172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document id="3E24067DB4BE249CDF2459082652CEAA" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629739267003" checkinUser="felipe" docAuthor="Valladares, Luis F. & Short, Andrew Edward Z." docDate="2011" docId="BC6CF939EE02FFF0FF44FDDEB650FBBE" docLanguage="en" docName="zt03021p068.pdf" docOrigin="Zootaxa 3021" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochthebius attritus LeConte 1878" docType="treatment" docVersion="4" lastPageNumber="67" masterDocId="40558141EE06FFF4FFD3FFE6B519FFE9" masterDocTitle="Revision of the Ochthebiinae of Venezuela (Coleoptera: Hydraenidae)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="67" updateTime="1699104834155" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="DE11636A7C31976B96328C6BAAA11BEB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="29059AFAFDE32BE3EF22AD26AB2E3CFB">
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<mods:title id="76DA56829D7B5449E82AED35586B0229">Revision of the Ochthebiinae of Venezuela (Coleoptera: Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="5AE93FEDEE4644DC2676D0C913059C81" type="personal">
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<mods:role id="114E90BF1F29E72FDD58E5F5FE6D9A50">
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<mods:roleTerm id="437F5C8649FDFF6D80E4B61238BA621B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="11327FFE437CC3B4B79EEC5DCC921B46">Valladares, Luis F.</mods:namePart>
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<mods:affiliation id="21F04461521155C80A2BF7578F2FE7DE">Departamento de Biodiversidad y Gestión Ambiental (Zoología), Facultad de Ciencias Biológicas y Ambientales, Universidad de León, 24071 León, SPAIN. E-mail: lfvald @ unileon. es</mods:affiliation>
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</mods:name>
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<mods:name id="02D174B4E33487B7032CBCD3A19B8F29" type="personal">
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<mods:role id="3B28D905F0131438CCA852D939B93D01">
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<mods:roleTerm id="5240C9E343BCFBEBDBB911B4ACBFC61B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7750D4903E0E08ABE661BDDA722CF5BD">Short, Andrew Edward Z.</mods:namePart>
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<mods:affiliation id="9C5836B1001B90AA32ADFA83FB64C585">Division of Entomology, Biodiversity Institute, and Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence KS 66045, USA. E-mail: aezshort @ ku. edu</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="9B874E7E27C773084996B01EC97D043E">text</mods:typeOfResource>
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<mods:titleInfo id="94C34BA122586F41EDB588C80FF4E38F">
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<mods:title id="8972D5A8C6447E2E61AD21F15E78C93D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="88EC6A2D3717523575CC2DA4F1F49D91">
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<mods:date id="FE87FC44EE96E0F171FBCC4A0D108FA3">2011</mods:date>
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<mods:detail id="A330FAF4DED63247916D45CA965874EE" type="pubDate">
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<mods:number id="B8487A18558EE7B6D43B7F991374F8E3">2011-09-12</mods:number>
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</mods:detail>
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<mods:detail id="D07C18C46A7814C69D05B0B0CB5F3C18" type="volume">
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<mods:number id="4785B282E9E56C9B0259B2CCB84DD63A">3021</mods:number>
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</mods:detail>
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<mods:start id="0766F114C92C07D79D0A12AB285B78BB">63</mods:start>
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<mods:end id="C5379FAE21F23A7E3C48CF68010DFF84">68</mods:end>
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</mods:extent>
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<mods:classification id="EE523AB0D892AE6CC4A2544164B20436">journal article</mods:classification>
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<mods:identifier id="D6C6FC76139A0215287A2CE007BF689E" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="BC6CF939EE02FFF0FF44FDDEB650FBBE" ID-DOI="http://doi.org/10.5281/zenodo.5285563" ID-Zenodo-Dep="5285563" LSID="urn:lsid:plazi:treatment:BC6CF939EE02FFF0FF44FDDEB650FBBE" httpUri="http://treatment.plazi.org/id/BC6CF939EE02FFF0FF44FDDEB650FBBE" lastPageNumber="67" pageId="4" pageNumber="67">
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<subSubSection id="7CDF1BA4EE02FFF0FF44FDDEB771FDA6" box="[151,616,567,592]" pageId="4" pageNumber="67" type="nomenclature">
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<paragraph id="347A482FEE02FFF0FF44FDDEB771FDA6" blockId="4.[151,616,567,625]" box="[151,616,567,592]" pageId="4" pageNumber="67">
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<heading id="6F32FF43EE02FFF0FF44FDDEB771FDA6" bold="true" box="[151,616,567,592]" fontSize="10" level="2" pageId="4" pageNumber="67" reason="2">
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<taxonomicName id="F3C533ACEE02FFF0FF44FDDEB771FDA6" ID-CoL="48F4B" authority="LeConte, 1878" authorityName="LeConte" authorityYear="1878" box="[151,616,567,592]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="attritus">
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<emphasis id="06B1943DEE02FFF0FF44FDDEB771FDA6" bold="true" box="[151,616,567,592]" pageId="4" pageNumber="67">
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<emphasis id="06B1943DEE02FFF0FF44FDDEB40CFDB9" bold="true" box="[151,277,568,592]" italics="true" pageId="4" pageNumber="67">Ochthebius</emphasis>
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(s. str.)
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<emphasis id="06B1943DEE02FFF0FEBCFDDEB4A7FDB9" bold="true" box="[367,446,568,592]" italics="true" pageId="4" pageNumber="67">attritus</emphasis>
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LeConte, 1878
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</emphasis>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="7CDF1BA4EE02FFF0FF44FDBCB499FD98" box="[151,384,602,625]" pageId="4" pageNumber="67" type="description">
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<paragraph id="347A482FEE02FFF0FF44FDBCB499FD98" blockId="4.[151,616,567,625]" box="[151,384,602,625]" pageId="4" pageNumber="67">
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<figureCitation id="ACFE54AAEE02FFF0FF44FDBCB5EBFD98" box="[151,242,602,625]" captionStart="FIGURE 1" captionStartId="1.[151,244,1801,1822]" captionTargetBox="[209,1380,974,1752]" captionTargetId="figure-438@1.[151,1436,954,1780]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus photographs of Venezuela Ochthebiinae species. A) Gymnochthebius fossatus (LeConte); B) Ochthebius (s. str.) lineatus LeConte; C) Ochthebius (s. str.) attritus LeConte." pageId="4" pageNumber="67">Figs. 1C</figureCitation>
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,
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<figureCitation id="ACFE54AAEE02FFF0FF2DFDBCB406FD98" box="[254,287,602,625]" captionStart="FIGURE 2" captionStartId="2.[151,244,1005,1026]" captionTargetBox="[205,1352,395,939]" captionTargetId="figure-214@2.[151,1436,367,982]" captionTargetPageId="2" captionText="FIGURE 2. Aedeagus in ventral (left) and lateral (right) view of Gymnochthebius fossatus (A); Ochthebius (s. str.) lineatus (B); Ochthebius (s. str.) attritus (C)." pageId="4" pageNumber="67">2C</figureCitation>
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,
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<figureCitation id="ACFE54AAEE02FFF0FEF8FDBCB454FD98" box="[299,333,602,625]" captionStart="FIGURE 3" captionStartId="2.[151,255,1675,1699]" captionTargetBox="[151,1436,1118,1650]" captionTargetId="figure-218@2.[151,1436,1113,1653]" captionTargetPageId="2" captionText="FIGURE 3. Distribution maps in Venezuela of: A) Ochthebius (s. str.) lineatus (circles) and Ochthebius (s. str.) attritus (star); B) Gymnochthebius fossatus." pageId="4" pageNumber="67">3A</figureCitation>
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,
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<figureCitation id="ACFE54AAEE02FFF0FE8AFDBCB464FD98" box="[345,381,602,625]" captionStart="FIGURE 4" captionStartId="3.[151,244,1984,2005]" captionTargetBox="[151,1435,470,1963]" captionTargetId="figure-182@3.[151,1436,469,1963]" captionTargetPageId="3" captionText="FIGURE 4. Photographs of selected collecting events of Ochthebiinae in Venezuela. A) Falcon State, Médanos de Coro, event VZ09- 0709-03Z; B) Sucre State: Finca Vuelta Larga, event VZ10-0129-03A; C) Guárico State: Río San Antonio, event VZ10-0204-02A." pageId="4" pageNumber="67">4A</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="7CDF1BA4EE02FFF0FF44FD79B1EFFD30" pageId="4" pageNumber="67" type="materials_examined">
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<paragraph id="347A482FEE02FFF0FF44FD79B1EFFD30" blockId="4.[151,1437,671,1111]" pageId="4" pageNumber="67">
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Material examined (25):
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<materialsCitation id="84AD4272EE02FFF0FE67FD79B1EBFD30" collectingDate="2010-07-09" collectionCode="MALUZ, MIZA, SEMC, CVL" collectorName="de Coro & A. Short & W. Shepard" country="Venezuela" elevation="8" latitude="11.436916" location="Medanos de Coro" longLatPrecision="1" longitude="-69.66853" pageId="4" pageNumber="67" specimenCount="25" stateProvince="Falcon">
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<emphasis id="06B1943DEE02FFF0FE67FD79B618FD5E" bold="true" box="[436,769,671,695]" pageId="4" pageNumber="67">
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<collectingCountry id="4CD208BFEE02FFF0FE67FD79B747FD5E" box="[436,606,671,695]" name="Venezuela" pageId="4" pageNumber="67">VENEZUELA</collectingCountry>
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:
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<collectingRegion id="F60186CDEE02FFF0FDB8FD79B7E2FD5E" box="[619,763,671,695]" country="Venezuela" name="Falcon" pageId="4" pageNumber="67">Falcón State</collectingRegion>
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:
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</emphasis>
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<location id="311A1EF4EE02FFF0FCDBFD46B6D2FD5E" LSID="urn:lsid:plazi:treatment:BC6CF939EE02FFF0FF44FDDEB650FBBE:311A1EF4EE02FFF0FCDBFD46B6D2FD5E" box="[776,971,672,695]" country="Venezuela" latitude="11.436916" longLatPrecision="1" longitude="-69.66853" name="Medanos de Coro" pageId="4" pageNumber="67" stateProvince="Falcon">
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Médanos
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<collectorName id="99302DF9EE02FFF0FCA1FD46B6D2FD5E" box="[882,971,672,695]" pageId="4" pageNumber="67">de Coro</collectorName>
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</location>
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,
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<geoCoordinate id="51F12EE8EE02FFF0FC04FD46B17FFD5E" box="[983,1126,672,695]" degrees="11" direction="north" minutes="26.215" orientation="latitude" pageId="4" pageNumber="67" precision="1" value="11.436916">11°26.215' N</geoCoordinate>
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,
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<geoCoordinate id="51F12EE8EE02FFF0FBA0FD46B01FFD5E" box="[1139,1286,672,695]" degrees="69" direction="west" minutes="40.112" orientation="longitude" pageId="4" pageNumber="67" precision="1" value="-69.66853">69°40.112' W</geoCoordinate>
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,
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<quantity id="F33DE5CAEE02FFF0FAC1FD46B027FD5E" box="[1298,1342,672,695]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="67" unit="m" value="8.0">
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||
<elevation id="BFE8AF1CEE02FFF0FAC1FD46B027FD5E" box="[1298,1342,672,695]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="67" unit="m" value="8.0">8 m</elevation>
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</quantity>
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, pond in dunes,
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<date id="407B6EEFEE02FFF0FF31FD24B448FD30" box="[226,337,706,729]" pageId="4" pageNumber="67" value="2010-07-09">
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<collectingDate id="503F9707EE02FFF0FF31FD24B448FD30" box="[226,337,706,729]" pageId="4" pageNumber="67" value="2010-07-09">9.vii.2010</collectingDate>
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||
</date>
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, leg.
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<collectorName id="99302DF9EE02FFF0FE5AFD24B4FAFD30" box="[393,483,706,729]" pageId="4" pageNumber="67">A. Short</collectorName>
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&
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<collectorName id="99302DF9EE02FFF0FDD5FD24B79DFD30" box="[518,644,706,729]" pageId="4" pageNumber="67">W. Shepard</collectorName>
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, VZ09-0709-03Z (
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||
<specimenCount id="22C383A6EE02FFF0FC9EFD24B688FD30" box="[845,913,706,729]" pageId="4" pageNumber="67" type="generic">25 ex.</specimenCount>
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,
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<collectionCode id="52D4D0EAEE02FFF0FC4FFD24B6E7FD30" box="[924,1022,706,729]" pageId="4" pageNumber="67">MALUZ</collectionCode>
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,
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<collectionCode id="52D4D0EAEE02FFF0FBDAFD24B149FD30" box="[1033,1104,706,729]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="67" type="Museum">MIZA</collectionCode>
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,
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<collectionCode id="52D4D0EAEE02FFF0FB89FD24B1BCFD30" box="[1114,1189,706,729]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="67" type="Museum">SEMC</collectionCode>
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,
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<collectionCode id="52D4D0EAEE02FFF0FB63FD24B1F3FD30" box="[1200,1258,706,729]" pageId="4" pageNumber="67">CVL</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="7CDF1BA4EE02FFF0FF14FD02B665FC25" pageId="4" pageNumber="67" type="diagnosis">
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<paragraph id="347A482FEE02FFF0FF14FD02B665FC25" blockId="4.[151,1437,671,1111]" pageId="4" pageNumber="67">
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<emphasis id="06B1943DEE02FFF0FF14FD02B425FD15" bold="true" box="[199,316,740,764]" inLineHeading="true" pageId="4" pageNumber="67" reason="1">Diagnosis.</emphasis>
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Habitus as in
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<figureCitation id="ACFE54AAEE02FFF0FE30FD03B723FD15" box="[483,570,741,764]" captionStart="FIGURE 1" captionStartId="1.[151,244,1801,1822]" captionTargetBox="[209,1380,974,1752]" captionTargetId="figure-438@1.[151,1436,954,1780]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus photographs of Venezuela Ochthebiinae species. A) Gymnochthebius fossatus (LeConte); B) Ochthebius (s. str.) lineatus LeConte; C) Ochthebius (s. str.) attritus LeConte." pageId="4" pageNumber="67">Fig. 1C</figureCitation>
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. Length:
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<quantity id="F33DE5CAEE02FFF0FD75FD03B651FD15" box="[678,840,741,764]" metricMagnitude="-3" metricUnit="m" metricValue="1.535" metricValueMax="1.56" metricValueMin="1.51" pageId="4" pageNumber="67" unit="mm" value="1.535" valueMax="1.56" valueMin="1.51">1.51–1.56 mm</quantity>
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. Color: Head shining black with greenish reflections; pronotum brown with greenish metallic reflections; elytra and legs testaceous. Pronotum with postocular tooth; median groove connected to anterior foveae forming an anterior transverse depression; posterior foveae also posteriorly united. Males with a variable tooth at the apex of the labrum. Females with anterior edge of the labrum straight. Aedeagus (
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<figureCitation id="ACFE54AAEE02FFF0FAA2FCABB5AFFC6E" captionStart="FIGURE 2" captionStartId="2.[151,244,1005,1026]" captionTargetBox="[205,1352,395,939]" captionTargetId="figure-214@2.[151,1436,367,982]" captionTargetPageId="2" captionText="FIGURE 2. Aedeagus in ventral (left) and lateral (right) view of Gymnochthebius fossatus (A); Ochthebius (s. str.) lineatus (B); Ochthebius (s. str.) attritus (C)." pageId="4" pageNumber="67">Fig. 2C</figureCitation>
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) in ventral view with the main piece narrow and the distal lobe L-shaped. The aedeagal morphology is quite variable in this species (
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<bibRefCitation id="505435DEEE02FFF0FE93FC74B4C3FC40" author="Perkins" box="[320,474,914,937]" firstAuthor="Perkins" pageId="4" pageNumber="67" pagination="3 - 554" refId="ref3183" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: Classification, Biogeography and inferred Phylogeny. (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554." type="journal article" year="1980">Perkins, 1980</bibRefCitation>
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||
). The aedeagus from Venezuelan specimens are similar to those from Florida and
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||
<collectingCountry id="4CD208BFEE02FFF0FA88FC74B085FC40" box="[1371,1436,914,937]" name="Brazil" pageId="4" pageNumber="67">Brazil</collectingCountry>
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and quite different to those from the Caribbean Islands (
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<collectingCountry id="4CD208BFEE02FFF0FD38FC53B669FC25" box="[747,880,949,972]" name="Puerto Rico" pageId="4" pageNumber="67">Puerto Rico</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="7CDF1BA4EE02FFF0FF14FC31B650FBBE" pageId="4" pageNumber="67" type="distribution">
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||
<paragraph id="347A482FEE02FFF0FF14FC31B659FBF8" blockId="4.[151,1437,671,1111]" pageId="4" pageNumber="67">
|
||
<emphasis id="06B1943DEE02FFF0FF14FC31B440FC06" bold="true" box="[199,345,983,1007]" inLineHeading="true" pageId="4" pageNumber="67" reason="1">Distribution.</emphasis>
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<collectingCountry id="4CD208BFEE02FFF0FE8CFC3EB4BAFC06" box="[351,419,984,1007]" name="Brazil" pageId="4" pageNumber="67">Brazil</collectingCountry>
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||
,
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<collectingCountry id="4CD208BFEE02FFF0FE7EFC3EB700FC06" box="[429,537,984,1007]" name="Colombia" pageId="4" pageNumber="67">Colombia</collectingCountry>
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,
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<collectingCountry id="4CD208BFEE02FFF0FDF0FC3EB747FC06" box="[547,606,984,1007]" name="Cuba" pageId="4" pageNumber="67">Cuba</collectingCountry>
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,
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||
<collectingCountry id="4CD208BFEE02FFF0FDBBFC3EB651FC06" box="[616,840,984,1007]" name="Dominican Republic" pageId="4" pageNumber="67">Dominican Republic</collectingCountry>
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||
,
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<collectingCountry id="4CD208BFEE02FFF0FC80FC3EB6BEFC06" box="[851,935,984,1007]" name="Mexico" pageId="4" pageNumber="67">Mexico</collectingCountry>
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||
,
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<collectingCountry id="4CD208BFEE02FFF0FC62FC3EB12BFC06" box="[945,1074,984,1007]" name="Puerto Rico" pageId="4" pageNumber="67">Puerto Rico</collectingCountry>
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||
,
|
||
<collectingCountry id="4CD208BFEE02FFF0FBEEFC3EB19FFC06" box="[1085,1158,984,1007]" name="United States of America" pageId="4" pageNumber="67">U.S.A.</collectingCountry>
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||
(
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<collectingRegion id="F60186CDEE02FFF0FB47FC3EB1FCFC06" box="[1172,1253,984,1007]" country="United States of America" name="Florida" pageId="4" pageNumber="67">Florida</collectingRegion>
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,
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<collectingRegion id="F60186CDEE02FFF0FB3CFC3EB037FC06" box="[1263,1326,984,1007]" country="United States of America" name="Texas" pageId="4" pageNumber="67">Texas</collectingRegion>
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) (
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||
<bibRefCitation id="505435DEEE02FFF0FA97FC3EB5D6FBF8" author="Perkins" firstAuthor="Perkins" pageId="4" pageNumber="67" pagination="3 - 554" refId="ref3183" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: Classification, Biogeography and inferred Phylogeny. (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554." type="journal article" year="1980">Perkins, 1980</bibRefCitation>
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||
;
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<bibRefCitation id="505435DEEE02FFF0FF09FC1CB470FBF8" author="Hansen" box="[218,361,1018,1041]" firstAuthor="Hansen" pageId="4" pageNumber="67" refId="ref3146" refString="Hansen, M. (1998) Hydraenidae (Coleoptera). In: Hansen, M. (ed.). World Catalogue of Insects 1: Apollo Books, Stenstrup, 168 pp." type="book" year="1998">Hansen 1998</bibRefCitation>
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). New record for
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<collectingCountry id="4CD208BFEE02FFF0FDF5FC1CB78DFBF8" box="[550,660,1018,1041]" name="Venezuela" pageId="4" pageNumber="67">Venezuela</collectingCountry>
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(map:
|
||
<figureCitation id="ACFE54AAEE02FFF0FD0CFC1CB62DFBF8" box="[735,820,1018,1041]" captionStart="FIGURE 3" captionStartId="2.[151,255,1675,1699]" captionTargetBox="[151,1436,1118,1650]" captionTargetId="figure-218@2.[151,1436,1113,1653]" captionTargetPageId="2" captionText="FIGURE 3. Distribution maps in Venezuela of: A) Ochthebius (s. str.) lineatus (circles) and Ochthebius (s. str.) attritus (star); B) Gymnochthebius fossatus." pageId="4" pageNumber="67">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="347A482FEE02FFF0FF14FBFAB650FBBE" blockId="4.[151,1437,671,1111]" pageId="4" pageNumber="67">
|
||
<emphasis id="06B1943DEE02FFF0FF14FBFAB43CFBDD" bold="true" box="[199,293,1052,1076]" inLineHeading="true" pageId="4" pageNumber="67" reason="1">Habitat.</emphasis>
|
||
The single collecting event for this species in
|
||
<collectingCountry id="4CD208BFEE02FFF0FCF6FBFBB68BFBDD" box="[805,914,1053,1076]" name="Venezuela" pageId="4" pageNumber="67">Venezuela</collectingCountry>
|
||
was in a small pool in the Médanos de Coro, a large expanse of sand dunes along the northwest coast (
|
||
<figureCitation id="ACFE54AAEE02FFF0FD35FBA6B625FBBE" box="[742,828,1088,1111]" captionStart="FIGURE 4" captionStartId="3.[151,244,1984,2005]" captionTargetBox="[151,1435,470,1963]" captionTargetId="figure-182@3.[151,1436,469,1963]" captionTargetPageId="3" captionText="FIGURE 4. Photographs of selected collecting events of Ochthebiinae in Venezuela. A) Falcon State, Médanos de Coro, event VZ09- 0709-03Z; B) Sucre State: Finca Vuelta Larga, event VZ10-0129-03A; C) Guárico State: Río San Antonio, event VZ10-0204-02A." pageId="4" pageNumber="67">Fig. 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |