treatments-xml/data/03/92/87/039287E6FF8FFFD1FE108DA5CAA947DA.xml

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<mods:title id="3753FECE3991C420AB8CB83C0A34A9A0">Campydoroides manautei gen. et sp. nov. from New Caledonia and a reappraisal of the suborder Campydorina (Nematoda)</mods:title>
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<treatment id="039287E6FF8FFFD1FE108DA5CAA947DA" ID-DOI="http://doi.org/10.5281/zenodo.5613679" ID-GBIF-Taxon="156828368" ID-Zenodo-Dep="5613679" LSID="urn:lsid:zoobank.org:act:CD624B5D-3F24-4D54-9C3C-C00615BB340E" httpUri="http://treatment.plazi.org/id/039287E6FF8FFFD1FE108DA5CAA947DA" lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8">
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<paragraph id="8B8436F0FF8FFFDAFE108DA5CC5C43BB" blockId="7.[363,1224,391,488]" box="[472,1115,391,418]" pageId="7" pageNumber="8">
<heading id="D0CC819CFF8FFFDAFE108DA5CC5C43BB" box="[472,1115,391,418]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C3B4D73FF8FFFDAFE108DA5CBA343B8" authority="Holovachov" authorityName="Holovachov" authorityYear="2019" box="[472,932,391,418]" class="Adenophorea" family="Campydoridae" genus="Campydoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="7" pageNumber="8" phylum="Nematoda" rank="species" species="manautei" status="gen. et sp. nov.">
<emphasis id="B94FEAE2FF8FFFDAFE108DA5CB0A43B8" bold="true" box="[472,781,391,418]" italics="true" pageId="7" pageNumber="8">Campydoroides manautei</emphasis>
Holovachov
</taxonomicName>
<taxonomicNameLabel id="A27C5799FF8FFFDAFC638DAACC5C43BB" box="[939,1115,392,418]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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<paragraph id="8B8436F0FF8FFFDAFD7C8DECCB8743F1" blockId="7.[363,1224,391,488]" box="[692,896,462,488]" pageId="7" pageNumber="8">
<figureCitation id="13002A75FF8FFFDAFD7C8DECCB1943F1" box="[692,798,462,488]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="8.[189,232,1886,1912]" captionStartId-1="9.[189,232,1921,1947]" captionStartId-2="10.[189,232,1921,1947]" captionTargetBox-0="[358,1224,285,1830]" captionTargetBox-1="[357,1245,287,1863]" captionTargetBox-2="[215,1372,319,1860]" captionTargetId-0="figure@8.[288,1299,265,1845]" captionTargetId-1="figure@9.[232,1356,265,1881]" captionTargetId-2="figure@10.[189,1399,293,1881]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Fig. 2. Campydoroides manautei Holovachov gen. et sp. nov. A–B. Paratype female (SMNH Type- 9167). C–F. Holotype (MNHN-BN511). A. Female pharyngeal region, median section. B. Female anterior end, median section. C–D. Female anterior end, surface view. E. Female reproductive system. F. Female tail. Scale bar: 20 µm." captionText-1="Fig. 3. Campydoroides manautei Holovachov gen. et sp. nov., non-type female from New Zealand (SMNH-175283). A. Female pharyngeal region, median section. B. Female anterior end, surface view. C. Female reproductive system. D. Female tail. Scale bar: 20 µm." captionText-2="Fig. 4. Campydoroides manautei Holovachov gen. et sp. nov., non-type female from New Zealand (SMNH-175283). A. Anterior end, median section. B. Anterior end, surface view. C. Vulval region. D. Base of pharynx. Scale bar: 20 µm." httpUri-0="https://zenodo.org/record/2649837/files/figure.png" httpUri-1="https://zenodo.org/record/2649839/files/figure.png" httpUri-2="https://zenodo.org/record/2649841/files/figure.png" pageId="7" pageNumber="8">Figs 2–4</figureCitation>
; 
<tableCitation id="C6B9034BFF8FFFDAFCE18DECCB8743F1" box="[809,896,462,488]" captionStart="Table 3" captionStartId="11.[189,258,268,294]" captionTargetBox="[197,1386,375,1288]" captionText="Table 3. Morphometrics of Campydoroides manautei Holovachov gen. et sp. nov. (all measurements are in µm, except for the ratios a, b, c, c’ and V)." httpUri="http://table.plazi.org/id/DF446678FF83FFD6FF758D2ECACB4350" pageId="7" pageNumber="8" tableUuid="DF446678FF83FFD6FF758D2ECACB4350">Table 3</tableCitation>
</paragraph>
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<subSubSection id="C321657BFF8FFFDAFF758E32C9A140FF" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8B8436F0FF8FFFDAFF758E32C9314033" blockId="7.[189,310,528,554]" box="[189,310,528,554]" pageId="7" pageNumber="8">
<heading id="D0CC819CFF8FFFDAFF758E32C9314033" bold="true" box="[189,310,528,554]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B94FEAE2FF8FFFDAFF758E32C9314033" bold="true" box="[189,310,528,554]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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</paragraph>
<paragraph id="8B8436F0FF8FFFDAFF758E1DC9A140FF" blockId="7.[189,1399,574,742]" pageId="7" pageNumber="8">
<taxonomicName id="4C3B4D73FF8FFFDAFF758E1DC9E94041" authority="Holovachov, 2019" authorityName="Holovachov" authorityYear="2019" box="[189,494,575,601]" class="Adenophorea" family="Campydoridae" genus="Campydoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="7" pageNumber="8" phylum="Nematoda" rank="species" species="manautei" status="sp. nov.">
<emphasis id="B94FEAE2FF8FFFDAFF758E1DC9E94041" box="[189,494,575,601]" italics="true" pageId="7" pageNumber="8">Campydoroides manautei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C5799FF8FFFDAFE318E1DCA5F4040" box="[505,600,575,601]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
is characterised by a 
<quantity id="4CC39B15FF8FFFDAFCAF8E1DCC114040" box="[871,1046,575,601]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="8.2" metricValueMin="4.9" pageId="7" pageNumber="8" unit="mm" value="0.655" valueMax="0.82" valueMin="0.49">0.49–0.82 mm</quantity>
long body, papilliform inner labial, outer labial and cephalic sensilla, amphidial opening a transverse slit, strongly cuticularised stoma with single robust dorsal tooth, muscular pharynx with distinct basal bulb, large cardia, didelphic, amphidelphic female gonads, straight vagina, and elongate conoid tail equal to 6.0–9.6 anal body diameters in length.
</paragraph>
</subSubSection>
<subSubSection id="C321657BFF8FFFDAFF758F2CCBD94163" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8B8436F0FF8FFFDAFF758F2CC9434131" blockId="7.[189,324,782,808]" box="[189,324,782,808]" pageId="7" pageNumber="8">
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<emphasis id="B94FEAE2FF8FFFDAFF758F2CC9434131" bold="true" box="[189,324,782,808]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8436F0FF8FFFDAFF758F1FCBD94163" blockId="7.[189,1398,829,891]" pageId="7" pageNumber="8">The new species is named in honour of Joseph Manauté, director of the Parc provincial de la Rivière bleue, for his indispensable support during the sampling in the Park.</paragraph>
</subSubSection>
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<emphasis id="B94FEAE2FF8FFFDAFF758F80C96B41A5" bold="true" box="[189,364,930,956]" pageId="7" pageNumber="8">Type material</emphasis>
</heading>
</paragraph>
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<emphasis id="B94FEAE2FF8FFFDAFF228FF3C95B41F2" bold="true" box="[234,348,977,1003]" pageId="7" pageNumber="8">Holotype</emphasis>
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• 
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; 
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, 
<collectingMunicipality id="6BE0AC8AFF8FFFDAFD658FD7CAE64616" box="[685,737,1013,1039]" pageId="7" pageNumber="8">Parc</collectingMunicipality>
provincial 
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bleue, station 
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/4680; 
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, 
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; 
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<date id="FF851030FF8FFFDAFDD3883ACAA8462B" box="[539,687,1048,1074]" pageId="7" pageNumber="8" value="2018-07-18">18 Jul. 2018</date>
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; 
<collectorName id="26CE5326FF8FFFDAFD73883ACB69462B" box="[699,878,1048,1074]" pageId="7" pageNumber="8">O. Holovachov</collectorName>
leg.; virgin forest, soil; 
<specimenCode id="DB9D9E8BFF8FFFDAFB408835CD44462B" box="[1160,1347,1047,1074]" collectionCode="MNHN-BN" pageId="7" pageNumber="8">MNHN-BN511</specimenCode>
.
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</paragraph>
<paragraph id="8B8436F0FF8FFFDAFF228878C960466D" blockId="7.[189,1399,1114,1317]" box="[234,359,1114,1140]" pageId="7" pageNumber="8">
<typeStatus id="54808852FF8FFFDAFF228878C960466D" box="[234,359,1114,1140]" pageId="7" pageNumber="8" type="paratype">
<emphasis id="B94FEAE2FF8FFFDAFF228878C960466D" bold="true" box="[234,359,1114,1140]" pageId="7" pageNumber="8">Paratypes</emphasis>
</typeStatus>
</paragraph>
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<collectingCountry id="F32C7660FF8FFFDAFF75885FC9AA468E" box="[189,429,1149,1175]" name="New Caledonia" pageId="7" pageNumber="8">NEW CALEDONIA</collectingCountry>
• 
<specimenCount id="9D3DFD79FF8FFFDAFE08885FCA07468E" box="[448,512,1149,1175]" pageId="7" pageNumber="8" type="female">4 ♀♀</specimenCount>
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<location id="8EE4602BFF8FFFDAFDC3885FCAA8468E" LSID="urn:lsid:plazi:treatment:039287E6FF8FFFD1FE108DA5CAA947DA:8EE4602BFF8FFFDAFDC3885FCAA8468E" box="[523,687,1149,1175]" country="New Caledonia" latitude="-22.07335" longLatPrecision="1" longitude="166.61658" municipality="Parc" name="Yate Province" pageId="7" pageNumber="8">Yate Province</location>
, 
<collectingMunicipality id="6BE0AC8AFF8FFFDAFD70885FCAEB468E" box="[696,748,1149,1175]" pageId="7" pageNumber="8">Parc</collectingMunicipality>
provincial 
<collectorName id="26CE5326FF8FFFDAFCA3885FCBFC468E" box="[875,1019,1149,1175]" pageId="7" pageNumber="8">de la Rivière</collectorName>
bleue, station 
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/4680; 
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, 
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; 
<collectingDate id="EFC1E9D8FF8FFFDAFDEB8882CB9E46A3" box="[547,921,1184,1211]" isEnumeration="true" pageId="7" pageNumber="8">
<date id="FF851030FF8FFFDAFDEB8882CAC246A3" box="[547,709,1184,1211]" pageId="7" pageNumber="8" value="2017-11-19">19 Nov. 2017</date>
and 
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; 
<collectorName id="26CE5326FF8FFFDAFC6F8882CC5B46A3" box="[935,1116,1184,1210]" pageId="7" pageNumber="8">O. Holovachov</collectorName>
leg.; virgin forest, soil; 
<specimenCode id="DB9D9E8BFF8FFFDAFF7588E6C97046C6" box="[189,375,1220,1247]" collectionCode="MNHN-BN" pageId="7" pageNumber="8">MNHN-BN511</specimenCode>
, 
<collectionCode id="ED2AAE35FF8FFFDAFE4B88E6C9DF46C7" box="[387,472,1220,1246]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="7" pageNumber="8">SMNH</collectionCode>
Type-9167
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• 
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, 
<collectingMunicipality id="6BE0AC8AFF8FFFDAFCB788E6CBB446C7" box="[895,947,1220,1246]" pageId="7" pageNumber="8">Parc</collectingMunicipality>
provincial 
<collectorName id="26CE5326FF8FFFDAFBF288E6CCD646C7" box="[1082,1233,1220,1246]" pageId="7" pageNumber="8">de la Rivière</collectorName>
bleue, station 
<specimenCode id="DB9D9E8BFF8FFFDAFF7588C5C9484718" box="[189,335,1255,1281]" collectionCode="HYNC" pageId="7" pageNumber="8">HYNC2378</specimenCode>
; 
<geoCoordinate id="EE0F5037FF8FFFDAFE9288C7C9F34718" box="[346,500,1253,1281]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-22.089567">22°05.374′ S</geoCoordinate>
, 
<geoCoordinate id="EE0F5037FF8FFFDAFDC988C7CAAA471B" box="[513,685,1253,1282]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="166.63641">166°38.185′ E</geoCoordinate>
; 
<collectingDate id="EFC1E9D8FF8FFFDAFD7288C5CB5B4718" box="[698,860,1255,1282]" pageId="7" pageNumber="8" value="2017-11-19">
<date id="FF851030FF8FFFDAFD7288C5CB5B4718" box="[698,860,1255,1282]" pageId="7" pageNumber="8" value="2017-11-19">19 Nov. 2017</date>
</collectingDate>
; 
<collectorName id="26CE5326FF8FFFDAFCA088C5CC1B4718" box="[872,1052,1255,1281]" pageId="7" pageNumber="8">O. Holovachov</collectorName>
leg.; virgin forest, wet soil at the bottom of a stream bed; 
<collectionCode id="ED2AAE35FF8FFFDAFDCE8929CA5C473C" box="[518,603,1291,1317]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="7" pageNumber="8">SMNH</collectionCode>
Type-9168, 
<collectionCode id="ED2AAE35FF8FFFDAFD278929CB43473C" box="[751,836,1291,1317]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="7" pageNumber="8">SMNH</collectionCode>
Type-9169
</materialsCitation>
.
</paragraph>
<paragraph id="8B8436F0FF8FFFDAFF22896EC91A47B7" blockId="7.[189,1399,1356,1454]" pageId="7" pageNumber="8">
<emphasis id="B94FEAE2FF8FFFDAFF22896ECA50477F" bold="true" box="[234,599,1356,1383]" pageId="7" pageNumber="8">Additional non-type material</emphasis>
<materialsCitation id="3B533CADFF8FFFDAFF758952C91E47B7" ID-GBIF-Occurrence="2236816406" collectingDate="1988-11-10" collectorName="D. Sturhan" country="New Zealand" location="NEW ZEALAND" pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="2">
<collectingCountry id="F32C7660FF8FFFDAFF758952C9934793" box="[189,404,1392,1418]" name="New Zealand" pageId="7" pageNumber="8">NEW ZEALAND</collectingCountry>
• 
<specimenCount id="9D3DFD79FF8FFFDAFE7C8952C9FD4793" box="[436,506,1392,1418]" pageId="7" pageNumber="8" type="female">2 ♀♀</specimenCount>
; Napier, Puketitiri, station Z236; 
<collectingDate id="EFC1E9D8FF8FFFDAFC5D8952CC394793" box="[917,1086,1392,1418]" pageId="7" pageNumber="8" value="1988-11-10">
<date id="FF851030FF8FFFDAFC5D8952CC394793" box="[917,1086,1392,1418]" pageId="7" pageNumber="8" value="1988-11-10">10 Nov. 1988</date>
</collectingDate>
; 
<collectorName id="26CE5326FF8FFFDAFB858952CCC84793" box="[1101,1231,1392,1418]" pageId="7" pageNumber="8">D. Sturhan</collectorName>
leg.; SMNH- 175283
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321657BFF8FFFD1FF7589F7CB3D43E3" lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8" type="description">
<paragraph id="8B8436F0FF8FFFDAFF7589F7C94947F6" blockId="7.[189,334,1493,1519]" box="[189,334,1493,1519]" pageId="7" pageNumber="8">
<heading id="D0CC819CFF8FFFDAFF7589F7C94947F6" bold="true" box="[189,334,1493,1519]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B94FEAE2FF8FFFDAFF7589F7C94947F6" bold="true" box="[189,334,1493,1519]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8436F0FF8FFFDAFF228A26C9424407" blockId="7.[189,1399,1540,2027]" box="[234,325,1540,1566]" pageId="7" pageNumber="8">
<emphasis id="B94FEAE2FF8FFFDAFF228A26C9424407" bold="true" box="[234,325,1540,1566]" pageId="7" pageNumber="8">Female</emphasis>
</paragraph>
<paragraph id="8B8436F0FF8FFFDAFF758A05CD7045F2" blockId="7.[189,1399,1540,2027]" pageId="7" pageNumber="8">
Body cylindrical, tapering slightly along pharyngeal region towards truncated anterior end, tapering more rapidly along tail towards tail tip; straight or ventrally curved upon fixation. Cuticle finely transversely striated along entire body, starting from base of outer labial sensilla and ending near tail tip; striations less than 
<quantity id="4CC39B15FF8FFFDAFEE58AB0C96844B5" box="[301,367,1682,1708]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="1.0">1 µm</quantity>
; lateral alae absent; any other 
<typeStatus id="54808852FF8FFFDAFD118AB0CB1044B5" box="[729,791,1682,1708]" pageId="7" pageNumber="8">types</typeStatus>
of cuticular ornamentation absent. Body pores and epidermal glands absent. Somatic sensilla distinct along anteriormost part of pharyngeal region (cervical setae, see below) and on tail, indistinct or absent along rest of body. Labial region truncated, lips indistinct (fused). Six inner labial sensilla small papilliform, located on anterior surface of lips. Six outer labial sensilla small papilliform, located on outer periphery of labial region, 1.5–5.0 µm from anterior end (
<quantity id="4CC39B15FF8FFFDAFF0E8B61C9024544" box="[198,261,1859,1886]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="5.0">5 µm</quantity>
in specimen with everted stoma). Four cephalic sensilla small papilliform, located short distance behind lip region, 
<quantity id="4CC39B15FF8FFFDAFE5F8B44C9F44598" box="[407,499,1894,1921]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="7" pageNumber="8" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 µm</quantity>
from anterior end, at level with dorsal tooth tip (in relaxed specimens, with inverted stoma, see below). Four pairs of cervical sensilla present, papilliform in shape, arranged in one dorsosublateral, one subdorsal and two subventral pairs at level of stoma base. Amphid stirrup-shaped with transverse slit-like opening 
<quantity id="4CC39B15FF8FFFDAFD838BF3CAAD45F2" box="[587,682,2001,2027]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="8" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 µm</quantity>
wide, located at level with middle of stoma. Oral opening
</paragraph>
<caption id="DF446678FF80FFD5FF758B7CCA3845FB" httpUri="https://zenodo.org/record/2649837/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1886,1912]" targetBox="[358,1224,285,1830]" targetPageId="8">
<paragraph id="8B8436F0FF80FFD5FF758B7CCA3845FB" blockId="8.[189,1399,1886,2018]" pageId="8" pageNumber="9">
<emphasis id="B94FEAE2FF80FFD5FF758B7CC9004561" bold="true" box="[189,263,1886,1912]" pageId="8" pageNumber="9">Fig. 2</emphasis>
. 
<taxonomicName id="4C3B4D73FF80FFD5FED18B7CCAE44561" authority="Holovachov" authorityName="Holovachov" authorityYear="2019" box="[281,739,1886,1912]" class="Adenophorea" family="Campydoridae" genus="Campydoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="8" pageNumber="9" phylum="Nematoda" rank="species" species="manautei" status="gen. et sp. nov.">
<emphasis id="B94FEAE2FF80FFD5FED18B7CCA4E4561" box="[281,585,1886,1912]" italics="true" pageId="8" pageNumber="9">Campydoroides manautei</emphasis>
Holovachov
</taxonomicName>
<taxonomicNameLabel id="A27C5799FF80FFD5FD278B7DCBAE4560" box="[751,937,1887,1913]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="B94FEAE2FF80FFD5FC7C8B7CCBEB4561" bold="true" box="[948,1004,1886,1913]" pageId="8" pageNumber="9">A–B</emphasis>
. Paratype female (SMNH Type- 9167). 
<emphasis id="B94FEAE2FF80FFD5FEDC8BA3C94D4585" bold="true" box="[276,330,1921,1948]" pageId="8" pageNumber="9">C–F</emphasis>
. Holotype (MNHN-BN511). 
<emphasis id="B94FEAE2FF80FFD5FD768BA3CAD44582" bold="true" box="[702,723,1921,1947]" pageId="8" pageNumber="9">A</emphasis>
. Female pharyngeal region, median section. 
<emphasis id="B94FEAE2FF80FFD5FB328BA0CD094585" bold="true" box="[1274,1294,1922,1948]" pageId="8" pageNumber="9">B</emphasis>
. Female anterior end, median section. 
<emphasis id="B94FEAE2FF80FFD5FDD28B86CA5445A6" bold="true" box="[538,595,1956,1983]" pageId="8" pageNumber="9">C–D</emphasis>
. Female anterior end, surface view. 
<emphasis id="B94FEAE2FF80FFD5FBC88B87CC1345A6" bold="true" box="[1024,1044,1957,1983]" pageId="8" pageNumber="9">E</emphasis>
. Female reproductive system. 
<emphasis id="B94FEAE2FF80FFD5FF758BEAC8C845FB" bold="true" box="[189,207,1992,2018]" pageId="8" pageNumber="9">F</emphasis>
. Female tail. Scale bar: 20 µm.
</paragraph>
</caption>
<caption id="DF446678FF81FFD4FF758BA3CBC745FB" httpUri="https://zenodo.org/record/2649839/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1921,1947]" targetBox="[357,1245,287,1863]" targetPageId="9">
<paragraph id="8B8436F0FF81FFD4FF758BA3CBC745FB" blockId="9.[189,1406,1921,2018]" pageId="9" pageNumber="10">
<emphasis id="B94FEAE2FF81FFD4FF758BA3C90F4582" bold="true" box="[189,264,1921,1947]" pageId="9" pageNumber="10">Fig. 3</emphasis>
. 
<taxonomicName id="4C3B4D73FF81FFD4FED28BA3CAE14582" authority="Holovachov" authorityName="Holovachov" authorityYear="2019" box="[282,742,1921,1947]" class="Adenophorea" family="Campydoridae" genus="Campydoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="9" pageNumber="10" phylum="Nematoda" rank="species" species="manautei" status="gen. et sp. nov.">
<emphasis id="B94FEAE2FF81FFD4FED28BA3CA4C4582" box="[282,587,1921,1947]" italics="true" pageId="9" pageNumber="10">Campydoroides manautei</emphasis>
Holovachov
</taxonomicName>
<taxonomicNameLabel id="A27C5799FF81FFD4FD3A8BA3CBB64582" box="[754,945,1921,1947]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, non-type female from New Zealand (SMNH-175283). 
<emphasis id="B94FEAE2FF81FFD4FE5E8B86C9AC45A7" bold="true" box="[406,427,1956,1982]" pageId="9" pageNumber="10">A</emphasis>
. Female pharyngeal region, median section. 
<emphasis id="B94FEAE2FF81FFD4FC768B86CBD545A7" bold="true" box="[958,978,1956,1982]" pageId="9" pageNumber="10">B</emphasis>
. Female anterior end, surface view. 
<emphasis id="B94FEAE2FF81FFD4FF758BE5C8D545F8" bold="true" box="[189,210,1991,2017]" pageId="9" pageNumber="10">C</emphasis>
. Female reproductive system. 
<emphasis id="B94FEAE2FF81FFD4FDF28BEACA4845FB" bold="true" box="[570,591,1992,2018]" pageId="9" pageNumber="10">D</emphasis>
. Female tail. Scale bar: 20 µm.
</paragraph>
</caption>
<caption id="DF446678FF82FFD7FF758BA3CA7845FB" httpUri="https://zenodo.org/record/2649841/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1921,1947]" targetBox="[215,1372,319,1860]" targetPageId="10">
<paragraph id="8B8436F0FF82FFD7FF758BA3CA7845FB" blockId="10.[189,1399,1921,2018]" pageId="10" pageNumber="11">
<emphasis id="B94FEAE2FF82FFD7FF758BA3C90F4582" bold="true" box="[189,264,1921,1947]" pageId="10" pageNumber="11">Fig. 4</emphasis>
. 
<taxonomicName id="4C3B4D73FF82FFD7FED28BA3CAE14582" authority="Holovachov" authorityName="Holovachov" authorityYear="2019" box="[282,742,1921,1947]" class="Adenophorea" family="Campydoridae" genus="Campydoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="11" phylum="Nematoda" rank="species" species="manautei" status="gen. et sp. nov.">
<emphasis id="B94FEAE2FF82FFD7FED28BA3CA4C4582" box="[282,587,1921,1947]" italics="true" pageId="10" pageNumber="11">Campydoroides manautei</emphasis>
Holovachov
</taxonomicName>
<taxonomicNameLabel id="A27C5799FF82FFD7FD3A8BA3CBB64582" box="[754,945,1921,1947]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, non-type female from New Zealand (SMNH-175283). 
<emphasis id="B94FEAE2FF82FFD7FE518B86C9A945A7" bold="true" box="[409,430,1956,1982]" pageId="10" pageNumber="11">A</emphasis>
. Anterior end, median section. 
<emphasis id="B94FEAE2FF82FFD7FCE78B86CB4445A7" bold="true" box="[815,835,1956,1982]" pageId="10" pageNumber="11">B</emphasis>
. Anterior end, surface view. 
<emphasis id="B94FEAE2FF82FFD7FB6D8B86CCBD45A7" bold="true" box="[1189,1210,1956,1982]" pageId="10" pageNumber="11">C</emphasis>
. Vulval region. 
<emphasis id="B94FEAE2FF82FFD7FF758BEAC8D545FB" bold="true" box="[189,210,1992,2018]" pageId="10" pageNumber="11">D</emphasis>
. Base of pharynx. Scale bar: 20 µm.
</paragraph>
</caption>
<caption id="DF446678FF83FFD6FF758D2ECACB4350" ID-Table-UUID="DF446678FF83FFD6FF758D2ECACB4350" httpUri="http://table.plazi.org/id/DF446678FF83FFD6FF758D2ECACB4350" pageId="11" pageNumber="12" targetBox="[197,1386,375,1288]" targetIsTable="true" targetPageId="11">
<paragraph id="8B8436F0FF83FFD6FF758D2ECACB4350" blockId="11.[189,1399,267,329]" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF758D2EC910433C" bold="true" box="[189,279,267,294]" pageId="11" pageNumber="12">Table 3</emphasis>
. Morphometrics of 
<taxonomicName id="4C3B4D73FF83FFD6FE368D29CBB8433C" authority="Holovachov" authorityName="Holovachov" authorityYear="2019" box="[510,959,267,293]" class="Adenophorea" family="Campydoridae" genus="Campydoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="11" pageNumber="12" phylum="Nematoda" rank="species" species="manautei" status="gen. et sp. nov.">
<emphasis id="B94FEAE2FF83FFD6FE368D29CB2D433C" box="[510,810,267,293]" italics="true" pageId="11" pageNumber="12">Campydoroides manautei</emphasis>
Holovachov
</taxonomicName>
<taxonomicNameLabel id="A27C5799FF83FFD6FC0D8D2ECC75433F" box="[965,1138,268,294]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(all measurements are in µm, except for the ratios 
<emphasis id="B94FEAE2FF83FFD6FDCB8D0DCA154351" box="[515,530,303,328]" italics="true" pageId="11" pageNumber="12">a</emphasis>
, 
<emphasis id="B94FEAE2FF83FFD6FDE88D0DCA284350" box="[544,559,303,329]" italics="true" pageId="11" pageNumber="12">b</emphasis>
, 
<emphasis id="B94FEAE2FF83FFD6FDF58D0DCA4D4351" box="[573,586,303,328]" italics="true" pageId="11" pageNumber="12">c</emphasis>
, 
<emphasis id="B94FEAE2FF83FFD6FD918D0DCA774350" box="[601,624,303,329]" italics="true" pageId="11" pageNumber="12">c’</emphasis>
and 
<emphasis id="B94FEAE2FF83FFD6FD618D0DCABC4351" box="[681,699,303,328]" italics="true" pageId="11" pageNumber="12">V</emphasis>
).
</paragraph>
</caption>
<paragraph id="8B8436F0FF83FFD6FF0D8DABCD084711" pageId="11" pageNumber="12">
<table id="F93BC450FF830022FF0D8D55CD6D4711" box="[197,1386,375,1288]" gridcols="4" gridrows="25" pageId="11" pageNumber="12">
<tr id="350B34B2FF830022FF0D8D55CD6D43AD" box="[197,1386,375,436]" gridrow="0" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8D55CA1043AD" box="[197,535,375,436]" gridcol="0" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF0D8DABC94143BA" bold="true" box="[197,326,393,419]" pageId="11" pageNumber="12">Character</emphasis>
</th>
<th id="76DA5DCEFF830022FDF68D55CB0C43AD" box="[574,779,375,436]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FDF68DABCB0C43BA" bold="true" box="[574,779,393,419]" pageId="11" pageNumber="12">Holotype female</emphasis>
</th>
<th id="76DA5DCEFF830022FCFB8D55CC4043AD" box="[819,1095,375,436]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FCFB8D55CC1143AD" bold="true" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FCFB8D55CC404388" bold="true" box="[819,1095,375,401]" pageId="11" pageNumber="12">New Caledonia (n = 7,</emphasis>
incl. holotype)
</emphasis>
</th>
<th id="76DA5DCEFF830022FBA38D55CD6D43AD" box="[1131,1386,375,436]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">New Zealand (n = 2)</th>
</tr>
<tr id="350B34B2FF830022FF0D8D9CCD6D43C0" box="[197,1386,446,473]" gridrow="1" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8D9CCA1043C0" box="[197,535,446,473]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">Body length</th>
<td id="76DA5DCEFF830022FDF68D9CCB0C43C0" box="[574,779,446,473]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">609</td>
<td id="76DA5DCEFF830022FCFB8D9CCC4043C0" box="[819,1095,446,473]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">537 ± 45 (492–609)</td>
<td id="76DA5DCEFF830022FBA38D9CCD6D43C0" box="[1131,1386,446,473]" gridcol="3" gridrow="1" pageId="11" pageNumber="12">786; 817</td>
</tr>
<tr id="350B34B2FF830022FF0D8DC3CD6D43E5" box="[197,1386,481,508]" gridrow="2" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8DC3CA1043E5" box="[197,535,481,508]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">Body diameter</th>
<td id="76DA5DCEFF830022FDF68DC3CB0C43E5" box="[574,779,481,508]" gridcol="1" gridrow="2" pageId="11" pageNumber="12">26</td>
<td id="76DA5DCEFF830022FCFB8DC3CC4043E5" box="[819,1095,481,508]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">24 ± 3 (19–28)</td>
<td id="76DA5DCEFF830022FBA38DC3CD6D43E5" box="[1131,1386,481,508]" gridcol="3" gridrow="2" pageId="11" pageNumber="12">25; 28</td>
</tr>
<tr id="350B34B2FF830022FF0D8E27CD6D4039" box="[197,1386,517,544]" gridrow="3" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8E27CA104039" box="[197,535,517,544]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Pharynx length</th>
<td id="76DA5DCEFF830022FDF68E27CB0C4039" box="[574,779,517,544]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">110</td>
<td id="76DA5DCEFF830022FCFB8E27CC404039" box="[819,1095,517,544]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">94 ± 9 (86–110)</td>
<td id="76DA5DCEFF830022FBA38E27CD6D4039" box="[1131,1386,517,544]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">140; 146</td>
</tr>
<tr id="350B34B2FF830022FF0D8E0ACD6D405A" box="[197,1386,552,579]" gridrow="4" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8E0ACA10405A" box="[197,535,552,579]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Tail length</th>
<td id="76DA5DCEFF830022FDF68E0ACB0C405A" box="[574,779,552,579]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">128</td>
<td id="76DA5DCEFF830022FCFB8E0ACC40405A" box="[819,1095,552,579]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">115 ± 9 (104–128)</td>
<td id="76DA5DCEFF830022FBA38E0ACD6D405A" box="[1131,1386,552,579]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">173; 178</td>
</tr>
<tr id="350B34B2FF830022FF0D8E69CD6D407F" box="[197,1386,587,614]" gridrow="5" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8E69CA10407F" box="[197,535,587,614]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">Anal body diameter</th>
<td id="76DA5DCEFF830022FDF68E69CB0C407F" box="[574,779,587,614]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">16</td>
<td id="76DA5DCEFF830022FCFB8E69CC40407F" box="[819,1095,587,614]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">16 ± 1 (15–17)</td>
<td id="76DA5DCEFF830022FBA38E69CD6D407F" box="[1131,1386,587,614]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">18; 19</td>
</tr>
<tr id="350B34B2FF830022FF0D8E4DCD6D4093" box="[197,1386,623,650]" gridrow="6" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8E4DCA104093" box="[197,535,623,650]" gridcol="0" gridrow="6" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF0D8E52C8D34090" box="[197,212,624,649]" italics="true" pageId="11" pageNumber="12">a</emphasis>
</th>
<td id="76DA5DCEFF830022FDF68E4DCB0C4093" box="[574,779,623,650]" gridcol="1" gridrow="6" pageId="11" pageNumber="12">23.9</td>
<td id="76DA5DCEFF830022FCFB8E4DCC404093" box="[819,1095,623,650]" gridcol="2" gridrow="6" pageId="11" pageNumber="12">22.7 ± 2 (19.5–25.5)</td>
<td id="76DA5DCEFF830022FBA38E4DCD6D4093" box="[1131,1386,623,650]" gridcol="3" gridrow="6" pageId="11" pageNumber="12">31.6; 29.6</td>
</tr>
<tr id="350B34B2FF830022FF0D8EB0CD6D40B7" box="[197,1386,658,686]" gridrow="7" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8EB0CA1040B7" box="[197,535,658,686]" gridcol="0" gridrow="7" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF0D8EB1C8D340B4" box="[197,212,659,685]" italics="true" pageId="11" pageNumber="12">b</emphasis>
</th>
<td id="76DA5DCEFF830022FDF68EB0CB0C40B7" box="[574,779,658,686]" gridcol="1" gridrow="7" pageId="11" pageNumber="12">5.6</td>
<td id="76DA5DCEFF830022FCFB8EB0CC4040B7" box="[819,1095,658,686]" gridcol="2" gridrow="7" pageId="11" pageNumber="12">5.7 ± 0.4 (4.9–6.1)</td>
<td id="76DA5DCEFF830022FBA38EB0CD6D40B7" box="[1131,1386,658,686]" gridcol="3" gridrow="7" pageId="11" pageNumber="12">5.6; 5.6</td>
</tr>
<tr id="350B34B2FF830022FF0D8E94CD6D40C8" box="[197,1386,694,721]" gridrow="8" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8E94CA1040C8" box="[197,535,694,721]" gridcol="0" gridrow="8" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF0D8E95C8D540C9" box="[197,210,695,720]" italics="true" pageId="11" pageNumber="12">c</emphasis>
</th>
<td id="76DA5DCEFF830022FDF68E94CB0C40C8" box="[574,779,694,721]" gridcol="1" gridrow="8" pageId="11" pageNumber="12">4.8</td>
<td id="76DA5DCEFF830022FCFB8E94CC4040C8" box="[819,1095,694,721]" gridcol="2" gridrow="8" pageId="11" pageNumber="12">4.7 ± 0.2 (4.3–5.0)</td>
<td id="76DA5DCEFF830022FBA38E94CD6D40C8" box="[1131,1386,694,721]" gridcol="3" gridrow="8" pageId="11" pageNumber="12">4.6; 4.6</td>
</tr>
<tr id="350B34B2FF830022FF0D8EF8CD6D40EC" box="[197,1386,730,757]" gridrow="9" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8EF8CA1040EC" box="[197,535,730,757]" gridcol="0" gridrow="9" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF0D8EF8C8DB40ED" box="[197,220,730,756]" italics="true" pageId="11" pageNumber="12">c’</emphasis>
</th>
<td id="76DA5DCEFF830022FDF68EF8CB0C40EC" box="[574,779,730,757]" gridcol="1" gridrow="9" pageId="11" pageNumber="12">8.0</td>
<td id="76DA5DCEFF830022FCFB8EF8CC4040EC" box="[819,1095,730,757]" gridcol="2" gridrow="9" pageId="11" pageNumber="12">7.1 ± 0.8 (6.0–8.0)</td>
<td id="76DA5DCEFF830022FBA38EF8CD6D40EC" box="[1131,1386,730,757]" gridcol="3" gridrow="9" pageId="11" pageNumber="12">9.6; 9.2</td>
</tr>
<tr id="350B34B2FF830022FF0D8EDFCD6D4101" box="[197,1386,765,792]" gridrow="10" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8EDFCA104101" box="[197,535,765,792]" gridcol="0" gridrow="10" pageId="11" pageNumber="12">
<emphasis id="B94FEAE2FF83FFD6FF0D8EDCC8D0410E" box="[197,215,766,791]" italics="true" pageId="11" pageNumber="12">V</emphasis>
(%)
</th>
<td id="76DA5DCEFF830022FDF68EDFCB0C4101" box="[574,779,765,792]" gridcol="1" gridrow="10" pageId="11" pageNumber="12">40.5</td>
<td id="76DA5DCEFF830022FCFB8EDFCC404101" box="[819,1095,765,792]" gridcol="2" gridrow="10" pageId="11" pageNumber="12">41.4 ± 1.1 (40.5–43.4)</td>
<td id="76DA5DCEFF830022FBA38EDFCD6D4101" box="[1131,1386,765,792]" gridcol="3" gridrow="10" pageId="11" pageNumber="12">42.1; 42.6</td>
</tr>
<tr id="350B34B2FF830022FF0D8F03CD6D4122" box="[197,1386,801,827]" gridrow="11" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8F03CA104122" box="[197,535,801,827]" gridcol="0" gridrow="11" pageId="11" pageNumber="12">Labial region diameter</th>
<td id="76DA5DCEFF830022FDF68F03CB0C4122" box="[574,779,801,827]" gridcol="1" gridrow="11" pageId="11" pageNumber="12">12</td>
<td id="76DA5DCEFF830022FCFB8F03CC404122" box="[819,1095,801,827]" gridcol="2" gridrow="11" pageId="11" pageNumber="12">12 (12–14)</td>
<td id="76DA5DCEFF830022FBA38F03CD6D4122" box="[1131,1386,801,827]" gridcol="3" gridrow="11" pageId="11" pageNumber="12">14; 14</td>
</tr>
<tr id="350B34B2FF830022FF0D8F66CD6D4146" box="[197,1386,836,863]" gridrow="12" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8F66CA104146" box="[197,535,836,863]" gridcol="0" gridrow="12" pageId="11" pageNumber="12">Amphid from anterior end</th>
<td id="76DA5DCEFF830022FDF68F66CB0C4146" box="[574,779,836,863]" gridcol="1" gridrow="12" pageId="11" pageNumber="12">7</td>
<td id="76DA5DCEFF830022FCFB8F66CC404146" box="[819,1095,836,863]" gridcol="2" gridrow="12" pageId="11" pageNumber="12">8 (7–9)</td>
<td id="76DA5DCEFF830022FBA38F66CD6D4146" box="[1131,1386,836,863]" gridcol="3" gridrow="12" pageId="11" pageNumber="12">9; 8</td>
</tr>
<tr id="350B34B2FF830022FF0D8F45CD6D419B" box="[197,1386,871,898]" gridrow="13" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8F45CA10419B" box="[197,535,871,898]" gridcol="0" gridrow="13" pageId="11" pageNumber="12">Stoma length</th>
<td id="76DA5DCEFF830022FDF68F45CB0C419B" box="[574,779,871,898]" gridcol="1" gridrow="13" pageId="11" pageNumber="12">17</td>
<td id="76DA5DCEFF830022FCFB8F45CC40419B" box="[819,1095,871,898]" gridcol="2" gridrow="13" pageId="11" pageNumber="12">16 (13–17)</td>
<td id="76DA5DCEFF830022FBA38F45CD6D419B" box="[1131,1386,871,898]" gridcol="3" gridrow="13" pageId="11" pageNumber="12">19; 17</td>
</tr>
<tr id="350B34B2FF830022FF0D8FA9CD6D41BF" box="[197,1386,907,934]" gridrow="14" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8FA9CA1041BF" box="[197,535,907,934]" gridcol="0" gridrow="14" pageId="11" pageNumber="12">Stoma width</th>
<td id="76DA5DCEFF830022FDF68FA9CB0C41BF" box="[574,779,907,934]" gridcol="1" gridrow="14" pageId="11" pageNumber="12">6</td>
<td id="76DA5DCEFF830022FCFB8FA9CC4041BF" box="[819,1095,907,934]" gridcol="2" gridrow="14" pageId="11" pageNumber="12">6 (5–6)</td>
<td id="76DA5DCEFF830022FBA38FA9CD6D41BF" box="[1131,1386,907,934]" gridcol="3" gridrow="14" pageId="11" pageNumber="12">6; 6</td>
</tr>
<tr id="350B34B2FF830022FF0D8F8CCD6D41D0" box="[197,1386,942,969]" gridrow="15" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8F8CCA1041D0" box="[197,535,942,969]" gridcol="0" gridrow="15" pageId="11" pageNumber="12">Nerve ring from anterior end</th>
<td id="76DA5DCEFF830022FDF68F8CCB0C41D0" box="[574,779,942,969]" gridcol="1" gridrow="15" pageId="11" pageNumber="12">59</td>
<td id="76DA5DCEFF830022FCFB8F8CCC4041D0" box="[819,1095,942,969]" gridcol="2" gridrow="15" pageId="11" pageNumber="12">47 ± 7 (37–59)</td>
<td id="76DA5DCEFF830022FBA38F8CCD6D41D0" box="[1131,1386,942,969]" gridcol="3" gridrow="15" pageId="11" pageNumber="12">75; 76</td>
</tr>
<tr id="350B34B2FF830022FF0D8FF0CD6D41F5" box="[197,1386,978,1004]" gridrow="16" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8FF0CA1041F5" box="[197,535,978,1004]" gridcol="0" gridrow="16" pageId="11" pageNumber="12">Basal bulb length</th>
<td id="76DA5DCEFF830022FDF68FF0CB0C41F5" box="[574,779,978,1004]" gridcol="1" gridrow="16" pageId="11" pageNumber="12">20</td>
<td id="76DA5DCEFF830022FCFB8FF0CC4041F5" box="[819,1095,978,1004]" gridcol="2" gridrow="16" pageId="11" pageNumber="12">21 ± 2 (17–23)</td>
<td id="76DA5DCEFF830022FBA38FF0CD6D41F5" box="[1131,1386,978,1004]" gridcol="3" gridrow="16" pageId="11" pageNumber="12">26; 23</td>
</tr>
<tr id="350B34B2FF830022FF0D8FD7CD6D4609" box="[197,1386,1013,1040]" gridrow="17" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8FD7CA104609" box="[197,535,1013,1040]" gridcol="0" gridrow="17" pageId="11" pageNumber="12">Basal bulb width</th>
<td id="76DA5DCEFF830022FDF68FD7CB0C4609" box="[574,779,1013,1040]" gridcol="1" gridrow="17" pageId="11" pageNumber="12">19</td>
<td id="76DA5DCEFF830022FCFB8FD7CC404609" box="[819,1095,1013,1040]" gridcol="2" gridrow="17" pageId="11" pageNumber="12">17 ± 1 (16–19)</td>
<td id="76DA5DCEFF830022FBA38FD7CD6D4609" box="[1131,1386,1013,1040]" gridcol="3" gridrow="17" pageId="11" pageNumber="12">20; 21</td>
</tr>
<tr id="350B34B2FF830022FF0D883BCD6D462D" box="[197,1386,1049,1076]" gridrow="18" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D883BCA10462D" box="[197,535,1049,1076]" gridcol="0" gridrow="18" pageId="11" pageNumber="12">Cardia length</th>
<td id="76DA5DCEFF830022FDF6883BCB0C462D" box="[574,779,1049,1076]" gridcol="1" gridrow="18" pageId="11" pageNumber="12">14</td>
<td id="76DA5DCEFF830022FCFB883BCC40462D" box="[819,1095,1049,1076]" gridcol="2" gridrow="18" pageId="11" pageNumber="12">9 ± 3 (6–14)</td>
<td id="76DA5DCEFF830022FBA3883BCD6D462D" box="[1131,1386,1049,1076]" gridcol="3" gridrow="18" pageId="11" pageNumber="12">14; 15</td>
</tr>
<tr id="350B34B2FF830022FF0D881ECD6D464E" box="[197,1386,1084,1111]" gridrow="19" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D881ECA10464E" box="[197,535,1084,1111]" gridcol="0" gridrow="19" pageId="11" pageNumber="12">Cardia width</th>
<td id="76DA5DCEFF830022FDF6881ECB0C464E" box="[574,779,1084,1111]" gridcol="1" gridrow="19" pageId="11" pageNumber="12">10</td>
<td id="76DA5DCEFF830022FCFB881ECC40464E" box="[819,1095,1084,1111]" gridcol="2" gridrow="19" pageId="11" pageNumber="12">11 ± 1 (10–12)</td>
<td id="76DA5DCEFF830022FBA3881ECD6D464E" box="[1131,1386,1084,1111]" gridcol="3" gridrow="19" pageId="11" pageNumber="12">12; 12</td>
</tr>
<tr id="350B34B2FF830022FF0D8842CD6D4662" box="[197,1386,1120,1147]" gridrow="20" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8842CA104662" box="[197,535,1120,1147]" gridcol="0" gridrow="20" pageId="11" pageNumber="12">Vagina length</th>
<td id="76DA5DCEFF830022FDF68842CB0C4662" box="[574,779,1120,1147]" gridcol="1" gridrow="20" pageId="11" pageNumber="12">7</td>
<td id="76DA5DCEFF830022FCFB8842CC404662" box="[819,1095,1120,1147]" gridcol="2" gridrow="20" pageId="11" pageNumber="12">6 (6–7)</td>
<td id="76DA5DCEFF830022FBA38842CD6D4662" box="[1131,1386,1120,1147]" gridcol="3" gridrow="20" pageId="11" pageNumber="12">7; 7</td>
</tr>
<tr id="350B34B2FF830022FF0D88A1CD6D4687" box="[197,1386,1155,1182]" gridrow="21" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D88A1CA104687" box="[197,535,1155,1182]" gridcol="0" gridrow="21" pageId="11" pageNumber="12">Rectum length</th>
<td id="76DA5DCEFF830022FDF688A1CB0C4687" box="[574,779,1155,1182]" gridcol="1" gridrow="21" pageId="11" pageNumber="12">15</td>
<td id="76DA5DCEFF830022FCFB88A1CC404687" box="[819,1095,1155,1182]" gridcol="2" gridrow="21" pageId="11" pageNumber="12">14 ± 1 (12–16)</td>
<td id="76DA5DCEFF830022FBA388A1CD6D4687" box="[1131,1386,1155,1182]" gridcol="3" gridrow="21" pageId="11" pageNumber="12">15; 15</td>
</tr>
<tr id="350B34B2FF830022FF0D8885CD6D46D8" box="[197,1386,1191,1217]" gridrow="22" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D8885CA1046D8" box="[197,535,1191,1217]" gridcol="0" gridrow="22" pageId="11" pageNumber="12">Cardia width</th>
<td id="76DA5DCEFF830022FDF68885CB0C46D8" box="[574,779,1191,1217]" gridcol="1" gridrow="22" pageId="11" pageNumber="12">10</td>
<td id="76DA5DCEFF830022FCFB8885CC4046D8" box="[819,1095,1191,1217]" gridcol="2" gridrow="22" pageId="11" pageNumber="12">11 ± 1 (10–12)</td>
<td id="76DA5DCEFF830022FBA38885CD6D46D8" box="[1131,1386,1191,1217]" gridcol="3" gridrow="22" pageId="11" pageNumber="12">12; 12</td>
</tr>
<tr id="350B34B2FF830022FF0D88E8CD6D46FC" box="[197,1386,1226,1253]" gridrow="23" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D88E8CA1046FC" box="[197,535,1226,1253]" gridcol="0" gridrow="23" pageId="11" pageNumber="12">Vagina length</th>
<td id="76DA5DCEFF830022FDF688E8CB0C46FC" box="[574,779,1226,1253]" gridcol="1" gridrow="23" pageId="11" pageNumber="12">7</td>
<td id="76DA5DCEFF830022FCFB88E8CC4046FC" box="[819,1095,1226,1253]" gridcol="2" gridrow="23" pageId="11" pageNumber="12">6 (6–7)</td>
<td id="76DA5DCEFF830022FBA388E8CD6D46FC" box="[1131,1386,1226,1253]" gridcol="3" gridrow="23" pageId="11" pageNumber="12">7; 7</td>
</tr>
<tr id="350B34B2FF830022FF0D88CFCD6D4711" box="[197,1386,1261,1288]" gridrow="24" pageId="11" pageNumber="12">
<th id="76DA5DCEFF830022FF0D88CFCA104711" box="[197,535,1261,1288]" gridcol="0" gridrow="24" pageId="11" pageNumber="12">Rectum length</th>
<td id="76DA5DCEFF830022FDF688CFCB0C4711" box="[574,779,1261,1288]" gridcol="1" gridrow="24" pageId="11" pageNumber="12">15</td>
<td id="76DA5DCEFF830022FCFB88CFCC404711" box="[819,1095,1261,1288]" gridcol="2" gridrow="24" pageId="11" pageNumber="12">14 ± 1 (12–16)</td>
<td id="76DA5DCEFF830022FBA388CFCD6D4711" box="[1131,1386,1261,1288]" gridcol="3" gridrow="24" pageId="11" pageNumber="12">15; 15</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8436F0FF83FFD1FF758966CC0B438A" blockId="11.[189,1399,1348,2027]" lastBlockId="12.[189,1398,267,403]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
apical, surrounded by a fringe of tiny projections, visible only when stoma is everted to the outside (
<figureCitation id="13002A75FF83FFD6FF0E894AC922479A" box="[198,293,1384,1411]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[358,1224,285,1830]" captionTargetId="figure@8.[288,1299,265,1845]" captionTargetPageId="8" captionText="Fig. 2. Campydoroides manautei Holovachov gen. et sp. nov. A–B. Paratype female (SMNH Type- 9167). C–F. Holotype (MNHN-BN511). A. Female pharyngeal region, median section. B. Female anterior end, median section. C–D. Female anterior end, surface view. E. Female reproductive system. F. Female tail. Scale bar: 20 µm." httpUri="https://zenodo.org/record/2649837/files/figure.png" pageId="11" pageNumber="12">Fig. 2A</figureCitation>
). Stoma conoid; cheilostom short and weakly cuticularised; gymnostom as long and broad as cheilostom, weakly cuticularised; stegostom with strongly cuticularised walls, distinctly subdivided into three sections by breaks in cuticularised walls, with large dorsal tooth projecting anteriad from its middle section. Tooth heavily cuticularised, 
<quantity id="4CC39B15FF83FFD6FDA789F7CACC47E9" box="[623,715,1493,1520]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="11" pageNumber="12" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 µm</quantity>
long, hollow inside, protrusible, its tip is 
<quantity id="4CC39B15FF83FFD6FB7489F7CD3247E9" box="[1212,1333,1493,1520]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="11" pageNumber="12" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 µm</quantity>
from stoma base. Pharyngeal tubes absent. Pharynx cylindrical anteriorly, with distinct basal bulb; anterior cylindrical part of pharynx with evenly distributed myofilaments, basal bulb strongly muscularised; pharyngeal lumen uniform in thickness along entire pharynx length; valves absent. Pharyngeal glands and their orifices indistinct. Cardia large, ovoid, partially embedded into intestinal tissue. Nerve ring encircling pharynx at 43–54% of its length. Secretory-excretory system indistinct/absent. Female reproductive system didelphic, amphidelphic, ovary branches reflexed antidromously, usually located on same side of intestine; symmetrical in younger females, asymmetrical in older females when large developing ovocyte or intrauterine egg is present in one of genital tubes. Anterior genital tube 
<quantity id="4CC39B15FF83FFD6FB378AD5CD70450B" box="[1279,1399,1783,1810]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="9.1" metricValueMin="2.1" pageId="11" pageNumber="12" unit="mm" value="56.0" valueMax="91.0" valueMin="21.0">21–91 µm</quantity>
long, equal to 4–13% of total body length, situated to either right (n = 4) or left (n = 3) of intestine; posterior genital tube 
<quantity id="4CC39B15FF83FFD6FE748B1DCA334543" box="[444,564,1855,1882]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="5.5" metricValueMin="2.3" pageId="11" pageNumber="12" unit="mm" value="39.0" valueMax="55.0" valueMin="23.0">23–55 µm</quantity>
long, equal to 4–9% of total body length, situated to either right (n = 3) or left (n = 4) of intestine. Vulva a transverse slit, located anterior to midbody. Vagina straight, equal to 0.2–0.3 body diameters in length, with developed sphincter muscle surrounding its proximal part; 
<emphasis id="B94FEAE2FF83FFD6FA8A8BABC99F45DF" italics="true" pageId="11" pageNumber="12">pars refringens vaginae</emphasis>
or epiptygmata absent. Spermatheca absent. Intrauterine egg measuring 64–77 × 
<quantity id="4CC39B15FF83FFD6FA828B8FC90C45F2" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="11" pageNumber="12" unit="mm" value="19.0" valueMax="21.0" valueMin="17.0">17– 21 µm</quantity>
. Rectum short, equal to 0.8–1.2 anal body diameters in length. Tail elongate, distinctly subdivided into conoid proximal part 
<quantity id="4CC39B15FF84FFD1FE3C8D29CA69433F" box="[500,622,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="4.2" metricValueMin="2.6" pageId="12" pageNumber="13" unit="mm" value="34.0" valueMax="42.0" valueMin="26.0">26–42 µm</quantity>
in length, and subcylindrical distal part 
<quantity id="4CC39B15FF84FFD1FB878D29CCD0433F" box="[1103,1239,267,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.06" metricValueMax="1.3599999999999999" metricValueMin="0.76" pageId="12" pageNumber="13" unit="mm" value="106.0" valueMax="136.0" valueMin="76.0">76–136 µm</quantity>
long. Caudal setae present, one subdorsal pair along proximal conoid part of tail, one subventral pair at level with junction between proximal and distal parts of tail, and single lateral setae on left side of distal cylindrical part of tail. Caudal glands present, opening via small terminal spinneret.
</paragraph>
<paragraph id="8B8436F0FF84FFD1FF228D9EC92D43CF" blockId="12.[189,826,444,507]" box="[234,298,444,470]" pageId="12" pageNumber="13">
<emphasis id="B94FEAE2FF84FFD1FF228D9EC92D43CF" bold="true" box="[234,298,444,470]" pageId="12" pageNumber="13">Male</emphasis>
</paragraph>
<paragraph id="8B8436F0FF84FFD1FF758DC3CB3D43E3" blockId="12.[189,826,444,507]" box="[189,826,480,507]" pageId="12" pageNumber="13">Not found. The species is presumed to be thelytokous.</paragraph>
</subSubSection>
<subSubSection id="C321657BFF84FFD1FF758E01CAA947DA" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8B8436F0FF84FFD1FF758E01C9284024" blockId="12.[189,303,547,573]" box="[189,303,547,573]" pageId="12" pageNumber="13">
<heading id="D0CC819CFF84FFD1FF758E01C9284024" bold="true" box="[189,303,547,573]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B94FEAE2FF84FFD1FF758E01C9284024" bold="true" box="[189,303,547,573]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8436F0FF84FFD1FF758E71CD6F40C3" blockId="12.[189,1399,595,731]" pageId="12" pageNumber="13">
The two female specimens collected in 
<collectingCountry id="F32C7660FF84FFD1FD6F8E71CB4F4074" box="[679,840,595,621]" name="New Zealand" pageId="12" pageNumber="13">New Zealand</collectingCountry>
match very well the morphology of the 
<typeStatus id="54808852FF84FFD1FA8C8E71CD704074" box="[1348,1399,595,621]" pageId="12" pageNumber="13">type</typeStatus>
specimens from 
<collectingCountry id="F32C7660FF84FFD1FE478E5ACA574088" box="[399,592,631,658]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
, except for the larger body size, higher values for some body measurements and longer tail (including higher 
<emphasis id="B94FEAE2FF84FFD1FD218EBECB0740AF" box="[745,768,668,694]" italics="true" pageId="12" pageNumber="13">c’</emphasis>
-ratio). Since no qualitative morphological differences were found between the two populations, they both are here considered to belong to the same species.
</paragraph>
<paragraph id="8B8436F0FF84FFD1FF758F25CB0D413C" blockId="12.[189,778,774,805]" box="[189,778,774,805]" pageId="12" pageNumber="13">
<heading id="D0CC819CFF84FFD1FF758F25CB0D413C" bold="true" box="[189,778,774,805]" fontSize="13" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B94FEAE2FF84FFD1FF758F25CB0D413C" bold="true" box="[189,778,774,805]" italics="true" pageId="12" pageNumber="13">
Notes on the systematics of 
<taxonomicName id="4C3B4D73FF84FFD1FD8F8F24CB0D413C" authorityName="Jairajpuri" authorityYear="1983" box="[583,778,774,805]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="subOrder" subOrder="Campydorina">Campydorina</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8436F0FF84FFD1FF758F1FCAA947DA" blockId="12.[189,1399,829,1475]" pageId="12" pageNumber="13">
The unique and poorly understood morphology of the genus 
<taxonomicName id="4C3B4D73FF84FFD1FC438F1FCC11414E" authorityName="Holovachov" authorityYear="2019" box="[907,1046,829,855]" class="Adenophorea" family="Campgdoridae" genus="Campydora" kingdom="Animalia" order="Dorylaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FC438F1FCC11414E" box="[907,1046,829,855]" italics="true" pageId="12" pageNumber="13">Campydora</emphasis>
</taxonomicName>
hindered understanding of its relationships with other nematodes and resulted in the taxonomic inflation from the genus to the suborder level (
<bibRefCitation id="EFAA4B01FF84FFD1FECF8FA4C9C741B9" author="Jairajpuri M. S." box="[263,448,902,928]" journalOrPublisher="Nematologia Mediterranea" pageId="12" pageNumber="13" pagination="33 - 42" part="11" refId="ref7198" refString="Jairajpuri M. S. 1983. Observations on Campydora (Nematoda: Dorylaimida). Nematologia Mediterranea 11: 33 - 42." title="Observations on Campydora (Nematoda: Dorylaimida)" type="journal article" year="1983">Jairajpuri 1983</bibRefCitation>
). This decision was followed in subsequent publications, although the suborder was transferred from the order 
<taxonomicName id="4C3B4D73FF84FFD1FDF68F88CAD541DD" authorityName="Pearse" authorityYear="1942" box="[574,722,938,964]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="order">Dorylaimida</taxonomicName>
to the order 
<taxonomicName id="4C3B4D73FF84FFD1FCB18F88CBE441DD" authorityName="Filipjev" authorityYear="1929" box="[889,995,938,964]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="order">Enoplida</taxonomicName>
based on the morphological and molecular evidence, and its relationships within the order 
<taxonomicName id="4C3B4D73FF84FFD1FCAA8FEDCBCB41F0" authorityName="Filipjev" authorityYear="1929" box="[866,972,975,1001]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="order">Enoplida</taxonomicName>
remained unresolved (
<bibRefCitation id="EFAA4B01FF84FFD1FB1E8FEDCD7541F0" author="Siddiqi M. R." box="[1238,1394,975,1001]" journalOrPublisher="Pakistan Journal of Nematology" pageId="12" pageNumber="13" pagination="79 - 110" part="1" refId="ref7911" refString="Siddiqi M. R. 1983. Phylogenetic relationships of the soil nematode orders Dorylaimida, Mononchida, Triplonchida and Alaimida, with a revised classification of the subclass Enoplia. Pakistan Journal of Nematology 1: 79 - 110." title="Phylogenetic relationships of the soil nematode orders Dorylaimida, Mononchida, Triplonchida and Alaimida, with a revised classification of the subclass Enoplia" type="journal article" year="1983">Siddiqi 1983</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FF758FD1C99D4614" author="Winiszewska G." box="[189,410,1011,1037]" journalOrPublisher="Annales Zoologici" pageId="12" pageNumber="13" pagination="391 - 394" part="51" refId="ref8572" refString="Winiszewska G. 2001. Some observations on Campydora Cobb, 1920 (Nematoda), with descriptions of male Campydora demonstrans Cobb, 1920. Annales Zoologici 51: 391 - 394." title="Some observations on Campydora Cobb, 1920 (Nematoda), with descriptions of male Campydora demonstrans Cobb" type="journal article" year="2001">Winiszewska 2001</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FE6A8FD1CA6A4614" author="Mullin P. G. &amp; Harris T. &amp; Powers T. O." box="[418,621,1011,1037]" journalOrPublisher="Nematology" pageId="12" pageNumber="13" pagination="699 - 711" part="5" refId="ref7618" refString="Mullin P. G., Harris T. S &amp; Powers T. O. 2003. Systematic status of Campydora Cobb, 1920 (Nematoda, Campydorina). Nematology 5: 699 - 711. https: // doi. org / 10.1163 / 156854103322746878" title="Systematic status of Campydora Cobb, 1920 (Nematoda, Campydorina)" type="journal article" year="2003">
Mullin 
<emphasis id="B94FEAE2FF84FFD1FE3D8FD6CA2B4614" box="[501,556,1011,1037]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2003
</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FDBD8FD1CB2A4614" author="Zhao Z. &amp; Li D. &amp; Buckley T. R." box="[629,813,1011,1037]" journalOrPublisher="Zootaxa" pageId="12" pageNumber="13" pagination="41 - 57" part="3208" refId="ref8632" refString="Zhao Z., Li D. &amp; Buckley T. R. 2012. Analysis of primary structure loops from Hairpins 35 and 48 of the Nematoda SSU rRNA gene provides further evidence that the genera Tripylina Brzeski, 1963, Trischistoma Cobb, 1913 and Rhabdolaimus de Man, 1880 are members of Enoplida. Zootaxa 3208: 41 - 57. https: // doi. org / 10.11646 / zootaxa. 3208.1.3" title="Analysis of primary structure loops from Hairpins 35 and 48 of the Nematoda SSU rRNA gene provides further evidence that the genera Tripylina Brzeski, 1963, Trischistoma Cobb, 1913 and Rhabdolaimus de Man, 1880 are members of Enoplida" type="journal article" year="2012">
Zhao 
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</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FCFD8FD1CBF74614" author="Smol N. &amp; Muthumbi A. &amp; Sharma J." box="[821,1008,1011,1037]" editor="Schmidt-Rhaesa A." journalOrPublisher="De Gryuter." pageId="12" pageNumber="13" pagination="193 - 250" part="2" refId="ref7994" refString="Smol N., Muthumbi A. &amp; Sharma J. 2014. Order Enoplida. In: Schmidt-Rhaesa A. (ed.) Handbook of Zoology. Gastrotricha, Cycloneuralia and Gnathifera 2: 193 - 250. De Gryuter. https: // doi. org / 10.1515 / 9783110274257.193" title="Order Enoplida" type="journal article" volumeTitle="Handbook of Zoology. Gastrotricha, Cycloneuralia and Gnathifera" year="2014">
Smol 
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</bibRefCitation>
).Subsequent phylogenetic studies using 18S rDNA sequence suggested close affinities between the genera 
<taxonomicName id="4C3B4D73FF84FFD1FBF9883ACCBB462B" authorityName="Holovachov" authorityYear="2019" box="[1073,1212,1048,1074]" class="Adenophorea" family="Campgdoridae" genus="Campydora" kingdom="Animalia" order="Dorylaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FBF9883ACCBB462B" box="[1073,1212,1048,1074]" italics="true" pageId="12" pageNumber="13">Campydora</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D73FF84FFD1FB05883AC962464F" authority="de Man, 1880" authorityName="de Man" authorityYear="1880" class="Adenophorea" family="Rhabdolaimidae" genus="Rhabdolaimus" kingdom="Animalia" order="Araeolaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FB05883ACD71462B" box="[1229,1398,1048,1074]" italics="true" pageId="12" pageNumber="13">Rhabdolaimus</emphasis>
de Man, 1880
</taxonomicName>
and 
<taxonomicName id="4C3B4D73FF84FFD1FE6B881ECB06464F" authority="de Man, 1888" authorityName="de Man" authorityYear="1888" box="[419,769,1084,1111]" class="Adenophorea" family="Rhabdolaimidae" genus="Syringolaimus" kingdom="Animalia" order="Araeolaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FE6B881ECA4B464F" box="[419,588,1084,1110]" italics="true" pageId="12" pageNumber="13">Syringolaimus</emphasis>
de Man, 1888
</taxonomicName>
, placing these three genera as a monophyletic clade within the unresolved order 
<taxonomicName id="4C3B4D73FF84FFD1FDDF8843CA864662" authorityName="Filipjev" authorityYear="1929" box="[535,641,1121,1147]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="order">Enoplida</taxonomicName>
(
<bibRefCitation id="EFAA4B01FF84FFD1FD5C8843CB7C4662" author="Meldal B. H. M. &amp; Debenham N. J. &amp; De Ley P. &amp; De Ley I. T. &amp; Vanfleteren J. R. &amp; Vierstraete A. R. &amp; Bert W. &amp; Borgonie G. &amp; Moens T. &amp; Tyler P. A. &amp; Austen M. C. &amp; Blaxter M. L. &amp; Rogers A. D. &amp; Lambshead P. J. G." box="[660,891,1121,1147]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="12" pageNumber="13" pagination="622 - 636" part="42" refId="ref7497" refString="Meldal B. H. M., Debenham N. J., De Ley P., De Ley I. T., Vanfleteren J. R., Vierstraete A. R., Bert W., Borgonie G., Moens T., Tyler P. A., Austen M. C., Blaxter M. L., Rogers A. D. &amp; Lambshead P. J. G. 2007. An improved molecular phylogeny of the Nematoda with special emphasis on marine taxa. Molecular Phylogenetics and Evolution 42: 622 - 636. https: // doi. org / 10.1016 / j. ympev. 2006.08.025" title="An improved molecular phylogeny of the Nematoda with special emphasis on marine taxa" type="journal article" year="2007">
Meldal 
<emphasis id="B94FEAE2FF84FFD1FD3C8840CB354662" box="[756,818,1121,1147]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2007
</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FC438843CCA24662" author="van Megen H. &amp; van den Elsen S. &amp; Holterman M. &amp; Karssen G. &amp; Mooyman P. &amp; Bongers T. &amp; Holovachov O. &amp; Bakker J. &amp; Helder J." box="[907,1189,1121,1147]" journalOrPublisher="Nematology" pageId="12" pageNumber="13" pagination="927 - 950" part="11" refId="ref7421" refString="van Megen H., van den Elsen S., Holterman M., Karssen G., Mooyman P., Bongers T., Holovachov O., Bakker J. &amp; Helder J. 2009. A phylogenetic tree of nematodes based on about 1,200 full length small subunit ribosomal DNA sequences. Nematology 11: 927 - 950. https: // doi. org / 10.1163 / 156854109 X 456862" title="A phylogenetic tree of nematodes based on about 1,200 full length small subunit ribosomal DNA sequences" type="journal article" year="2009">
van Megen 
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Bik 
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; 
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Shokoohi 
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</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FE7688A7CA5B46B9" author="Smythe A. B." box="[446,604,1157,1184]" journalOrPublisher="Integrative and Comparative Biology" pageId="12" pageNumber="13" pagination="228 - 240" part="55" refId="ref8047" refString="Smythe A. B. 2015. Evolution of feeding structures in the marine nematode order Enoplida. Integrative and Comparative Biology 55: 228 - 240. https: // doi. org / 10.1093 / icb / icv 043" title="Evolution of feeding structures in the marine nematode order Enoplida" type="journal article" year="2015">Smythe 2015</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FDA088A7CB3F4686" author="Leduc D. &amp; Zhao Z. Q. &amp; Verdon V. &amp; Xu Y." box="[616,824,1157,1183]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="12" pageNumber="13" pagination="29 - 36" part="122" refId="ref7284" refString="Leduc D., Zhao Z. Q., Verdon V. &amp; Xu Y. 2018. Phylogenetic position of the enigmatic deep-sea nematode order Rhaptothyreida: a molecular analysis. Molecular Phylogenetics and Evolution 122: 29 - 36. https: // doi. org / 10.1016 / j. ympev. 2018.01.018" title="Phylogenetic position of the enigmatic deep-sea nematode order Rhaptothyreida: a molecular analysis" type="journal article" year="2018">
Leduc 
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</bibRefCitation>
). This grouping can be further substantiated by at least one morphological character: the presence of a well-developed basal bulb, not found in any other 
<taxonomicName id="4C3B4D73FF84FFD1FF7588ECC92046F1" authorityName="Filipjev" authorityYear="1929" box="[189,295,1230,1256]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="order">Enoplida</taxonomicName>
(the genera 
<taxonomicName id="4C3B4D73FF84FFD1FE7588ECCB3646F1" authority="de Man, 1922" authorityName="de Man" authorityYear="1922" box="[445,817,1230,1257]" class="Adenophorea" family="Eurystominidae" genus="Polygastrophora" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FE7588ECCA8446F1" box="[445,643,1230,1256]" italics="true" pageId="12" pageNumber="13">Polygastrophora</emphasis>
de Man, 1922
</taxonomicName>
and 
<taxonomicName id="4C3B4D73FF84FFD1FCA488ECCC6446F1" authority="Cobb, 1920" authorityName="Cobb" authorityYear="1920" box="[876,1123,1230,1256]" class="Adenophorea" family="Enchelidiidae" genus="Belbolla" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FCA488ECCBC846F1" box="[876,975,1230,1256]" italics="true" pageId="12" pageNumber="13">Belbolla</emphasis>
Cobb, 1920
</taxonomicName>
have the posterior part of the pharynx modified into a series of bulb-like swellings that are of independent origin). Placement of the genus 
<taxonomicName id="4C3B4D73FF84FFD1FE918935CA054728" authorityName="de Man" authorityYear="1888" box="[345,514,1303,1329]" class="Adenophorea" family="Rhabdolaimidae" genus="Syringolaimus" kingdom="Animalia" order="Araeolaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FE918935CA054728" box="[345,514,1303,1329]" italics="true" pageId="12" pageNumber="13">Syringolaimus</emphasis>
</taxonomicName>
in the suborder 
<taxonomicName id="4C3B4D73FF84FFD1FD008935CB6F4728" authorityName="Jairajpuri" authorityYear="1983" box="[712,872,1303,1329]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="subOrder" subOrder="Campydorina">Campydorina</taxonomicName>
is further supported by the monorchic male reproductive system similar to the reproductive system found in the species from the genera 
<taxonomicName id="4C3B4D73FF84FFD1FB24891ECD70474F" authorityName="Holovachov" authorityYear="2019" box="[1260,1399,1340,1366]" class="Adenophorea" family="Campgdoridae" genus="Campydora" kingdom="Animalia" order="Dorylaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FB24891ECD70474F" box="[1260,1399,1340,1366]" italics="true" pageId="12" pageNumber="13">Campydora</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D73FF84FFD1FF388943C9554763" authorityName="Hoeppli &amp; Chu" authorityYear="1934" box="[240,338,1377,1402]" class="Adenophorea" family="Rhabdolaimidae" genus="Rogerus" kingdom="Animalia" order="Araeolaimida" pageId="12" pageNumber="13" phylum="Nematoda" rank="genus">
<emphasis id="B94FEAE2FF84FFD1FF388943C9554763" box="[240,338,1377,1402]" italics="true" pageId="12" pageNumber="13">Rogerus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAA4B01FF84FFD1FEA98942CA674763" author="Hoeppli R. &amp; Chu H. J." box="[353,608,1376,1402]" journalOrPublisher="The Hong Kong Naturalist Supplement" pageId="12" pageNumber="13" pagination="15 - 29" part="1" refId="ref7090" refString="Hoeppli R. &amp; Chu H. J. 1932. Free-living nematodes from hot springs in China and Formosa. The Hong Kong Naturalist Supplement 1: 15 - 29." title="Free-living nematodes from hot springs in China and Formosa" type="journal article" year="1932">Hoeppli &amp; Chu 1932</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FDA58942CB494763" author="Winiszewska G." box="[621,846,1376,1402]" journalOrPublisher="Annales Zoologici" pageId="12" pageNumber="13" pagination="391 - 394" part="51" refId="ref8572" refString="Winiszewska G. 2001. Some observations on Campydora Cobb, 1920 (Nematoda), with descriptions of male Campydora demonstrans Cobb, 1920. Annales Zoologici 51: 391 - 394." title="Some observations on Campydora Cobb, 1920 (Nematoda), with descriptions of male Campydora demonstrans Cobb" type="journal article" year="2001">Winiszewska 2001</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FC928942CC274763" author="Tchesunov A. V." box="[858,1056,1376,1402]" journalOrPublisher="Zootaxa" pageId="12" pageNumber="13" pagination="478 - 500" part="4306" refId="ref8280" refString="Tchesunov A. V. 2017. Free-living nematodes of the genus Syringolaimus de Man, 1888 (Nematoda, Enoplida, Ironidae) from the Angola Basin, South-East Atlantic Abyss. Zootaxa 4306: 478 - 500. https: // doi. org / 10.11646 / zootaxa. 4306.4.2" title="Free-living nematodes of the genus Syringolaimus de Man, 1888 (Nematoda, Enoplida, Ironidae) from the Angola Basin, South-East Atlantic Abyss" type="journal article" year="2017">Tchesunov 2017</bibRefCitation>
), while it is diorchic in other genera of 
<taxonomicName id="4C3B4D73FF84FFD1FEF989A4CA2A4786" authority="Siddiqi, 1983" authorityName="Siddiqi" authorityYear="1983" box="[305,557,1413,1439]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="subOrder" subOrder="Ironina">
<emphasis id="B94FEAE2FF84FFD1FEF989A4C98F4786" box="[305,392,1414,1439]" italics="true" pageId="12" pageNumber="13">Ironina</emphasis>
<bibRefCitation id="EFAA4B01FF84FFD1FE4789A7CA2A4786" author="Siddiqi M. R." box="[399,557,1413,1439]" journalOrPublisher="Pakistan Journal of Nematology" pageId="12" pageNumber="13" pagination="79 - 110" part="1" refId="ref7911" refString="Siddiqi M. R. 1983. Phylogenetic relationships of the soil nematode orders Dorylaimida, Mononchida, Triplonchida and Alaimida, with a revised classification of the subclass Enoplia. Pakistan Journal of Nematology 1: 79 - 110." title="Phylogenetic relationships of the soil nematode orders Dorylaimida, Mononchida, Triplonchida and Alaimida, with a revised classification of the subclass Enoplia" type="journal article" year="1983">Siddiqi, 1983</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFAA4B01FF84FFD1FDF489A7CB334786" author="Shokoohi E. &amp; Mehrabi-Nasab A. &amp; Abolafia J." box="[572,820,1413,1439]" journalOrPublisher="Nematology" pageId="12" pageNumber="13" pagination="835 - 849" part="15" refId="ref7858" refString="Shokoohi E., Mehrabi-Nasab A. &amp; Abolafia J. 2013. Studies on Ironus Bastian, 1865 (Enoplida: Ironidae) from Iran with a comment on its phylogenetic position. Nematology 15: 835 - 849. https: // doi. org / 10.1163 / 15685411 - 00002722" title="Studies on Ironus Bastian, 1865 (Enoplida: Ironidae) from Iran with a comment on its phylogenetic position" type="journal article" year="2013">
Shokoohi 
<emphasis id="B94FEAE2FF84FFD1FD7D89A4CAE84786" box="[693,751,1413,1439]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2013
</bibRefCitation>
; 
<bibRefCitation id="EFAA4B01FF84FFD1FC8889A7CC1B47B9" author="Chen Y. Z. &amp; Guo Y. Q." box="[832,1052,1413,1440]" journalOrPublisher="Zootaxa" pageId="12" pageNumber="13" pagination="151 - 175" part="4018" refId="ref6402" refString="Chen Y. Z. &amp; Guo Y. Q. 2015. Three new and two known free-living marine nematode species of the family Ironidae from the East China Sea. Zootaxa 4018: 151 - 175. https: // doi. org / 10.11646 / zootaxa. 4018.2.1" title="Three new and two known free-living marine nematode species of the family Ironidae from the East China Sea" type="journal article" year="2015">Chen &amp; Guo 2015</bibRefCitation>
). The revised classification of the suborder 
<taxonomicName id="4C3B4D73FF84FFD1FE9E898BC9F147DA" authorityName="Jairajpuri" authorityYear="1983" box="[342,502,1449,1475]" class="Adenophorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="12" pageNumber="13" phylum="Nematoda" rank="subOrder" subOrder="Campydorina">Campydorina</taxonomicName>
is given below.
</paragraph>
</subSubSection>
</treatment>
</document>