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<document ID-DOI="http://dx.doi.org/10.3897/zse.96.48041" ID-GBIF-Dataset="c4e83292-cb81-45a8-86ee-67e9dfe303f7" ID-GBIF-Taxon="167407582" ID-Pensoft-Pub="1860-0743-2-537" ID-Pensoft-UUID="E9BEBC6C66D650EF89DADA67C9017BCD" ID-ZooBank="E1A270E298E04F349BFBDCC49CCFAE47" ModsDocID="1860-0743-2-537" checkinTime="1599019535613" checkinUser="pensoft" docAuthor="Lee, Bee Yan, Richer De Forges, Bertrand &amp; Ng, Peter K. L." docDate="2020" docId="43DBA09EA87956C1BFC226B428EF0FB4" docLanguage="en" docName="ZoosystEvol 96(2): 537-569" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.48041" docTitle="Scyramathia hertwigi Doflein, in Chun 1900" docType="treatment" docVersion="3" id="E9BEBC6C66D650EF89DADA67C9017BCD" lastPageNumber="537" masterDocId="E9BEBC6C66D650EF89DADA67C9017BCD" masterDocTitle="Revision of the deep-water spider crab genus, Scyramathia A. Milne-Edwards, 1880, with the description of a new species from the Mediterranean and notes on Rochinia A. Milne-Edwards, 1875, and Anamathia Smith, 1885 (Crustacea, Decapoda, Brachyura, Epialtidae)" masterLastPageNumber="569" masterPageNumber="537" pageId="0" pageNumber="537" updateTime="1643450818311" updateUser="ExternalLinkService">
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<mods:title>Revision of the deep-water spider crab genus, Scyramathia A. Milne-Edwards, 1880, with the description of a new species from the Mediterranean and notes on Rochinia A. Milne-Edwards, 1875, and Anamathia Smith, 1885 (Crustacea, Decapoda, Brachyura, Epialtidae)</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lee, Bee Yan</mods:namePart>
<mods:affiliation>Tropical Marine Science Institute, National University of Singapore, 18 Kent Ridge Road, Singapore 119227, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5806-6999</mods:nameIdentifier>
<mods:nameIdentifier type="email">beeyan06@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Richer De Forges, Bertrand</mods:namePart>
<mods:affiliation>c / o Museum National d'Histoire Naturelle, Institut de Systematique, Evolution, Biodiversite, ISYEB - UMR 7205 - CNRS, MNHN, UPMC, EPHE, Departement Systematique et Evolution, 57 rue Cuvier, CP 26, F- 75005, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
</mods:name>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>96</mods:number>
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<mods:number>2</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zse.96.48041</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.48041</mods:identifier>
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<treatment ID-GBIF-Taxon="167407582" LSID="urn:lsid:plazi:treatment:43DBA09EA87956C1BFC226B428EF0FB4" httpUri="http://treatment.plazi.org/id/43DBA09EA87956C1BFC226B428EF0FB4" lastPageNumber="537" pageId="0" pageNumber="537">
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<taxonomicName LSID="43DBA09E-A879-56C1-BFC2-26B428EF0FB4" authority="Doflein, in Chun, 1900" authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="hertwigi">Scyramathia hertwigi Doflein, in Chun, 1900</taxonomicName>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Scyramathia hertwigi Doflein, in Chun 1900, lectotype male (33.7 x 27.0 mm) (ZMB 13699), South Atlantic Ocean. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure10" httpUri="https://binary.pensoft.net/fig/447927" pageId="0" pageNumber="537">Figs 10A-C</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dorsal view of P 5. A, B. Scyramathia carpenteri (Norman, in Thomson 1873); A. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; B. ovigerous female (35.8 x 29.0 mm) (NHM 1978.99), Atlantic Ocean; C, D. S. hertwigi Doflein, in Chun 1900; C. male (34.6 x 27.3 mm) (NHM 1924.7.19.1 - 3), South Africa; D. female (24.5 x 18.4 mm) (SAM-A 47872), South Africa; E, S. umbonata (Stimpson, 1871), female (36.7 x 27.9 mm) (NSMT-Cr 7178), off Suriname; F-J. S. tenuipes sp. nov.; F. holotype male (33.9 x 26.3 mm) (ICMD 288), Mediterranean Sea; G. paratype male (29.7 x 22.0 mm) (ICMD 000707), Mediterranean Sea; H. paratype male (32.0 x 25.4 mm) (ZRC 2019.1636), Mediterranean Sea,; I. paratype ovigerous female (31.2 x 24.6 mm) (ICMD 000708), Mediterranean Sea,; J. paratype ovigerous female (32.2 x 25.8 mm) (ZRC 2019.1635), Mediterranean Sea." figureDoi="10.3897/zse.96.48041.figure14" httpUri="https://binary.pensoft.net/fig/447931" pageId="0" pageNumber="537">, 14C, D</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. A-D. Scyramathia hertwigi Doflein, in Chun 1900, paralectotype male (24.7 x 19.0 mm) (ZMB 13700), Agulhasstrom [= Agulhas current], left G 1; E-H. Scyramathia tenuipes sp. nov., holotype male (33.9 x 26.3 mm) (ICMD 288), Mediterranean Sea, left G 1. A, E. ventral view; B, F. ventral view of distal portion; C, G. dorsal view; D, H. dorsal view of distal portion. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48041.figure15" httpUri="https://binary.pensoft.net/fig/447932" pageId="0" pageNumber="537">, 15A-D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="537" type="reference_group">
<paragraph pageId="0" pageNumber="537">
<taxonomicName authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia Hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="Hertwigi">Scyramathia Hertwigi</taxonomicName>
: Doflein, in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun 1900</bibRefCitation>
: 172 (fig.); Doflein, in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B20" refString="Chun, C, 1902-1903. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographien, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 3 Karten und 482 Abbildungen im Text. Zweite umgearbeitete und stark vermehrte Auflage. Gustav Fischer, Jena, ix + 592 pp. [46 pls., 3 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographien, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 3 Karten und 482 Abbildungen im Text. Zweite umgearbeitete und stark vermehrte Auflage. Gustav Fischer, Jena, ix + 592 pp. [46 pls., 3 maps]" year="1902 - 1903">Chun 1902-1903</bibRefCitation>
: 172 (fig.).
</paragraph>
<paragraph pageId="0" pageNumber="537">
<taxonomicName authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia Hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="Hertwigi">Scyramathia Hertwigi</taxonomicName>
Doflein, 1904: 81-84, pl. 27 fig. 3, pl. 28 fig. 1 (type locality: off Cape Point and on Agulhas Bank).
</paragraph>
<paragraph pageId="0" pageNumber="537">
<taxonomicName authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="hertwigi">Scyramathia hertwigi</taxonomicName>
:
<bibRefCitation author="Stebbing, TRR" journalOrPublisher="Bulletin of Marine Science" pageId="0" pageNumber="537" refId="B119" refString="Stebbing, TRR, 1902. South African Crustacea. Part II. Marine Investigations in South Africa, Cape of Good Hope, Department of Agriculture, Cape Town 2: 1-92. [pls. [Crustacea] 5-16]" title="South African Crustacea. Part II. Marine Investigations in South Africa, Cape of Good Hope, Department of Agriculture, Cape Town 2: 1 - 92. [pls. [Crustacea] 5 - 16]" year="1902">Stebbing 1902</bibRefCitation>
: 7, pl. 6;
<bibRefCitation author="Stebbing, TRR" journalOrPublisher="Bulletin of Marine Science" pageId="0" pageNumber="537" refId="B120" refString="Stebbing, TRR, 1905. South African Crustacea. Part III. Marine Investigations in South Africa, Cape of Good Hope, Department of Agriculture, Cape Town 4: 21-123. [pls. [Crustacea] 17-21, 21 bis, 22-26]" title="South African Crustacea. Part III. Marine Investigations in South Africa, Cape of Good Hope, Department of Agriculture, Cape Town 4: 21 - 123. [pls. [Crustacea] 17 - 21, 21 bis, 22 - 26]" year="1905">Stebbing 1905</bibRefCitation>
: 25;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.15558" author="Stebbing, TRR" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="537" pagination="281 - 593" refId="B121" refString="Stebbing, TRR, 1910. General Catalogue of South African Crustacea (Part V. of S. A. Crustacea, for the Marine Investigations in South Africa). Annals of the South African Museum 6 (4): 281 - 593, DOI: https://doi.org/10.5962/bhl.part.15558" title="General Catalogue of South African Crustacea (Part V. of S. A. Crustacea, for the Marine Investigations in South Africa)." url="https://doi.org/10.5962/bhl.part.15558" volume="6" year="1910">Stebbing 1910</bibRefCitation>
: 289;
<bibRefCitation author="Barnard, KH" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="537" pagination="1 - 837" refId="B8" refString="Barnard, KH, 1950. Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum 38: 1 - 837" title="Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps)." volume="38" year="1950">Barnard 1950</bibRefCitation>
: 50, 51, fig. 11b, c;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.1.10" author="Low, MEY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="113 - 123" refId="B59" refString="Low, MEY, Lee, BY, Ng, PKL, 2017. The Brachyura (Crustacea: Decapoda) of the Deutschen Tiefsee-Expedition auf dem Dampfer 'Valdivia' 1898-1899: dates of publications and authorship. Zootaxa 4341 (1): 113 - 123, DOI: https://doi.org/10.11646/zootaxa.4341.1.10" title="The Brachyura (Crustacea: Decapoda) of the Deutschen Tiefsee-Expedition auf dem Dampfer ' Valdivia' 1898 - 1899: dates of publications and authorship." url="https://doi.org/10.11646/zootaxa.4341.1.10" volume="4341" year="2017">Low et al. 2017</bibRefCitation>
: 116, fig. 2A, tables 2, 3.
</paragraph>
<paragraph pageId="0" pageNumber="537">
<taxonomicName baseAuthorityName="C. W. Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Rochinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rochinia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">Rochinia carpenteri</taxonomicName>
:
<bibRefCitation author="Monod, T" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire (IFAN)" pageId="0" pageNumber="537" pagination="1 - 674" refId="B82" refString="Monod, T, 1956. Hippidea et Brachyura ouest-africains. Memoires de l'Institut Francais d'Afrique Noire (IFAN) 45: 1 - 674" title="Hippidea et Brachyura ouest-africains." volume="45" year="1956">Monod 1956</bibRefCitation>
: 32 (list), 515. [not
<taxonomicName class="Gymnolaemata" family="Epialtidae" genus="Amathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Bryozoa" rank="species" species="carpenteri">Amathia carpenteri</taxonomicName>
Norman, in
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16330" author="Thomson, CW" journalOrPublisher="Deep-Sea Research II" pageId="0" pageNumber="537" refId="B129" refString="Thomson, CW, 1873. The depths of the sea. An account of the general results of the dredging cruises of H.M.SS. &quot;Porcupine&quot; and &quot;Lightning&quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F.R.S., J. Gwyn Jeffreys, F.R.S., and Dr. Wyville Thomson, F.R.S. Macmillan and Co., London: i-xxi + 1-527. [pls. 1-8] https://doi.org/10.5962/bhl.title.16330" title="The depths of the sea. An account of the general results of the dredging cruises of H. M. SS. &quot; Porcupine &quot; and &quot; Lightning &quot; during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London: i-xxi + 1 - 527. [pls. 1 - 8]" url="https://doi.org/10.5962/bhl.title.16330" year="1873">Thomson 1873</bibRefCitation>
].
</paragraph>
<paragraph pageId="0" pageNumber="537">
<taxonomicName baseAuthorityName="Doflein" baseAuthorityYear="1904" class="Malacostraca" family="Epialtidae" genus="Rochinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rochinia hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="hertwigi">Rochinia hertwigi</taxonomicName>
:
<bibRefCitation author="Serene, R" journalOrPublisher="Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand" pageId="0" pageNumber="537" pagination="1 - 186" refId="B113" refString="Serene, R, Lohavanijaya, P, 1973. The Brachyura (Crustacea: Decapoda) collected by the Naga Expedition, including a review of the Homolidae. Naga Report. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand 4 (4): 1 - 186" title="The Brachyura (Crustacea: Decapoda) collected by the Naga Expedition, including a review of the Homolidae. Naga Report." volume="4" year="1973">
<normalizedToken originalValue="Serène">Serene</normalizedToken>
and Lohavanijaya 1973
</bibRefCitation>
: 56 (key);
<bibRefCitation author="MacPherson, E" journalOrPublisher="Resultados Expeditiones Cientificas" pageId="0" pageNumber="537" pagination="3 - 79" refId="B61" refString="MacPherson, E, 1983. Crustaceos Decapodos capturados en las costas de Namibia. Resultados Expeditiones Cientificas 11: 3 - 79" title="Crustaceos Decapodos capturados en las costas de Namibia." volume="11" year="1983">MacPherson 1983</bibRefCitation>
: 32-35, figs 19, 20a-c;
<bibRefCitation author="Griffin, DJG" journalOrPublisher="Bergens Museums Aarbok Naturvidenskabelig raekke" pageId="0" pageNumber="537" refId="B44" refString="Griffin, DJG, Tranter, HA, 1986. The DecapodaBrachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie 39: [i]-[vii], 1-335." title="The DecapodaBrachyura of the Siboga expedition. Part VIII: Majidae. Siboga Expeditie Monografie 39: [i] - [vii], 1 - 335." year="1986">Griffin and Tranter 1986</bibRefCitation>
: 175 (key), 180;
<bibRefCitation DOI="https://doi.org/10.2992/0097-4463(2005)74[151:MFDCBF]2.0.CO;2" author="Casadio, S" journalOrPublisher="Proceedings of the Royal Society of London" pageId="0" pageNumber="537" refId="B16" refString="Casadio, S, Feldmann, RM, Parras, A, Schweitzer, CE, 2005. Miocene fossil Decapoda (Crustacea: Brachyura) from Patagonia, Argentina, and their paleoecological setting. Annals of Carnegie Museum 74(3): 151-188. https://doi.org/10.2992/0097-4463(2005)74[151:MFDCBF]2.0.CO;2" title="Miocene fossil Decapoda (Crustacea: Brachyura) from Patagonia, Argentina, and their paleoecological setting. Annals of Carnegie Museum 74 (3): 151 - 188." url="https://doi.org/10.2992/0097-4463(2005)74[151:MFDCBF]2.0.CO;2" year="2005">
<normalizedToken originalValue="Casadío">Casadio</normalizedToken>
et al. 2005
</bibRefCitation>
: 159 (list);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1610.1.5" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="61 - 68" refId="B86" refString="Ng, PKL, Richer de Forges, B, 2007. A new species of deep-water spider crab of the genus Rochinia A. Milne-Edwards, 1875, from Guam (Crustacea: Brachyura: Majidae). Zootaxa 1610: 61 - 68, DOI: https://doi.org/10.11646/zootaxa.1610.1.5" title="A new species of deep-water spider crab of the genus Rochinia A. Milne-Edwards, 1875, from Guam (Crustacea: Brachyura: Majidae)." url="https://doi.org/10.11646/zootaxa.1610.1.5" volume="1610" year="2007">Ng and Richer de Forges 2007</bibRefCitation>
: 62 (list);
<bibRefCitation author="Ng, PKL" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="537" refId="B85" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology 17 (supplement): 1-286." title="Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology 17 (supplement): 1 - 286." year="2008">Ng et al. 2008</bibRefCitation>
: 105 (list);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4418.3.1" author="Tavares, MS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="201 - 227" refId="B126" refString="Tavares, MS, Santana, W, 2018. Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae). Zootaxa 4418 (3): 201 - 227, DOI: https://doi.org/10.11646/zootaxa.4418.3.1" title="Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae)." url="https://doi.org/10.11646/zootaxa.4418.3.1" volume="4418" year="2018">Tavares and Santana 2018</bibRefCitation>
: 223 (list);
<bibRefCitation author="Griffiths, CL" editor="Atkinson, LJ" journalOrPublisher="Malachite Marketing and Media, Pretoria" pageId="0" pageNumber="537" pagination="133 - 226" refId="B45" refString="Griffiths, CL, Landschoff, J, Atkinson, LJ, 2018. Phylum Arthropoda. In: Atkinson, LJ, Sink, KJ, Eds., Field Guide to the Offshore Marine Invertebrates of South Africa. Malachite Marketing and Media, Pretoria: 133 - 226" title="Phylum Arthropoda." volumeTitle="Field Guide to the Offshore Marine Invertebrates of South Africa." year="2018">Griffiths et al. 2018</bibRefCitation>
: 199.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="537" type="materials_examined">
<paragraph pageId="0" pageNumber="537">Materials examined.</paragraph>
<paragraph pageId="0" pageNumber="537">
<emphasis bold="true" italics="true" pageId="0" pageNumber="537">Lectotype</emphasis>
: ♂ (33.7
<normalizedToken originalValue="×">x</normalizedToken>
27.0 mm) (ZMB 13699), Atlantic, South Atlantic Ocean, Agulhasstrom [= Agulhas current], coll. Deutsche Tiefsee Expedition, 2 November 1898.
<emphasis bold="true" italics="true" pageId="0" pageNumber="537">Paralectotypes</emphasis>
: 3 ♂♂ (24.7
<normalizedToken originalValue="×">x</normalizedToken>
19.0 mm, 23.6
<normalizedToken originalValue="×">x</normalizedToken>
17.7 mm, 18.7
<normalizedToken originalValue="×">x</normalizedToken>
13.7 mm), 2 ♀♀ (28.3
<normalizedToken originalValue="×">x</normalizedToken>
22.6 mm, 26.4
<normalizedToken originalValue="×">x</normalizedToken>
20.2 mm) (ZMB 13700), stn 103, Agulhasstrom [= Agulhas current], 500 m, coll. Deutsche Tiefsee Expedition, 2 November 1898; 8 ♂♂ (2 largest: 22.8
<normalizedToken originalValue="×">x</normalizedToken>
17.9 mm, 21.6
<normalizedToken originalValue="×">x</normalizedToken>
16.5 mm; 2 smallest: 16.2
<normalizedToken originalValue="×">x</normalizedToken>
12.3 mm, 16.0
<normalizedToken originalValue="×">x</normalizedToken>
11.9 mm), 2 ovigerous ♀♀ (25.9
<normalizedToken originalValue="×">x</normalizedToken>
20.5 mm, 24.3
<normalizedToken originalValue="×">x</normalizedToken>
19.2 mm), 8 ♀♀ (2 largest: 25.1
<normalizedToken originalValue="×">x</normalizedToken>
19.4 mm, 22.6
<normalizedToken originalValue="×">x</normalizedToken>
17.3 mm; 2 smallest: 20.1
<normalizedToken originalValue="×">x</normalizedToken>
15.2 mm, 14.3
<normalizedToken originalValue="×">x</normalizedToken>
10.3 mm) (SMF 23085), stn 103, Atlantic, South Atlantic, Agulhasstrom [= Agulhas current],
<geoCoordinate degrees="35" direction="south" minutes="10" orientation="latitude" precision="925" value="-35.166668">35°10'S</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="2" orientation="longitude" precision="925" value="23.033333">23°2'E</geoCoordinate>
, 500 m, coll. FS
<normalizedToken originalValue="Valdivia”">'Valdivia&quot;</normalizedToken>
, 2 November 1898.
<emphasis bold="true" italics="true" pageId="0" pageNumber="537">Other material</emphasis>
: South Africa • 1 ♂ (29.2
<normalizedToken originalValue="×">x</normalizedToken>
22.9 mm) (SAM-A39580), west grounds, I + J trawls, coll. UCT Ecological Survey, 22 November 1946; 2 ♂♂ (33.2
<normalizedToken originalValue="×">x</normalizedToken>
27.7 mm, 31.9
<normalizedToken originalValue="×">x</normalizedToken>
27.1 mm) (SAM-A47887), west coast, 333 m, coll. 17 January 2007; 1 ♂ (19.0
<normalizedToken originalValue="×">x</normalizedToken>
14.2 mm), 1 ovigerous ♀ (34.4
<normalizedToken originalValue="×">x</normalizedToken>
27.1 mm) (SAM-A12149), West of Slangkop, Cape Peninsula, 457 m, coll. P Henry, 16 February 1965; 1 ♂ (47.0
<normalizedToken originalValue="×">x</normalizedToken>
38.0 mm) (SAM-A40027), stn A10121-082-035-4197,
<geoCoordinate degrees="34" direction="south" minutes="36.6" orientation="latitude" precision="92" value="-34.61">34°36.6'S</geoCoordinate>
,
<geoCoordinate degrees="25" direction="east" minutes="15.1" orientation="longitude" precision="92" value="25.251667">25°15.1'E</geoCoordinate>
, 480 m, coll. R.S. Africana South Coast Hare Biomass Survey, 31 May 1990; 1 ♂ (23.7
<normalizedToken originalValue="×">x</normalizedToken>
18.5 mm), 1 ovigerous ♀ (36.0
<normalizedToken originalValue="×">x</normalizedToken>
28.5 mm), 1 ♀ (24.5
<normalizedToken originalValue="×">x</normalizedToken>
18.4 mm) (SAM-A47872), stn NAN401 T027,
<geoCoordinate degrees="34" direction="south" minutes="20.2" orientation="latitude" precision="92" value="-34.336666">34°20.2'S</geoCoordinate>
,
<geoCoordinate degrees="18" direction="east" minutes="12.7" orientation="longitude" precision="92" value="18.211666">18°12.7'E</geoCoordinate>
, 290 m, coll. L Atkinson, Nansen West Coast Survey, 15 January 2007; 1 ♂ (40.7
<normalizedToken originalValue="×">x</normalizedToken>
31.1 mm) (AM P34653), off Cape Town,
<geoCoordinate degrees="34" direction="south" minutes="21" orientation="latitude" precision="925" value="-34.35">34°21'S</geoCoordinate>
,
<geoCoordinate degrees="17" direction="east" minutes="57" orientation="longitude" precision="925" value="17.95">17°57'E</geoCoordinate>
, coll. 18 December 1929; 3 ♂♂ (34.6
<normalizedToken originalValue="×">x</normalizedToken>
27.3 mm, 29.3
<normalizedToken originalValue="×">x</normalizedToken>
23.1 mm, 22.0
<normalizedToken originalValue="×">x</normalizedToken>
17.0 mm) (NHM 1924.7.19.1-3), stn 103, Cape, no other data. SOUTH ATLANTIC • 1 ♀ (21.5
<normalizedToken originalValue="×">x</normalizedToken>
18.9 mm) (ZMB 26204), stn 153/1968,
<geoCoordinate degrees="28" direction="south" minutes="22" orientation="latitude" precision="925" value="-28.366667">28°22'S</geoCoordinate>
,
<geoCoordinate degrees="14" direction="east" minutes="24" orientation="longitude" precision="925" value="14.4">14°24'E</geoCoordinate>
, 470-460 m, coll. E Haeckel, 9 May 1968.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="537" type="diagnosis">
<paragraph pageId="0" pageNumber="537">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="537">
Carapace pyriform, covered with short setae. Pseudorostral spine short, diverging. Supraorbital eave with sharp preorbital spine; postorbital lobe fused to hepatic plate on large size specimens. Carapace with plates: 1 hepatic plate pointing upwards, 1 short mesogastric spine above 1 oblong mesogastric plate, 1 large protogastric granule, 1 oblong cardiac plate, 1 large epibranchial plate, 1 mesobranchial plate, 1 posterior blunt spine, 1 lateral branchial plate curved upwards (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Scyramathia hertwigi Doflein, in Chun 1900, lectotype male (33.7 x 27.0 mm) (ZMB 13699), South Atlantic Ocean. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure10" httpUri="https://binary.pensoft.net/fig/447927" pageId="0" pageNumber="537">10A</figureCitation>
). Plates less distinct on juvenile specimens. Antennal flagellum equal to or slightly longer than pseudorostral spines. Basal antennal article with convex outer margin, with blunt distal angle of article. Buccal frame covered by third maxilliped, distal angle of buccal frame raised, distinct. Pterygostomial region plate-like with 3 or 4 granules on outer margin (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Scyramathia hertwigi Doflein, in Chun 1900, lectotype male (33.7 x 27.0 mm) (ZMB 13699), South Atlantic Ocean. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure10" httpUri="https://binary.pensoft.net/fig/447927" pageId="0" pageNumber="537">10B</figureCitation>
). Chelipeds slender, propodus longer than fingers, slightly carinate; carpus with granules on carina margin; merus triangular in cross-section, granulated on edge. Ambulatory legs slender; merus cylindrical; male P2 merus length 0.8-1.2 times carapace length, female P2 merus length 0.4-0.8 times carapace length, male P2 merus length 10.9-14.9 times width, female P2 merus length about 8.5 times width; male P5 merus length 0.4-0.6 times carapace length, female P5 merus length 0.4-0.5 times carapace length, male P5 merus length 4-6.8 times width, female P5 merus length 4.6-5.8 times width (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Scyramathia hertwigi Doflein, in Chun 1900, lectotype male (33.7 x 27.0 mm) (ZMB 13699), South Atlantic Ocean. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure10" httpUri="https://binary.pensoft.net/fig/447927" pageId="0" pageNumber="537">10A</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dorsal view of P 5. A, B. Scyramathia carpenteri (Norman, in Thomson 1873); A. male (41.3 x 34.0 mm) (NHM 1983.449), North Sea; B. ovigerous female (35.8 x 29.0 mm) (NHM 1978.99), Atlantic Ocean; C, D. S. hertwigi Doflein, in Chun 1900; C. male (34.6 x 27.3 mm) (NHM 1924.7.19.1 - 3), South Africa; D. female (24.5 x 18.4 mm) (SAM-A 47872), South Africa; E, S. umbonata (Stimpson, 1871), female (36.7 x 27.9 mm) (NSMT-Cr 7178), off Suriname; F-J. S. tenuipes sp. nov.; F. holotype male (33.9 x 26.3 mm) (ICMD 288), Mediterranean Sea; G. paratype male (29.7 x 22.0 mm) (ICMD 000707), Mediterranean Sea; H. paratype male (32.0 x 25.4 mm) (ZRC 2019.1636), Mediterranean Sea,; I. paratype ovigerous female (31.2 x 24.6 mm) (ICMD 000708), Mediterranean Sea,; J. paratype ovigerous female (32.2 x 25.8 mm) (ZRC 2019.1635), Mediterranean Sea." figureDoi="10.3897/zse.96.48041.figure14" httpUri="https://binary.pensoft.net/fig/447931" pageId="0" pageNumber="537">14C, D</figureCitation>
). Male thoracic sternum wide, concave on sternites 1-4; regions well defined; sternites 3, 4 widest, lateral margins slightly constricted medially. Male pleon with triangular telson; surface of somites smooth (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Scyramathia hertwigi Doflein, in Chun 1900, lectotype male (33.7 x 27.0 mm) (ZMB 13699), South Atlantic Ocean. A. overall dorsal view; B. overall ventral view; C. lateral view of carapace." figureDoi="10.3897/zse.96.48041.figure10" httpUri="https://binary.pensoft.net/fig/447927" pageId="0" pageNumber="537">10B</figureCitation>
). G1 straight with flattened sharp tip (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. A-D. Scyramathia hertwigi Doflein, in Chun 1900, paralectotype male (24.7 x 19.0 mm) (ZMB 13700), Agulhasstrom [= Agulhas current], left G 1; E-H. Scyramathia tenuipes sp. nov., holotype male (33.9 x 26.3 mm) (ICMD 288), Mediterranean Sea, left G 1. A, E. ventral view; B, F. ventral view of distal portion; C, G. dorsal view; D, H. dorsal view of distal portion. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48041.figure15" httpUri="https://binary.pensoft.net/fig/447932" pageId="0" pageNumber="537">15A-D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.48041.figure10" httpUri="https://binary.pensoft.net/fig/447927" pageId="0" pageNumber="537" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="537">
<emphasis bold="true" pageId="0" pageNumber="537">Figure 10.</emphasis>
<taxonomicName authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="hertwigi">
<emphasis italics="true" pageId="0" pageNumber="537">Scyramathia hertwigi</emphasis>
</taxonomicName>
Doflein, in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun 1900</bibRefCitation>
, lectotype male (33.7
<normalizedToken originalValue="×">x</normalizedToken>
27.0 mm) (ZMB 13699), South Atlantic Ocean.
<emphasis bold="true" pageId="0" pageNumber="537">A.</emphasis>
overall dorsal view;
<emphasis bold="true" pageId="0" pageNumber="537">B.</emphasis>
overall ventral view;
<emphasis bold="true" pageId="0" pageNumber="537">C.</emphasis>
lateral view of carapace.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="537" type="remarks">
<paragraph pageId="0" pageNumber="537">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="537">
The author and year of publication for
<taxonomicName authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="hertwigi">
<emphasis italics="true" pageId="0" pageNumber="537">Scyramathia hertwigi</emphasis>
</taxonomicName>
has traditionally been cited as
<bibRefCitation author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein (1904)</bibRefCitation>
.
<bibRefCitation author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein (1904</bibRefCitation>
: 81), however, cited a figure of this species in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun (1900)</bibRefCitation>
. Hence, the authorship and date of publication for
<taxonomicName genus="S." lsidName="S. hertwigi" pageId="0" pageNumber="537" rank="species" species="hertwigi">
<emphasis italics="true" pageId="0" pageNumber="537">S. hertwigi</emphasis>
</taxonomicName>
should be &quot;Doflein in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun 1900</bibRefCitation>
&quot; (see
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.1.10" author="Low, MEY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="113 - 123" refId="B59" refString="Low, MEY, Lee, BY, Ng, PKL, 2017. The Brachyura (Crustacea: Decapoda) of the Deutschen Tiefsee-Expedition auf dem Dampfer 'Valdivia' 1898-1899: dates of publications and authorship. Zootaxa 4341 (1): 113 - 123, DOI: https://doi.org/10.11646/zootaxa.4341.1.10" title="The Brachyura (Crustacea: Decapoda) of the Deutschen Tiefsee-Expedition auf dem Dampfer ' Valdivia' 1898 - 1899: dates of publications and authorship." url="https://doi.org/10.11646/zootaxa.4341.1.10" volume="4341" year="2017">Low et al. 2017</bibRefCitation>
). The figure by Doflein that is in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun (1900)</bibRefCitation>
was stated to be drawn to scale (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4341.1.10" author="Low, MEY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="113 - 123" refId="B59" refString="Low, MEY, Lee, BY, Ng, PKL, 2017. The Brachyura (Crustacea: Decapoda) of the Deutschen Tiefsee-Expedition auf dem Dampfer 'Valdivia' 1898-1899: dates of publications and authorship. Zootaxa 4341 (1): 113 - 123, DOI: https://doi.org/10.11646/zootaxa.4341.1.10" title="The Brachyura (Crustacea: Decapoda) of the Deutschen Tiefsee-Expedition auf dem Dampfer ' Valdivia' 1898 - 1899: dates of publications and authorship." url="https://doi.org/10.11646/zootaxa.4341.1.10" volume="4341" year="2017">Low et al. 2017</bibRefCitation>
).
<bibRefCitation author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein (1904)</bibRefCitation>
, when describing the species, did not specify the number of specimens examined, but mentioned that many male and female specimens were examined. The specimen that was figured by Doflein, in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun (1900)</bibRefCitation>
, is currently deposited in ZMB. The measurement of the male specimen deposited in the Museum
<normalizedToken originalValue="für">fuer</normalizedToken>
Naturkunde, Berlin (ZMB) is very similar to that drawn by Doflein, in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun (1900)</bibRefCitation>
and we are confident they are the same specimen. The location where this specimen was collected matches that listed in
<bibRefCitation author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein (1904)</bibRefCitation>
. Since no type was designated in
<bibRefCitation author="Chun, C" journalOrPublisher="Universitetsforlaget Oslo, Denmark" pageId="0" pageNumber="537" refId="B19" refString="Chun, C, 1900. Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" title="Aus den Tiefen des Weltmeeres. Schilderungen von der Deutschen Tiefsee-Expedition. Mit 6 Chromolithographieen, 8 Heliogravueren, 32 als Tafeln gedruckten Vollbildern, 2 Karten und 390 Abbildungen im Text. Gustav Fischer, Jena, vi + 549 pp. [46 pls., 2 maps]" year="1900">Chun (1900)</bibRefCitation>
or
<bibRefCitation author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein (1904)</bibRefCitation>
, this male specimen (ZMB 13699) is herein designated as the lectotype of
<taxonomicName genus="S." lsidName="S. hertwigi" pageId="0" pageNumber="537" rank="species" species="hertwigi">
<emphasis italics="true" pageId="0" pageNumber="537">S. hertwigi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="537">
The Atlas by
<bibRefCitation author="Doflein, F" journalOrPublisher="Marine Geology" pageId="0" pageNumber="537" refId="B33" refString="Doflein, F, 1904. Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Gustav Fischer, Jena, volume 6: 1-6, i-xiv, 1-314. [figs 1-58]" title="Brachyura. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Gustav Fischer, Jena, volume 6: 1 - 6, i-xiv, 1 - 314. [figs 1 - 58]" year="1904">Doflein (1904</bibRefCitation>
: pl. 27 figs 1-7) has figures that show a series of sizes of the carapace with plates, in particular the horse-shoe shaped epibranchial plate. In this species, the pseudorostral spines are always proportionately shorter than those in
<taxonomicName genus="S." lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
. The differences between
<taxonomicName genus="S." lsidName="S. carpenteri" pageId="0" pageNumber="537" rank="species" species="carpenteri">
<emphasis italics="true" pageId="0" pageNumber="537">S. carpenteri</emphasis>
</taxonomicName>
and
<taxonomicName genus="S." lsidName="S. hertwigi" pageId="0" pageNumber="537" rank="species" species="hertwigi">
<emphasis italics="true" pageId="0" pageNumber="537">S. hertwigi</emphasis>
</taxonomicName>
were discussed by
<bibRefCitation author="MacPherson, E" journalOrPublisher="Resultados Expeditiones Cientificas" pageId="0" pageNumber="537" pagination="3 - 79" refId="B61" refString="MacPherson, E, 1983. Crustaceos Decapodos capturados en las costas de Namibia. Resultados Expeditiones Cientificas 11: 3 - 79" title="Crustaceos Decapodos capturados en las costas de Namibia." volume="11" year="1983">MacPherson (1983</bibRefCitation>
: 35), and we agree with his interpretation. There are also slight variations amongst the specimens from South Atlantic (SMF 23085), which tend to have less distinct plates on the carapace, and slightly shorter and stouter ambulatory legs while a large specimen (AM P34653) from South Africa has distinct plates on carapace, and somewhat longer and slender ambulatory legs. These differences, however, are not substantial and should not be a reason to treat them as different.
</paragraph>
<paragraph pageId="0" pageNumber="537">
The specimens of &quot;
<taxonomicName baseAuthorityName="Norman, in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia carpenteri" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="carpenteri">
<emphasis italics="true" pageId="0" pageNumber="537">Scyramathia carpenteri</emphasis>
</taxonomicName>
&quot; from off Cape Bajador, West Africa (= off Boujdour, Western Sahara) reported by
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4418.3.1" author="Tavares, MS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="537" pagination="201 - 227" refId="B126" refString="Tavares, MS, Santana, W, 2018. Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae). Zootaxa 4418 (3): 201 - 227, DOI: https://doi.org/10.11646/zootaxa.4418.3.1" title="Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae)." url="https://doi.org/10.11646/zootaxa.4418.3.1" volume="4418" year="2018">Tavares and Santana (2018</bibRefCitation>
: 204) may not be that species. This location overlaps with the distribution range of
<taxonomicName genus="S." lsidName="S. tenuipes" pageId="0" pageNumber="537" rank="species" species="tenuipes">
<emphasis italics="true" pageId="0" pageNumber="537">S. tenuipes</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="537">sp. nov.</emphasis>
, and it is possible their specimens belong to this species instead. Their material will need to be re-examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="537" type="distribution">
<paragraph pageId="0" pageNumber="537">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="537">
<taxonomicName authorityName="Doflein, in Chun" authorityYear="1900" class="Malacostraca" family="Epialtidae" genus="Scyramathia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scyramathia hertwigi" order="Decapoda" pageId="0" pageNumber="537" phylum="Arthropoda" rank="species" species="hertwigi">
<emphasis italics="true" pageId="0" pageNumber="537">Scyramathia hertwigi</emphasis>
</taxonomicName>
is known from its type locality, off Cape Point and on Agulhas Bank, South Africa, as well as parts of the south Atlantic along the African coast. It is the only species that enters the southwestern-most edge of the Indian Ocean.
</paragraph>
</subSubSection>
</treatment>
</document>