189 lines
20 KiB
XML
189 lines
20 KiB
XML
<document id="0D60F35059DAEB068A5695E7017267C0" ID-DOI="10.5281/zenodo.191026" ID-GBIF-Dataset="c3a9826c-c202-4c3f-b252-53f18f333335" ID-ISSN="1175-5326" ID-Zenodo-Dep="191026" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461122839876" checkinUser="plazi" docAuthor="Parra, Luis E., Villagrán-Mella, Romina & Marquet, Pablo A." docDate="2009" docId="432A5E1EFFA12D4DFF0AAAF8A4ABF920" docLanguage="en" docName="zt02273p068.pdf" docOrigin="Zootaxa 2273" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hasodima ediliacarmenae Parra, sp. nov." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="BF132666FFA02D4FFF9DAF4AA006FFD6" masterDocTitle="Phylogeny of the genera Euclidiodes and Hasodima (Lepidoptera: Geometridae) and description of two new species from the Fray Jorge relict forest in northern Chile" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698596215224" updateUser="plazi">
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<mods:title id="6DCF412546DCD8DD124B4E9C45ADDE7D">Phylogeny of the genera Euclidiodes and Hasodima (Lepidoptera: Geometridae) and description of two new species from the Fray Jorge relict forest in northern Chile</mods:title>
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<mods:namePart id="7A080EA9BDA339EF16A158572ED98B13">Parra, Luis E.</mods:namePart>
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<mods:namePart id="8F5C71D0D433515CA9AF45FDDEA7B3FF">Villagrán-Mella, Romina</mods:namePart>
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<mods:roleTerm id="7CD21E5B980465995CFD4721B703B203">Author</mods:roleTerm>
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<mods:namePart id="943438CD2B10127FF3170221F689E0F9">Marquet, Pablo A.</mods:namePart>
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<mods:typeOfResource id="5429071741CD10D682DB9F85DB72AEB9">text</mods:typeOfResource>
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<mods:title id="EAB1A79FE2F120CC2BA04D97E7FC78D6">Zootaxa</mods:title>
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<mods:part id="FF6C6955CA3A4D03A16271A8D94FFD17">
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<mods:date id="FFF4212E84C8EB86B8424F875CC4AF55">2009</mods:date>
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<mods:detail id="5888996CB5D5C4A8DEEC1D592C87BF9E" type="volume">
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<mods:number id="7399F1ACF2BA0748A4B6F1337025BE1B">2273</mods:number>
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<mods:identifier id="10227684FF19013B655D7A4C12CBF6E6" type="DOI">10.5281/zenodo.191026</mods:identifier>
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<mods:identifier id="2DDC45422E0C48FE924EA4044F146739" type="ISSN">1175-5326</mods:identifier>
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<treatment id="432A5E1EFFA12D4DFF0AAAF8A4ABF920" ID-DOI="http://doi.org/10.5281/zenodo.6214938" ID-GBIF-Taxon="119625977" ID-Zenodo-Dep="6214938" LSID="urn:lsid:plazi:treatment:432A5E1EFFA12D4DFF0AAAF8A4ABF920" httpUri="http://treatment.plazi.org/id/432A5E1EFFA12D4DFF0AAAF8A4ABF920" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
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<subSubSection id="8399BC83FFA12D4EFF0AAAF8A25BF901" pageId="1" pageNumber="60" type="nomenclature">
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<paragraph id="CB3CEF08FFA12D4EFF0AAAF8A28EFA1A" blockId="1.[151,648,1458,1484]" box="[151,648,1458,1484]" pageId="1" pageNumber="60">
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<heading id="90745864FFA12D4EFF0AAAF8A28EFA1A" bold="true" box="[151,648,1458,1484]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
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<emphasis id="F9F7331AFFA12D4EFF0AAAF8A28EFA1A" bold="true" box="[151,648,1458,1484]" pageId="1" pageNumber="60">
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<taxonomicName id="0C83948BFFA12D4EFF0AAAF8A225FA1A" authority="Parra" authorityName="Parra" box="[151,547,1458,1484]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="ediliacarmenae" status="sp. nov.">
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<emphasis id="F9F7331AFFA12D4EFF0AAAF8A1D5FA1A" bold="true" box="[151,467,1458,1484]" italics="true" pageId="1" pageNumber="60">Hasodima ediliacarmenae</emphasis>
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Parra
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</taxonomicName>
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<taxonomicNameLabel id="E2C48E61FFA12D4EFDB6AAF8A28EFA1A" box="[555,648,1458,1484]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="CB3CEF08FFA12D4EFF0AAAB1A25BF901" blockId="1.[151,1437,1531,2022]" pageId="1" pageNumber="60">
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<emphasis id="F9F7331AFFA12D4EFF0AAAB1A0D8F9C3" bold="true" box="[151,222,1531,1557]" pageId="1" pageNumber="60">
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<typeStatus id="143851AAFFA12D4EFF0AAAB1A0D8F9C3" box="[151,222,1531,1557]" pageId="1" pageNumber="60">Types</typeStatus>
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</emphasis>
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.
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<typeStatus id="143851AAFFA12D4EFF72AAB6A15BF9C0" box="[239,349,1532,1558]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
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(male), Fray Jorge,
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<date id="BF3DC9C8FFA12D4EFDD3AAB6A321F9C0" box="[590,807,1532,1558]" pageId="1" pageNumber="60">31 April to 1 May</date>
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, 2005, Romina Villagrán- Mella coll. (UCCC-ZUC). Allotype (female), Fray Jorge,
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<date id="BF3DC9C8FFA12D4EFD9EA968A2EFF9EA" box="[515,745,1570,1596]" pageId="1" pageNumber="60">31 April to 1 Mayo</date>
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, 2005, Romina Villagrán- Mella coll., Museo de Zoología, Universidad de Concepción (UCCC-MZUC).
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<typeStatus id="143851AAFFA12D4EFD49A903A354F9B5" box="[724,850,1609,1635]" pageId="1" pageNumber="60" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="DD852481FFA12D4EFCC3A903A3C4F9B5" box="[862,962,1609,1635]" pageId="1" pageNumber="60" type="male">4 males</specimenCount>
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, Fray Jorge,
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<date id="BF3DC9C8FFA12D4EFBF8A903A54CF9B5" box="[1125,1354,1609,1635]" pageId="1" pageNumber="60">31 April to 1 May</date>
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, 2005, Romina Villagrán- Mella coll.;
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<specimenCount id="DD852481FFA12D4EFD9BA93AA251F95C" box="[518,599,1648,1674]" pageId="1" pageNumber="60" type="male">1 male</specimenCount>
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, Fray Jorge, 4 to
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<date id="BF3DC9C8FFA12D4EFC82A93AA328F95C" box="[799,814,1648,1674]" pageId="1" pageNumber="60">5</date>
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May, 2005, Romina Villagrán- Mella coll.;
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<specimenCount id="DD852481FFA12D4EFAB2A93AA59EF95C" box="[1327,1432,1648,1674]" pageId="1" pageNumber="60" type="female">1 female</specimenCount>
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, Fray Jorge,
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<date id="BF3DC9C8FFA12D4EFEB1A9DCA205F966" box="[300,515,1686,1712]" pageId="1" pageNumber="60">30 to 31 October</date>
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, 2005, Romina Villagrán- Mella coll.;
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<specimenCount id="DD852481FFA12D4EFC76A9DCA444F966" box="[1003,1090,1686,1712]" pageId="1" pageNumber="60" type="male">1 male</specimenCount>
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, 6 to
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<date id="BF3DC9C8FFA12D4EFB17A9DCA49FF966" box="[1162,1177,1686,1712]" pageId="1" pageNumber="60">7</date>
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June, 2006, Romina Villagrán- Mella coll. (UCCC-MZUC)
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</paragraph>
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</subSubSection>
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<subSubSection id="8399BC83FFA12D4EFF58A9A9A3BAF84E" pageId="1" pageNumber="60" type="diagnosis">
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<paragraph id="CB3CEF08FFA12D4EFF58A9A9A3BAF84E" blockId="1.[151,1437,1531,2022]" pageId="1" pageNumber="60">
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<emphasis id="F9F7331AFFA12D4EFF58A9A9A143F92B" bold="true" box="[197,325,1763,1789]" pageId="1" pageNumber="60">Diagnosis.</emphasis>
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<taxonomicName id="0C83948BFFA12D4EFED3A9AEA281F92B" box="[334,647,1764,1789]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="ediliacarmenae">
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<emphasis id="F9F7331AFFA12D4EFED3A9AEA281F92B" box="[334,647,1764,1789]" italics="true" pageId="1" pageNumber="60">Hasodima ediliacarmenae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="0C83948BFFA12D4EFD5FA9AEA340F92B" box="[706,838,1764,1789]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="bartletti">
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<emphasis id="F9F7331AFFA12D4EFD5FA9AEA340F92B" box="[706,838,1764,1789]" italics="true" pageId="1" pageNumber="60">H. bartletti</emphasis>
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</taxonomicName>
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are very similar, being distinguished by the shape of the uncus, pseudouncus in male genitalia and the shape of the bursa copulatrix on the females. In
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<taxonomicName id="0C83948BFFA12D4EFAE2A841A156F89C" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="ediliacarmenae">
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<emphasis id="F9F7331AFFA12D4EFAE2A841A156F89C" italics="true" pageId="1" pageNumber="60">H. ediliacarmenae</emphasis>
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</taxonomicName>
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the uncus and pseudouncus are digitiform, while in
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<taxonomicName id="0C83948BFFA12D4EFC5AA87BA44AF89C" box="[967,1100,1841,1866]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="bartletti">
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<emphasis id="F9F7331AFFA12D4EFC5AA87BA44AF89C" box="[967,1100,1841,1866]" italics="true" pageId="1" pageNumber="60">H. bartletti</emphasis>
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</taxonomicName>
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the subterminal zone of the uncus is enlarged and the pseudouncus is lanceolate. Female genitalia with anterior part of corpus bursae subpyriform in the new species whilst being spherical in
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<taxonomicName id="0C83948BFFA12D4EFCAEA835A3B3F84E" box="[819,949,1919,1944]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="bartletti">
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<emphasis id="F9F7331AFFA12D4EFCAEA835A3B3F84E" box="[819,949,1919,1944]" italics="true" pageId="1" pageNumber="60">H. bartletti</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="8399BC83FFA12D4DFF58A8EFA59AF9E3" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60" type="description">
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<paragraph id="CB3CEF08FFA12D4DFF58A8EFA15BFE31" blockId="1.[151,1437,1531,2022]" lastBlockId="2.[151,1437,152,487]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
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<emphasis id="F9F7331AFFA12D4EFF58A8EFA165F869" bold="true" box="[197,355,1957,1983]" pageId="1" pageNumber="60">Description.</emphasis>
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Adult (
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<figureCitation id="53B8F38DFFA12D4EFE5EA8EFA20DF869" box="[451,523,1957,1983]" captionStart="FIGURE 1" captionStartId="2.[151,257,990,1014]" captionTargetBox="[217,1369,545,964]" captionTargetId="figure@2.[217,1369,545,965]" captionTargetPageId="2" captionText="FIGURE 1. Adult specimens of the genera Hasodima and Euclidiodes. (a) H. ediliacarmenae Parra sp. nov., (b) E. frayjorgeana Parra sp. nov. Scale 1 cm." httpUri="https://zenodo.org/record/191027/files/figure.png" pageId="1" pageNumber="60">Fig. 1</figureCitation>
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a). Head brown-green; brown-green, antennae bipectinate. Thorax browngreen; legs similarly colored, tibial spur formula 0-2-4; posterior tibias with hair tuft. Forewings brown-green; antemedial and postmedial lines fine, dark brown-green and close to inner margin of the forewings, antemedial line proximally bordered by a whitish band reaching the basal edge of the wing; ground colour of terminal area lighter than the rest of the wing terminal dots brown; discal dot small, dark brown-green; postmedial line oblique; basal area with dark brown bar perpendicular to the body axis; underside of wings lighter brown-green, with discal dot and diffuse bands. Hindwings ochre, the terminal area dark brown; discal dot and postmedial line barely perceptible; underside of hindwing ochre; postmedial line and discal dot less apparent; terminal area dark brown. Abdomen conlorous to the wings, with dorsal tufts of scales between the third and seventh abdominal segment. Female. Similar to the male, with filiform antennae, posterior tibias without hair tuft.
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</paragraph>
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<caption id="9FFCBF80FFA22D4DFF0AAC94A238FBCE" httpUri="https://zenodo.org/record/191027/files/figure.png" pageId="2" pageNumber="61" targetBox="[217,1369,545,964]" targetPageId="2">
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<paragraph id="CB3CEF08FFA22D4DFF0AAC94A238FBCE" blockId="2.[151,1437,990,1049]" pageId="2" pageNumber="61">
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<emphasis id="F9F7331AFFA22D4DFF0AAC94A11BFC20" bold="true" box="[151,285,990,1014]" pageId="2" pageNumber="61">FIGURE 1.</emphasis>
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Adult specimens of the genera
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<taxonomicName id="0C83948BFFA22D4DFDE2AC95A2F6FC20" box="[639,752,991,1014]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="F9F7331AFFA22D4DFDE2AC95A2F6FC20" box="[639,752,991,1014]" italics="true" pageId="2" pageNumber="61">Hasodima</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="0C83948BFFA22D4DFCBBAC95A3AEFC20" box="[806,936,991,1014]" class="Insecta" family="Geometridae" genus="Euclidiodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="F9F7331AFFA22D4DFCBBAC95A3AEFC20" box="[806,936,991,1014]" italics="true" pageId="2" pageNumber="61">Euclidiodes</emphasis>
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</taxonomicName>
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. (a)
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<taxonomicName id="0C83948BFFA22D4DFC43AC95A4EBFC20" authority="Parra" authorityName="Parra" box="[990,1261,991,1014]" class="Insecta" family="Geometridae" genus="Hasodima" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="ediliacarmenae" status="sp. nov.">
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<emphasis id="F9F7331AFFA22D4DFC43AC95A4ADFC20" box="[990,1195,991,1014]" italics="true" pageId="2" pageNumber="61">H. ediliacarmenae</emphasis>
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Parra
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</taxonomicName>
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<emphasis id="F9F7331AFFA22D4DFB68AC94A548FC20" bold="true" box="[1269,1358,990,1014]" pageId="2" pageNumber="61">
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<taxonomicNameLabel id="E2C48E61FFA22D4DFB68AC94A548FC20" box="[1269,1358,990,1014]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, (b)
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<taxonomicName id="0C83948BFFA22D4DFA19AC95A166FBCE" authority="Parra" authorityName="Parra" class="Insecta" family="Geometridae" genus="Euclidiodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="frayjorgeana" status="sp. nov.">
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<emphasis id="F9F7331AFFA22D4DFA19AC95A127FBCF" italics="true" pageId="2" pageNumber="61">E. frayjorgeana</emphasis>
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Parra
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</taxonomicName>
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<emphasis id="F9F7331AFFA22D4DFEFBAB4BA1BAFBCF" bold="true" box="[358,444,1025,1049]" pageId="2" pageNumber="61">
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<taxonomicNameLabel id="E2C48E61FFA22D4DFEFBAB4BA1BAFBCF" box="[358,444,1025,1049]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Scale 1 cm.
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</paragraph>
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</caption>
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<paragraph id="CB3CEF08FFA22D4DFF58AB01A293FA9B" blockId="2.[151,1437,1099,1782]" pageId="2" pageNumber="61">
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Male genitalia (
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<figureCitation id="53B8F38DFFA22D4DFE19AB01A1C8FBB3" box="[388,462,1099,1125]" captionStart="FIGURE 2. H" captionStartId="3.[151,255,1529,1553]" captionTargetBox="[219,1376,643,1475]" captionTargetId="figure@3.[195,1392,622,1503]" captionTargetPageId="3" captionText="FIGURE 2. H. ediliacarmenae Parra sp. nov. (a) ventral view of male genitalia, (b) lateral view of aedeagus, (c) ventral view of female genitalia. Scale 1 mm." httpUri="https://zenodo.org/record/191028/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
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a–b). Uncus digitiform, apically rounded; pseudouncus digitiform; socii present, with setae; apical cleavage of valves subequal to the width of the valvula; costal margin of valva strongly sclerotized, subapically with digitiform projection; one cuspidiform projection in the center with numerous hairs; gnathos “V” shaped apically widened dentate and tapering tip; juxta subrectangular with two lateral processes in the form of a hook, 1/3 length of uncus; furca acicular, 4 times larger than lateral processes; region of the sacculus hollow; saccus subquadrangular; aedeagus straight, at ductus ejaculatorius constricted; vesica with a group of long and fine spines.
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</paragraph>
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<paragraph id="CB3CEF08FFA22D4DFF58AA10A30CF9D8" blockId="2.[151,1437,1099,1782]" pageId="2" pageNumber="61">
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Female genitalia (
|
||
<figureCitation id="53B8F38DFFA22D4DFE3AAA10A1F2FAA2" box="[423,500,1370,1396]" captionStart="FIGURE 2. H" captionStartId="3.[151,255,1529,1553]" captionTargetBox="[219,1376,643,1475]" captionTargetId="figure@3.[195,1392,622,1503]" captionTargetPageId="3" captionText="FIGURE 2. H. ediliacarmenae Parra sp. nov. (a) ventral view of male genitalia, (b) lateral view of aedeagus, (c) ventral view of female genitalia. Scale 1 mm." httpUri="https://zenodo.org/record/191028/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
|
||
c). Corpus bursae subpyriform, ductus bursae sclerotized with longitudinal furrows; signum large, peduncle and ensiform, five times smaller than the corpus bursae; peduncle the half of the ensiform body of signum; anterior apophyses 1/3 of the posterior apophyses; ductus seminalis arising from intersection between the corpus bursae and the ductus bursae; antrum quadrangular with sclerotized lateral areas, 16 times smaller than the corpus bursae.
|
||
</paragraph>
|
||
<paragraph id="CB3CEF08FFA22D4DFF58A951A59AF9E3" blockId="2.[151,1437,1099,1782]" box="[197,1436,1563,1589]" pageId="2" pageNumber="61">
|
||
Length of forewing. Measured from the base to the apex. Male:
|
||
<quantity id="0C7B42EDFFA22D4DFC30A951A42DF9E3" box="[941,1067,1563,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="2" pageNumber="61" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14–15 mm</quantity>
|
||
(n = 6); female:
|
||
<quantity id="0C7B42EDFFA22D4DFB70A951A546F9E3" box="[1261,1344,1563,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="61" unit="mm" value="15.0">15 mm</quantity>
|
||
(n = 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8399BC83FFA22D4DFF5BA90BA317F919" pageId="2" pageNumber="61" type="distribution">
|
||
<paragraph id="CB3CEF08FFA22D4DFF5BA90BA0D8F954" blockId="2.[151,1437,1099,1782]" pageId="2" pageNumber="61">
|
||
<emphasis id="F9F7331AFFA22D4DFF5BA90BA161F98D" bold="true" box="[198,359,1601,1627]" pageId="2" pageNumber="61">Distribution.</emphasis>
|
||
This species has only been collected in Fray Jorge National Park, in Coquimbo Region of
|
||
<collectingCountry id="B394AF98FFA22D4DFF0AA922A0DDF954" box="[151,219,1640,1666]" name="Chile" pageId="2" pageNumber="61">Chile</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CB3CEF08FFA22D4DFF58A9C5A317F919" blockId="2.[151,1437,1099,1782]" pageId="2" pageNumber="61">
|
||
<emphasis id="F9F7331AFFA22D4DFF58A9C5A169F97F" bold="true" box="[197,367,1679,1705]" pageId="2" pageNumber="61">Flight period.</emphasis>
|
||
Specimens of this species have been collected during the austral fall and spring, during the months of April, May, June and October, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8399BC83FFA22D4DFF58A996A4ABF920" box="[197,1197,1756,1782]" pageId="2" pageNumber="61" type="etymology">
|
||
<paragraph id="CB3CEF08FFA22D4DFF58A996A4ABF920" blockId="2.[151,1437,1099,1782]" box="[197,1197,1756,1782]" pageId="2" pageNumber="61">
|
||
<emphasis id="F9F7331AFFA22D4DFF58A996A157F920" bold="true" box="[197,337,1756,1782]" pageId="2" pageNumber="61">Etymology.</emphasis>
|
||
This species is dedicated to Edilia del Carmen, the wife of Luis E. Parra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |