181 lines
20 KiB
XML
181 lines
20 KiB
XML
<document ENCODING="UTF-8" ID-GBIF-Dataset="db3b4f9d-87ca-4661-a002-c868b7417878" ModsDocID="z01416p001" checkinTime="1247405641513" checkinUser="thomas" docAuthor="Alexandre P. Marceniuk & Naércio A. Menezes" docDate="2007" docId="FC70B862DF0D334CB558CD1B971EC57F" docLanguage="en" docName="2007_Marceniuk_Menezes_gg1.xml" docOrigin="Zootaxa 1416" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:FFC65592-D8DB-41BE-AEAC-A41EAB6C6185" docTitle="Carlarius Marceniuk & Menezes, 2007, new genus" docType="treatment" docVersion="12" lastPageNumber="37" masterDocId="E3B79C8FF6B0B87BE406EA4948E96A3B" masterDocTitle="Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera." masterLastPageNumber="126" masterPageNumber="1" pageNumber="36" updateTime="1677103743125" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Alexandre P. Marceniuk</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Naércio A. Menezes</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2007</mods:date>
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<mods:detail type="volume">
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<mods:number>1416</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>126</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.zoobank.org/urn:lsid:zoobank.org:pub:FFC65592-D8DB-41BE-AEAC-A41EAB6C6185</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ZooBank-PUB">z01416p001</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6236843" ID-GBIF-Taxon="100130212" ID-Zenodo-Dep="6236843" LSID="urn:lsid:plazi:treatment:FC70B862DF0D334CB558CD1B971EC57F" httpUri="http://treatment.plazi.org/id/FC70B862DF0D334CB558CD1B971EC57F" lastPageNumber="37" pageNumber="36">
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<subSubSection type="nomenclature">
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<paragraph pageNumber="36">
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<taxonomicName ID-CoL="3HNC" ID-ENA="591312" class="Actinopterygii" family="Ariidae" genus="Carlarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Carlarius</taxonomicName>
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<taxonomicNameLabel rank="genus">new genus</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection type="description">
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<paragraph pageNumber="36">(fig. 29)</paragraph>
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</subSubSection>
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<subSubSection type="nomenclature">
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<paragraph pageNumber="36">
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Type species:
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:B92897AF-B325-4D6B-8032-63BE476D80F3" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius heudelotii Valenciennes 1840:73" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="heudelotii">Arius heudelotii</taxonomicName>
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Valenciennes, 1840.
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</paragraph>
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</subSubSection>
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<subSubSection type="diagnosis">
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<paragraph pageNumber="36">
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Diagnosis. Three shared characters distinguishes
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Carlarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Carlarius</taxonomicName>
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from the remaining ariid genera: (1) mesethmoid wide at its median portion (shared with
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Amphiarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Amphiarius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:E0FC1BFA-4268-4955-BBF3-891B2146C227" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius caelatus Valenciennes 1840:66" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="caelatus">Arius caelatus</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:FB0B3553-B6DA-48F8-BA0A-EE430A96E0DB" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius madagascariensis Vaillant 1894:77 [10]" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="madagascariensis">A. madagascariensis</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:347CE897-3860-4DA6-85B8-033B2318B81A" class="Actinopterygii" family="Ariidae" genus="Aspistor" kingdom="Animalia" lsidName-ZBK="Aspistor Jordan & Evermann 1898:2763" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Aspistor</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:4453B6E6-6001-42AB-8D66-002DF12E8C99" class="Actinopterygii" family="Ariidae" genus="Cryptarius" kingdom="Animalia" lsidName-ZBK="Cryptarius Kailola 2004:134" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Cryptarius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:4781B086-6A2E-487B-8D97-1D1A856A235A" class="Actinopterygii" family="Ariidae" genus="Galeichthys" kingdom="Animalia" lsidName-ZBK="Galeichthys Valenciennes 1840:28" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Galeichthys</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:CC7B154F-0B6B-4A8C-A4DD-2EBF83B9C00A" class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" lsidName-ZBK="Genidens Castelnau 1855:33" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Genidens</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:3F1F6AF2-6F38-42C0-9C82-C1468F19FAF0" class="Actinopterygii" family="Ariidae" genus="Hemiarius" kingdom="Animalia" lsidName-ZBK="Hemiarius Bleeker 1862:7, 29" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Hemiarius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:F589893C-76A9-4820-A8B9-B794837B3FB9" class="Actinopterygii" family="Ariidae" genus="Nedystoma" kingdom="Animalia" lsidName-ZBK="Nedystoma Ogilby 1898:32" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Nedystoma</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:5BD940A7-0AA1-4F57-B0A5-9D295A651224" class="Actinopterygii" family="Ariidae" genus="Nemapteryx" kingdom="Animalia" lsidName-ZBK="Nemapteryx Ogilby 1908:3, 10" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Nemapteryx</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:A465F2E2-3CB7-445A-A792-A01BE7E8F95E" class="Actinopterygii" family="Ariidae" genus="Neoarius" kingdom="Animalia" lsidName-ZBK="Neoarius Castelnau 1878:237" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Neoarius</taxonomicName>
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,
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Netuma" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Netuma</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" class="Actinopterygii" family="Ariidae" genus="Notarius" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Notarius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:1AE6B6D0-01A8-43E7-9505-E6855D1FC99A" class="Actinopterygii" family="Ariidae" genus="Plicofollis" kingdom="Animalia" lsidName-ZBK="Plicofollis Kailola 2004:141" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Plicofollis</taxonomicName>
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,
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Potamosilurus" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Potamosilurus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageNumber="36" phylum="Arthropoda" rank="species" species="assimilis">Sciades assimilis</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageNumber="36" phylum="Arthropoda" rank="species" species="bonillai">S. bonillai</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageNumber="36" phylum="Arthropoda" rank="species" species="felis">S. felis</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageNumber="36" phylum="Arthropoda" rank="species" species="platypogon">S. platypogon</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageNumber="36" phylum="Arthropoda" rank="species" species="sagor">S. sagor</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageNumber="36" phylum="Arthropoda" rank="species" species="seemanni">S. seemanni</taxonomicName>
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); (2) tooth plates associated with vomer absent [shared with
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Amphiarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Amphiarius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:830D572B-79ED-4DBD-84EC-A10E3E7E748C" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius Valenciennes 1840:53" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Arius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:A8968D59-B441-4D26-BC5F-B90032C1F1D1" class="Actinopterygii" family="Ariidae" genus="Batrachocephalus" kingdom="Animalia" lsidName-ZBK="Batrachocephalus Bleeker 1846:176" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Batrachocephalus</taxonomicName>
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,
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Cathorops" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Cathorops</taxonomicName>
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(with exception of
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Cathorops" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="dasycephalus">C. dasycephalus</taxonomicName>
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),
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:57F8F227-2C96-4E3F-B4D1-3CDE8E5DE40F" class="Actinopterygii" family="Ariidae" genus="Cephalocassis" kingdom="Animalia" lsidName-ZBK="Cephalocassis Bleeker 1857:473" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Cephalocassis</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:8E024180-3250-46F3-A751-1D583938D54E" class="Actinopterygii" family="Ariidae" genus="Cinetodus" kingdom="Animalia" lsidName-ZBK="Cinetodus Ogilby 1898:32" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Cinetodus</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:4453B6E6-6001-42AB-8D66-002DF12E8C99" class="Actinopterygii" family="Ariidae" genus="Cryptarius" kingdom="Animalia" lsidName-ZBK="Cryptarius Kailola 2004:134" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Cryptarius</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:3EF9A59B-2473-44CF-BB0F-3846968A6AB8" family="Ariidae" genus="Doiichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Doiichthys Weber 1913:532" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Doiichthys</taxonomicName>
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,
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Genidens" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="genidens">Genidens genidens</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:B824B602-4F87-4771-B334-61385645D6C4" class="Actinopterygii" family="Ariidae" genus="Ketengus" kingdom="Animalia" lsidName-ZBK="Ketengus Bleeker 1847:167" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Ketengus</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:F589893C-76A9-4820-A8B9-B794837B3FB9" class="Actinopterygii" family="Ariidae" genus="Nedystoma" kingdom="Animalia" lsidName-ZBK="Nedystoma Ogilby 1898:32" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Nedystoma</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:7EFF0E2C-627F-4985-9DCA-5178DA9034F3" class="Actinopterygii" family="Ariidae" genus="Osteogeneiosus" kingdom="Animalia" lsidName-ZBK="Osteogeneiosus Bleeker 1846:173" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Osteogeneiosus</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:D469DDD6-F17E-4DEB-99B7-0C54A27687E2" authority="Ogilby, 1898" authorityName="Ogilby" authorityYear="1898" class="Actinopterygii" family="Ariidae" genus="Pachyula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-ZBK="Pachyula Ogilby 1898:33" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Pachyula</taxonomicName>
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,
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Plicofollis" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="platystomus">Plicofollis platystomus</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:AAD7809B-5174-4F77-B9AC-2359BB91181F" class="Actinopterygii" family="Ariidae" genus="Potamarius" kingdom="Animalia" lsidName-ZBK="Potamarius Hubbs & Miller 1960:101" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Potamarius</taxonomicName>
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and
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Potamosilurus" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="velutinus">Potamosilurus velutinus</taxonomicName>
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]; (3) epioccipital dorsally exposed [shared with
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Amphiarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="rugispinis">Amphiarius rugispinis</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:347CE897-3860-4DA6-85B8-033B2318B81A" class="Actinopterygii" family="Ariidae" genus="Aspistor" kingdom="Animalia" lsidName-ZBK="Aspistor Jordan & Evermann 1898:2763" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Aspistor</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:8E024180-3250-46F3-A751-1D583938D54E" class="Actinopterygii" family="Ariidae" genus="Cinetodus" kingdom="Animalia" lsidName-ZBK="Cinetodus Ogilby 1898:32" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Cinetodus</taxonomicName>
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,
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:D469DDD6-F17E-4DEB-99B7-0C54A27687E2" authority="Ogilby, 1898" authorityName="Ogilby" authorityYear="1898" class="Actinopterygii" family="Ariidae" genus="Pachyula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-ZBK="Pachyula Ogilby 1898:33" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Pachyula</taxonomicName>
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and
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:1AE6B6D0-01A8-43E7-9505-E6855D1FC99A" class="Actinopterygii" family="Ariidae" genus="Plicofollis" kingdom="Animalia" lsidName-ZBK="Plicofollis Kailola 2004:141" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Plicofollis</taxonomicName>
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(with exception of
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Plicofollis" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="platystomus">P. platystomus</taxonomicName>
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and
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Plicofollis" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="tenuispinis">P. tenuispinis</taxonomicName>
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)].
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</paragraph>
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<paragraph pageNumber="36">Supplementary morphological characters. Cephalic shield conspicuously granulated visible under the skin; a moderately developed fenestra limited by frontal and lateral ethmoid bones visible under the skin; medial groove of neurocranium very distinct, limited by frontal bones and/or on supraoccipital; posterior cranial fontanel of moderate size, long and narrow; fenestra limited by supraoccipital, pterotic and sphenotic absent; fossa limited by pterotic, supracleithrum and extrascapular moderately large; epioccipital invading dorsal portion of cephalic shield; occipital process triangular, moderately long and wide, progressively narrower toward its posterior part; anterior and median nuchal plates fused and indistinct, forming a structure of semi-lunar aspect; vomerine tooth plates absent; accessory tooth plates well developed oval shaped, with conical teeth; maxillary barbel fleshy and cylindrical; two pairs of mental barbels; base of adipose fin moderately long, about half the length of anal-fin base; lateral line not bifurcated at caudal region, reaching base of caudal-fin upper lobe; cleithrum wide, with second dorsal process on its upper portion; posterior cleithral process moderately long and distinct from second dorsal process of cleithrum.</paragraph>
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</subSubSection>
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<subSubSection type="discussion">
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<paragraph lastPageNumber="37" pageNumber="36">
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Remarks.
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Carlarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Carlarius</taxonomicName>
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is established to accommodate all ariid species from the occidental portion of the African continent that in previous classifications were included in
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:830D572B-79ED-4DBD-84EC-A10E3E7E748C" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius Valenciennes 1840:53" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Arius</taxonomicName>
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(Fowler, 1936; Taylor, 1986, 1990; Burgess, 1989; Daget, 1992) and were recognized by Marceniuk (2003) as a monophyletic lineage. They can
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<pageStartToken pageNumber="37">be</pageStartToken>
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easily differentiated from all members of
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:830D572B-79ED-4DBD-84EC-A10E3E7E748C" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius Valenciennes 1840:53" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="genus">Arius</taxonomicName>
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by having the lateral line not bifurcated at caudal region, reaching caudal-fin upper lobe and the epioccipital bone exposed on the dorsal part of the cranium (see also diagnosis of the genus).
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<taxonomicName class="Actinopterygii" family="Ariidae" genus="Carlarius" kingdom="Animalia" order="Siluriformes" pageNumber="36" phylum="Chordata" rank="species" species="gigas">C. gigas</taxonomicName>
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(Boulenger, 1911) was not examined and its inclusion in the genus is based on diagnostic characters from the literature.
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</paragraph>
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</subSubSection>
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<subSubSection type="distribution">
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<paragraph pageNumber="37">Distribution and habitat. Western Africa, brackish and fresh waters.</paragraph>
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</subSubSection>
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<subSubSection type="etymology">
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<paragraph pageNumber="37">
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Etymology. The first part of the generic name honors Dr. Carl Ferraris, Research Associate at the California Academy of Sciences for his contribution to the knowledge of Siluriformes fishes and continuous support and encouragement throughout the development of this work. The second part is from the frequently used generic name
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:830D572B-79ED-4DBD-84EC-A10E3E7E748C" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" lsidName-ZBK="Arius Valenciennes 1840:53" order="Siluriformes" pageNumber="37" phylum="Chordata" rank="genus">Arius</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |