992 lines
228 KiB
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992 lines
228 KiB
XML
<document id="27D493EAB97FA680375FB9D96C083169" ID-CLB-Dataset="30623" ID-DOI="10.11646/zootaxa.4387.3.8" ID-GBIF-Dataset="7548c751-764f-4dc4-95a5-614d17d80c8f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187739" ID-ZooBank="DA75D56E-22BB-487E-8BA3-42C5C98F9E53" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059648244" checkinUser="plazi" docAuthor="Yamasaki, Hiroshi, Neuhaus, Birger & George, Kai Horst" docDate="2018" docId="03CD87E02454FFC83D862F1009C7FDAF" docLanguage="en" docName="zootaxa.4387.3.8.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinoderes multiporus Yamasaki & Neuhaus & George 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="552" masterDocId="FFF4FF982456FFC33D11283B0B42FFCF" masterDocTitle="New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Mediterranean seamounts and from the deep-sea floor in the Northeast Atlantic Ocean, including notes on two undescribed species" masterLastPageNumber="566" masterPageNumber="541" pageNumber="543" updateTime="1698526505608" updateUser="ExternalLinkService">
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<mods:title id="BCA2E52D9FABFCEF7EF37751369B9906">New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Mediterranean seamounts and from the deep-sea floor in the Northeast Atlantic Ocean, including notes on two undescribed species</mods:title>
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<mods:namePart id="D6942CF189BD05E0E61C0CB636C20494">Yamasaki, Hiroshi</mods:namePart>
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<mods:namePart id="DF69B65C06D9F1B4436623956B2CBF2A">Neuhaus, Birger</mods:namePart>
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<mods:namePart id="E07F18D0F169E523C2687E4DFE0AA582">George, Kai Horst</mods:namePart>
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<mods:typeOfResource id="B7526FF2736057513E83BE501DAD2647">text</mods:typeOfResource>
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<mods:title id="85AAD46D0DFB7D1CA223D15CAE5592BC">Zootaxa</mods:title>
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<mods:part id="AF0695D4F862076140B0A48FB294FE1A">
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<mods:date id="554DED2851CB7E08B026EE35148CE2FF">2018</mods:date>
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<mods:number id="0A6608F7F581A5798ACCC65EE3A8B023">2018-02-28</mods:number>
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<mods:number id="AC1FE27E3A0AB5C3F021932FDB89A48D">4387</mods:number>
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<treatment id="03CD87E02454FFC83D862F1009C7FDAF" ID-DOI="http://doi.org/10.5281/zenodo.5952903" ID-GBIF-Taxon="141264089" ID-Zenodo-Dep="5952903" LSID="urn:lsid:plazi:treatment:03CD87E02454FFC83D862F1009C7FDAF" httpUri="http://treatment.plazi.org/id/03CD87E02454FFC83D862F1009C7FDAF" lastPageId="11" lastPageNumber="552" pageId="2" pageNumber="543">
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<subSubSection id="C37E657D2454FFC13D862F100AD1F8A9" pageId="2" pageNumber="543" type="nomenclature">
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<paragraph id="8BDB36F62454FFC13D862F10095CF88A" blockId="2.[151,542,1835,1894]" box="[151,542,1835,1861]" pageId="2" pageNumber="543">
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<heading id="D093819A2454FFC13D862F10095CF88A" bold="true" box="[151,542,1835,1861]" fontSize="11" level="1" pageId="2" pageNumber="543" reason="1">
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<emphasis id="B910EAE42454FFC13D862F10095CF88A" bold="true" box="[151,542,1835,1861]" pageId="2" pageNumber="543">
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<taxonomicName id="4C644D752454FFC13D862F100AFBF88A" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[151,441,1835,1861]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="2" pageNumber="543" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
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<emphasis id="B910EAE42454FFC13D862F100AFBF88A" bold="true" box="[151,441,1835,1861]" italics="true" pageId="2" pageNumber="543">Echinoderes multiporus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A223579F2454FFC13CD02F10095CF88A" box="[449,542,1835,1861]" pageId="2" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDB36F62454FFC13D862F760AD1F8A9" blockId="2.[151,542,1835,1894]" box="[151,403,1869,1894]" pageId="2" pageNumber="543">
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(
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<figureCitation id="135F2A732454FFC13D8E2F760A47F8A9" box="[159,261,1869,1894]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[151,250,1845,1867]" captionStartId-1="5.[151,250,1853,1875]" captionStartId-2="6.[151,250,1847,1869]" captionStartId-3="8.[151,250,1829,1851]" captionStartId-4="9.[151,250,1884,1906]" captionStartId-5="10.[151,250,1829,1851]" captionTargetBox-0="[232,1343,197,1832]" captionTargetBox-1="[233,1358,193,1842]" captionTargetBox-2="[236,1354,187,1837]" captionTargetBox-3="[162,1426,843,1776]" captionTargetBox-4="[161,1426,209,1855]" captionTargetBox-5="[247,1350,193,1818]" captionTargetId-0="figure@4.[232,1343,193,1832]" captionTargetId-1="figure@5.[233,1358,193,1842]" captionTargetId-2="figure@6.[236,1354,187,1837]" captionTargetId-3="figure@8.[161,1426,843,1776]" captionTargetId-4="figure@9.[161,1426,209,1856]" captionTargetId-5="figure@10.[247,1350,193,1818]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." captionText-1="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." captionText-2="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." captionText-3="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." captionText-4="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." captionText-5="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri-0="https://zenodo.org/record/1187743/files/figure.png" httpUri-1="https://zenodo.org/record/1187745/files/figure.png" httpUri-2="https://zenodo.org/record/1187747/files/figure.png" httpUri-3="https://zenodo.org/record/1187749/files/figure.png" httpUri-4="https://zenodo.org/record/1187751/files/figure.png" httpUri-5="https://zenodo.org/record/1187753/files/figure.png" pageId="2" pageNumber="543">Figs 2–7</figureCitation>
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;
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<tableCitation id="C6E6034D2454FFC13C012F760A32F8A9" box="[272,368,1869,1894]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Measurements for adult Echinoderes multiporus sp. nov. (in micrometers or as percentage). Abbreviations: (ac), acicular spine; LTAS, length of lateral terminal accessory spine; LTS, length of lateral terminal spine; LV, length of lateroventral spine/tube; MD, length of middorsal spine; ML, length of midlateral tube; MSW, maximum sternal width; N, number of specimens measured; S, segment length; SD, standard deviation; SW, standard width; TL, trunk length; (tu), tube; VL, length of ventrolateral tube." pageId="2" pageNumber="543">Tables 2</tableCitation>
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,
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<tableCitation id="C6E6034D2454FFC13C6D2F750AC8F8A9" box="[380,394,1870,1894]" captionStart="TABLE 3" captionStartId="11.[151,239,653,676]" captionText="TABLE 3. Summary of locations of cuticular structures and spines in Echinoderes multiporus sp. nov. Character appearing with variation marked as bold and underlined. Abbreviations: ac, acicular spine; (f), female condition of sexually dimorphiccharacter; gco1, type-1 glandular celloutlet; gco2, type-2 glandular cell outlet; LA, lateral accessory; LD, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; LV, lateroventral; (m), male condition of sexually dimorphic character; MD, middorsal; ML, midlateral; pa, papilla; pe, penile spine; PD, paradorsal; SD, subdorsal; si, sieve plate; SL, sublateral; ss, sensory spot; tu, tube; VL, ventrolateral; VM, ventromedial." pageId="2" pageNumber="543">3</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E657D2454FFC13D862FAE0E22F839" pageId="2" pageNumber="543" type="diagnosis">
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<paragraph id="8BDB36F62454FFC13D862FAE0E22F839" blockId="2.[151,1436,1941,2038]" pageId="2" pageNumber="543">
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<emphasis id="B910EAE42454FFC13D862FAE0A53F861" bold="true" box="[151,273,1941,1966]" pageId="2" pageNumber="543">Diagnosis.</emphasis>
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<taxonomicName id="4C644D752454FFC13C082FAE0AE6F862" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[281,420,1941,1965]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="2" pageNumber="543" phylum="Cephalorhyncha" rank="genus">
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<emphasis id="B910EAE42454FFC13C082FAE0AE6F862" box="[281,420,1941,1965]" italics="true" pageId="2" pageNumber="543">Echinoderes</emphasis>
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</taxonomicName>
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with middorsal acicular spines on segments 4, 6, and 8; ventrolateral tubes on segment 2; lateroventral tubes on segment 5; lateroventral acicular spines on segments 6–9; midlateral tubes on segment 10; type-2 glandular cell outlets in subdorsal position on segment 2 and in laterodorsal position on segments 4–9.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E657D2455FFC03DD628AC09E5FF1A" pageId="3" pageNumber="544" type="etymology">
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<paragraph id="8BDB36F62455FFC03DD628AC09E5FF1A" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
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<emphasis id="B910EAE42455FFC03DD628AC0A0CFF7F" bold="true" box="[199,334,151,176]" pageId="3" pageNumber="544">Etymology.</emphasis>
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The species name is composed of the Latin
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<emphasis id="B910EAE42455FFC03E5928A308C0FF7F" box="[840,898,152,176]" italics="true" pageId="3" pageNumber="544">multi</emphasis>
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(many) and Latin
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<emphasis id="B910EAE42455FFC0394728A20FD4FF7F" box="[1110,1174,153,176]" italics="true" pageId="3" pageNumber="544">porus</emphasis>
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(pore), referring to the type-2 glandular cell outlets on most segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E657D2455FFC03DD628E40E3AFDD7" pageId="3" pageNumber="544" type="materials_examined">
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<paragraph id="8BDB36F62455FFC03DD628E40F89FED2" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
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<emphasis id="B910EAE42455FFC03DD628E40AF4FF37" bold="true" box="[199,438,223,248]" pageId="3" pageNumber="544">Material examined.</emphasis>
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Holotype: Adult male (ZMB 11590), collected at station 1127 on the Eratosthenes Seamount (
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<figureCitation id="135F2A732455FFC03C07293F0AD0FED3" box="[278,402,260,285]" captionStart="FIGURE1" captionStartId="2.[151,250,1247,1269]" captionTargetBox="[191,1396,253,1229]" captionTargetId="figure@2.[191,1396,253,1233]" captionTargetPageId="2" captionText="FIGURE1. Map of the sampling localities.A, map of northeast Atlantic Ocean and Mediterranean Sea; B, enlarged map of the Sedlo seamount and surrounding deep-sea floor; C, enlarged map of the Anaximenes Seamount; D, enlarged map of the Eratosthenes Seamount." httpUri="https://zenodo.org/record/1187741/files/figure.png" pageId="3" pageNumber="544">Fig. 1A, D</figureCitation>
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;
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<tableCitation id="C6E6034D2455FFC03C8F293F0AB2FED3" box="[414,496,260,285]" captionStart="TABLE 1" captionStartId="1.[151,239,820,843]" captionText="TABLE 1. Data of sampling stations." pageId="3" pageNumber="544">Table 1</tableCitation>
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), mounted as glycerol-paraffin slide on a Cobb aluminum frame.
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</paragraph>
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<paragraph id="8BDB36F62455FFC03DD629130A26FE62" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
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<typeStatus id="54DF88542455FFC03DD629130A7BFE8F" box="[199,313,296,320]" pageId="3" pageNumber="544" type="paratype">Paratypes</typeStatus>
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: four adult females and two adult males (
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<collectionCode id="ED75AE332455FFC03E0F2913081EFE8F" box="[798,860,296,320]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="544">ZMB</collectionCode>
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11591a–11591f), collected at station 1095 on the Eratosthenes Seamount; one adult female and one adult male (
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<collectionCode id="ED75AE332455FFC03E47297708D6FEAB" box="[854,916,332,356]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="544">ZMB</collectionCode>
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11592a, 11592b), collected at station 1100 on the Eratosthenes Seamount (
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<figureCitation id="135F2A732455FFC03CF22954092AFE48" box="[483,616,367,392]" captionStart="FIGURE1" captionStartId="2.[151,250,1247,1269]" captionTargetBox="[191,1396,253,1229]" captionTargetId="figure@2.[191,1396,253,1233]" captionTargetPageId="2" captionText="FIGURE1. Map of the sampling localities.A, map of northeast Atlantic Ocean and Mediterranean Sea; B, enlarged map of the Sedlo seamount and surrounding deep-sea floor; C, enlarged map of the Anaximenes Seamount; D, enlarged map of the Eratosthenes Seamount." httpUri="https://zenodo.org/record/1187741/files/figure.png" pageId="3" pageNumber="544">Fig. 1A, D</figureCitation>
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;
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<tableCitation id="C6E6034D2455FFC03F682954098DFE47" box="[633,719,367,392]" captionStart="TABLE 1" captionStartId="1.[151,239,820,843]" captionText="TABLE 1. Data of sampling stations." pageId="3" pageNumber="544">Table 1</tableCitation>
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). All
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<typeStatus id="54DF88542455FFC03E0D294B08C5FE47" box="[796,903,368,392]" pageId="3" pageNumber="544" type="paratype">paratypes</typeStatus>
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mounted as glycerol-paraffin slides on Cobb aluminum frames.
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</paragraph>
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<paragraph id="8BDB36F62455FFC03DD6298C0E2EFE3A" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
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Additional material: one adult female and three adult males, collected at station 1095 on the Eratosthenes Seamount (
|
||
<figureCitation id="135F2A732455FFC03C0729E70AD0FE3B" box="[278,402,476,501]" captionStart="FIGURE1" captionStartId="2.[151,250,1247,1269]" captionTargetBox="[191,1396,253,1229]" captionTargetId="figure@2.[191,1396,253,1233]" captionTargetPageId="2" captionText="FIGURE1. Map of the sampling localities.A, map of northeast Atlantic Ocean and Mediterranean Sea; B, enlarged map of the Sedlo seamount and surrounding deep-sea floor; C, enlarged map of the Anaximenes Seamount; D, enlarged map of the Eratosthenes Seamount." httpUri="https://zenodo.org/record/1187741/files/figure.png" pageId="3" pageNumber="544">Fig. 1A, D</figureCitation>
|
||
;
|
||
<tableCitation id="C6E6034D2455FFC03C8F29E70AB2FE3B" box="[414,496,476,501]" captionStart="TABLE 1" captionStartId="1.[151,239,820,843]" captionText="TABLE 1. Data of sampling stations." pageId="3" pageNumber="544">Table 1</tableCitation>
|
||
), mounted on an aluminum stub for SEM observations (ZMB
|
||
<quantity id="4C9C9B132455FFC039BC29E70E40FE3A" box="[1197,1282,476,501]" metricMagnitude="1" metricUnit="kg" metricValue="1.1591" pageId="3" pageNumber="544" unit="g" value="11591.0">11591g</quantity>
|
||
–11591j).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD629C40E3AFDD7" blockId="3.[151,1437,151,1977]" box="[199,1400,511,536]" pageId="3" pageNumber="544">
|
||
<emphasis id="B910EAE42455FFC03DD629C40A27FDD7" bold="true" box="[199,357,511,536]" pageId="3" pageNumber="544">Type locality.</emphasis>
|
||
Eratosthenes Seamount (33°49'49.80"N, 32°47'49.20"E),
|
||
<quantity id="4C9C9B132455FFC03EE32A3B0F7BFDD8" box="[1010,1081,512,536]" metricMagnitude="2" metricUnit="m" metricValue="9.91" pageId="3" pageNumber="544" unit="m" value="991.0">991 m</quantity>
|
||
depth (
|
||
<figureCitation id="135F2A732455FFC0399F29C40E4BFDD8" box="[1166,1289,511,536]" captionStart="FIGURE1" captionStartId="2.[151,250,1247,1269]" captionTargetBox="[191,1396,253,1229]" captionTargetId="figure@2.[191,1396,253,1233]" captionTargetPageId="2" captionText="FIGURE1. Map of the sampling localities.A, map of northeast Atlantic Ocean and Mediterranean Sea; B, enlarged map of the Sedlo seamount and surrounding deep-sea floor; C, enlarged map of the Anaximenes Seamount; D, enlarged map of the Eratosthenes Seamount." httpUri="https://zenodo.org/record/1187741/files/figure.png" pageId="3" pageNumber="544">Fig. 1A, D</figureCitation>
|
||
;
|
||
<tableCitation id="C6E6034D2455FFC0380429C40E25FDD7" box="[1301,1383,511,536]" captionStart="TABLE 1" captionStartId="1.[151,239,820,843]" captionText="TABLE 1. Data of sampling stations." pageId="3" pageNumber="544">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E657D2455FFCB3DD62A1F0B95FE63" lastPageId="8" lastPageNumber="549" pageId="3" pageNumber="544" type="description">
|
||
<paragraph id="8BDB36F62455FFC03DD62A1F0AE0FD4A" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
<emphasis id="B910EAE42455FFC03DD62A1F0A1BFDF2" bold="true" box="[199,345,548,573]" pageId="3" pageNumber="544">Description.</emphasis>
|
||
Adult with head, neck, and eleven trunk segments (
|
||
<figureCitation id="135F2A732455FFC03EAF2A1F0F01FDF3" box="[958,1091,548,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC039432A1F0F34FDF3" box="[1106,1142,548,572]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC039972A1F0F8DFDF3" box="[1158,1231,548,572]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, B</figureCitation>
|
||
). See
|
||
<tableCitation id="C6E6034D2455FFC0380B2A1F0E33FDF3" box="[1306,1393,548,573]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Measurements for adult Echinoderes multiporus sp. nov. (in micrometers or as percentage). Abbreviations: (ac), acicular spine; LTAS, length of lateral terminal accessory spine; LTS, length of lateral terminal spine; LV, length of lateroventral spine/tube; MD, length of middorsal spine; ML, length of midlateral tube; MSW, maximum sternal width; N, number of specimens measured; S, segment length; SD, standard deviation; SW, standard width; TL, trunk length; (tu), tube; VL, length of ventrolateral tube." pageId="3" pageNumber="544">Table 2</tableCitation>
|
||
for measurements.
|
||
<tableCitation id="C6E6034D2455FFC03C572A7C0AD8FDAF" box="[326,410,583,608]" captionStart="TABLE 3" captionStartId="11.[151,239,653,676]" captionText="TABLE 3. Summary of locations of cuticular structures and spines in Echinoderes multiporus sp. nov. Character appearing with variation marked as bold and underlined. Abbreviations: ac, acicular spine; (f), female condition of sexually dimorphiccharacter; gco1, type-1 glandular celloutlet; gco2, type-2 glandular cell outlet; LA, lateral accessory; LD, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; LV, lateroventral; (m), male condition of sexually dimorphic character; MD, middorsal; ML, midlateral; pa, papilla; pe, penile spine; PD, paradorsal; SD, subdorsal; si, sieve plate; SL, sublateral; ss, sensory spot; tu, tube; VL, ventrolateral; VM, ventromedial." pageId="3" pageNumber="544">Table 3</tableCitation>
|
||
indicates the positions of cuticular structures (sensory spots, glandular cell outlets, spines, tubes, and sieve plates).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62AB409C0FCDA" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">Head consisting of retractable mouth cone and introvert. Mouth cone with inner oral styles and nine outer oral styles. Introvert composed of one ring of primary scalids, several rings of spinoscalids, and one ring of trichoscalids. Exact number and arrangement of inner and outer oral styles and scalids not traceable because of withdrawn head in all examined specimens.</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62B24086BFC92" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Neck with 16 placids (
|
||
<figureCitation id="135F2A732455FFC03CD72B240904FCF8" box="[454,582,799,824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03F422B1B0934FCF7" box="[595,630,800,824]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03F932B1B0988FCF7" box="[642,714,800,824]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C, D</figureCitation>
|
||
). Midventral placid broadest. Remaining placids similar in size. Two trichoscalid plates present ventrally and four dorsally.
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62B530F3FFB97" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 1 consisting of complete cuticular ring. This and following nine segments with thick pachycyclus at anterior margin of each segment (
|
||
<figureCitation id="135F2A732455FFC03F0A2BB709DCFC6B" box="[539,670,908,933]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03FBA2BB709B1FC6B" box="[683,755,908,932]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, B</figureCitation>
|
||
). Pachycyclus interrupted middorsally in segments 2–10 as well as at tergosternal junctions in segments 3–10. Sensory spots located in subdorsal and laterodorsal position (
|
||
<figureCitation id="135F2A732455FFC03D8E2BEF0A64FC23" box="[159,294,980,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C272BEF0A1BFC23" box="[310,345,980,1004]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C782BEF0ACCFC23" box="[361,398,980,1004]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C</figureCitation>
|
||
). A couple of hairs present beside each sensory spot (
|
||
<figureCitation id="135F2A732455FFC039022BEF0F30FC23" box="[1043,1138,980,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">Fig. 3D</figureCitation>
|
||
). Several additional hairs sparsely distributed. Type-1 glandular cell outlets present in middorsal and lateroventral position (
|
||
<figureCitation id="135F2A732455FFC039F82BCC0E2AFBC0" box="[1257,1384,1015,1040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038652BC30ED5FBDF" box="[1396,1431,1016,1040]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03D862C270B9DFBFB" box="[151,223,1052,1076]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C, D</figureCitation>
|
||
). Posterior part of this and following nine segments with primary pectinate fringe (
|
||
<figureCitation id="135F2A732455FFC039832C270E56FBFB" box="[1170,1300,1052,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038312C270E27FBFB" box="[1312,1381,1052,1076]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038622C270EDAFBFB" box="[1395,1432,1052,1076]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03D862C040BFFFB97" box="[151,189,1087,1112]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="3" pageNumber="544">5A</figureCitation>
|
||
). Pectinate fringe teeth of primary pectinate fringe on this segment medium in length.
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62C5F0960FA9B" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 2 with complete cuticular ring as segment 1. Ventrolateral tubes present (
|
||
<figureCitation id="135F2A732455FFC039792C5F0F87FBB3" box="[1128,1221,1124,1149]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC039C32C5F0FB6FBB3" box="[1234,1268,1124,1148]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038102C5F0E64FBB3" box="[1281,1318,1124,1148]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038232C5F0E1AFBB2" box="[1330,1368,1124,1149]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="3" pageNumber="544">5A</figureCitation>
|
||
). Few cuticular hairs rising from perforation sites in central to posterior area of this and following seven segments (
|
||
<figureCitation id="135F2A732455FFC0387B2CBC0B9EFB0B" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">Figs 3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03DFB2C970A70FB0B" box="[234,306,1196,1220]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C2E2C970A26FB0A" box="[319,356,1196,1221]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="3" pageNumber="544">5A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C602C970AF5FB0A" box="[369,439,1196,1221]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6A–C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CD52C970973FB0B" box="[452,561,1196,1220]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="3" pageNumber="544">7A, B, D</figureCitation>
|
||
). Sensory spots present in middorsal, laterodorsal and ventromedial position (
|
||
<figureCitation id="135F2A732455FFC03D8E2CF40A5FFB27" box="[159,285,1231,1256]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C382CEB0A2FFB27" box="[297,365,1232,1256]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3C–E</figureCitation>
|
||
). Type-1 glandular cell outlet present in middorsal and ventromedial position (
|
||
<figureCitation id="135F2A732455FFC039F12CF40E1DFB27" box="[1248,1375,1231,1256]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC0387A2CEB0ED2FB27" box="[1387,1424,1232,1256]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C</figureCitation>
|
||
). Type-2 glandular cell outlets in subdorsal position (
|
||
<figureCitation id="135F2A732455FFC03FC72CCF0870FAC3" box="[726,818,1268,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E2C2CCF08C1FAC3" box="[829,899,1268,1292]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, C</figureCitation>
|
||
). Primary pectinate fringe on this and following six segments similar to fringe on segment 1, but with slightly longer pectinate fringe teeth in midlateral to ventrolateral area (
|
||
<figureCitation id="135F2A732455FFC03C782D070A84FA9B" box="[361,454,1340,1365]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CC32D070957FA9B" box="[466,533,1340,1364]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62D5B0917FA0F" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 3 and following eight segments consisting of one tergal and two sternal plates. Sensory spots present in subdorsal and sublateral position (
|
||
<figureCitation id="135F2A732455FFC03F212DBF09EFFA53" box="[560,685,1412,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03FA92DBF099BFA53" box="[696,729,1412,1436]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03FF52DBF0869FA53" box="[740,811,1412,1436]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C, D</figureCitation>
|
||
). Type-1 glandular cell outlets situated in middorsal and ventromedial position (
|
||
<figureCitation id="135F2A732455FFC03C8A2D9C095BFA0F" box="[411,537,1447,1472]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03F352D93090BFA0F" box="[548,585,1448,1472]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62DF70901F9E3" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 4 with middorsal acicular spine (
|
||
<figureCitation id="135F2A732455FFC03F8D2DF709BAFA2B" box="[668,760,1484,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E122DF70867FA2B" box="[771,805,1484,1508]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E212DF70834FA2B" box="[816,886,1484,1508]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, C</figureCitation>
|
||
). No sensory spots present. Type-1 glandular cell outlets present paradorsal and ventromedial position (
|
||
<figureCitation id="135F2A732455FFC03E022DD408DFF9C7" box="[787,925,1519,1544]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03EBF2DCB0891F9C7" box="[942,979,1520,1544]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4C</figureCitation>
|
||
). Type-2 glandular cell outlets present laterodorsally (
|
||
<figureCitation id="135F2A732455FFC03C512E2F0ADEF9E3" box="[320,412,1556,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CB62E2F0AA6F9E3" box="[423,484,1556,1580]" captionStart="FIGURE 3" captionStartId="5.[151,250,1853,1875]" captionTargetBox="[233,1358,193,1842]" captionTargetId="figure@5.[233,1358,193,1842]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes multiporus sp. nov., males (A, ZMB 11591g; B, C, ZMB 11591i) and a female (E–F, ZMB 11591j), scanning electron micrographs.A, entire animal, lateral view (right side); B, neck and segment 1, frontal view; C, segments 1– 4, ventral view; D, close-up of laterodorsal sensory spots on segments 1 and 2; E, segments 2–4, lateral view (left side); F, close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="3" pageNumber="544">3E, F</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CFE2E2F0974F9E3" box="[495,566,1556,1580]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62E030AA9F957" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 5 with lateroventral tubes (
|
||
<figureCitation id="135F2A732455FFC03F4F2E0C09FEF99F" box="[606,700,1591,1616]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03FD92E0309AFF99F" box="[712,749,1592,1616]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03FE92E0C085FF99F" box="[760,797,1591,1616]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="3" pageNumber="544">5A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E382E0C080FF99F" box="[809,845,1591,1616]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6B</figureCitation>
|
||
). Sensory spots absent. Type-1 glandular cell outlets present in middorsal and ventromedial position (
|
||
<figureCitation id="135F2A732455FFC03FA92E670857F9BB" box="[696,789,1628,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E302E670807F9BA" box="[801,837,1628,1653]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6B</figureCitation>
|
||
). Type-2 glandular cell outlets present in laterodorsal position (
|
||
<figureCitation id="135F2A732455FFC03C112E440A1EF957" box="[256,348,1663,1688]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C762EBB0AEEF957" box="[359,428,1664,1688]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CA92E440A9CF957" box="[440,478,1663,1688]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62E9F0BF8F8E7" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 6 with middorsal and lateroventral acicular spines (
|
||
<figureCitation id="135F2A732455FFC03E7C2E9F08A9F973" box="[877,1003,1700,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03EE72E9F0F59F973" box="[1014,1051,1700,1724]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC039372E9F0F09F972" box="[1062,1099,1700,1725]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="3" pageNumber="544">5A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC039472E9F0FDEF972" box="[1110,1180,1700,1725]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6A–C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC039B62E9F0FAFF973" box="[1191,1261,1700,1724]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="3" pageNumber="544">7A, B</figureCitation>
|
||
). Sensory spots present in midlateral position (
|
||
<figureCitation id="135F2A732455FFC03F062EFC09E8F92F" box="[535,682,1735,1760]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03FD12EFC09A7F92F" box="[704,741,1735,1760]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6A</figureCitation>
|
||
). Type-1 glandular cell outlets present paradorsally and ventromedially (
|
||
<figureCitation id="135F2A732455FFC03C5E2ED70A8EF8CB" box="[335,460,1772,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CC62ED70AB9F8CA" box="[471,507,1772,1797]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6B</figureCitation>
|
||
). Type-2 glandular cell outlets present in laterodorsal position (
|
||
<figureCitation id="135F2A732455FFC039AD2ED70E5AF8CB" box="[1212,1304,1772,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038322ED70E2AF8CB" box="[1315,1384,1772,1796]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A, C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC038622ED70BEFF8E7" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62455FFC03DD62F0F09E4F877" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="544">
|
||
Segment 7 with lateroventral acicular spines (
|
||
<figureCitation id="135F2A732455FFC03FCF2F0F087CF883" box="[734,830,1844,1869]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E5F2F0F0833F883" box="[846,881,1844,1868]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E902F0F0889F883" box="[897,971,1844,1869]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6B, D</figureCitation>
|
||
). Sensory spots present in ventromedial position (
|
||
<figureCitation id="135F2A732455FFC03C102F6C0A1CF8BF" box="[257,350,1879,1904]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C7B2F6C0ACFF8BF" box="[362,397,1879,1904]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03C8B2F630AFCF8BF" box="[410,446,1880,1904]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="3" pageNumber="544">7B</figureCitation>
|
||
). Midlateral sensory spots confirmed for six out of twelve specimens. Type-1 glandular cell outlets present middorsally and ventromedially (
|
||
<figureCitation id="135F2A732455FFC03FE12F470830F85B" box="[752,882,1916,1941]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03E6E2F47088AF85B" box="[895,968,1916,1941]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6C, D</figureCitation>
|
||
). Type-2 glandular cell outlets present in laterodorsal position (
|
||
<figureCitation id="135F2A732455FFC03C9A2F9B0AA5F877" box="[395,487,1952,1977]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="3" pageNumber="544">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03CE22F9B0955F877" box="[499,535,1952,1976]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="3" pageNumber="544">4A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03F322F9B092AF877" box="[547,616,1952,1977]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="3" pageNumber="544">6A, C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732455FFC03F652F9B09D8F877" box="[628,666,1952,1976]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="3" pageNumber="544">7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1B667E2452FFC73D862F0E0A12F82B" httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="4" pageNumber="545" startId="4.[151,250,1845,1867]" targetBox="[232,1343,197,1832]" targetPageId="4">
|
||
<paragraph id="8BDB36F62452FFC73D862F0E0A12F82B" blockId="4.[151,1436,1844,2020]" pageId="4" pageNumber="545">
|
||
<emphasis id="B910EAE42452FFC73D862F0E0A57F885" bold="true" box="[151,277,1845,1867]" pageId="4" pageNumber="545">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C644D752452FFC73C0F2F0F0951F885" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[286,531,1844,1866]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="4" pageNumber="545" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE42452FFC73C0F2F0F0951F885" box="[286,531,1844,1866]" italics="true" pageId="4" pageNumber="545">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F2452FFC73F0D2F0E092DF885" box="[540,623,1845,1866]" pageId="4" pageNumber="545" rank="species">
|
||
<emphasis id="B910EAE42452FFC73F0D2F0E092DF885" bold="true" box="[540,623,1845,1866]" pageId="4" pageNumber="545">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, camera lucida drawings.
|
||
<emphasis id="B910EAE42452FFC73E942F0E08D5F885" bold="true" box="[901,919,1845,1866]" pageId="4" pageNumber="545">A</emphasis>
|
||
,
|
||
<emphasis id="B910EAE42452FFC73EB62F0E08FAF885" bold="true" box="[935,952,1845,1866]" pageId="4" pageNumber="545">B</emphasis>
|
||
, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively;
|
||
<emphasis id="B910EAE42452FFC73FF62F6809BBF8A6" bold="true" box="[743,761,1875,1897]" pageId="4" pageNumber="545">C</emphasis>
|
||
,
|
||
<emphasis id="B910EAE42452FFC73E192F680858F8A7" bold="true" box="[776,794,1875,1896]" pageId="4" pageNumber="545">D</emphasis>
|
||
, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1B667E2453FFC63D862F060E17F823" httpUri="https://zenodo.org/record/1187745/files/figure.png" pageId="5" pageNumber="546" startId="5.[151,250,1853,1875]" targetBox="[233,1358,193,1842]" targetPageId="5">
|
||
<paragraph id="8BDB36F62453FFC63D862F060E17F823" blockId="5.[151,1436,1852,2028]" pageId="5" pageNumber="546">
|
||
<emphasis id="B910EAE42453FFC63D862F060A51F89D" bold="true" box="[151,275,1853,1875]" pageId="5" pageNumber="546">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C644D752453FFC63C0B2F07094EF89D" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[282,524,1852,1874]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="5" pageNumber="546" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE42453FFC63C0B2F07094EF89D" box="[282,524,1852,1874]" italics="true" pageId="5" pageNumber="546">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F2453FFC63F022F060921F89D" box="[531,611,1853,1874]" pageId="5" pageNumber="546" rank="species">
|
||
<emphasis id="B910EAE42453FFC63F022F060921F89D" bold="true" box="[531,611,1853,1874]" pageId="5" pageNumber="546">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, males (
|
||
<emphasis id="B910EAE42453FFC63FAA2F06098FF89D" bold="true" box="[699,717,1853,1874]" pageId="5" pageNumber="546">A</emphasis>
|
||
, ZMB 11591g;
|
||
<emphasis id="B910EAE42453FFC63E602F0608C0F89D" bold="true" box="[881,898,1853,1874]" pageId="5" pageNumber="546">B</emphasis>
|
||
,
|
||
<emphasis id="B910EAE42453FFC63E9E2F0608E3F89C" bold="true" box="[911,929,1853,1875]" pageId="5" pageNumber="546">C</emphasis>
|
||
, ZMB 11591i) and a female (
|
||
<emphasis id="B910EAE42453FFC639C32F060FBFF89D" bold="true" box="[1234,1277,1853,1874]" pageId="5" pageNumber="546">E–F</emphasis>
|
||
, ZMB 11591j), scanning electron micrographs.
|
||
<emphasis id="B910EAE42453FFC63CCB2F600AAEF8BF" bold="true" box="[474,492,1883,1904]" pageId="5" pageNumber="546">A</emphasis>
|
||
, entire animal, lateral view (right side);
|
||
<emphasis id="B910EAE42453FFC63E952F6708D7F8BE" bold="true" box="[900,917,1884,1905]" pageId="5" pageNumber="546">B</emphasis>
|
||
, neck and segment 1, frontal view;
|
||
<emphasis id="B910EAE42453FFC639EE2F600E53F8BE" bold="true" box="[1279,1297,1883,1905]" pageId="5" pageNumber="546">C</emphasis>
|
||
, segments 1– 4, ventral view;
|
||
<emphasis id="B910EAE42453FFC63C532F410A16F840" bold="true" box="[322,340,1914,1935]" pageId="5" pageNumber="546">D</emphasis>
|
||
, close-up of laterodorsal sensory spots on segments 1 and 2;
|
||
<emphasis id="B910EAE42453FFC63EC92F4108ADF840" bold="true" box="[984,1007,1914,1935]" pageId="5" pageNumber="546">E,</emphasis>
|
||
segments 2–4, lateral view (left side);
|
||
<emphasis id="B910EAE42453FFC638992F410EDBF840" bold="true" box="[1416,1433,1914,1935]" pageId="5" pageNumber="546">F,</emphasis>
|
||
close-up of a laterodorsal type-2 glandular cell outlet on segment 4. Abbreviations: mds, middorsal acicular spine; mvp, midventral placid; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows indicate sensory spots, white arrowhead type-1 glandular cell outlet, and black arrowheads type-2 glandular cell outlets.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1B667E2450FFC53D862F0C0F60F829" httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="6" pageNumber="547" startId="6.[151,250,1847,1869]" targetBox="[236,1354,187,1837]" targetPageId="6">
|
||
<paragraph id="8BDB36F62450FFC53D862F0C0F60F829" blockId="6.[151,1437,1847,2022]" pageId="6" pageNumber="547">
|
||
<emphasis id="B910EAE42450FFC53D862F0C0A54F883" bold="true" box="[151,278,1847,1869]" pageId="6" pageNumber="547">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C644D752450FFC53C312F0C0957F882" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[288,533,1847,1869]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="6" pageNumber="547" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE42450FFC53C312F0C0957F882" box="[288,533,1847,1869]" italics="true" pageId="6" pageNumber="547">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F2450FFC53F0E2F0C0931F883" box="[543,627,1847,1868]" pageId="6" pageNumber="547" rank="species">
|
||
<emphasis id="B910EAE42450FFC53F0E2F0C0931F883" bold="true" box="[543,627,1847,1868]" pageId="6" pageNumber="547">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Nomarski photomicrographs of the holotypic male (ZMB 11590).
|
||
<emphasis id="B910EAE42450FFC538512F0C0E10F883" bold="true" box="[1344,1362,1847,1868]" pageId="6" pageNumber="547">A</emphasis>
|
||
, entire animal, dorsal view;
|
||
<emphasis id="B910EAE42450FFC53C602F6D0AC0F8A4" bold="true" box="[369,386,1878,1899]" pageId="6" pageNumber="547">B</emphasis>
|
||
, entire animal, ventral view;
|
||
<emphasis id="B910EAE42450FFC53FA52F6E0984F8A4" bold="true" box="[692,710,1877,1899]" pageId="6" pageNumber="547">C</emphasis>
|
||
, neck and segments 1–6, dorsal view;
|
||
<emphasis id="B910EAE42450FFC5394D2F6D0F2CF8A4" bold="true" box="[1116,1134,1878,1899]" pageId="6" pageNumber="547">D</emphasis>
|
||
, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1B667E2451FFC43D8628A30922FEE0" pageId="7" pageNumber="548">
|
||
<paragraph id="8BDB36F62451FFC43D8628A30922FEE0" blockId="7.[151,1436,151,303]" pageId="7" pageNumber="548">
|
||
<emphasis id="B910EAE42451FFC43D8628A30A48FF60" bold="true" box="[151,266,152,175]" pageId="7" pageNumber="548">TABLE 2.</emphasis>
|
||
Measurements for adult
|
||
<taxonomicName id="4C644D752451FFC43F0D28AC0859FF60" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[540,795,151,175]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="7" pageNumber="548" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE42451FFC43F0D28AC0859FF60" box="[540,795,151,175]" italics="true" pageId="7" pageNumber="548">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F2451FFC43E3528A30839FF60" box="[804,891,152,175]" pageId="7" pageNumber="548" rank="species">
|
||
<emphasis id="B910EAE42451FFC43E3528A30839FF60" bold="true" box="[804,891,152,175]" pageId="7" pageNumber="548">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(in micrometers or as percentage). Abbreviations: (ac), acicular spine; LTAS, length of lateral terminal accessory spine; LTS, length of lateral terminal spine; LV, length of lateroventral spine/tube; MD, length of middorsal spine; ML, length of midlateral tube; MSW, maximum sternal width; N, number of specimens measured; S, segment length; SD, standard deviation; SW, standard width; TL, trunk length; (tu), tube; VL, length of ventrolateral tube.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB36F62451FFC43D8E29710FE6F9F7" pageId="7" pageNumber="548">
|
||
<table id="F964C4562451003C3D8E29710FE6F9F7" box="[159,1188,330,1592]" colsContinueIn="14.[159,1211,946,1919]" gridcols="5" gridrows="31" pageId="7" pageNumber="548">
|
||
<tr id="355434B42451003C3D8E29710FE6FEAF" box="[159,1188,330,352]" gridrow="0" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E29710A56FEAF" box="[159,276,330,352]" gridcol="0" gridrow="0" pageId="7" pageNumber="548">Character</th>
|
||
<th id="76855DC82451003C3C8A29710AEFFEAF" box="[411,429,330,352]" gridcol="1" gridrow="0" pageId="7" pageNumber="548">N</th>
|
||
<th id="76855DC82451003C3F33297109DEFEAF" box="[546,668,330,352]" gridcol="2" gridrow="0" pageId="7" pageNumber="548">Range</th>
|
||
<th id="76855DC82451003C3E53297108C6FEAF" box="[834,900,330,352]" gridcol="3" gridrow="0" pageId="7" pageNumber="548">Mean</th>
|
||
<th id="76855DC82451003C397229710FE6FEAF" box="[1123,1188,330,352]" gridcol="4" gridrow="0" pageId="7" pageNumber="548">SD</th>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E294F0FE6FE45" box="[159,1188,372,394]" gridrow="1" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E294F0A56FE45" box="[159,276,372,394]" gridcol="0" gridrow="1" pageId="7" pageNumber="548">TL</th>
|
||
<td id="76855DC82451003C3C8A294F0AEFFE45" box="[411,429,372,394]" gridcol="1" gridrow="1" pageId="7" pageNumber="548">9</td>
|
||
<td id="76855DC82451003C3F33294F09DEFE45" box="[546,668,372,394]" gridcol="2" gridrow="1" pageId="7" pageNumber="548">198–229</td>
|
||
<td id="76855DC82451003C3E53294F08C6FE45" box="[834,900,372,394]" gridcol="3" gridrow="1" pageId="7" pageNumber="548">210.0</td>
|
||
<td id="76855DC82451003C3972294F0FE6FE45" box="[1123,1188,372,394]" gridcol="4" gridrow="1" pageId="7" pageNumber="548">11.02</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E29A60FE6FE7C" box="[159,1188,413,435]" gridrow="2" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E29A60A56FE7C" box="[159,276,413,435]" gridcol="0" gridrow="2" pageId="7" pageNumber="548">MSW-8</th>
|
||
<td id="76855DC82451003C3C8A29A60AEFFE7C" box="[411,429,413,435]" gridcol="1" gridrow="2" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F3329A609DEFE7C" box="[546,668,413,435]" gridcol="2" gridrow="2" pageId="7" pageNumber="548">43–51</td>
|
||
<td id="76855DC82451003C3E5329A608C6FE7C" box="[834,900,413,435]" gridcol="3" gridrow="2" pageId="7" pageNumber="548">46.8</td>
|
||
<td id="76855DC82451003C397229A60FE6FE7C" box="[1123,1188,413,435]" gridcol="4" gridrow="2" pageId="7" pageNumber="548">3.29</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E29FD0FE6FE13" box="[159,1188,454,476]" gridrow="3" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E29FD0A56FE13" box="[159,276,454,476]" gridcol="0" gridrow="3" pageId="7" pageNumber="548">MSW-8/TL</th>
|
||
<td id="76855DC82451003C3C8A29FD0AEFFE13" box="[411,429,454,476]" gridcol="1" gridrow="3" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F3329FD09DEFE13" box="[546,668,454,476]" gridcol="2" gridrow="3" pageId="7" pageNumber="548">20.5–24.0%</td>
|
||
<td id="76855DC82451003C3E5329FD08C6FE13" box="[834,900,454,476]" gridcol="3" gridrow="3" pageId="7" pageNumber="548">22.2%</td>
|
||
<td id="76855DC82451003C397229FD0FE6FE13" box="[1123,1188,454,476]" gridcol="4" gridrow="3" pageId="7" pageNumber="548">1.47%</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E29CB0FE6FDC9" box="[159,1188,496,518]" gridrow="4" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E29CB0A56FDC9" box="[159,276,496,518]" gridcol="0" gridrow="4" pageId="7" pageNumber="548">SW-10</th>
|
||
<td id="76855DC82451003C3C8A29CB0AEFFDC9" box="[411,429,496,518]" gridcol="1" gridrow="4" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F3329CB09DEFDC9" box="[546,668,496,518]" gridcol="2" gridrow="4" pageId="7" pageNumber="548">35–39</td>
|
||
<td id="76855DC82451003C3E5329CB08C6FDC9" box="[834,900,496,518]" gridcol="3" gridrow="4" pageId="7" pageNumber="548">37.5</td>
|
||
<td id="76855DC82451003C397229CB0FE6FDC9" box="[1123,1188,496,518]" gridcol="4" gridrow="4" pageId="7" pageNumber="548">1.60</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2A220FE6FDE0" box="[159,1188,537,559]" gridrow="5" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2A220A56FDE0" box="[159,276,537,559]" gridcol="0" gridrow="5" pageId="7" pageNumber="548">SW-10/TL</th>
|
||
<td id="76855DC82451003C3C8A2A220AEFFDE0" box="[411,429,537,559]" gridcol="1" gridrow="5" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332A2209DEFDE0" box="[546,668,537,559]" gridcol="2" gridrow="5" pageId="7" pageNumber="548">16.1–19.4%</td>
|
||
<td id="76855DC82451003C3E532A2208C6FDE0" box="[834,900,537,559]" gridcol="3" gridrow="5" pageId="7" pageNumber="548">17.8%</td>
|
||
<td id="76855DC82451003C39722A220FE6FDE0" box="[1123,1188,537,559]" gridcol="4" gridrow="5" pageId="7" pageNumber="548">1.26%</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2A790FE6FD97" box="[159,1188,578,600]" gridrow="6" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2A790A56FD97" box="[159,276,578,600]" gridcol="0" gridrow="6" pageId="7" pageNumber="548">S1</th>
|
||
<td id="76855DC82451003C3C8A2A790AEFFD97" box="[411,429,578,600]" gridcol="1" gridrow="6" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332A7909DEFD97" box="[546,668,578,600]" gridcol="2" gridrow="6" pageId="7" pageNumber="548">30–34</td>
|
||
<td id="76855DC82451003C3E532A7908C6FD97" box="[834,900,578,600]" gridcol="3" gridrow="6" pageId="7" pageNumber="548">31.8</td>
|
||
<td id="76855DC82451003C39722A790FE6FD97" box="[1123,1188,578,600]" gridcol="4" gridrow="6" pageId="7" pageNumber="548">1.60</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2A570FE6FD4D" box="[159,1188,620,642]" gridrow="7" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2A570A56FD4D" box="[159,276,620,642]" gridcol="0" gridrow="7" pageId="7" pageNumber="548">S2</th>
|
||
<td id="76855DC82451003C3C8A2A570AEFFD4D" box="[411,429,620,642]" gridcol="1" gridrow="7" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332A5709DEFD4D" box="[546,668,620,642]" gridcol="2" gridrow="7" pageId="7" pageNumber="548">23–26</td>
|
||
<td id="76855DC82451003C3E532A5708C6FD4D" box="[834,900,620,642]" gridcol="3" gridrow="7" pageId="7" pageNumber="548">24.4</td>
|
||
<td id="76855DC82451003C39722A570FE6FD4D" box="[1123,1188,620,642]" gridcol="4" gridrow="7" pageId="7" pageNumber="548">1.28</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2AAE0FE6FD64" box="[159,1188,661,683]" gridrow="8" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2AAE0A56FD64" box="[159,276,661,683]" gridcol="0" gridrow="8" pageId="7" pageNumber="548">S3</th>
|
||
<td id="76855DC82451003C3C8A2AAE0AEFFD64" box="[411,429,661,683]" gridcol="1" gridrow="8" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332AAE09DEFD64" box="[546,668,661,683]" gridcol="2" gridrow="8" pageId="7" pageNumber="548">20–21</td>
|
||
<td id="76855DC82451003C3E532AAE08C6FD64" box="[834,900,661,683]" gridcol="3" gridrow="8" pageId="7" pageNumber="548">20.6</td>
|
||
<td id="76855DC82451003C39722AAE0FE6FD64" box="[1123,1188,661,683]" gridcol="4" gridrow="8" pageId="7" pageNumber="548">0.60</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2A850FE6FD1B" box="[159,1188,702,724]" gridrow="9" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2A850A56FD1B" box="[159,276,702,724]" gridcol="0" gridrow="9" pageId="7" pageNumber="548">S4</th>
|
||
<td id="76855DC82451003C3C8A2A850AEFFD1B" box="[411,429,702,724]" gridcol="1" gridrow="9" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332A8509DEFD1B" box="[546,668,702,724]" gridcol="2" gridrow="9" pageId="7" pageNumber="548">20–23</td>
|
||
<td id="76855DC82451003C3E532A8508C6FD1B" box="[834,900,702,724]" gridcol="3" gridrow="9" pageId="7" pageNumber="548">21.8</td>
|
||
<td id="76855DC82451003C39722A850FE6FD1B" box="[1123,1188,702,724]" gridcol="4" gridrow="9" pageId="7" pageNumber="548">0.94</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2AD30FE6FD31" box="[159,1188,744,766]" gridrow="10" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2AD30A56FD31" box="[159,276,744,766]" gridcol="0" gridrow="10" pageId="7" pageNumber="548">S5</th>
|
||
<td id="76855DC82451003C3C8A2AD30AEFFD31" box="[411,429,744,766]" gridcol="1" gridrow="10" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332AD309DEFD31" box="[546,668,744,766]" gridcol="2" gridrow="10" pageId="7" pageNumber="548">21–22</td>
|
||
<td id="76855DC82451003C3E532AD308C6FD31" box="[834,900,744,766]" gridcol="3" gridrow="10" pageId="7" pageNumber="548">21.9</td>
|
||
<td id="76855DC82451003C39722AD30FE6FD31" box="[1123,1188,744,766]" gridcol="4" gridrow="10" pageId="7" pageNumber="548">0.70</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2B2A0FE6FCE8" box="[159,1188,785,807]" gridrow="11" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2B2A0A56FCE8" box="[159,276,785,807]" gridcol="0" gridrow="11" pageId="7" pageNumber="548">S6</th>
|
||
<td id="76855DC82451003C3C8A2B2A0AEFFCE8" box="[411,429,785,807]" gridcol="1" gridrow="11" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332B2A09DEFCE8" box="[546,668,785,807]" gridcol="2" gridrow="11" pageId="7" pageNumber="548">21–24</td>
|
||
<td id="76855DC82451003C3E532B2A08C6FCE8" box="[834,900,785,807]" gridcol="3" gridrow="11" pageId="7" pageNumber="548">23.0</td>
|
||
<td id="76855DC82451003C39722B2A0FE6FCE8" box="[1123,1188,785,807]" gridcol="4" gridrow="11" pageId="7" pageNumber="548">0.93</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2B010FE6FC9F" box="[159,1188,826,848]" gridrow="12" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2B010A56FC9F" box="[159,276,826,848]" gridcol="0" gridrow="12" pageId="7" pageNumber="548">S7</th>
|
||
<td id="76855DC82451003C3C8A2B010AEFFC9F" box="[411,429,826,848]" gridcol="1" gridrow="12" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332B0109DEFC9F" box="[546,668,826,848]" gridcol="2" gridrow="12" pageId="7" pageNumber="548">26–27</td>
|
||
<td id="76855DC82451003C3E532B0108C6FC9F" box="[834,900,826,848]" gridcol="3" gridrow="12" pageId="7" pageNumber="548">26.3</td>
|
||
<td id="76855DC82451003C39722B010FE6FC9F" box="[1123,1188,826,848]" gridcol="4" gridrow="12" pageId="7" pageNumber="548">0.54</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2B5F0FE6FCB5" box="[159,1188,868,890]" gridrow="13" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2B5F0A56FCB5" box="[159,276,868,890]" gridcol="0" gridrow="13" pageId="7" pageNumber="548">S8</th>
|
||
<td id="76855DC82451003C3C8A2B5F0AEFFCB5" box="[411,429,868,890]" gridcol="1" gridrow="13" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332B5F09DEFCB5" box="[546,668,868,890]" gridcol="2" gridrow="13" pageId="7" pageNumber="548">28–30</td>
|
||
<td id="76855DC82451003C3E532B5F08C6FCB5" box="[834,900,868,890]" gridcol="3" gridrow="13" pageId="7" pageNumber="548">29.3</td>
|
||
<td id="76855DC82451003C39722B5F0FE6FCB5" box="[1123,1188,868,890]" gridcol="4" gridrow="13" pageId="7" pageNumber="548">0.87</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2BB60FE6FC6C" box="[159,1188,909,931]" gridrow="14" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2BB60A56FC6C" box="[159,276,909,931]" gridcol="0" gridrow="14" pageId="7" pageNumber="548">S9</th>
|
||
<td id="76855DC82451003C3C8A2BB60AEFFC6C" box="[411,429,909,931]" gridcol="1" gridrow="14" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332BB609DEFC6C" box="[546,668,909,931]" gridcol="2" gridrow="14" pageId="7" pageNumber="548">29–33</td>
|
||
<td id="76855DC82451003C3E532BB608C6FC6C" box="[834,900,909,931]" gridcol="3" gridrow="14" pageId="7" pageNumber="548">31.1</td>
|
||
<td id="76855DC82451003C39722BB60FE6FC6C" box="[1123,1188,909,931]" gridcol="4" gridrow="14" pageId="7" pageNumber="548">1.75</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2B8D0FE6FC03" box="[159,1188,950,972]" gridrow="15" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2B8D0A56FC03" box="[159,276,950,972]" gridcol="0" gridrow="15" pageId="7" pageNumber="548">S10</th>
|
||
<td id="76855DC82451003C3C8A2B8D0AEFFC03" box="[411,429,950,972]" gridcol="1" gridrow="15" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332B8D09DEFC03" box="[546,668,950,972]" gridcol="2" gridrow="15" pageId="7" pageNumber="548">36–41</td>
|
||
<td id="76855DC82451003C3E532B8D08C6FC03" box="[834,900,950,972]" gridcol="3" gridrow="15" pageId="7" pageNumber="548">38.2</td>
|
||
<td id="76855DC82451003C39722B8D0FE6FC03" box="[1123,1188,950,972]" gridcol="4" gridrow="15" pageId="7" pageNumber="548">2.11</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2BDB0FE6FC39" box="[159,1188,992,1014]" gridrow="16" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2BDB0A56FC39" box="[159,276,992,1014]" gridcol="0" gridrow="16" pageId="7" pageNumber="548">S11</th>
|
||
<td id="76855DC82451003C3C8A2BDB0AEFFC39" box="[411,429,992,1014]" gridcol="1" gridrow="16" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332BDB09DEFC39" box="[546,668,992,1014]" gridcol="2" gridrow="16" pageId="7" pageNumber="548">26–33</td>
|
||
<td id="76855DC82451003C3E532BDB08C6FC39" box="[834,900,992,1014]" gridcol="3" gridrow="16" pageId="7" pageNumber="548">30.8</td>
|
||
<td id="76855DC82451003C39722BDB0FE6FC39" box="[1123,1188,992,1014]" gridcol="4" gridrow="16" pageId="7" pageNumber="548">2.74</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2C320FE6FBD0" box="[159,1188,1033,1055]" gridrow="17" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2C320A56FBD0" box="[159,276,1033,1055]" gridcol="0" gridrow="17" pageId="7" pageNumber="548">MD4 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2C320AEFFBD0" box="[411,429,1033,1055]" gridcol="1" gridrow="17" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332C3209DEFBD0" box="[546,668,1033,1055]" gridcol="2" gridrow="17" pageId="7" pageNumber="548">32–40</td>
|
||
<td id="76855DC82451003C3E532C3208C6FBD0" box="[834,900,1033,1055]" gridcol="3" gridrow="17" pageId="7" pageNumber="548">36.0</td>
|
||
<td id="76855DC82451003C39722C320FE6FBD0" box="[1123,1188,1033,1055]" gridcol="4" gridrow="17" pageId="7" pageNumber="548">3.21</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2C090FE6FB87" box="[159,1188,1074,1096]" gridrow="18" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2C090A56FB87" box="[159,276,1074,1096]" gridcol="0" gridrow="18" pageId="7" pageNumber="548">MD6 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2C090AEFFB87" box="[411,429,1074,1096]" gridcol="1" gridrow="18" pageId="7" pageNumber="548">6</td>
|
||
<td id="76855DC82451003C3F332C0909DEFB87" box="[546,668,1074,1096]" gridcol="2" gridrow="18" pageId="7" pageNumber="548">63–68</td>
|
||
<td id="76855DC82451003C3E532C0908C6FB87" box="[834,900,1074,1096]" gridcol="3" gridrow="18" pageId="7" pageNumber="548">65.6</td>
|
||
<td id="76855DC82451003C39722C090FE6FB87" box="[1123,1188,1074,1096]" gridcol="4" gridrow="18" pageId="7" pageNumber="548">2.03</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2C670FE6FBBD" box="[159,1188,1116,1138]" gridrow="19" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2C670A56FBBD" box="[159,276,1116,1138]" gridcol="0" gridrow="19" pageId="7" pageNumber="548">MD8 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2C670AEFFBBD" box="[411,429,1116,1138]" gridcol="1" gridrow="19" pageId="7" pageNumber="548">6</td>
|
||
<td id="76855DC82451003C3F332C6709DEFBBD" box="[546,668,1116,1138]" gridcol="2" gridrow="19" pageId="7" pageNumber="548">79–86</td>
|
||
<td id="76855DC82451003C3E532C6708C6FBBD" box="[834,900,1116,1138]" gridcol="3" gridrow="19" pageId="7" pageNumber="548">82.1</td>
|
||
<td id="76855DC82451003C39722C670FE6FBBD" box="[1123,1188,1116,1138]" gridcol="4" gridrow="19" pageId="7" pageNumber="548">2.86</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2CBE0FE6FB54" box="[159,1188,1157,1179]" gridrow="20" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2CBE0A56FB54" box="[159,276,1157,1179]" gridcol="0" gridrow="20" pageId="7" pageNumber="548">VL2 (tu)</th>
|
||
<td id="76855DC82451003C3C8A2CBE0AEFFB54" box="[411,429,1157,1179]" gridcol="1" gridrow="20" pageId="7" pageNumber="548">8</td>
|
||
<td id="76855DC82451003C3F332CBE09DEFB54" box="[546,668,1157,1179]" gridcol="2" gridrow="20" pageId="7" pageNumber="548">11–17</td>
|
||
<td id="76855DC82451003C3E532CBE08C6FB54" box="[834,900,1157,1179]" gridcol="3" gridrow="20" pageId="7" pageNumber="548">14.6</td>
|
||
<td id="76855DC82451003C39722CBE0FE6FB54" box="[1123,1188,1157,1179]" gridcol="4" gridrow="20" pageId="7" pageNumber="548">2.14</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2C950FE6FB0B" box="[159,1188,1198,1220]" gridrow="21" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2C950A56FB0B" box="[159,276,1198,1220]" gridcol="0" gridrow="21" pageId="7" pageNumber="548">LV5 (tu)</th>
|
||
<td id="76855DC82451003C3C8A2C950AEFFB0B" box="[411,429,1198,1220]" gridcol="1" gridrow="21" pageId="7" pageNumber="548">6</td>
|
||
<td id="76855DC82451003C3F332C9509DEFB0B" box="[546,668,1198,1220]" gridcol="2" gridrow="21" pageId="7" pageNumber="548">14–19</td>
|
||
<td id="76855DC82451003C3E532C9508C6FB0B" box="[834,900,1198,1220]" gridcol="3" gridrow="21" pageId="7" pageNumber="548">16.0</td>
|
||
<td id="76855DC82451003C39722C950FE6FB0B" box="[1123,1188,1198,1220]" gridcol="4" gridrow="21" pageId="7" pageNumber="548">2.54</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2CE30FE6FB21" box="[159,1188,1240,1262]" gridrow="22" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2CE30A56FB21" box="[159,276,1240,1262]" gridcol="0" gridrow="22" pageId="7" pageNumber="548">LV6 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2CE30AEFFB21" box="[411,429,1240,1262]" gridcol="1" gridrow="22" pageId="7" pageNumber="548">9</td>
|
||
<td id="76855DC82451003C3F332CE309DEFB21" box="[546,668,1240,1262]" gridcol="2" gridrow="22" pageId="7" pageNumber="548">31–36</td>
|
||
<td id="76855DC82451003C3E532CE308C6FB21" box="[834,900,1240,1262]" gridcol="3" gridrow="22" pageId="7" pageNumber="548">33.0</td>
|
||
<td id="76855DC82451003C39722CE30FE6FB21" box="[1123,1188,1240,1262]" gridcol="4" gridrow="22" pageId="7" pageNumber="548">1.55</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2D3A0FE6FAD8" box="[159,1188,1281,1303]" gridrow="23" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2D3A0A56FAD8" box="[159,276,1281,1303]" gridcol="0" gridrow="23" pageId="7" pageNumber="548">LV7 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2D3A0AEFFAD8" box="[411,429,1281,1303]" gridcol="1" gridrow="23" pageId="7" pageNumber="548">9</td>
|
||
<td id="76855DC82451003C3F332D3A09DEFAD8" box="[546,668,1281,1303]" gridcol="2" gridrow="23" pageId="7" pageNumber="548">39–44</td>
|
||
<td id="76855DC82451003C3E532D3A08C6FAD8" box="[834,900,1281,1303]" gridcol="3" gridrow="23" pageId="7" pageNumber="548">41.8</td>
|
||
<td id="76855DC82451003C39722D3A0FE6FAD8" box="[1123,1188,1281,1303]" gridcol="4" gridrow="23" pageId="7" pageNumber="548">1.73</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2D110FE6FA8F" box="[159,1188,1322,1344]" gridrow="24" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2D110A56FA8F" box="[159,276,1322,1344]" gridcol="0" gridrow="24" pageId="7" pageNumber="548">LV8 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2D110AEFFA8F" box="[411,429,1322,1344]" gridcol="1" gridrow="24" pageId="7" pageNumber="548">9</td>
|
||
<td id="76855DC82451003C3F332D1109DEFA8F" box="[546,668,1322,1344]" gridcol="2" gridrow="24" pageId="7" pageNumber="548">49–57</td>
|
||
<td id="76855DC82451003C3E532D1108C6FA8F" box="[834,900,1322,1344]" gridcol="3" gridrow="24" pageId="7" pageNumber="548">52.3</td>
|
||
<td id="76855DC82451003C39722D110FE6FA8F" box="[1123,1188,1322,1344]" gridcol="4" gridrow="24" pageId="7" pageNumber="548">2.47</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2D6F0FE6FAA5" box="[159,1188,1364,1386]" gridrow="25" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2D6F0A56FAA5" box="[159,276,1364,1386]" gridcol="0" gridrow="25" pageId="7" pageNumber="548">LV9 (ac)</th>
|
||
<td id="76855DC82451003C3C8A2D6F0AEFFAA5" box="[411,429,1364,1386]" gridcol="1" gridrow="25" pageId="7" pageNumber="548">9</td>
|
||
<td id="76855DC82451003C3F332D6F09DEFAA5" box="[546,668,1364,1386]" gridcol="2" gridrow="25" pageId="7" pageNumber="548">51–60</td>
|
||
<td id="76855DC82451003C3E532D6F08C6FAA5" box="[834,900,1364,1386]" gridcol="3" gridrow="25" pageId="7" pageNumber="548">55.4</td>
|
||
<td id="76855DC82451003C39722D6F0FE6FAA5" box="[1123,1188,1364,1386]" gridcol="4" gridrow="25" pageId="7" pageNumber="548">2.95</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2D460FE6FA5C" box="[159,1188,1405,1427]" gridrow="26" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2D460A56FA5C" box="[159,276,1405,1427]" gridcol="0" gridrow="26" pageId="7" pageNumber="548">ML10 (tu)</th>
|
||
<td id="76855DC82451003C3C8A2D460AEFFA5C" box="[411,429,1405,1427]" gridcol="1" gridrow="26" pageId="7" pageNumber="548">9</td>
|
||
<td id="76855DC82451003C3F332D4609DEFA5C" box="[546,668,1405,1427]" gridcol="2" gridrow="26" pageId="7" pageNumber="548">14–18</td>
|
||
<td id="76855DC82451003C3E532D4608C6FA5C" box="[834,900,1405,1427]" gridcol="3" gridrow="26" pageId="7" pageNumber="548">16.3</td>
|
||
<td id="76855DC82451003C39722D460FE6FA5C" box="[1123,1188,1405,1427]" gridcol="4" gridrow="26" pageId="7" pageNumber="548">1.46</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2D9D0FE6FA73" box="[159,1188,1446,1468]" gridrow="27" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2D9D0A56FA73" box="[159,276,1446,1468]" gridcol="0" gridrow="27" pageId="7" pageNumber="548">LTS</th>
|
||
<td id="76855DC82451003C3C8A2D9D0AEFFA73" box="[411,429,1446,1468]" gridcol="1" gridrow="27" pageId="7" pageNumber="548">8</td>
|
||
<td id="76855DC82451003C3F332D9D09DEFA73" box="[546,668,1446,1468]" gridcol="2" gridrow="27" pageId="7" pageNumber="548">130–165</td>
|
||
<td id="76855DC82451003C3E532D9D08C6FA73" box="[834,900,1446,1468]" gridcol="3" gridrow="27" pageId="7" pageNumber="548">143.8</td>
|
||
<td id="76855DC82451003C39722D9D0FE6FA73" box="[1123,1188,1446,1468]" gridcol="4" gridrow="27" pageId="7" pageNumber="548">11.29</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2DEB0FE6FA29" box="[159,1188,1488,1510]" gridrow="28" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2DEB0A56FA29" box="[159,276,1488,1510]" gridcol="0" gridrow="28" pageId="7" pageNumber="548">LTAS</th>
|
||
<td id="76855DC82451003C3C8A2DEB0AEFFA29" box="[411,429,1488,1510]" gridcol="1" gridrow="28" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332DEB09DEFA29" box="[546,668,1488,1510]" gridcol="2" gridrow="28" pageId="7" pageNumber="548">41–49</td>
|
||
<td id="76855DC82451003C3E532DEB08C6FA29" box="[834,900,1488,1510]" gridcol="3" gridrow="28" pageId="7" pageNumber="548">45.8</td>
|
||
<td id="76855DC82451003C39722DEB0FE6FA29" box="[1123,1188,1488,1510]" gridcol="4" gridrow="28" pageId="7" pageNumber="548">2.95</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2DC20FE6F9C0" box="[159,1188,1529,1551]" gridrow="29" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2DC20A56F9C0" box="[159,276,1529,1551]" gridcol="0" gridrow="29" pageId="7" pageNumber="548">LTS/TL</th>
|
||
<td id="76855DC82451003C3C8A2DC20AEFF9C0" box="[411,429,1529,1551]" gridcol="1" gridrow="29" pageId="7" pageNumber="548">8</td>
|
||
<td id="76855DC82451003C3F332DC209DEF9C0" box="[546,668,1529,1551]" gridcol="2" gridrow="29" pageId="7" pageNumber="548">64.0–71.9%</td>
|
||
<td id="76855DC82451003C3E532DC208C6F9C0" box="[834,900,1529,1551]" gridcol="3" gridrow="29" pageId="7" pageNumber="548">68.5%</td>
|
||
<td id="76855DC82451003C39722DC20FE6F9C0" box="[1123,1188,1529,1551]" gridcol="4" gridrow="29" pageId="7" pageNumber="548">2.91%</td>
|
||
</tr>
|
||
<tr id="355434B42451003C3D8E2E190FE6F9F7" box="[159,1188,1570,1592]" gridrow="30" pageId="7" pageNumber="548">
|
||
<th id="76855DC82451003C3D8E2E190A56F9F7" box="[159,276,1570,1592]" gridcol="0" gridrow="30" pageId="7" pageNumber="548">LTAS/TL</th>
|
||
<td id="76855DC82451003C3C8A2E190AEFF9F7" box="[411,429,1570,1592]" gridcol="1" gridrow="30" pageId="7" pageNumber="548">5</td>
|
||
<td id="76855DC82451003C3F332E1909DEF9F7" box="[546,668,1570,1592]" gridcol="2" gridrow="30" pageId="7" pageNumber="548">20.8–23.0%</td>
|
||
<td id="76855DC82451003C3E532E1908C6F9F7" box="[834,900,1570,1592]" gridcol="3" gridrow="30" pageId="7" pageNumber="548">21.8%</td>
|
||
<td id="76855DC82451003C39722E190FE6F9F7" box="[1123,1188,1570,1592]" gridcol="4" gridrow="30" pageId="7" pageNumber="548">1.04%</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62451FFC43DD62E430861F917" blockId="7.[151,1436,1655,2005]" pageId="7" pageNumber="548">
|
||
Segment 8 similar to segment 6, except for the presence of ventromedial papillae in females (
|
||
<figureCitation id="135F2A732451FFC439F62E4C0E25F95F" box="[1255,1383,1655,1680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC438622E430BEEF97B" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="7" pageNumber="548">4A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43DAA2EA70B9FF97A" box="[187,221,1692,1717]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="7" pageNumber="548">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43DFC2EA70A1EF97B" box="[237,348,1692,1717]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="7" pageNumber="548">6A, C, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43C7A2EA70AF0F97B" box="[363,434,1692,1716]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="7" pageNumber="548">7A–C</figureCitation>
|
||
). Papillae visible only in SEM, but appearing as funnel-shaped subcuticular structures without large pore visible below papillae in LM (
|
||
<figureCitation id="135F2A732451FFC43FAF2E840854F917" box="[702,790,1727,1752]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="7" pageNumber="548">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62451FFC43DD62EDF0FC9F842" blockId="7.[151,1436,1655,2005]" pageId="7" pageNumber="548">
|
||
Segment 9 with lateroventral acicular spines (
|
||
<figureCitation id="135F2A732451FFC43FE12EDF083EF933" box="[752,892,1764,1789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43E9E2EDF08F3F932" box="[911,945,1764,1789]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="7" pageNumber="548">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43ED52EDF08ABF932" box="[964,1001,1764,1789]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="7" pageNumber="548">6D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43EEA2EDE0F05F933" box="[1019,1095,1764,1788]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="7" pageNumber="548">7D, E</figureCitation>
|
||
). Paradorsal, midlateral, and ventrolateral sensory spots present (
|
||
<figureCitation id="135F2A732451FFC43F392F3C09EAF8EF" box="[552,680,1799,1824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43FA22F3C0997F8EF" box="[691,725,1799,1824]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="7" pageNumber="548">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43FF02F3C0865F8D0" box="[737,807,1799,1824]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="7" pageNumber="548">6C, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43E232F33081AF8EF" box="[818,856,1800,1824]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="7" pageNumber="548">7D</figureCitation>
|
||
). Type-1 glandular cell outlets present in paradorsal and ventromedial position (
|
||
<figureCitation id="135F2A732451FFC43CDF2F170912F88B" box="[462,592,1836,1861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43F4D2F17093DF88A" box="[604,639,1836,1861]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="7" pageNumber="548">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43F9D2F170997F88B" box="[652,725,1836,1861]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="7" pageNumber="548">6C, D</figureCitation>
|
||
). Type-2 glandular cell outlets present laterodorsally (
|
||
<figureCitation id="135F2A732451FFC4382B2F170BECF8A8" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2B, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43DA82F6B0B9CF8A7" box="[185,222,1872,1896]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="7" pageNumber="548">4A</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43DFB2F740A4FF8A7" box="[234,269,1871,1896]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="7" pageNumber="548">6C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43C082F6B0A20F8A7" box="[281,354,1872,1896]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="7" pageNumber="548">7A, D</figureCitation>
|
||
). Small rounded sieve plates present in lateral accessory position (
|
||
<figureCitation id="135F2A732451FFC4395D2F740F8FF8A8" box="[1100,1229,1871,1896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2A, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC439C82F6B0FB9F8A7" box="[1241,1275,1872,1896]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="7" pageNumber="548">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC438192F740E6CF8A7" box="[1288,1326,1871,1896]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="7" pageNumber="548">6D</figureCitation>
|
||
). Primary pectinate fringe with quite shorter pectinate fringe teeth than fringe on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BDB36F62451FFCB3DD62FA309DBFF1A" blockId="7.[151,1436,1655,2005]" lastBlockId="8.[151,1437,151,752]" lastPageId="8" lastPageNumber="549" pageId="7" pageNumber="548">
|
||
Segment 10 with midlateral tubes (
|
||
<figureCitation id="135F2A732451FFC43F652FAC0843F860" box="[628,769,1943,1968]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2B, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43E052FA30874F87F" box="[788,822,1944,1968]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="7" pageNumber="548">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43E5B2FA308CCF87F" box="[842,910,1944,1968]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="7" pageNumber="548">7D–F</figureCitation>
|
||
). No cuticular hairs present. Subdorsal and ventrolateral sensory spots present (
|
||
<figureCitation id="135F2A732451FFC43F592F870992F81B" box="[584,720,1980,2005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="7" pageNumber="548">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43FF22F870844F81A" box="[739,774,1980,2005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="7" pageNumber="548">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A732451FFC43E0B2F8608CCF81B" box="[794,910,1980,2004]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="7" pageNumber="548">7D, F, G</figureCitation>
|
||
). Two type-1 glandular cell outlets aligned middorsally (
|
||
<figureCitation id="135F2A73245EFFCB3C3D28AC0AE5FF7F" box="[300,423,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Fig. 2B, D</figureCitation>
|
||
). Additional pair of type-1 glandular cell outlets present in ventromedial position. Primary pectinate fringe similar to preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BDB36F6245EFFCB3DD628DB0B95FE63" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="549">
|
||
Segment 11 with lateral terminal spines (
|
||
<figureCitation id="135F2A73245EFFCB3FB028E40866FF37" box="[673,804,223,248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3E2328DB0838FF38" box="[818,890,224,248]" captionStart="FIGURE 4" captionStartId="6.[151,250,1847,1869]" captionTargetBox="[236,1354,187,1837]" captionTargetId="figure@6.[236,1354,187,1837]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, entire animal, dorsal view; B, entire animal, ventral view; C, neck and segments 1–6, dorsal view; D, segments 1–6, ventral view. Abbreviations: lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mdp, middorsal placid; mds, middorsal acicular spine; mlt, midlateral tube; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187747/files/figure.png" pageId="8" pageNumber="549">4A, B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3E9928E408E9FF37" box="[904,939,223,248]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="8" pageNumber="549">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3EAB28DB0F41FF37" box="[954,1027,224,248]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="8" pageNumber="549">7D–G</figureCitation>
|
||
). Two pairs of thin and long penile spines present in males, and one pair of lateral terminal accessory spines present in females (
|
||
<figureCitation id="135F2A73245EFFCB39A7293F0E75FED3" box="[1206,1335,260,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3852293F0E24FED2" box="[1347,1382,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1829,1851]" captionTargetBox="[162,1426,843,1776]" captionTargetId="figure@8.[161,1426,843,1776]" captionTargetPageId="8" captionText="FIGURE 5. Echinoderes multiporus sp. nov., Nomarski photomicrographs of a paratypic female (ZMB 11591a). A, segments 2–7, ventral view; B, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets." httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="8" pageNumber="549">5B</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3862293F0ED5FED2" box="[1395,1431,260,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1884,1906]" captionTargetBox="[161,1426,209,1855]" captionTargetId="figure@9.[161,1426,209,1856]" captionTargetPageId="9" captionText="FIGURE 6. Echinoderes multiporus sp. nov., Nomarski photomicrographs of the holotypic male (ZMB 11590). A, segments 5–8, dorsal view; B, segments 5–8, ventral view; C, segments 6–10, dorsal view; D, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets." httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="8" pageNumber="549">6D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3D8629130BA2FE8F" box="[151,224,296,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="8" pageNumber="549">7D–G</figureCitation>
|
||
). Subdorsal and ventromedial sensory spots present (
|
||
<figureCitation id="135F2A73245EFFCB3E21291C08F2FE8F" box="[816,944,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Figs 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3EAB291308BEFE8F" box="[954,1020,296,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="8" pageNumber="549">7F, G</figureCitation>
|
||
). Type-1 glandular cell outlet present middorsally (
|
||
<figureCitation id="135F2A73245EFFCB3C3F29770AECFEAB" box="[302,430,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Fig. 2B, D</figureCitation>
|
||
). Posterior edges of sternal plates rounded with thin pectinate fringe teeth (
|
||
<figureCitation id="135F2A73245EFFCB380529770ED5FEAB" box="[1300,1431,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Figs 2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3D86294B0BFEFE47" box="[151,188,368,392]" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="8" pageNumber="549">7G</figureCitation>
|
||
). Posterior edge of tergal plate protruding subdorsally, forming long pointed tergal extensions (
|
||
<figureCitation id="135F2A73245EFFCB39F929540E25FE48" box="[1256,1383,367,392]" captionStart="FIGURE 2" captionStartId="4.[151,250,1845,1867]" captionTargetBox="[232,1343,197,1832]" captionTargetId="figure@4.[232,1343,193,1832]" captionTargetPageId="4" captionText="FIGURE 2. Echinoderes multiporus sp. nov., camera lucida drawings. A, B, Holotype, male (ZMB 11590), entire animal, ventral and dorsal views of segments 1–11, respectively; C, D, paratype, female (ZMB 11591a), segments 8–11, ventral and dorsal views, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla; pe, penile spine; si, sieve plate; ss, sensory spot; te, tergal extension; vlt, ventrolateral tube." httpUri="https://zenodo.org/record/1187743/files/figure.png" pageId="8" pageNumber="549">Figs 2B, D</figureCitation>
|
||
,
|
||
<figureCitation id="135F2A73245EFFCB3862294B0B88FE63" captionStart="FIGURE 7" captionStartId="10.[151,250,1829,1851]" captionTargetBox="[247,1350,193,1818]" captionTargetId="figure@10.[247,1350,193,1818]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes multiporus sp. nov., males (A, D, ZMB 11591g; B, E, G, ZMB 11591i) and a female (C, F, ZMB 11591j), scanning electron micrographs. A, segments 5–9, laterodorsal view (right side); B, segments 6–8, ventral view; C, anterior part of segment 8, ventral view; D, segments 9–11, laterodorsal view (right side); E, segments 10 and 11, lateral view (right side); F, segments 10 and 11, lateral view (left side); G, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandularcell outlets." httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="8" pageNumber="549">7D, F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E657D245EFFC83DD6298C09C7FDAF" lastPageId="11" lastPageNumber="552" pageId="8" pageNumber="549" type="discussion">
|
||
<paragraph id="8BDB36F6245EFFCB3DD6298C0F76FD3F" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="549">
|
||
<emphasis id="B910EAE4245EFFCB3DD6298C0A79FE1F" bold="true" box="[199,315,439,464]" pageId="8" pageNumber="549">Remarks.</emphasis>
|
||
Based on the spine and tube pattern having middorsal acicular spines only on segments 4, 6, and 8, lateroventral acicular spines on segments 6–9, ventrolateral tubes on segment 2, lateroventral tubes on segment 5, and midlateral tubes on segment 10, and lacking lateral accessory tubes on segment 8,
|
||
<taxonomicName id="4C644D75245EFFCB39762A3B0E31FDD7" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[1127,1395,512,536]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245EFFCB39762A3B0E31FDD7" box="[1127,1395,512,536]" italics="true" pageId="8" pageNumber="549">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245EFFCB386B29C40B8AFDF2" pageId="8" pageNumber="549" rank="species">
|
||
<emphasis id="B910EAE4245EFFCB386B29C40B8AFDF2" bold="true" pageId="8" pageNumber="549">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C644D75245EFFCB3C792A1E098AFDF3" authority="Higgins, 1982" authorityName="Higgins" authorityYear="1982" box="[360,712,548,573]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="bermudensis">
|
||
<emphasis id="B910EAE4245EFFCB3C792A1E095AFDF3" box="[360,536,548,572]" italics="true" pageId="8" pageNumber="549">E. bermudensis</emphasis>
|
||
<bibRefCitation id="EFF54B07245EFFCB3F332A1F098AFDF3" box="[546,712,548,573]" pageId="8" pageNumber="549" refId="ref8690" refString="Higgins, R. P. (1982) Three new species of Kinorhyncha from Bermuda. Transactions of the American Microscopical Society, 101, 305 - 316. https: // doi. org / 10.2307 / 3225748" type="journal article">Higgins, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C644D75245EFFCB3FC72A1E0FD2FDF2" authority="Landers & Sorensen, 2016" authorityName="Landers & Sorensen" authorityYear="2016" box="[726,1168,548,573]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="joyceae">
|
||
<emphasis id="B910EAE4245EFFCB3FC72A1E080CFDF3" box="[726,846,549,572]" italics="true" pageId="8" pageNumber="549">E. joyceae</emphasis>
|
||
<bibRefCitation id="EFF54B07245EFFCB3E492A1F0FD2FDF2" box="[856,1168,548,573]" pageId="8" pageNumber="549" refId="ref8860" refString="Landers, S. C. & Sorensen, M. V. (2016) Two new species of Echinoderes (Kinorhyncha, Cyclorhagida), E. romanoi sp. n. and E. joyceae sp. n., from the Gulf of Mexico. Zookeys, 594, 51 - 71. https: // doi. org / 10.3897 / zookeys. 594.8623" type="journal article">Landers & Sørensen, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C644D75245EFFCB39C32A1F0E1FFDF3" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[1234,1373,548,572]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="B910EAE4245EFFCB39C32A1F0E1FFDF3" box="[1234,1373,548,572]" italics="true" pageId="8" pageNumber="549">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
apex Yamasaki
|
||
<emphasis id="B910EAE4245EFFCB3C1F2A720A13FDAF" box="[270,337,584,608]" italics="true" pageId="8" pageNumber="549">et al.,</emphasis>
|
||
2018 (
|
||
<bibRefCitation id="EFF54B07245EFFCB3CB72A7C0904FDAF" box="[422,582,583,608]" pageId="8" pageNumber="549" refId="ref8690" refString="Higgins, R. P. (1982) Three new species of Kinorhyncha from Bermuda. Transactions of the American Microscopical Society, 101, 305 - 316. https: // doi. org / 10.2307 / 3225748" type="journal article">Higgins 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF54B07245EFFCB3F442A7C08CAFDAF" box="[597,904,583,608]" pageId="8" pageNumber="549" refId="ref8860" refString="Landers, S. C. & Sorensen, M. V. (2016) Two new species of Echinoderes (Kinorhyncha, Cyclorhagida), E. romanoi sp. n. and E. joyceae sp. n., from the Gulf of Mexico. Zookeys, 594, 51 - 71. https: // doi. org / 10.3897 / zookeys. 594.8623" type="journal article">Landers & Sørensen 2016</bibRefCitation>
|
||
; Yamasaki
|
||
<emphasis id="B910EAE4245EFFCB391F2A720F00FDAF" box="[1038,1090,584,608]" italics="true" pageId="8" pageNumber="549">et al</emphasis>
|
||
. in press). However, the new species can be easily distinguished from the above mentioned three species by the presence of type-2 glandular cell outlets on segments 2 and 3–9 (type-2 glandular cell outlets are present on segments 2, 6, and
|
||
<quantity id="4C9C9B13245EFFCB39D22AAB0FADFD67" box="[1219,1263,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="549" unit="in" value="8.0">8 in</quantity>
|
||
<taxonomicName id="4C644D75245EFFCB39E72AAA0E29FD68" authorityName="Landers & Sorensen" authorityYear="2016" box="[1270,1387,656,680]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="joyceae">
|
||
<emphasis id="B910EAE4245EFFCB39E72AAA0E45FD67" box="[1270,1287,657,680]" italics="true" pageId="8" pageNumber="549">E</emphasis>
|
||
.
|
||
<emphasis id="B910EAE4245EFFCB38072AAB0E29FD68" box="[1302,1387,656,679]" italics="true" pageId="8" pageNumber="549">joyceae</emphasis>
|
||
</taxonomicName>
|
||
and the undescribed species, but absent in
|
||
<taxonomicName id="4C644D75245EFFCB3F5F2A8E09BEFD03" authorityName="Higgins" authorityYear="1982" box="[590,764,692,716]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="bermudensis">
|
||
<emphasis id="B910EAE4245EFFCB3F5F2A8E09BEFD03" box="[590,764,692,716]" italics="true" pageId="8" pageNumber="549">E. bermudensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C644D75245EFFCB3E072A8F0F61FD03" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[790,1059,692,716]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245EFFCB3E072A8F0F61FD03" box="[790,1059,692,716]" italics="true" pageId="8" pageNumber="549">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245EFFCB393D2A8F0FC5FD02" box="[1068,1159,692,717]" pageId="8" pageNumber="549" rank="species">
|
||
<emphasis id="B910EAE4245EFFCB393D2A8F0FC5FD02" bold="true" box="[1068,1159,692,717]" pageId="8" pageNumber="549">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the only
|
||
<taxonomicName id="4C644D75245EFFCB38002A8F0EDEFD03" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[1297,1436,692,716]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="B910EAE4245EFFCB38002A8F0EDEFD03" box="[1297,1436,692,716]" italics="true" pageId="8" pageNumber="549">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
species with the laterodorsal type-2 glandular cell outlets on most of the segments.
|
||
</paragraph>
|
||
<caption id="DF1B667E245EFFCB3D862F1E0A31F87A" httpUri="https://zenodo.org/record/1187749/files/figure.png" pageId="8" pageNumber="549" startId="8.[151,250,1829,1851]" targetBox="[162,1426,843,1776]" targetPageId="8">
|
||
<paragraph id="8BDB36F6245EFFCB3D862F1E0A31F87A" blockId="8.[151,1437,1829,1973]" pageId="8" pageNumber="549">
|
||
<emphasis id="B910EAE4245EFFCB3D862F1E0A51F8F5" bold="true" box="[151,275,1829,1851]" pageId="8" pageNumber="549">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C644D75245EFFCB3C0B2F1E094EF8F4" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[282,524,1829,1851]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="549" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245EFFCB3C0B2F1E094EF8F4" box="[282,524,1829,1851]" italics="true" pageId="8" pageNumber="549">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245EFFCB3F022F1E0921F8F5" box="[531,611,1829,1850]" pageId="8" pageNumber="549" rank="species">
|
||
<emphasis id="B910EAE4245EFFCB3F022F1E0921F8F5" bold="true" box="[531,611,1829,1850]" pageId="8" pageNumber="549">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Nomarski photomicrographs of a paratypic female (ZMB 11591a).
|
||
<emphasis id="B910EAE4245EFFCB380F2F1E0E72F8F5" bold="true" box="[1310,1328,1829,1850]" pageId="8" pageNumber="549">A</emphasis>
|
||
, segments 2–7, ventral view;
|
||
<emphasis id="B910EAE4245EFFCB3C4D2F7F0A2FF896" bold="true" box="[348,365,1860,1881]" pageId="8" pageNumber="549">B</emphasis>
|
||
, segments 8–11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; lvt, lateroventral tube; pa, papilla; ppf, primary pectinate fringe; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1B667E245FFFCA3D862F6708E2F823" httpUri="https://zenodo.org/record/1187751/files/figure.png" pageId="9" pageNumber="550" startId="9.[151,250,1884,1906]" targetBox="[161,1426,209,1855]" targetPageId="9">
|
||
<paragraph id="8BDB36F6245FFFCA3D862F6708E2F823" blockId="9.[151,1437,1884,2028]" pageId="9" pageNumber="550">
|
||
<emphasis id="B910EAE4245FFFCA3D862F670A56F8BE" bold="true" box="[151,276,1884,1906]" pageId="9" pageNumber="550">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C644D75245FFFCA3C0A2F67094CF8BD" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[283,526,1884,1906]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="9" pageNumber="550" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245FFFCA3C0A2F67094CF8BD" box="[283,526,1884,1906]" italics="true" pageId="9" pageNumber="550">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245FFFCA3F042F670925F8BE" box="[533,615,1884,1905]" pageId="9" pageNumber="550" rank="species">
|
||
<emphasis id="B910EAE4245FFFCA3F042F670925F8BE" bold="true" box="[533,615,1884,1905]" pageId="9" pageNumber="550">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Nomarski photomicrographs of the holotypic male (ZMB 11590).
|
||
<emphasis id="B910EAE4245FFFCA380C2F670E6DF8BE" bold="true" box="[1309,1327,1884,1905]" pageId="9" pageNumber="550">A</emphasis>
|
||
, segments 5–8, dorsal view;
|
||
<emphasis id="B910EAE4245FFFCA3C4B2F400A29F85F" bold="true" box="[346,363,1915,1936]" pageId="9" pageNumber="550">B</emphasis>
|
||
, segments 5–8, ventral view;
|
||
<emphasis id="B910EAE4245FFFCA3FA12F410980F85F" bold="true" box="[688,706,1914,1936]" pageId="9" pageNumber="550">C</emphasis>
|
||
, segments 6–10, dorsal view;
|
||
<emphasis id="B910EAE4245FFFCA391D2F400F5CF85F" bold="true" box="[1036,1054,1915,1936]" pageId="9" pageNumber="550">D</emphasis>
|
||
, segments 7–11, ventrolateral view. Abbreviations: lvs, lateroventral acicular spine; lvt, lateroventral tube; mds, middorsal acicular spine; pe, penile spine; si, sieve plate. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowheads, type-1 glandular cell outlets; black arrowheads, type-2 glandular cell outlets.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1B667E245CFFC93D862F1E0E71F83D" httpUri="https://zenodo.org/record/1187753/files/figure.png" pageId="10" pageNumber="551" startId="10.[151,250,1829,1851]" targetBox="[247,1350,193,1818]" targetPageId="10">
|
||
<paragraph id="8BDB36F6245CFFC93D862F1E0E71F83D" blockId="10.[151,1436,1828,2034]" pageId="10" pageNumber="551">
|
||
<emphasis id="B910EAE4245CFFC93D862F1E0A56F8F5" bold="true" box="[151,276,1829,1851]" pageId="10" pageNumber="551">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C644D75245CFFC93C0D2F1F0952F8F5" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[284,528,1828,1850]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="551" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245CFFC93C0D2F1F0952F8F5" box="[284,528,1828,1850]" italics="true" pageId="10" pageNumber="551">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245CFFC93F092F1E0928F8F5" box="[536,618,1829,1850]" pageId="10" pageNumber="551" rank="species">
|
||
<emphasis id="B910EAE4245CFFC93F092F1E0928F8F5" bold="true" box="[536,618,1829,1850]" pageId="10" pageNumber="551">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, males (
|
||
<emphasis id="B910EAE4245CFFC93FD42F1E0995F8F5" bold="true" box="[709,727,1829,1850]" pageId="10" pageNumber="551">A</emphasis>
|
||
,
|
||
<emphasis id="B910EAE4245CFFC93FF72F1E09BAF8F5" bold="true" box="[742,760,1829,1850]" pageId="10" pageNumber="551">D</emphasis>
|
||
, ZMB 11591g;
|
||
<emphasis id="B910EAE4245CFFC93EB12F1E08F3F8F5" bold="true" box="[928,945,1829,1850]" pageId="10" pageNumber="551">B</emphasis>
|
||
,
|
||
<emphasis id="B910EAE4245CFFC93EAE2F1E0892F8F5" bold="true" box="[959,976,1829,1850]" pageId="10" pageNumber="551">E</emphasis>
|
||
,
|
||
<emphasis id="B910EAE4245CFFC93ECF2F1E08ADF8F4" bold="true" box="[990,1007,1829,1851]" pageId="10" pageNumber="551">G</emphasis>
|
||
, ZMB 11591i) and a female (
|
||
<emphasis id="B910EAE4245CFFC938392F1E0E78F8F4" bold="true" box="[1320,1338,1829,1851]" pageId="10" pageNumber="551">C</emphasis>
|
||
,
|
||
<emphasis id="B910EAE4245CFFC938582F1E0E14F8F5" bold="true" box="[1353,1366,1829,1850]" pageId="10" pageNumber="551">F</emphasis>
|
||
, ZMB 11591j), scanning electron micrographs.
|
||
<emphasis id="B910EAE4245CFFC93F2F2F780912F897" bold="true" box="[574,592,1859,1880]" pageId="10" pageNumber="551">A</emphasis>
|
||
, segments 5–9, laterodorsal view (right side);
|
||
<emphasis id="B910EAE4245CFFC939262F7F0F0AF896" bold="true" box="[1079,1096,1860,1881]" pageId="10" pageNumber="551">B</emphasis>
|
||
, segments 6–8, ventral view;
|
||
<emphasis id="B910EAE4245CFFC938922F780ED7F896" bold="true" box="[1411,1429,1859,1881]" pageId="10" pageNumber="551">C</emphasis>
|
||
, anterior part of segment 8, ventral view;
|
||
<emphasis id="B910EAE4245CFFC93F292F590908F8B8" bold="true" box="[568,586,1890,1911]" pageId="10" pageNumber="551">D</emphasis>
|
||
, segments 9–11, laterodorsal view (right side);
|
||
<emphasis id="B910EAE4245CFFC939212F590F03F8B8" bold="true" box="[1072,1089,1890,1911]" pageId="10" pageNumber="551">E</emphasis>
|
||
, segments 10 and 11, lateral view (right side);
|
||
<emphasis id="B910EAE4245CFFC93C092FBA0A67F859" bold="true" box="[280,293,1921,1942]" pageId="10" pageNumber="551">F</emphasis>
|
||
, segments 10 and 11, lateral view (left side);
|
||
<emphasis id="B910EAE4245CFFC93E152FBB0857F859" bold="true" box="[772,789,1920,1942]" pageId="10" pageNumber="551">G</emphasis>
|
||
, segments 10 and 11, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral acicular spine; mds, middorsal acicular spine; mlt, midlateral tube; pa, papilla, pe, penile spine. Digits after abbreviations indicate the corresponding segment number. Black arrows, sensory spots; white arrowhead, type-1 glandular cell outlet; black arrowheads, type-2 glandular cell outlets.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB36F6245DFFC83DD628AC09C7FDAF" blockId="11.[151,1436,151,608]" pageId="11" pageNumber="552">
|
||
The presence of only two pairs of penile spines found in males of
|
||
<taxonomicName id="4C644D75245DFFC83EA528A30F83FF7F" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[948,1217,152,176]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245DFFC83EA528A30F83FF7F" box="[948,1217,152,176]" italics="true" pageId="11" pageNumber="552">Echinoderes multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245DFFC839D928AC0E61FF7F" box="[1224,1315,151,176]" pageId="11" pageNumber="552" rank="species">
|
||
<emphasis id="B910EAE4245DFFC839D928AC0E61FF7F" bold="true" box="[1224,1315,151,176]" pageId="11" pageNumber="552">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is also the rare character among the congeners. Males of
|
||
<taxonomicName id="4C644D75245DFFC83F8828870866FF1B" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[665,804,188,212]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="B910EAE4245DFFC83F8828870866FF1B" box="[665,804,188,212]" italics="true" pageId="11" pageNumber="552">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
species usually have three pairs of penile spine, whereas two pairs of penile spines have been found in the following ten species together with
|
||
<taxonomicName id="4C644D75245DFFC8399E28DA0E69FF37" authority="Yamasaki & Neuhaus & George, 2018" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[1167,1323,224,248]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="multiporus" status="sp. nov.">
|
||
<emphasis id="B910EAE4245DFFC8399E28DA0E69FF37" box="[1167,1323,224,248]" italics="true" pageId="11" pageNumber="552">E. multiporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223579F245DFFC8382728E40ED6FF37" box="[1334,1428,223,248]" pageId="11" pageNumber="552" rank="species">
|
||
<emphasis id="B910EAE4245DFFC8382728E40ED6FF37" bold="true" box="[1334,1428,223,248]" pageId="11" pageNumber="552">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B910EAE4245DFFC83D86293F0A6BFED3" box="[151,297,260,284]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C644D75245DFFC83D86293F0A64FED3" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[151,294,260,284]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">Echinoderes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C644D75245DFFC83C3E293E0933FED3" authority="Adrianov et al., 2002" authorityName="Adrianov et al." authorityYear="2002" box="[303,625,260,285]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="aureus">
|
||
<emphasis id="B910EAE4245DFFC83C3E293E0A38FED3" box="[303,378,261,284]" italics="true" pageId="11" pageNumber="552">aureus</emphasis>
|
||
Adrianov
|
||
<emphasis id="B910EAE4245DFFC83CE0293E0963FED3" box="[497,545,260,284]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
., 2002
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B910EAE4245DFFC83F6D293F0833FED3" box="[636,881,260,284]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C644D75245DFFC83F6D293F0849FED3" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[636,779,260,284]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">Echinoderes</taxonomicName>
|
||
. astridae
|
||
</emphasis>
|
||
<bibRefCitation id="EFF54B07245DFFC83E66293F0F64FED3" box="[887,1062,260,284]" pageId="11" pageNumber="552" refId="ref9246" refString="Neuhaus, B., Pardos, F., Sorensen, M. V. & Higgins, R. P. (2014) New species of Centroderes (Kinorhyncha, Cyclorhagida) from the Northwest Atlantic Ocean, life cycle, and ground pattern of the genus. Zootaxa, 3901 (1), 1 ¯ 69. https: // doi. org / 10.11646 / zootaxa. 3901.1.1" type="journal article">Sørensen, 2014</bibRefCitation>
|
||
;
|
||
<emphasis id="B910EAE4245DFFC83920293F0E6DFED3" box="[1073,1327,260,284]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C644D75245DFFC83920293F0F82FED3" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[1073,1216,260,284]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">Echinoderes</taxonomicName>
|
||
. cavernus
|
||
</emphasis>
|
||
Sørensen
|
||
<emphasis id="B910EAE4245DFFC83D8629120B88FE8F" box="[151,202,296,320]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
., 2000;
|
||
<taxonomicName id="4C644D75245DFFC83C38291309A0FE8F" authority="Higgins, 1985" authorityName="Higgins" authorityYear="1985" box="[297,738,295,320]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="kristenseni">
|
||
<emphasis id="B910EAE4245DFFC83C3829130974FE8F" box="[297,566,296,320]" italics="true" pageId="11" pageNumber="552">Echinoderes kristenseni</emphasis>
|
||
<bibRefCitation id="EFF54B07245DFFC83F2E291C09A0FE8F" box="[575,738,295,320]" pageId="11" pageNumber="552" refId="ref8730" refString="Higgins, R. P. (1985) The genus Echinoderes (Kinorhyncha: Cyclorhagida) from the English Channel. Journal of the Marine Biological Association of the United Kingdom, 65, 785 - 800. https: // doi. org / 10.1017 / S 0025315400052590" type="journal article">Higgins, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C644D75245DFFC83FE129130F15FE8F" authority="Chang" authorityName="Chang" box="[752,1111,295,320]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="lanceolatus">
|
||
<emphasis id="B910EAE4245DFFC83FE129130F47FE8F" box="[752,1029,296,320]" italics="true" pageId="11" pageNumber="552">Echinoderes lanceolatus</emphasis>
|
||
Chang
|
||
</taxonomicName>
|
||
& Song, 2002;
|
||
<taxonomicName id="4C644D75245DFFC838002913094DFEAA" authority="Higgins, 1967" authorityName="Higgins" authorityYear="1967" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="newcaledoniensis">
|
||
<emphasis id="B910EAE4245DFFC8380029130A1EFEAB" italics="true" pageId="11" pageNumber="552">Echinoderes newcaledoniensis</emphasis>
|
||
<bibRefCitation id="EFF54B07245DFFC83C792977094DFEAA" box="[360,527,332,357]" pageId="11" pageNumber="552" refId="ref8502" refString="Higgins, R. P. (1967) The Kinorhyncha of New Caledonia. In: Expedition Francaise sur les Recifs coralliens de la Nouvelle- Caledonie. 2. Fondation Singer-Polignac, Paris, pp. 75 - 90." type="book chapter">Higgins, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B910EAE4245DFFC83F0E29770858FEAB" box="[543,794,332,356]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C644D75245DFFC83F0E297709ECFEAB" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[543,686,332,356]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">Echinoderes</taxonomicName>
|
||
. marthae
|
||
</emphasis>
|
||
<bibRefCitation id="EFF54B07245DFFC83E3429770898FEAB" box="[805,986,332,356]" pageId="11" pageNumber="552" refId="ref9246" refString="Neuhaus, B., Pardos, F., Sorensen, M. V. & Higgins, R. P. (2014) New species of Centroderes (Kinorhyncha, Cyclorhagida) from the Northwest Atlantic Ocean, life cycle, and ground pattern of the genus. Zootaxa, 3901 (1), 1 ¯ 69. https: // doi. org / 10.11646 / zootaxa. 3901.1.1" type="journal article">Sørensen, 2014</bibRefCitation>
|
||
;
|
||
<taxonomicName id="4C644D75245DFFC83EFB29770B99FE47" authority="(Omer-Cooper, 1957)" authorityName="Omer-Cooper" authorityYear="1957" baseAuthorityName="Omer-Cooper" baseAuthorityYear="1957" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="maxwelli">
|
||
<emphasis id="B910EAE4245DFFC83EFB29770FA4FEAB" box="[1002,1254,332,356]" italics="true" pageId="11" pageNumber="552">Echinoderes maxwelli</emphasis>
|
||
(Omer-Cooper, 1957)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C644D75245DFFC83DF8294B09C2FE47" authority="(Blake, 1930)" authorityName="Blake" authorityYear="1930" baseAuthorityName="Blake" baseAuthorityYear="1930" box="[233,640,367,392]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="remanei">
|
||
<emphasis id="B910EAE4245DFFC83DF8294B0A9AFE48" box="[233,472,368,392]" italics="true" pageId="11" pageNumber="552">Echinoderes remanei</emphasis>
|
||
(Blake, 1930)
|
||
</taxonomicName>
|
||
; and
|
||
<emphasis id="B910EAE4245DFFC83FAE294B0810FE47" box="[703,850,368,392]" italics="true" pageId="11" pageNumber="552">
|
||
<taxonomicName id="4C644D75245DFFC83FAE294B080CFE47" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[703,846,368,392]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">Echinoderes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C644D75245DFFC83E4B294A0F36FE47" authority="Lundbye et al., 2011" authorityName="Lundbye et al." authorityYear="2011" box="[858,1140,367,392]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="species" species="rex">
|
||
<emphasis id="B910EAE4245DFFC83E4B294A083FFE47" box="[858,893,369,392]" italics="true" pageId="11" pageNumber="552">rex</emphasis>
|
||
<bibRefCitation id="EFF54B07245DFFC83E9729540F36FE47" box="[902,1140,367,392]" pageId="11" pageNumber="552" refId="ref8929" refString="Lundbye, H., Rho, H. S. & Sorensen, M. V. (2011) Echinoderes rex n. sp. (Kinorhyncha: Cyclorhagida), the largest Echinoderes species found so far. Scientia Marina, 75, 41 - 51. https: // doi. org / 10.3989 / scimar. 2011.75 n 1041" type="journal article">
|
||
Lundbye
|
||
<emphasis id="B910EAE4245DFFC83EE2294A0F64FE47" box="[1011,1062,368,392]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFF54B07245DFFC839A229540E12FE47" box="[1203,1360,367,392]" pageId="11" pageNumber="552" refId="ref8465" refString="Higgins, R. P. (1964) Redescription of the kinorhynch Echinoderes remanei (Blake, 1930) Karling, 1954. Transactions of the American Microscopical Society, 83, 243 - 247." type="journal article">Higgins 1964</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF54B07245DFFC8384C29540EDAFE47" box="[1373,1432,367,392]" pageId="11" pageNumber="552" refId="ref8502" refString="Higgins, R. P. (1967) The Kinorhyncha of New Caledonia. In: Expedition Francaise sur les Recifs coralliens de la Nouvelle- Caledonie. 2. Fondation Singer-Polignac, Paris, pp. 75 - 90." type="book chapter">1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF54B07245DFFC83D8629AF0B90FE62" box="[151,210,404,429]" pageId="11" pageNumber="552" refId="ref8730" refString="Higgins, R. P. (1985) The genus Echinoderes (Kinorhyncha: Cyclorhagida) from the English Channel. Journal of the Marine Biological Association of the United Kingdom, 65, 785 - 800. https: // doi. org / 10.1017 / S 0025315400052590" type="journal article">1985</bibRefCitation>
|
||
; Sørensen
|
||
<emphasis id="B910EAE4245DFFC83C4329AE0AC7FE63" box="[338,389,404,428]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
. 2000; Adrianov
|
||
<emphasis id="B910EAE4245DFFC83F4429AE09C8FE63" box="[597,650,404,428]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
. 2002; Chang & Song 2002;
|
||
<bibRefCitation id="EFF54B07245DFFC83EF729AF0F96FE63" box="[998,1236,404,429]" pageId="11" pageNumber="552" refId="ref8929" refString="Lundbye, H., Rho, H. S. & Sorensen, M. V. (2011) Echinoderes rex n. sp. (Kinorhyncha: Cyclorhagida), the largest Echinoderes species found so far. Scientia Marina, 75, 41 - 51. https: // doi. org / 10.3989 / scimar. 2011.75 n 1041" type="journal article">
|
||
Lundbye
|
||
<emphasis id="B910EAE4245DFFC8394529AE0FCBFE63" box="[1108,1161,404,428]" italics="true" pageId="11" pageNumber="552">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF54B07245DFFC839F229AF0ED3FE63" box="[1251,1425,404,428]" pageId="11" pageNumber="552" refId="ref9246" refString="Neuhaus, B., Pardos, F., Sorensen, M. V. & Higgins, R. P. (2014) New species of Centroderes (Kinorhyncha, Cyclorhagida) from the Northwest Atlantic Ocean, life cycle, and ground pattern of the genus. Zootaxa, 3901 (1), 1 ¯ 69. https: // doi. org / 10.11646 / zootaxa. 3901.1.1" type="journal article">Sørensen 2014</bibRefCitation>
|
||
). Despite the presence of only two pairs of penile spines in these species, the species do not agree in most of the other morphological characters, i.e., the shape and the patterns of spines, tubes, and type-2 glandular cell outlets. It can be supposed that only two pairs of penile spines evolved repeatedly independently within the genus. However, the morphological evolution of
|
||
<taxonomicName id="4C644D75245DFFC83F1C2A1F09DAFDF3" authorityName="Yamasaki & Neuhaus & George" authorityYear="2018" box="[525,664,548,572]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="552" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="B910EAE4245DFFC83F1C2A1F09DAFDF3" box="[525,664,548,572]" italics="true" pageId="11" pageNumber="552">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
is still open to question due to the lack of knowledge about the phylogenetic relationships within the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |