominican Republic, La Vega, S of La Vega." httpUri="https://zenodo.org/record/194871/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
a, b, d), blue or green, or absent (
<figureCitation id="0FC150D1D67AFF89FD647111FCAEFE8F" box="[746,828,301,327]" captionStart="FIGURE 1" captionStartId="17.[151,255,1606,1630]" captionTargetBox="[229,1344,194,1567]" captionTargetId="figure@17.[227,1360,175,1582]" captionTargetPageId="17" captionText="FIGURE 1. Head, dorsal view of right half. (a) Leptobasis candelaria, male, Mexico, Veracruz, E of Córdoba; (b) L. guanacaste, male paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. mauffrayi, male holotype, Peru, Manu, Pakitza; (d) L. vacillans, male, Dominican Republic, La Vega, S of La Vega." httpUri="https://zenodo.org/record/194871/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
c, 26d). Frons rounded; location of most posterior point of head at level of eyes. Posterior lobe of prothorax smoothly convex, bi- or trilobate, projected medially or not projected in both sexes (
<figureCitation id="0FC150D1D67AFF89FE737144FDC2FE5A" box="[509,592,376,402]" captionStart="FIGURE 2" captionStartId="17.[151,255,1702,1726]" captionText="FIGURE 2. Female pronotum, dorsal view. (a) Leptobasis buchholzi, paratype, Venezuela, Guárico, Pericoco [redrawn from a draft by J. De Marmels]; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Mexico, Campeche, Laguna Zoh; (e) L. mauffrayi, allotype, Peru, Manu, Pakitza; (f) L. melinogaster, USA, Texas, Kleberg, King Ranch; (g) L. raineyi, Trinidad, St. George, Indian Walk Ride; (h) L. vacillans, Cuba, Cienfuegos, E of Pepito Rey." pageId="3" pageNumber="4">Figs. 2</figureCitation>
,
<figureCitation id="0FC150D1D67AFF89FDD27144FDFCFE5A" box="[604,622,376,402]" captionStart="FIGURE 4" captionStartId="18.[151,257,1493,1517]" captionText="FIGURE 4. Male pronotum and mesostigmal plates, dorsal view. (a) Leptobasis guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (b) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (c) L. vacillans, Dominican Republic, Jarabacoa." pageId="3" pageNumber="4">4</figureCitation>
). Pterothorax with (
<figureCitation id="0FC150D1D67AFF89FCD87144FC24FE5A" box="[854,950,376,402]" captionStart="FIGURE 24" captionStartId="31.[151,254,1798,1822]" captionTargetBox="[151,1436,184,1775]" captionTargetId="figure@31.[151,1436,184,1777]" captionTargetPageId="31" captionText="FIGURE 24. Adults of Leptobasis lucifer, USA, Florida, Collier, Corkscrew Swamp Sanctuary. (a) male, photographed by Dennis R. Paulson; (b) female, photographed by Netta Smith." httpUri="https://zenodo.org/record/194885/files/figure.png" pageId="3" pageNumber="4">Figs. 24</figureCitation>
a, 26a) or without (
<figureCitation id="0FC150D1D67AFF89FB1D7144FB60FE5A" box="[1171,1266,376,402]" captionStart="FIGURE 26" captionStartId="34.[151,256,151,175]" captionText="FIGURE 26. Adults of Leptobasis vacillans. (a) mature male, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber; (b) immature male, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber; (c) mature female, Colombia, Antioquía, Municipio de la Pintada, Farallón de la Paz, cañón del río Cauca, photographed by Cornelio A. Bota Sierra; (d) teneral female, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber." pageId="3" pageNumber="4">Figs. 26</figureCitation>
b-d) dark middorsal and humeral stripes; dark metapleural stripe absent; pale antehumeral stripe present and mesepimeron and metepisternum mostly pale; pterothorax entirely orange in immature specimens (
<figureCitation id="0FC150D1D67AFF89FB3371FEFA88FE14" box="[1213,1306,450,476]" captionStart="FIGURE 26" captionStartId="34.[151,256,151,175]" captionText="FIGURE 26. Adults of Leptobasis vacillans. (a) mature male, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber; (b) immature male, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber; (c) mature female, Colombia, Antioquía, Municipio de la Pintada, Farallón de la Paz, cañón del río Cauca, photographed by Cornelio A. Bota Sierra; (d) teneral female, Costa Rica, Limón, SW of Beverly, photographed by William A. Haber." pageId="3" pageNumber="4">Fig. 26</figureCitation>
d). Female lacking well defined mesepisternal carinae between mesostigmal plates and medial carina (
<figureCitation id="0FC150D1D67AFF89FA8971D4FACEFDCA" box="[1287,1372,488,514]" captionStart="FIGURE 3" captionStartId="18.[151,254,1365,1389]" captionTargetBox="[235,1351,210,1294]" captionTargetId="figure@18.[227,1360,194,1344]" captionTargetPageId="18" captionText="FIGURE 3. Female mesostigmal plates, dorsal view. (a) Leptobasis buchholzi, right plate, paratype, Venezuela, Guárico, Pericoco [redrawn from a draft by J. De Marmels]; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Mexico, Campeche, Laguna Zoh; (e) L. mauffrayi, allotype, Peru, Manu, Pakitza." httpUri="https://zenodo.org/record/194872/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
b–e). Metatibial spurs shorter than twice intervening spaces (
<figureCitation id="0FC150D1D67AFF89FCB37231FC15FDEF" box="[829,903,525,551]" captionStart="FIGURE 5" captionStartId="19.[151,256,1573,1597]" captionTargetBox="[163,1418,187,1542]" captionTargetId="figure@19.[151,1436,172,1552]" captionTargetPageId="19" captionText="FIGURE 5. Male metafemur. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. vacillans, Dominican Republic, Jarabacoa." httpUri="https://zenodo.org/record/194873/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
c); pretarsus with supplementary tooth very small, forming a right angle with claw. CuA extending 2–6 cells distal to vein descending from subnodus (
<figureCitation id="0FC150D1D67AFF89FF117264FE80FDBA" box="[159,274,600,626]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="20.[151,257,1619,1643]" captionStartId-1="21.[151,255,1900,1924]" captionStartId-2="22.[151,257,1594,1618]" captionTargetBox-0="[240,1342,201,1587]" captionTargetBox-1="[154,1420,184,1871]" captionTargetBox-2="[162,1417,186,1562]" captionTargetId-0="figure@20.[237,1349,194,1598]" captionTargetId-1="figure@21.[151,1436,170,1879]" captionTargetId-2="figure@22.[158,1429,176,1573]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURE 6. Wings. (a) Leptobasis buchholzi, male, Colombia, Puerto Colombia; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga." captionText-1="FIGURE 7. Wings. (a) Leptobasis lucifer, female, Mexico, Campeche, Laguna Zoh; (b) L. mauffrayi, male paratype, Peru, Manu, Pakitza; (c) L. melinogaster, male, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve." captionText-2="FIGURE 8. Wings. (a) Leptobasis raineyi, male, Trinidad, St. Andrew, 1 mi W of Cumuto; (b) L. vacillans, male, Dominican Republic, La Altagracia, Nisibon." httpUri-0="https://zenodo.org/record/194874/files/figure.png" httpUri-1="https://zenodo.org/record/194875/files/figure.png" httpUri-2="https://zenodo.org/record/194876/files/figure.png" pageId="3" pageNumber="4">Figs. 6–8</figureCitation>
), occasionally
<quantity id="5002E1B1D67AFF89FE4D7264FD9CFDBA" box="[451,526,600,626]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="3" pageNumber="4" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
<taxonomicName id="50FA37D7D67AFF89FD987264FD33FDB9" box="[534,673,600,625]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67AFF89FD987264FD33FDB9" box="[534,673,600,625]" italics="true" pageId="3" pageNumber="4">L. vacillans</emphasis>
</taxonomicName>
; CuP linking CuA to posterior wing margin (
<figureCitation id="0FC150D1D67AFF89FB337264FABDFDBA" box="[1213,1327,600,626]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="20.[151,257,1619,1643]" captionStartId-1="21.[151,255,1900,1924]" captionStartId-2="22.[151,257,1594,1618]" captionTargetBox-0="[240,1342,201,1587]" captionTargetBox-1="[154,1420,184,1871]" captionTargetBox-2="[162,1417,186,1562]" captionTargetId-0="figure@20.[237,1349,194,1598]" captionTargetId-1="figure@21.[151,1436,170,1879]" captionTargetId-2="figure@22.[158,1429,176,1573]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURE 6. Wings. (a) Leptobasis buchholzi, male, Colombia, Puerto Colombia; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga." captionText-1="FIGURE 7. Wings. (a) Leptobasis lucifer, female, Mexico, Campeche, Laguna Zoh; (b) L. mauffrayi, male paratype, Peru, Manu, Pakitza; (c) L. melinogaster, male, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve." captionText-2="FIGURE 8. Wings. (a) Leptobasis raineyi, male, Trinidad, St. Andrew, 1 mi W of Cumuto; (b) L. vacillans, male, Dominican Republic, La Altagracia, Nisibon." httpUri-0="https://zenodo.org/record/194874/files/figure.png" httpUri-1="https://zenodo.org/record/194875/files/figure.png" httpUri-2="https://zenodo.org/record/194876/files/figure.png" pageId="3" pageNumber="4">Figs. 6–8</figureCitation>
). Genital ligula distal segment with inner fold proximal to flexure; lacking lateral lobes; with apex entire to bifid; with a lateral chitinized flap-like process distal to flexure directed posteriorly on each side (
<figureCitation id="0FC150D1D67AFF89FB2F729EFAB6FD74" box="[1185,1316,674,700]" captionStart-0="FIGURES 9 - 12" captionStart-1="FIGURES 13 - 16" captionStartId-0="23.[151,269,1661,1685]" captionStartId-1="24.[151,274,1852,1876]" captionTargetBox-0="[236,1337,209,1625]" captionTargetBox-1="[155,1435,186,1820]" captionTargetId-0="figure@23.[226,1360,194,1639]" captionTargetId-1="figure@24.[151,1436,176,1831]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 9 - 12. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 9: Leptobasis buchholzi, Colombia, Puerto Colombia. 10: L. candelaria, Belize, Toledo, Monkey River. 11: L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; 12: L. lucifer, Mexico, Campeche, Laguna Zoh." captionText-1="FIGURES 13 - 16. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 13: Leptobasis mauffrayi, paratype, Peru, Huanuco, N of Cucharas; 14: L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; 15: L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; 16: L. vacillans, Dominican Republic, La Altagracia, Nisibon." httpUri-0="https://zenodo.org/record/194877/files/figure.png" httpUri-1="https://zenodo.org/record/194878/files/figure.png" pageId="3" pageNumber="4">Figs. 9–16</figureCitation>
). Posterodorsal margin of male S10 with (
<figureCitation id="0FC150D1D67AFF89FD9272F4FDEFFD2A" box="[540,637,712,738]" captionStart="FIGURE 21" captionStartId="28.[151,255,1395,1419]" captionTargetBox="[153,1429,211,1340]" captionTargetId="figure@28.[151,1436,194,1374]" captionTargetPageId="28" captionText="FIGURE 21. Male S 10, lateral view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194882/files/figure.png" pageId="3" pageNumber="4">Figs. 21</figureCitation>
a, d, f) or without (
<figureCitation id="0FC150D1D67AFF89FCD372F4FC2CFD2A" box="[861,958,712,738]" captionStart="FIGURE 21" captionStartId="28.[151,255,1395,1419]" captionTargetBox="[153,1429,211,1340]" captionTargetId="figure@28.[151,1436,194,1374]" captionTargetPageId="28" captionText="FIGURE 21. Male S 10, lateral view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194882/files/figure.png" pageId="3" pageNumber="4">Figs. 21</figureCitation>
b, c, e, g–h) a pair of lobe-like processes on sides of medial cleft. Male cercus shorter than or subequal to S10, with a short ventro-basal branch on medial surface; male paraproct entire and shorter to longer than cercus (
<figureCitation id="0FC150D1D67AFF89FC70732EFBCAFCE4" box="[1022,1112,786,812]" captionStart="FIGURE 21" captionStartId="28.[151,255,1395,1419]" captionTargetBox="[153,1429,211,1340]" captionTargetId="figure@28.[151,1436,194,1374]" captionTargetPageId="28" captionText="FIGURE 21. Male S 10, lateral view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194882/files/figure.png" pageId="3" pageNumber="4">Fig. 21</figureCitation>
). Female usually lacking a vulvar spine on S8 (
<figureCitation id="0FC150D1D67AFF89FE0A7304FE76FC9A" box="[388,484,824,850]" captionStart="FIGURE 19" captionStartId="26.[151,255,1251,1275]" captionTargetBox="[153,1435,199,1216]" captionTargetId="figure@26.[151,1436,194,1230]" captionTargetPageId="26" captionText="FIGURE 19. Female S 7 – 10, lateral view. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, allotype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. lucifer, Mexico, Campeche, Laguna Zoh; (d) L. mauffrayi, allotype, Peru, Madre de Dios, Pakitza; (e) L. raineyi, Trinidad, St. George, Indian Walk Ride; (f) L. vacillans, Mexico, Morelos, La Fuente." httpUri="https://zenodo.org/record/194880/files/figure.png" pageId="3" pageNumber="4">Figs. 19</figureCitation>
a–e), although a small spine is present in most females of
<taxonomicName id="50FA37D7D67AFF89FB1C7304FA89FC99" box="[1170,1307,824,849]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67AFF89FB1C7304FA89FC99" box="[1170,1307,824,849]" italics="true" pageId="3" pageNumber="4">L. vacillans</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67AFF89FAA27304FA13FC9A" box="[1324,1409,824,850]" captionStart="FIGURE 19" captionStartId="26.[151,255,1251,1275]" captionTargetBox="[153,1435,199,1216]" captionTargetId="figure@26.[151,1436,194,1230]" captionTargetPageId="26" captionText="FIGURE 19. Female S 7 – 10, lateral view. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, allotype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. lucifer, Mexico, Campeche, Laguna Zoh; (d) L. mauffrayi, allotype, Peru, Madre de Dios, Pakitza; (e) L. raineyi, Trinidad, St. George, Indian Walk Ride; (f) L. vacillans, Mexico, Morelos, La Fuente." httpUri="https://zenodo.org/record/194880/files/figure.png" pageId="3" pageNumber="4">Fig. 19</figureCitation>
f); postero-dorsal margin of female S9 with denticles; ovipositor extending distally to tip of cerci (
<figureCitation id="0FC150D1D67AFF89FB747361FACEFCBF" box="[1274,1372,861,887]" captionStart="FIGURE 19" captionStartId="26.[151,255,1251,1275]" captionTargetBox="[153,1435,199,1216]" captionTargetId="figure@26.[151,1436,194,1230]" captionTargetPageId="26" captionText="FIGURE 19. Female S 7 – 10, lateral view. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, allotype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. lucifer, Mexico, Campeche, Laguna Zoh; (d) L. mauffrayi, allotype, Peru, Madre de Dios, Pakitza; (e) L. raineyi, Trinidad, St. George, Indian Walk Ride; (f) L. vacillans, Mexico, Morelos, La Fuente." httpUri="https://zenodo.org/record/194880/files/figure.png" pageId="3" pageNumber="4">Figs. 19</figureCitation>
,
<figureCitation id="0FC150D1D67AFF89FAE77361FA1AFCBF" box="[1385,1416,861,887]" captionStart="FIGURE 23" captionStartId="30.[151,254,1524,1548]" captionTargetBox="[151,1436,194,1501]" captionTargetId="figure@30.[151,1436,194,1503]" captionTargetPageId="30" captionText="FIGURE 23. Adults of Leptobasis. (a) L. guanacaste immature male, Costa Rica, Guanacaste, Río Cañas, photographed by William A. Haber; (b) L. guanacaste female, Costa Rica, Guanacaste, Río Cañas, photographed by William A. Haber." httpUri="https://zenodo.org/record/194884/files/figure.png" pageId="3" pageNumber="4">23</figureCitation>
b, 24b, 25a, c, 26c, d).
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67AFF89FF48739BFE18FA04" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="97454C54D67AFF89FF48739BFE18FA04" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="4">
<emphasis id="A58E9046D67AFF89FF48739BFED4FC09" bold="true" box="[198,326,935,961]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="50FA37D7D67AFF89FEDF7394FE42FC09" box="[337,464,936,961]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FEDF7394FE42FC09" box="[337,464,936,961]" italics="true" pageId="3" pageNumber="4">Leptobasis</emphasis>
</taxonomicName>
shares the combination of a rounded frons, CuP reaching hind margin of wing, CuA relatively short, and supplementary pretarsal claw vestigial or reduced and forming a right angle with claw with
<taxonomicName id="50FA37D7D67AFF89FE9C73CEFD85FBC4" authority="Garrison" authorityName="Garrison" box="[274,535,1010,1036]" class="Insecta" family="Coenagrionidae" genus="Dolonagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FE9C73CEFE3BFBC3" box="[274,425,1010,1035]" italics="true" pageId="3" pageNumber="4">Dolonagrion</emphasis>
Garrison
</taxonomicName>
&amp; von Ellenrieder 2008,
<taxonomicName id="50FA37D7D67AFF89FCCA73CEFB37FBC4" authority="Sjostedt 1918" authorityName="Sjostedt" authorityYear="1918" box="[836,1189,1010,1036]" class="Insecta" family="Coenagrionidae" genus="Mesoleptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FCCA73CEFC68FBC3" box="[836,1018,1010,1035]" italics="true" pageId="3" pageNumber="4">Mesoleptobasis</emphasis>
<bibRefCitation id="F36B31A5D67AFF89FB8C73CEFB37FBC4" author="Sjostedt" box="[1026,1189,1010,1036]" pageId="3" pageNumber="16" refString="Sjostedt, Y. (1918) Wissenschaftliche Ergebnisse der Schwedischen Entomologischen Reise des Herrn Dr. A. Roman in Amazonas 1914 - 1915. 1, Odonata. Arkiv vor Zoologi, 11, 1 - 54." type="journal article" year="1918">Sjöstedt 1918</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="50FA37D7D67AFF89FB6A73CEFF4BFBFA" authority="Selys 1876" authorityName="Selys" authorityYear="1876" class="Insecta" family="Coenagrionidae" genus="Telagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FB6A73CEFAC7FBC3" box="[1252,1365,1010,1035]" italics="true" pageId="3" pageNumber="4">Telagrion</emphasis>
<bibRefCitation id="F36B31A5D67AFF89FAD273CEFF4BFBFA" author="Selys" pageId="3" pageNumber="16" refString="Selys, E., (1876) Synopsis des Agrionines. 5 e legion: Agrion (suite). Le genre Agrion. Bulletin de l'Academie Royale de Belgique, 41, 952 - 991 (242 - 322 reprint)." type="journal article" year="1876">Selys 1876</bibRefCitation>
</taxonomicName>
. It differs from all of them by the presence on the distal segment of the genital ligula of a pair of chitinized, flap-like, movable processes directed posteriorly (
<figureCitation id="0FC150D1D67AFF89FC077401FB83FB9F" box="[905,1041,1085,1111]" captionStart-0="FIGURES 9 - 12" captionStart-1="FIGURES 13 - 16" captionStartId-0="23.[151,269,1661,1685]" captionStartId-1="24.[151,274,1852,1876]" captionTargetBox-0="[236,1337,209,1625]" captionTargetBox-1="[155,1435,186,1820]" captionTargetId-0="figure@23.[226,1360,194,1639]" captionTargetId-1="figure@24.[151,1436,176,1831]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 9 - 12. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 9: Leptobasis buchholzi, Colombia, Puerto Colombia. 10: L. candelaria, Belize, Toledo, Monkey River. 11: L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; 12: L. lucifer, Mexico, Campeche, Laguna Zoh." captionText-1="FIGURES 13 - 16. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 13: Leptobasis mauffrayi, paratype, Peru, Huanuco, N of Cucharas; 14: L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; 15: L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; 16: L. vacillans, Dominican Republic, La Altagracia, Nisibon." httpUri-0="https://zenodo.org/record/194877/files/figure.png" httpUri-1="https://zenodo.org/record/194878/files/figure.png" pageId="3" pageNumber="4">Figs. 9–16</figureCitation>
). Branched male cerci lacking a transverse membranous dorsal depression (
<figureCitation id="0FC150D1D67AFF89FD2E745EFC94FBB4" box="[672,774,1122,1148]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="3" pageNumber="4">Figs. 20</figureCitation>
,
<figureCitation id="0FC150D1D67AFF89FC9D745EFCA7FBB4" box="[787,821,1122,1148]" captionStart="FIGURE 22" captionStartId="29.[151,254,1497,1521]" captionTargetBox="[158,1426,211,1460]" captionTargetId="figure@29.[151,1436,194,1476]" captionTargetPageId="29" captionText="FIGURE 22. Male S 10. (a) Leptobasis guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (b) L. lucifer, Costa Rica, Heredia, Finca La Selva; (c) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (d) L. melinogaster, USA, Texas, Santa Anna; (e) L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; (f) L. vacillans, Puerto Rico, Mayagüez, Mayagüez. (a) posterior view; (b – f) dorsal view." httpUri="https://zenodo.org/record/194883/files/figure.png" pageId="3" pageNumber="4">22</figureCitation>
), and tip of female ovipositor extending distally to postero-dorsal margin of S10 for a distance shorter than length of S10 (
<figureCitation id="0FC150D1D67AFF89FC7D74B4FBDEFB6A" box="[1011,1100,1160,1186]" captionStart="FIGURE 19" captionStartId="26.[151,255,1251,1275]" captionTargetBox="[153,1435,199,1216]" captionTargetId="figure@26.[151,1436,194,1230]" captionTargetPageId="26" captionText="FIGURE 19. Female S 7 – 10, lateral view. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, allotype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. lucifer, Mexico, Campeche, Laguna Zoh; (d) L. mauffrayi, allotype, Peru, Madre de Dios, Pakitza; (e) L. raineyi, Trinidad, St. George, Indian Walk Ride; (f) L. vacillans, Mexico, Morelos, La Fuente." httpUri="https://zenodo.org/record/194880/files/figure.png" pageId="3" pageNumber="4">Fig. 19</figureCitation>
), further distinguish it from
<taxonomicName id="50FA37D7D67AFF89FF197491FEA3FB0E" box="[151,305,1197,1222]" class="Insecta" family="Coenagrionidae" genus="Dolonagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FF197491FEA3FB0E" box="[151,305,1197,1222]" italics="true" pageId="3" pageNumber="4">Dolonagrion</emphasis>
</taxonomicName>
, which has entire cerci with a membranous dorsal depression and ovipositor surpassing tip of cerci for a distance longer than S10 length (Garrison &amp; von Ellenrieder 2008). It further differs from
<taxonomicName id="50FA37D7D67AFF89FF1974C4FEDCFAD9" box="[151,334,1272,1297]" class="Insecta" family="Coenagrionidae" genus="Mesoleptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FF1974C4FEDCFAD9" box="[151,334,1272,1297]" italics="true" pageId="3" pageNumber="4">Mesoleptobasis</emphasis>
</taxonomicName>
in that the vein descending from the quadrangle does not form a straight line to wing margin (
<figureCitation id="0FC150D1D67AFF89FF117521FE80FAFF" box="[159,274,1309,1335]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="20.[151,257,1619,1643]" captionStartId-1="21.[151,255,1900,1924]" captionStartId-2="22.[151,257,1594,1618]" captionTargetBox-0="[240,1342,201,1587]" captionTargetBox-1="[154,1420,184,1871]" captionTargetBox-2="[162,1417,186,1562]" captionTargetId-0="figure@20.[237,1349,194,1598]" captionTargetId-1="figure@21.[151,1436,170,1879]" captionTargetId-2="figure@22.[158,1429,176,1573]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURE 6. Wings. (a) Leptobasis buchholzi, male, Colombia, Puerto Colombia; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga." captionText-1="FIGURE 7. Wings. (a) Leptobasis lucifer, female, Mexico, Campeche, Laguna Zoh; (b) L. mauffrayi, male paratype, Peru, Manu, Pakitza; (c) L. melinogaster, male, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve." captionText-2="FIGURE 8. Wings. (a) Leptobasis raineyi, male, Trinidad, St. Andrew, 1 mi W of Cumuto; (b) L. vacillans, male, Dominican Republic, La Altagracia, Nisibon." httpUri-0="https://zenodo.org/record/194874/files/figure.png" httpUri-1="https://zenodo.org/record/194875/files/figure.png" httpUri-2="https://zenodo.org/record/194876/files/figure.png" pageId="3" pageNumber="4">Figs. 6–8</figureCitation>
), versus forming a straight line in the latter (Garrison &amp; von
<bibRefCitation id="F36B31A5D67AFF89FC667521FB21FAFF" author="Ellenrieder" box="[1000,1203,1309,1335]" pageId="3" pageNumber="15" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2009) Redefinition of Mesoleptobasis Sjostedt 1918 with the inclusion of Metaleptobasis [sic. Mesoleptobasis] cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa, 2145, 47 - 68." type="journal article" year="2009">Ellenrieder 2009</bibRefCitation>
).
<taxonomicName id="50FA37D7D67AFF89FB467521FADAFAFE" box="[1224,1352,1309,1334]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FB467521FADAFAFE" box="[1224,1352,1309,1334]" italics="true" pageId="3" pageNumber="4">Leptobasis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="50FA37D7D67AFF89FF56757FFED8FA94" box="[216,330,1347,1372]" class="Insecta" family="Coenagrionidae" genus="Telagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FF56757FFED8FA94" box="[216,330,1347,1372]" italics="true" pageId="3" pageNumber="4">Telagrion</emphasis>
</taxonomicName>
by the absence of paired ental medio-longitudinal ridges and of an outer-lateral patch of thin setae on the distal segment of the genital ligula (
<figureCitation id="0FC150D1D67AFF89FC887554FC01FA4A" box="[774,915,1384,1410]" captionStart-0="FIGURES 9 - 12" captionStart-1="FIGURES 13 - 16" captionStartId-0="23.[151,269,1661,1685]" captionStartId-1="24.[151,274,1852,1876]" captionTargetBox-0="[236,1337,209,1625]" captionTargetBox-1="[155,1435,186,1820]" captionTargetId-0="figure@23.[226,1360,194,1639]" captionTargetId-1="figure@24.[151,1436,176,1831]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 9 - 12. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 9: Leptobasis buchholzi, Colombia, Puerto Colombia. 10: L. candelaria, Belize, Toledo, Monkey River. 11: L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; 12: L. lucifer, Mexico, Campeche, Laguna Zoh." captionText-1="FIGURES 13 - 16. Genital ligula, (a) lateral view; (b) ectal view; (c) latero-ental view. 13: Leptobasis mauffrayi, paratype, Peru, Huanuco, N of Cucharas; 14: L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; 15: L. raineyi, Trinidad, St. Andrew, 1 mile W of Cumuto; 16: L. vacillans, Dominican Republic, La Altagracia, Nisibon." httpUri-0="https://zenodo.org/record/194877/files/figure.png" httpUri-1="https://zenodo.org/record/194878/files/figure.png" pageId="3" pageNumber="4">Figs. 9–16</figureCitation>
), which are present in the latter (see von
<bibRefCitation id="F36B31A5D67AFF89FF1975B1FE68FA6F" author="Ellenrieder" box="[151,506,1421,1447]" pageId="3" pageNumber="14" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2008) Dolonagrion nov. gen. for Telagrion fulvellum from South America (Odonata: Coenagrionidae). International Journal of Odonatology, 11 (2), 173 - 183." type="journal article" year="2008">Ellenrieder &amp; Garrison 2008</bibRefCitation>
), and inner margin of female cerci straight or convex in dorsal view, versus concave in the latter.
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67AFF89FF4875EBFCBCF9DF" pageId="3" pageNumber="4" type="distribution">
<paragraph id="97454C54D67AFF89FF4875EBFCBCF9DF" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="4">
<emphasis id="A58E9046D67AFF89FF4875EBFEF5FA39" bold="true" box="[198,359,1495,1521]" pageId="3" pageNumber="4">Distribution.</emphasis>
<taxonomicName id="50FA37D7D67AFF89FEFC75E4FE63FA39" box="[370,497,1496,1521]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FEFC75E4FE63FA39" box="[370,497,1496,1521]" italics="true" pageId="3" pageNumber="4">Leptobasis</emphasis>
</taxonomicName>
occurs from southern Texas (
<collectingCountry id="EFED0CC4D67AFF89FCD175E4FC0DFA3A" box="[863,927,1496,1522]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
), the Antilles, and Central
<collectingCountry id="EFED0CC4D67AFF89FB6375E4FAC0FA3A" box="[1261,1362,1496,1522]" name="United States of America" pageId="3" pageNumber="4">America</collectingCountry>
south through
<collectingCountry id="EFED0CC4D67AFF89FF7775C1FEE3F9DF" box="[249,369,1533,1559]" name="Venezuela" pageId="3" pageNumber="4">Venezuela</collectingCountry>
and into southern
<collectingCountry id="EFED0CC4D67AFF89FDC275C1FD10F9DF" box="[588,642,1533,1559]" name="Peru" pageId="3" pageNumber="4">Peru</collectingCountry>
(
<figureCitation id="0FC150D1D67AFF89FD1F75C1FCB0F9DF" box="[657,802,1533,1559]" captionStart-0="FIGURE 27" captionStart-1="FIGURE 28" captionStartId-0="34.[151,254,1739,1763]" captionStartId-1="35.[151,254,1459,1483]" captionTargetBox-0="[160,1423,522,1711]" captionTargetBox-1="[160,1425,210,1419]" captionTargetId-0="figure@34.[151,1436,514,1718]" captionTargetId-1="figure@35.[151,1436,194,1438]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURE 27. Distribution of Leptobasis buchholzi, L. candelaria, L. guanacaste, L. lucifer, and L. melinogaster." captionText-1="FIGURE 28. Distribution of Leptobasis mauffrayi, L. raineyi, and L. vacillans." httpUri-0="https://zenodo.org/record/194886/files/figure.png" httpUri-1="https://zenodo.org/record/194887/files/figure.png" pageId="3" pageNumber="4">Figs. 27–28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67AFF8EFF48761EFBFEFF34" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="biology_ecology">
<paragraph id="97454C54D67AFF89FF48761EFC07F93F" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="4">
<emphasis id="A58E9046D67AFF89FF48761EFEB8F9F4" bold="true" box="[198,298,1570,1596]" pageId="3" pageNumber="4">Biology.</emphasis>
Adults perch in shady areas on littoral vegetation, grasses or sedges, along margins of ponds, lakes, swamps, and marshes, banks of rivers, and temporary streams; they are rarely seen flying over open water.
<bibRefCitation id="F36B31A5D67AFF89FF6A7651FE15F94F" author="Daigle" box="[228,391,1645,1671]" pageId="3" pageNumber="14" refString="Daigle, J. J. (2009) Corkscrew Chrysobasis Soiree. Argia, 21 (1), 9 - 10." type="journal article" year="2009">Daigle (2009)</bibRefCitation>
reported
<taxonomicName id="50FA37D7D67AFF89FE787651FDF2F94E" box="[502,608,1645,1670]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lucifer">
<emphasis id="A58E9046D67AFF89FE787651FDF2F94E" box="[502,608,1645,1670]" italics="true" pageId="3" pageNumber="4">L. lucifer</emphasis>
</taxonomicName>
flying around fern patches among cypress trees in swamps, with males perching on vines and leaf tips, and females ovipositing in the stem of
<taxonomicName id="50FA37D7D67AFF89FB8876AFFB45F964" box="[1030,1239,1683,1708]" class="Arachnida" family="Thomisidae" genus="Lycopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rubellus">
<emphasis id="A58E9046D67AFF89FB8876AFFB45F964" box="[1030,1239,1683,1708]" italics="true" pageId="3" pageNumber="4">Lycopus rubellus</emphasis>
</taxonomicName>
. The only larva described so far is that of
<taxonomicName id="50FA37D7D67AFF89FE5F7684FDCFF919" box="[465,605,1720,1745]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67AFF89FE5F7684FDCFF919" box="[465,605,1720,1745]" italics="true" pageId="3" pageNumber="4">L. vacillans</emphasis>
</taxonomicName>
(
<bibRefCitation id="F36B31A5D67AFF89FDE37684FC8EF91A" author="Geijskes" box="[621,796,1720,1746]" pageId="3" pageNumber="15" refString="Geijskes, D. C. (1941) Notes on Odonata of Surinam. II. Six mostly new zygopterous nymphs from coastland waters. Annals of the Entomological Society of America, 35, 719 - 134." type="journal volume" year="1941">Geijskes 1941</bibRefCitation>
,
<bibRefCitation id="F36B31A5D67AFF89FCA47684FC73F91A" author="Needham" box="[810,993,1720,1746]" pageId="3" pageNumber="15" refString="Needham, J. G. (1941) Life history notes on some West Indian Coenagrionine dragonflies (Odonata). Journal of Agriculture, University of Puerto Rico, 25 (3), 1 - 18." type="journal article" year="1941">Needham 1941</bibRefCitation>
) although
<bibRefCitation id="F36B31A5D67AFF89FBED7684FABAF91A" author="Paulson" box="[1123,1320,1720,1746]" pageId="3" pageNumber="16" refString="Paulson, D. R. (2009 b) Leptobasis guanacaste, a new damselfly from Costa Rica (Odonata: Coenagrionidae). Zootaxa, 2239, 62 - 68." type="journal article" year="2009" yearSuffix="b">Paulson (2009b)</bibRefCitation>
reared
<taxonomicName id="50FA37D7D67AFF89FA0A7684FE8EF93E" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67AFF89FA0A7684FE8EF93E" italics="true" pageId="3" pageNumber="4">L. guanacaste</emphasis>
</taxonomicName>
and provided diagnostic differences between the two.
</paragraph>
<paragraph id="97454C54D67AFF8EFF48773EFBFEFF34" blockId="3.[151,1437,151,2007]" lastBlockId="4.[151,1436,152,252]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Adults of
<taxonomicName id="50FA37D7D67AFF89FEB3773FFE2EF8D4" box="[317,444,1795,1820]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FEB3773FFE2EF8D4" box="[317,444,1795,1820]" italics="true" pageId="3" pageNumber="4">Leptobasis</emphasis>
</taxonomicName>
undergo ontogenetic changes in color and pattern; tenerals and immatures are often pale orange (with a dark mid-dorsal stripe in
<taxonomicName id="50FA37D7D67AFF89FD337714FCF5F889" box="[701,871,1832,1857]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67AFF89FD337714FCF5F889" box="[701,871,1832,1857]" italics="true" pageId="3" pageNumber="4">L. guanacaste</emphasis>
</taxonomicName>
) and, in some species, may develop dark middorsal (
<taxonomicName id="50FA37D7D67AFF89FF7E7771FE07F8AE" box="[240,405,1869,1894]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67AFF89FF7E7771FE07F8AE" box="[240,405,1869,1894]" italics="true" pageId="3" pageNumber="4">L. guanacaste</emphasis>
</taxonomicName>
) and humeral thoracic stripes against a pale green to orange background (
<taxonomicName id="50FA37D7D67AFF89FA8D7771FA06F8AE" box="[1283,1428,1869,1894]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="buchholzi">
<emphasis id="A58E9046D67AFF89FA8D7771FA06F8AE" box="[1283,1428,1869,1894]" italics="true" pageId="3" pageNumber="4">L. buchholzi</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67AFF89FF19774FFEA6F844" box="[151,308,1907,1932]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67AFF89FF19774FFEA6F844" box="[151,308,1907,1932]" italics="true" pageId="3" pageNumber="4">L. candelaria</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67AFF89FECC774FFE3EF844" box="[322,428,1907,1932]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lucifer">
<emphasis id="A58E9046D67AFF89FECC774FFE3EF844" box="[322,428,1907,1932]" italics="true" pageId="3" pageNumber="4">L. lucifer</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67AFF89FE34774FFDD8F844" box="[442,586,1907,1932]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="A58E9046D67AFF89FE34774FFDD8F844" box="[442,586,1907,1932]" italics="true" pageId="3" pageNumber="4">L. mauffrayi</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67AFF89FDD6774FFC9DF844" box="[600,783,1907,1932]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="melinogaster">
<emphasis id="A58E9046D67AFF89FDD6774FFC9DF844" box="[600,783,1907,1932]" italics="true" pageId="3" pageNumber="4">L. melinogaster</emphasis>
</taxonomicName>
,
<taxonomicName id="50FA37D7D67AFF89FC92774FFC37F844" box="[796,933,1907,1932]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67AFF89FC92774FFC37F844" box="[796,933,1907,1932]" italics="true" pageId="3" pageNumber="4">L. vacillans</emphasis>
</taxonomicName>
). In
<taxonomicName id="50FA37D7D67AFF89FC52774FFBDFF844" box="[988,1101,1907,1932]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="raineyi">
<emphasis id="A58E9046D67AFF89FC52774FFBDFF844" box="[988,1101,1907,1932]" italics="true" pageId="3" pageNumber="4">L. raineyi</emphasis>
</taxonomicName>
these stripes are represented as orange against a pale blue background, but a similar pattern can exist in other
<taxonomicName id="50FA37D7D67AFF89FBB177A4FB2CF879" box="[1087,1214,1944,1969]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A58E9046D67AFF89FBB177A4FB2CF879" box="[1087,1214,1944,1969]" italics="true" pageId="3" pageNumber="4">Leptobasis</emphasis>
</taxonomicName>
species mentioned above. The dynamics of these color changes are not understood and these differences led
<bibRefCitation id="F36B31A5D67AFF89FB267781FAD9F81F" author="Calvert" box="[1192,1355,1981,2007]" pageId="3" pageNumber="14" refString="Calvert, P. P. (1902) Odonata. In: Godman, F. D. &amp; O. Salvin (Eds.), Biologia Centrali Americana - Insecta Neuroptera. R. H. Porter and Dulau, London, pp. 73 - 128." type="book chapter" year="1902">Calvert (1902</bibRefCitation>
,
<bibRefCitation id="F36B31A5D67AFF89FAD97781FA01F81F" author="Calvert" box="[1367,1427,1981,2007]" pageId="3" pageNumber="14" refString="Calvert, P. P. (1909) Contributions to a knowledge of the Odonata of the Neotropical region, exclusive of Mexico and Central America. Annals of the Carnegie Museum, 6 (3), 73 - 280." type="journal article" year="1909">1909</bibRefCitation>
) to apply three varietal names to
<taxonomicName id="50FA37D7D67DFF8EFD8070A4FD04FF79" box="[526,662,152,177]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A58E9046D67DFF8EFD8070A4FD04FF79" box="[526,662,152,177]" italics="true" pageId="4" pageNumber="5">L. vacillans</emphasis>
</taxonomicName>
.
<bibRefCitation id="F36B31A5D67DFF8EFD2B70A4FCF6FF7A" author="Paulson" box="[677,868,152,178]" pageId="4" pageNumber="16" refString="Paulson, D. R. (2009 a) Dragonflies and damselflies of the West. Princeton University Press, Princeton, New Jersey, 535 pp." type="book" year="2009" yearSuffix="a">Paulson (2009a)</bibRefCitation>
suggests that immature colored adults may span the dry season in the tropics; he found adults and larvae of
<taxonomicName id="50FA37D7D67DFF8EFCEE7081FB9BFF1E" box="[864,1033,189,214]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67DFF8EFCEE7081FB9BFF1E" box="[864,1033,189,214]" italics="true" pageId="4" pageNumber="5">L. guanacaste</emphasis>
</taxonomicName>
at temporary pools in
<collectingCountry id="EFED0CC4D67DFF8EFA977081FA0EFF1F" box="[1305,1436,189,215]" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
during the wet season, and observed mature and immature-colored females in pairs.
</paragraph>
</subSubSection>
</treatment>
</document>
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