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<mods:title id="93A79A06D0F30B183031C1FFB27C6E50">Taxonomical notes on the family Ptilocodiidae (Anthomedusae) from the central and southern of South China Sea, with a new genus and a new species</mods:title>
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<mods:affiliation id="09D571F78069E9C3C80283689B539AF4">South China Sea Fisheries Research Institute, Chinese Academy of Fishery Science, Guangzhou 510300, China &amp; Key Laboratory of Fishery Ecology and Environment of Guangdong Province, Guangzhou 510300, China</mods:affiliation>
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<mods:affiliation id="FC8FF4B71A678C6C7C58B7D14791D02B">College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China</mods:affiliation>
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<heading id="D0B1B65E1F21C038FF4DA3ABC0B248EC" box="[136,645,1490,1514]" fontSize="10" level="2" pageId="1" pageNumber="237" reason="2">
Family
<taxonomicName id="4C467AB11F21C038FF1AA3ABC02E48EC" ID-CoL="7NKS8" authority="Coward, 1909" authorityName="Coward" authorityYear="1909" box="[223,537,1490,1514]" class="Hydrozoa" family="Ptilocodiidae" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="family">
Ptilocodiidae
<bibRefCitation id="EFD77CC31F21C038FEBFA3ABC02E48EC" author="Coward" box="[378,537,1490,1514]" pageId="1" pageNumber="237" pagination="729 - 735" refId="ref3510" refString="Coward, W. E. 1909. On Ptilocodium repens a new gymnoblastic hydroid epizoic on a pennatulid. Verslag van de Gewone Vergaderingen der Wis-En Natuurkundige Afdeeling, 17 (2): 729 - 735." type="journal article" year="1909">Coward, 1909</bibRefCitation>
</taxonomicName>
emend.
</heading>
</paragraph>
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<treatmentCitation id="0AE727231F21C038FF4DA38DC3FE4B0F" author="Coward" box="[136,457,1524,1546]" page="729" pageId="1" pageNumber="237" year="1909">
<taxonomicName id="4C467AB11F21C038FF4DA38DC3FE4B0F" ID-CoL="7NKS8" authority="Coward, 1909: 729" authorityName="Coward" authorityPageNumber="729" authorityYear="1909" box="[136,457,1524,1546]" class="Hydrozoa" family="Ptilocodiidae" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="family">
Ptilocodiidae
<bibRefCitation id="EFD77CC31F21C038FEC8A38DC3FE4B0F" author="Coward" box="[269,457,1524,1546]" pageId="1" pageNumber="237" pagination="729 - 735" refId="ref3510" refString="Coward, W. E. 1909. On Ptilocodium repens a new gymnoblastic hydroid epizoic on a pennatulid. Verslag van de Gewone Vergaderingen der Wis-En Natuurkundige Afdeeling, 17 (2): 729 - 735." type="journal article" year="1909">Coward, 1909: 729</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AE727231F21C038FE17A38DC0CB4B0C" author="Bouillon" box="[466,764,1524,1546]" page="151" pageId="1" pageNumber="237" year="2006">
<bibRefCitation id="EFD77CC31F21C038FE17A38DC0CB4B0C" author="Bouillon" box="[466,764,1524,1546]" pageId="1" pageNumber="237" pagination="1 - 591" refId="ref3466" refString="Bouillon, J., Gravili, C., Pages, F., Gili, J-M., Boero, F. 2006. An introduction to Hydrozoa. Memoires du Museum national d'Histoire Naturelle, 194: 1 - 591." type="journal article" year="2006">
Bouillon
<emphasis id="B932DD201F21C038FDE9A38CC0604B0F" box="[556,599,1524,1546]" italics="true" pageId="1" pageNumber="237">et al</emphasis>
., 2006: 151–153
</bibRefCitation>
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;
<treatmentCitation id="0AE727231F21C038FCC0A38DC6324B0F" author="Schuchert" box="[773,1029,1524,1546]" page="470" pageId="1" pageNumber="237" year="2009">
<bibRefCitation id="EFD77CC31F21C038FCC0A38DC6324B0F" author="Schuchert" box="[773,1029,1524,1546]" pageId="1" pageNumber="237" pagination="441 - 507" refId="ref4008" refString="Schuchert, P. 2009. The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 5. Revue Suisse de Zoologie, 116 (3 - 4): 441 - 507." type="journal article" year="2009">Schuchert, 2009: 470–471</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE727231F21C038FBCAA38CC7354B0C" author="Xu &amp; Huang &amp; Lin &amp; Guo &amp; Wang &amp; The Superclass Hydrozoa of the Phylum Cnidaria in China" box="[1039,1282,1524,1546]" page="264" pageId="1" pageNumber="237" year="2014">
<bibRefCitation id="EFD77CC31F21C038FBCAA38CC7354B0C" author="Xu &amp; Huang &amp; Lin &amp; Guo &amp; Wang &amp; The Superclass Hydrozoa of the Phylum Cnidaria in China" box="[1039,1282,1524,1546]" pageId="1" pageNumber="237" refId="ref4372" refString="Xu, Z. Z., Huang, J. Q., Lin, M., Guo, D. H., Wang, C. G. 2014. The Superclass Hydrozoa of the Phylum Cnidaria in China. China Ocean Press, Beijing. 945 pp." type="book" year="2014">
Xu
<emphasis id="B932DD201F21C038FBF7A38CC66B4B0F" box="[1074,1116,1524,1546]" italics="true" pageId="1" pageNumber="237">et al</emphasis>
., 2014: 264–265
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C35C52B91F21C038FF7BA050C7764B81" pageId="1" pageNumber="237" type="diagnosis">
<paragraph id="8BF901321F21C038FF7BA050C7764B81" blockId="1.[136,1459,1577,2087]" pageId="1" pageNumber="237">Diagnosis. Hydroid. Hydrorhiza stolonal, reticular, or encrusting, covered by naked coenosarc; hydranths sessile, naked and polymorphic; gastrozooid tubular, without tentacles; dactylozooids with 4 or more capitate tentacles, sometimes filiform; gonophores on gonozooids or gastro-gonozooids; developing into fixed sporosacs, eumedusoids or free medusae.</paragraph>
</subSubSection>
<subSubSection id="C35C52B91F21C038FF78A0E8C0854A51" pageId="1" pageNumber="237" type="description">
<paragraph id="8BF901321F21C038FF78A0E8C0854A51" blockId="1.[136,1459,1577,2087]" pageId="1" pageNumber="237">Medusa. Mature medusa with more or less bell-shaped umbrella, with or without radial exumbrellar furrows; with marginal nematocyst ring from which usually arise several didermic centripetal nematocyst bands (tracks) or exumbrellar rows of refringent spots; with four marginal tentacles or tentacles absent; four radial canals and circular canal; manubrium with 4 perradial mouth arms ending in nematocyst clusters or 4 simple unbranched perradial oral tentacles arising above mouth rim unarmed terminal nematocyst clusters; gonads four interradial or eight adradial masses on manubrium, or completely perradial gonads; with or without ocelli.</paragraph>
</subSubSection>
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<paragraph id="8BF901321F21C03BFF78A118C7334C02" blockId="1.[136,1459,1577,2087]" lastBlockId="2.[136,1451,236,503]" lastPageId="2" lastPageNumber="238" pageId="1" pageNumber="237">
Remarks. According to the original report (
<bibRefCitation id="EFD77CC31F21C038FD55A118C1024A7F" author="Du" box="[656,821,1889,1913]" pageId="1" pageNumber="237" pagination="506 - 519" refId="ref3687" refString="Du, F. Y., Xu, Z. Z., Huang, J. Q., Guo, D. H. 2012. Studies on the medusae (Cnidaria) from the Beibu Gulf in the northern South China Sea, with description of three new species. Acta Zootaxonomica Sinica, 37 (3): 506 - 519." type="journal article" year="2012">
Du
<emphasis id="B932DD201F21C038FD7FA11BC0DD4A7F" box="[698,746,1889,1913]" italics="true" pageId="1" pageNumber="237">et al</emphasis>
., 2012
</bibRefCitation>
) and by reexamining its shape, position and attachment of oral tentacles,
<taxonomicName id="4C467AB11F21C038FEE4A1FDC1284A9A" authority="Xu, Huang &amp; Du, 2012" authorityName="Xu, Huang &amp; Du" authorityYear="2012" box="[289,799,1924,1948]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="species" species="perradialis">
<emphasis id="B932DD201F21C038FEE4A1FDC0144A9A" box="[289,547,1924,1948]" italics="true" pageId="1" pageNumber="237">Tregoubovia perradialis</emphasis>
Xu, Huang &amp; Du, 2012
</taxonomicName>
is obvious different from other species of
<taxonomicName id="4C467AB11F21C038FB19A1FDC2F64AB8" authority="Picard, 1958" authorityName="Picard" authorityYear="1958" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F21C038FB19A1FDC7554A9A" box="[1244,1378,1924,1948]" italics="true" pageId="1" pageNumber="237">Tregoubovia</emphasis>
Picard, 1958
</taxonomicName>
. In
<taxonomicName id="4C467AB11F21C038FF34A1DEC3B14AB8" authorityName="Xu, Huang &amp; Du" authorityYear="2012" box="[241,390,1958,1982]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="species" species="perradialis">
<emphasis id="B932DD201F21C038FF34A1DEC3B14AB8" box="[241,390,1958,1982]" italics="true" pageId="1" pageNumber="237">T. perradialis</emphasis>
</taxonomicName>
, the oral tentacles are arising above mouth rim with ring nematocysts, while the oral arms of
<taxonomicName id="4C467AB11F21C038FF4DA1B0C3394AE7" authorityName="Picard" authorityYear="1958" box="[136,270,1993,2017]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F21C038FF4DA1B0C3394AE7" box="[136,270,1993,2017]" italics="true" pageId="1" pageNumber="237">Tregoubovia</emphasis>
</taxonomicName>
are extending from the perradial corners of mouth margin, and armed with terminal nematocyst clusters. The position of gonads is on perradial position of manubrium in
<taxonomicName id="4C467AB11F21C038FD3BA194C1B94502" authorityName="Xu, Huang &amp; Du" authorityYear="2012" box="[766,910,2028,2052]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="species" species="perradialis">
<emphasis id="B932DD201F21C038FD3BA194C1B94502" box="[766,910,2028,2052]" italics="true" pageId="1" pageNumber="237">T. perradialis</emphasis>
</taxonomicName>
, and interradial in
<taxonomicName id="4C467AB11F21C038FB96A195C6EE4502" authorityName="Picard" authorityYear="1958" box="[1107,1241,2028,2052]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F21C038FB96A195C6EE4502" box="[1107,1241,2028,2052]" italics="true" pageId="1" pageNumber="237">Tregoubovia</emphasis>
</taxonomicName>
. According to these,
<taxonomicName id="4C467AB11F21C038FF4DAE76C32D4520" authorityName="Xu, Huang &amp; Du" authorityYear="2012" box="[136,282,2062,2086]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="species" species="perradialis">
<emphasis id="B932DD201F21C038FF4DAE76C32D4520" box="[136,282,2062,2086]" italics="true" pageId="1" pageNumber="237">T. perradialis</emphasis>
</taxonomicName>
should be removed from the genus
<taxonomicName id="4C467AB11F21C038FD65AE77C1114520" authorityName="Picard" authorityYear="1958" box="[672,806,2062,2086]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F21C038FD65AE77C1114520" box="[672,806,2062,2086]" italics="true" pageId="1" pageNumber="237">Tregoubovia</emphasis>
</taxonomicName>
, and a new genus
<taxonomicName id="4C467AB11F21C038FC2AAE77C79B4521" authority="Guo, Xu &amp; Huang" authorityName="Guo, Xu &amp; Huang" authorityYear="2017" box="[1007,1452,2062,2087]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregouboviopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="237" phylum="Cnidaria" rank="genus" status="gen.">
<emphasis id="B932DD201F21C038FC2AAE77C6A94520" box="[1007,1182,2062,2086]" italics="true" pageId="1" pageNumber="237">Tregouboviopsis</emphasis>
Guo, Xu &amp; Huang,
<emphasis id="B932DD201F21C038FAB8AE76C79B4521" bold="true" box="[1405,1452,2063,2087]" pageId="1" pageNumber="237">gen.</emphasis>
</taxonomicName>
<emphasis id="B932DD201F22C03BFF4DA694C2804C03" bold="true" box="[136,183,237,261]" pageId="2" pageNumber="238">nov.</emphasis>
is proposed to accommodate it. The diagnosis was therefore adapted to fit the new scope of the family.
</paragraph>
<paragraph id="8BF901321F22C03BFF78A776C2DA4C6A" blockId="2.[136,1451,236,503]" pageId="2" pageNumber="238">
Thus, the family
<taxonomicName id="4C467AB11F22C03BFEBCA776C0314C21" authorityName="Coward" authorityYear="1909" box="[377,518,271,295]" class="Hydrozoa" family="Ptilocodiidae" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="family">Ptilocodiidae</taxonomicName>
comprises six genera now:
<taxonomicName id="4C467AB11F22C03BFCF2A776C6424C21" authority="Stechow, 1909" authorityName="Stechow" authorityYear="1909" box="[823,1141,271,295]" class="Hydrozoa" family="Ptilocodiidae" genus="Hydrichthella" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFCF2A776C1FC4C21" box="[823,971,271,295]" italics="true" pageId="2" pageNumber="238">Hydrichthella</emphasis>
<bibRefCitation id="EFD77CC31F22C03BFC11A776C6424C21" author="Stechow" box="[980,1141,271,295]" pageId="2" pageNumber="238" pagination="1 - 111" refId="ref4047" refString="Stechow, E. 1909. Beitrae zur Naturgeschichte Ostasiens: Hydroidpolypen der japanischen Ostkuste I Athecata und Plumularidae. A bhandlungen der Mathematisch-Physikalischen K 1 asse der Koniglichen Bayerischen Akademie der Wissenschaften, 1 (6): 1 - 111." type="journal article" year="1909">Stechow, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C467AB11F22C03BFB44A776C7904C21" authority="Coward, 1909" authorityName="Coward" authorityYear="1909" box="[1153,1447,271,295]" class="Hydrozoa" family="Ptilocodiidae" genus="Ptilocodium" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB44A776C7344C21" box="[1153,1283,271,295]" italics="true" pageId="2" pageNumber="238">Ptilocodium</emphasis>
<bibRefCitation id="EFD77CC31F22C03BFAC9A776C7904C21" author="Coward" box="[1292,1447,271,295]" pageId="2" pageNumber="238" pagination="729 - 735" refId="ref3510" refString="Coward, W. E. 1909. On Ptilocodium repens a new gymnoblastic hydroid epizoic on a pennatulid. Verslag van de Gewone Vergaderingen der Wis-En Natuurkundige Afdeeling, 17 (2): 729 - 735." type="journal article" year="1909">Coward, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C467AB11F22C03BFF4DA74BC3F74C4C" authority="Bouillon, 1967" authorityName="Bouillon" authorityYear="1967" box="[136,448,306,330]" class="Hydrozoa" family="Ptilocodiidae" genus="Thecocodium" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFF4DA74BC32F4C4C" box="[136,280,306,330]" italics="true" pageId="2" pageNumber="238">Thecocodium</emphasis>
Bouillon, 1967
</taxonomicName>
,
<taxonomicName id="4C467AB11F22C03BFE0EA74BC0EC4C4C" authority="Bouillon, 1980" authorityName="Bouillon" authorityYear="1980" box="[459,731,306,330]" class="Hydrozoa" family="Ptilocodiidae" genus="Hansiella" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFE0EA74BC0044C4C" box="[459,563,306,330]" italics="true" pageId="2" pageNumber="238">Hansiella</emphasis>
Bouillon, 1980
</taxonomicName>
,
<taxonomicName id="4C467AB11F22C03BFD20A74BC1CF4C4F" authority="Picard, 1958" authorityName="Picard" authorityYear="1958" box="[741,1016,305,330]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFD20A74BC15C4C4C" box="[741,875,306,330]" italics="true" pageId="2" pageNumber="238">Tregoubovia</emphasis>
Picard, 1958
</taxonomicName>
and
<taxonomicName id="4C467AB11F22C03BFBEEA74BC2804C6A" authority="Guo, Xu &amp; Huang" authorityName="Guo, Xu &amp; Huang" authorityYear="2017" class="Hydrozoa" family="Ptilocodiidae" genus="Tregouboviopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="237" phylum="Cnidaria" rank="genus" status="gen.">
<emphasis id="B932DD201F22C03BFBEEA74BC6ED4C4C" box="[1067,1242,306,330]" italics="true" pageId="2" pageNumber="238">Tregouboviopsis</emphasis>
Guo, Xu &amp; Huang,
<taxonomicNameLabel id="A201605B1F22C03BFF4DA72DC2804C6A" box="[136,183,340,364]" pageId="2" pageNumber="238" rank="genus">
<emphasis id="B932DD201F22C03BFF4DA72DC2804C6A" bold="true" box="[136,183,340,364]" pageId="2" pageNumber="238">gen.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B932DD201F22C03BFF7BA72DC2DA4C6A" bold="true" box="[190,237,340,364]" pageId="2" pageNumber="238">nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35C52B91F22C03BFF78A70EC3DF4F2A" pageId="2" pageNumber="238" type="multiple">
<paragraph id="8BF901321F22C03BFF78A70EC2CE4CB4" blockId="2.[136,1451,236,503]" pageId="2" pageNumber="238">
For a recent revision of the family
<taxonomicName id="4C467AB11F22C03BFDEFA70EC0804C89" authorityName="Coward" authorityYear="1909" box="[554,695,375,399]" class="Hydrozoa" family="Ptilocodiidae" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="family">Ptilocodiidae</taxonomicName>
see
<bibRefCitation id="EFD77CC31F22C03BFD21A70EC1FC4C89" author="Bouillon" box="[740,971,375,399]" pageId="2" pageNumber="238" pagination="1 - 591" refId="ref3466" refString="Bouillon, J., Gravili, C., Pages, F., Gili, J-M., Boero, F. 2006. An introduction to Hydrozoa. Memoires du Museum national d'Histoire Naturelle, 194: 1 - 591." type="journal article" year="2006">
Bouillon
<emphasis id="B932DD201F22C03BFC82A701C1424C89" box="[839,885,375,399]" italics="true" pageId="2" pageNumber="238">et al</emphasis>
. (2006)
</bibRefCitation>
,
<bibRefCitation id="EFD77CC31F22C03BFC10A70EC6B84C89" author="Schuchert" box="[981,1167,375,399]" pageId="2" pageNumber="238" pagination="441 - 507" refId="ref4008" refString="Schuchert, P. 2009. The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 5. Revue Suisse de Zoologie, 116 (3 - 4): 441 - 507." type="journal article" year="2009">Schuchert (2009)</bibRefCitation>
,
<bibRefCitation id="EFD77CC31F22C03BFB5CA70EC7084C89" author="Du" box="[1177,1343,375,399]" pageId="2" pageNumber="238" pagination="506 - 519" refId="ref3687" refString="Du, F. Y., Xu, Z. Z., Huang, J. Q., Guo, D. H. 2012. Studies on the medusae (Cnidaria) from the Beibu Gulf in the northern South China Sea, with description of three new species. Acta Zootaxonomica Sinica, 37 (3): 506 - 519." type="journal article" year="2012">
Du
<emphasis id="B932DD201F22C03BFB7AA701C6DA4C89" box="[1215,1261,375,399]" italics="true" pageId="2" pageNumber="238">et al</emphasis>
. (2012)
</bibRefCitation>
and
<bibRefCitation id="EFD77CC31F22C03BFAB7A70EC2C24CB4" author="Xu &amp; Huang &amp; Lin &amp; Guo &amp; Wang &amp; The Superclass Hydrozoa of the Phylum Cnidaria in China" pageId="2" pageNumber="238" refId="ref4372" refString="Xu, Z. Z., Huang, J. Q., Lin, M., Guo, D. H., Wang, C. G. 2014. The Superclass Hydrozoa of the Phylum Cnidaria in China. China Ocean Press, Beijing. 945 pp." type="book" year="2014">
Xu
<emphasis id="B932DD201F22C03BFA5DA701C2AA4CB4" italics="true" pageId="2" pageNumber="238">et al</emphasis>
. (2014)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BF901321F22C03BFF78A7C4C3374CF1" blockId="2.[136,1451,236,503]" pageId="2" pageNumber="238">
From the Chinese waters, two genera,
<taxonomicName id="4C467AB11F22C03BFD92A7C5C0DC4CD2" authorityName="Stechow" authorityYear="1909" box="[599,747,444,468]" class="Hydrozoa" family="Ptilocodiidae" genus="Hydrichthella" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFD92A7C5C0DC4CD2" box="[599,747,444,468]" italics="true" pageId="2" pageNumber="238">Hydrichthella</emphasis>
</taxonomicName>
as hydroid and free eumedusoid, and
<taxonomicName id="4C467AB11F22C03BFB44A7C5C7074CD2" box="[1153,1328,444,468]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregouboviopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB44A7C5C7074CD2" box="[1153,1328,444,468]" italics="true" pageId="2" pageNumber="238">Tregouboviopsis</emphasis>
</taxonomicName>
as medusa, are known.
</paragraph>
<paragraph id="8BF901321F22C03BFF4DA46FC3DF4F2A" blockId="2.[136,1450,534,984]" box="[136,488,534,556]" pageId="2" pageNumber="238">
Key to the genera of
<taxonomicName id="4C467AB11F22C03BFE98A46FC3D34F2A" authorityName="Coward" authorityYear="1909" box="[349,484,534,556]" class="Hydrozoa" family="Ptilocodiidae" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="family">Ptilocodiidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C52B91F22C03BFF4DA43AC79D4ED1" pageId="2" pageNumber="238" type="key">
<key id="FFD756031F22C03BFF4DA43AC79D4ED1" pageId="2" pageNumber="238">
<keyStep id="30B219971F22C03BFF4DA43AC79E4F7E" pageId="2" pageNumber="238">
<paragraph id="8BF901321F22C03BFF4DA43AC79E4F5E" blockId="2.[136,1450,534,984]" box="[136,1449,578,600]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF4DA43AC79E4F5E" box="[136,1449,578,600]" pageId="2" pageNumber="238">1. Hydroids............................................................................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BF901321F22C03BFF7DA41BC79E4F7E" blockId="2.[136,1450,534,984]" box="[184,1449,610,632]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF7DA41BC79E4F7E" box="[184,1449,610,632]" pageId="2" pageNumber="238">Medusae ...............................................................................................................................................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B219971F22C03BFF4DA4FAC79E4FBE" pageId="2" pageNumber="238">
<paragraph id="8BF901321F22C03BFF4DA4FAC79E4F91" blockId="2.[136,1450,534,984]" box="[136,1449,642,664]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF4DA4FAC79E4F91" box="[136,1449,642,664]" pageId="2" pageNumber="238">
2. Dactylozooids with two types .................................................................................................................
<taxonomicName id="4C467AB11F22C03BFB4EA4FBC79E4F91" ID-CoL="7PKYZ" authority="Stechow, 1909" authorityName="Stechow" authorityYear="1909" box="[1163,1449,642,664]" class="Hydrozoa" family="Ptilocodiidae" genus="Hydrichthella" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB4EA4FBC79E4F91" bold="true" box="[1163,1449,642,664]" pageId="2" pageNumber="238">
<emphasis id="B932DD201F22C03BFB4EA4FBC7234F9E" bold="true" box="[1163,1300,642,664]" italics="true" pageId="2" pageNumber="238">Hydrichthella</emphasis>
<bibRefCitation id="EFD77CC31F22C03BFADEA4FBC79E4F91" author="Stechow" box="[1307,1449,642,664]" pageId="2" pageNumber="238" pagination="1 - 111" refId="ref4047" refString="Stechow, E. 1909. Beitrae zur Naturgeschichte Ostasiens: Hydroidpolypen der japanischen Ostkuste I Athecata und Plumularidae. A bhandlungen der Mathematisch-Physikalischen K 1 asse der Koniglichen Bayerischen Akademie der Wissenschaften, 1 (6): 1 - 111." type="journal article" year="1909">Stechow, 1909</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF901321F22C03BFF7DA4DBC79E4FBE" blockId="2.[136,1450,534,984]" box="[184,1449,674,696]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF7DA4DBC79E4FBE" box="[184,1449,674,696]" pageId="2" pageNumber="238">Dactylozooids with one type................................................................................................................................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B219971F22C03BFF4DA4BAC79E4FF1" pageId="2" pageNumber="238">
<paragraph id="8BF901321F22C03BFF4DA4BAC79E4FD1" blockId="2.[136,1450,534,984]" box="[136,1449,706,728]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF4DA4BAC79E4FD1" box="[136,1449,706,728]" pageId="2" pageNumber="238">
3. Hydrorhiza crust-like, not covered by visible perisarc...............................................................................
<taxonomicName id="4C467AB11F22C03BFB58A4BBC79E4FD1" ID-CoL="7PWY9" authority="Coward, 1909" authorityName="Coward" authorityYear="1909" box="[1181,1449,706,728]" class="Hydrozoa" family="Ptilocodiidae" genus="Ptilocodium" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB58A4BBC79E4FD1" bold="true" box="[1181,1449,706,728]" pageId="2" pageNumber="238">
<emphasis id="B932DD201F22C03BFB58A4BBC7224FDE" bold="true" box="[1181,1301,706,728]" italics="true" pageId="2" pageNumber="238">Ptilocodium</emphasis>
<bibRefCitation id="EFD77CC31F22C03BFADEA4BBC79E4FD1" author="Coward" box="[1307,1449,706,728]" pageId="2" pageNumber="238" pagination="729 - 735" refId="ref3510" refString="Coward, W. E. 1909. On Ptilocodium repens a new gymnoblastic hydroid epizoic on a pennatulid. Verslag van de Gewone Vergaderingen der Wis-En Natuurkundige Afdeeling, 17 (2): 729 - 735." type="journal article" year="1909">Coward, 1909</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF901321F22C03BFF7DA49BC79E4FF1" blockId="2.[136,1450,534,984]" box="[184,1449,738,760]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF7DA49BC79E4FF1" box="[184,1449,738,760]" pageId="2" pageNumber="238">
Hydrorhiza a network of perisarc-protected tube-like stolons.................................................................
<taxonomicName id="4C467AB11F22C03BFB4FA49BC79E4FF1" ID-CoL="7Q4B6" authority="Bouillon, 1967" authorityName="Bouillon" authorityYear="1967" box="[1162,1449,738,760]" class="Hydrozoa" family="Ptilocodiidae" genus="Thecocodium" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB4FA49BC79E4FF1" bold="true" box="[1162,1449,738,760]" pageId="2" pageNumber="238">
<emphasis id="B932DD201F22C03BFB4FA49BC7264FFE" bold="true" box="[1162,1297,738,760]" italics="true" pageId="2" pageNumber="238">Thecocodium</emphasis>
Bouillon, 1967
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B219971F22C03BFF4DA57AC79E4E3E" pageId="2" pageNumber="238">
<paragraph id="8BF901321F22C03BFF4DA57AC79E4E1E" blockId="2.[136,1450,534,984]" box="[136,1449,770,792]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF4DA57AC79E4E1E" box="[136,1449,770,792]" pageId="2" pageNumber="238">4. Without marginal tentacles................................................................................................................................................................... 5</keyLead>
</paragraph>
<paragraph id="8BF901321F22C03BFF7DA55BC79E4E3E" blockId="2.[136,1450,534,984]" box="[184,1449,802,824]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF7DA55BC79E4E3E" box="[184,1449,802,824]" pageId="2" pageNumber="238">With marginal tentacles........................................................................................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B219971F22C03BFF4DA53BC79E4E9E" pageId="2" pageNumber="238">
<paragraph id="8BF901321F22C03BFF4DA53BC79D4E51" blockId="2.[136,1450,534,984]" box="[136,1450,834,856]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF4DA53BC79D4E51" box="[136,1450,834,856]" pageId="2" pageNumber="238">
5. Four perradial oral arms, ending swollen with nematocyst clusters; gonads interradial..............................
<taxonomicName id="4C467AB11F22C03BFB62A53BC79D4E51" ID-CoL="7Q4X6" authority="Picard, 1958" authorityName="Picard" authorityYear="1958" box="[1191,1450,834,856]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregoubovia" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB62A53BC79D4E51" bold="true" box="[1191,1450,834,856]" pageId="2" pageNumber="238">
<emphasis id="B932DD201F22C03BFB62A53BC7144E5E" bold="true" box="[1191,1315,834,856]" italics="true" pageId="2" pageNumber="238">Tregoubovia</emphasis>
Picard, 1958
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF901321F22C03BFF7DA51AC79E4E9E" blockId="2.[136,1450,534,984]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF7DA51AC79E4E9E" pageId="2" pageNumber="238">
Four perradial simple oral tentacles, arising above mouth rim, without terminal nematocyst clusters; gonads perradial ...................... ...................................................................................................................................
<emphasis id="B932DD201F22C03BFC25A5FBC79E4E9E" bold="true" box="[992,1449,898,920]" pageId="2" pageNumber="238">
<taxonomicName id="4C467AB11F22C03BFC25A5FBC74E4E9E" ID-CoL="7Q4X7" authority="Guo, Xu &amp; Huang" authorityName="Guo, Xu &amp; Huang" authorityYear="2017" box="[992,1401,898,920]" class="Hydrozoa" family="Ptilocodiidae" genus="Tregouboviopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="237" phylum="Cnidaria" rank="genus" status="gen.">
<emphasis id="B932DD201F22C03BFC25A5FBC6B64E9E" bold="true" box="[992,1153,898,920]" italics="true" pageId="2" pageNumber="238">Tregouboviopsis</emphasis>
Guo, Xu &amp; Huang,
<taxonomicNameLabel id="A201605B1F22C03BFA8AA5FAC74E4E9E" box="[1359,1401,899,920]" pageId="2" pageNumber="238" rank="genus">gen.</taxonomicNameLabel>
</taxonomicName>
nov.
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B219971F22C03BFF4DA5DBC79D4ED1" pageId="2" pageNumber="238">
<paragraph id="8BF901321F22C03BFF4DA5DBC79D4EB1" blockId="2.[136,1450,534,984]" box="[136,1450,930,952]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF4DA5DBC79D4EB1" box="[136,1450,930,952]" pageId="2" pageNumber="238">
6. Four interradial gonads ...........................................................................................................................
<taxonomicName id="4C467AB11F22C03BFB4FA5DBC79D4EB1" ID-CoL="7Q4B6" authority="Bouillon, 1967" authorityName="Bouillon" authorityYear="1967" box="[1162,1450,930,952]" class="Hydrozoa" family="Ptilocodiidae" genus="Thecocodium" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB4FA5DBC79D4EB1" bold="true" box="[1162,1450,930,952]" pageId="2" pageNumber="238">
<emphasis id="B932DD201F22C03BFB4FA5DBC7264EBE" bold="true" box="[1162,1297,930,952]" italics="true" pageId="2" pageNumber="238">Thecocodium</emphasis>
Bouillon, 1967
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF901321F22C03BFF7DA5BBC79D4ED1" blockId="2.[136,1450,534,984]" box="[184,1450,962,984]" pageId="2" pageNumber="238">
<keyLead id="30B7A2071F22C03BFF7DA5BBC79D4ED1" box="[184,1450,962,984]" pageId="2" pageNumber="238">
Eight adradial gonads....................................................................................................................................
<taxonomicName id="4C467AB11F22C03BFB74A5BBC79D4ED1" ID-CoL="7PJBD" authority="Bouillon, 1980" authorityName="Bouillon" authorityYear="1980" box="[1201,1450,962,984]" class="Hydrozoa" family="Ptilocodiidae" genus="Hansiella" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="238" phylum="Cnidaria" rank="genus">
<emphasis id="B932DD201F22C03BFB74A5BBC79D4ED1" bold="true" box="[1201,1450,962,984]" pageId="2" pageNumber="238">
<emphasis id="B932DD201F22C03BFB74A5BBC7264EDE" bold="true" box="[1201,1297,962,984]" italics="true" pageId="2" pageNumber="238">Hansiella</emphasis>
Bouillon, 1980
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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