893 lines
85 KiB
XML
893 lines
85 KiB
XML
<document id="8D8054F12EF91B866D8E4D5AB7847E10" ID-CLB-Dataset="22593" ID-DOI="10.3897/zookeys.34.309" ID-GBIF-Dataset="09605303-c39a-4100-b5b4-fe409cccf3ab" ID-ISSN="1313–2970" ID-Zenodo-Dep="576611" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1588011467026" checkinUser="plazi" docAuthor="Gasca-Álvarez, Héctor & Amat-García, Germán" docDate="2010" docId="F34FE44E1F37FFEBCD84FB22566DFB0C" docLanguage="en" docName="ZK_article_2145.pdf" docOrigin="ZooKeys 34 (34)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@577d001f" docStyleName="zookeys.2008.journal_article" docTitle="Phileurini Burmeister 1847" docType="key" docVersion="7" lastPageNumber="176" masterDocId="0F769C361F27FFFCCD08FFFF523EFF96" masterDocTitle="Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia" masterLastPageNumber="192" masterPageNumber="153" pageNumber="169" updateTime="1698837288339" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="841CF4EF93CA248CB6ABDF81BD1B7C3C">Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia</mods:title>
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<mods:name id="6404D4753FD4D8CBF22FB244C05EB28A" type="personal">
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<mods:namePart id="04319D2C9DE12A107A87863DD00989E7">Gasca-Álvarez, Héctor</mods:namePart>
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<mods:affiliation id="4AF334612A3F07E4CAAB7B58D8360D75">Researcher Corporación Sentido Natural ,, Colombia</mods:affiliation>
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<mods:namePart id="EB0D9EA0B2DDD36F6C6D9EB6BFEB4337">Amat-García, Germán</mods:namePart>
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<mods:affiliation id="6BE0BF77BB0DFE3C24211A0D1AAAC5DB">Universidad Nacional de Colombia, Bogotá ,, Colombia</mods:affiliation>
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<mods:title id="C20AF7FC9AE688802269DD81BF0A8CC5">ZooKeys</mods:title>
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<mods:part id="4A9FA4CC356BCE2B193D9A2FBD48409B">
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<mods:date id="0DF341429CDA1EAE68B6962C6A64C5D4">2010</mods:date>
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<mods:number id="29A24BE64CA8B489F83F088F8E248AD4">2010-01-28</mods:number>
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<mods:number id="4708F55CE0A256F5E3677D60C19737D5">34</mods:number>
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<mods:identifier id="E8F59B1669898D67FEEE3CBE3DBD0A1A" type="DOI">10.3897/zookeys.34.309</mods:identifier>
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<mods:identifier id="A5A732D772992A71937BF9507F6389FF" type="ISSN">1313–2970</mods:identifier>
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<treatment id="F34FE44E1F37FFEBCD84FB22566DFB0C" ID-DOI="http://doi.org/10.5281/zenodo.3789632" ID-GBIF-Taxon="163794550" ID-Zenodo-Dep="3789632" LSID="urn:lsid:plazi:treatment:F34FE44E1F37FFEBCD84FB22566DFB0C" httpUri="http://treatment.plazi.org/id/F34FE44E1F37FFEBCD84FB22566DFB0C" lastPageId="23" lastPageNumber="176" pageId="16" pageNumber="169">
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<subSubSection id="33FC06D31F37FFEACD84FB2251AAFE7C" lastPageId="22" lastPageNumber="175" pageId="16" pageNumber="169" type="nomenclature">
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<paragraph id="7B5955581F37FFECCD84FB225115FB6E" blockId="16.[140,811,1245,1272]" box="[140,811,1245,1272]" pageId="16" pageNumber="169">
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<heading id="2011E2341F37FFECCD84FB225115FB6E" bold="true" box="[140,811,1245,1272]" fontSize="11" level="1" pageId="16" pageNumber="169" reason="1">
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<emphasis id="4992894A1F37FFECCD84FB225115FB6E" bold="true" box="[140,811,1245,1272]" pageId="16" pageNumber="169">
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Key to the genera of adult
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<taxonomicName id="BCE62EDB1F37FFECCCFFFB225045FB6E" ID-CoL="8RYTV" authorityName="Burmeister" authorityYear="1847" box="[503,635,1245,1272]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="169" phylum="Arthropoda" rank="tribe" tribe="Phileurini">Phileurini</taxonomicName>
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of
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<collectingCountry id="03F115C81F37FFECCFADFB225115FB6E" box="[677,811,1245,1272]" name="Colombia" pageId="16" pageNumber="169">Colombia</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="33FC06D31F37FFEDCD84FACF566DFBC5" lastPageId="17" lastPageNumber="170" pageId="16" pageNumber="169" type="key">
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<keyStep id="C0124DFD1F37FFECCD84FACF566DFAFB" pageId="16" pageNumber="169">
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<paragraph id="7B5955581F37FFECCD84FACF566DFADC" blockId="16.[140,1108,1328,1672]" box="[140,1107,1328,1355]" pageId="16" pageNumber="169">
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<keyLead id="C017F66D1F37FFECCD84FACF566DFADC" box="[140,1107,1328,1355]" pageId="16" pageNumber="169">
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1. Outer side of mandibles tridentate.............................................................
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<emphasis id="4992894A1F37FFECC94CFACF566DFADC" bold="true" box="[1092,1107,1328,1354]" pageId="16" pageNumber="169">2</emphasis>
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</keyLead>
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</paragraph>
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||
<paragraph id="7B5955581F37FFECCD84FAAB566DFAFB" blockId="16.[140,1108,1328,1672]" box="[140,1107,1363,1390]" pageId="16" pageNumber="169">
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||
<keyLead id="C017F66D1F37FFECCD84FAAB566DFAFB" box="[140,1107,1363,1390]" pageId="16" pageNumber="169">
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||
– Outer side of mandibles simply curved ......................................................
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||
<emphasis id="4992894A1F37FFECC94CFAAC566DFAFB" bold="true" box="[1092,1107,1363,1389]" pageId="16" pageNumber="169">3</emphasis>
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||
</keyLead>
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||
</paragraph>
|
||
</keyStep>
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||
<keyStep id="C0124DFD1F37FFECCD84FA88566AF9D7" pageId="16" pageNumber="169">
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||
<paragraph id="7B5955581F37FFECCD84FA88566DFA41" blockId="16.[140,1108,1328,1672]" pageId="16" pageNumber="169">
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||
<keyLead id="C017F66D1F37FFECCD84FA88566DFA41" pageId="16" pageNumber="169">
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2. Body dorsoventrally flattened. Frons with 2 tubercles. Protibia with 3 small teeth. Apical margin of metatibia with 3 teeth. Pygidium strongly punctate... ....................................................................................
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<taxonomicName id="BCE62EDB1F37FFECCE52FA43566DFA41" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[858,1107,1468,1495]" class="Insecta" family="Dynastidae" genus="Metaphileurus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="169" phylum="Arthropoda" rank="genus">
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<emphasis id="4992894A1F37FFECCE52FA43566DFA41" bold="true" box="[858,1107,1468,1495]" pageId="16" pageNumber="169">
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<emphasis id="4992894A1F37FFECCE52FA43563BFA41" bold="true" box="[858,1029,1468,1495]" italics="true" pageId="16" pageNumber="169">Metaphileurus</emphasis>
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Kolbe
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</emphasis>
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||
</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="7B5955581F37FFECCD84FA1E566AF9D7" blockId="16.[140,1108,1328,1672]" pageId="16" pageNumber="169">
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<keyLead id="C017F66D1F37FFECCD84FA1E566AF9D7" pageId="16" pageNumber="169">
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– Body not dorsoventrally flattened. Frons with tubercle or posteriorly curved horn. Protibia with 4 teeth, basal tooth reduced. Apical margin of metatibia with 5 teeth. Pygidium convex, finely wrinkled.......
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<taxonomicName id="BCE62EDB1F37FFECCE4CF9D9566AF9D7" authority="Kolbe" authorityName="Kolbe" box="[836,1108,1574,1601]" class="Insecta" family="Dynastidae" genus="Oryctophileurus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="169" phylum="Arthropoda" rank="genus">
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||
<emphasis id="4992894A1F37FFECCE4CF9D9566AF9D7" bold="true" box="[836,1108,1574,1601]" pageId="16" pageNumber="169">
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||
<emphasis id="4992894A1F37FFECCE4CF9D9563BF9D7" bold="true" box="[836,1029,1574,1601]" italics="true" pageId="16" pageNumber="169">Oryctophileurus</emphasis>
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Kolbe
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="C0124DFD1F37FFECCD84F9B5566DF911" pageId="16" pageNumber="169">
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<paragraph id="7B5955581F37FFECCD84F9B5566DF9F5" blockId="16.[140,1108,1328,1672]" box="[140,1107,1609,1636]" pageId="16" pageNumber="169">
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||
<keyLead id="C017F66D1F37FFECCD84F9B5566DF9F5" box="[140,1107,1609,1636]" pageId="16" pageNumber="169">
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||
3. Apex of metatibia truncate, without teeth ..................................................
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<emphasis id="4992894A1F37FFECC94CF9B6566DF9F5" bold="true" box="[1092,1107,1609,1635]" pageId="16" pageNumber="169">4</emphasis>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph id="7B5955581F37FFECCD84F991566DF911" blockId="16.[140,1108,1328,1672]" box="[140,1107,1645,1672]" pageId="16" pageNumber="169">
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||
<keyLead id="C017F66D1F37FFECCD84F991566DF911" box="[140,1107,1645,1672]" pageId="16" pageNumber="169">
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||
– Apex of metatibia acute or with 2–3 teeth..................................................
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||
<emphasis id="4992894A1F37FFECC94CF992566DF911" bold="true" box="[1092,1107,1645,1671]" pageId="16" pageNumber="169">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
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||
<keyStep id="C0124DFD1F36FFEDCD84FF58566AFEBC" pageId="17" pageNumber="170">
|
||
<paragraph id="7B5955581F36FFEDCD84FF58566DFF72" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
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||
<keyLead id="C017F66D1F36FFEDCD84FF58566DFF72" pageId="17" pageNumber="170">
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4. Frons with small tubercle. Prosternal process moderately elevated, anterior region convex, densely setose. Protibia quadridentate.....
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<taxonomicName id="BCE62EDB1F36FFEDCE59FF36566DFF72" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[849,1107,201,228]" class="Insecta" family="Dynastidae" genus="Haplophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="4992894A1F36FFEDCE59FF36566DFF72" bold="true" box="[849,1107,201,228]" pageId="17" pageNumber="170">
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||
<emphasis id="4992894A1F36FFEDCE59FF365639FF72" bold="true" box="[849,1031,201,228]" italics="true" pageId="17" pageNumber="170">Haplophileurus</emphasis>
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Kolbe
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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||
<paragraph id="7B5955581F36FFEDCD84FF11566AFEBC" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
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||
<keyLead id="C017F66D1F36FFEDCD84FF11566AFEBC" pageId="17" pageNumber="170">
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||
– Frons with 2 tubercles or horns placed near lateral margins (
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||
<figureCitation id="E3DD49DD1F36FFEDCE9CFF1251D6FE91" box="[916,1000,237,263]" captionStart="Figure 28" captionStartId="17.[140,212,1654,1676]" captionText="Figure 28. Head of Archophileurus sp. (scale bar: 2 mm)." pageId="17" pageNumber="170">Fig. 28</figureCitation>
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). Prosternal process variable, punctate. Protibia tridentate......
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<taxonomicName id="BCE62EDB1F36FFEDCE58FEEF566AFEBC" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[848,1108,272,299]" class="Insecta" family="Dynastidae" genus="Archophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
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||
<emphasis id="4992894A1F36FFEDCE58FEEF566AFEBC" bold="true" box="[848,1108,272,299]" pageId="17" pageNumber="170">
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<emphasis id="4992894A1F36FFEDCE58FEEF563AFEBD" bold="true" box="[848,1028,272,299]" italics="true" pageId="17" pageNumber="170">Archophileurus</emphasis>
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Kolbe
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="C0124DFD1F36FFEDCD84FECC566DFEE7" pageId="17" pageNumber="170">
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||
<paragraph id="7B5955581F36FFEDCD84FECC566DFEDB" blockId="17.[140,1108,167,1108]" box="[140,1107,307,334]" pageId="17" pageNumber="170">
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||
<keyLead id="C017F66D1F36FFEDCD84FECC566DFEDB" box="[140,1107,307,334]" pageId="17" pageNumber="170">
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||
5. Apex of metatibia tridentate.......................................................................
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<emphasis id="4992894A1F36FFEDC94CFECC566DFEDB" bold="true" box="[1092,1107,307,333]" pageId="17" pageNumber="170">6</emphasis>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph id="7B5955581F36FFEDCD84FEA8566DFEE7" blockId="17.[140,1108,167,1108]" box="[140,1107,343,369]" pageId="17" pageNumber="170">
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||
<keyLead id="C017F66D1F36FFEDCD84FEA8566DFEE7" box="[140,1107,343,369]" pageId="17" pageNumber="170">
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||
– Apex of metatibia with acute tooth or with 1–2 large teeth ........................
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||
<emphasis id="4992894A1F36FFEDC94CFEA8566DFEE7" bold="true" box="[1092,1107,343,369]" pageId="17" pageNumber="170">7</emphasis>
|
||
</keyLead>
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||
</paragraph>
|
||
</keyStep>
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||
<keyStep id="C0124DFD1F36FFEDCD84FE85566DFDB7" pageId="17" pageNumber="170">
|
||
<paragraph id="7B5955581F36FFEDCD84FE85566DFE4C" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FE85566DFE4C" pageId="17" pageNumber="170">
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||
6.
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<collectingRegion id="B9229BBA1F36FFEDCDFEFE855364FE02" box="[246,346,378,404]" country="Myanmar" name="Chin" pageId="17" pageNumber="170">Mentum</collectingRegion>
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trapezoidal short. Pronotum with longitudinal, broad furrow, with or without small tubercle in middle of apical margin. Protibia with 3 teeth ... ...................................................................................
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<taxonomicName id="BCE62EDB1F36FFEDCE5FFE3F566DFE4C" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[855,1107,448,475]" class="Insecta" family="Dynastidae" genus="Amblyoproctus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
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||
<emphasis id="4992894A1F36FFEDCE5FFE3F566DFE4C" bold="true" box="[855,1107,448,475]" pageId="17" pageNumber="170">
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||
<emphasis id="4992894A1F36FFEDCE5FFE3F563BFE4D" bold="true" box="[855,1029,448,475]" italics="true" pageId="17" pageNumber="170">Amblyoproctus</emphasis>
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Kolbe
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||
</emphasis>
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph id="7B5955581F36FFEDCD84FE1B566DFDB7" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FE1B566DFDB7" pageId="17" pageNumber="170">
|
||
–
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||
<collectingRegion id="B9229BBA1F36FFEDCDFEFE1B5366FE68" box="[246,344,484,510]" country="Myanmar" name="Chin" pageId="17" pageNumber="170">Mentum</collectingRegion>
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||
trapezoidal elongated. Pronotum with anterior margin rugose, convex, or with pronotal cavity and 4 subequal tubercles ..........
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<taxonomicName id="BCE62EDB1F36FFEDCE5CFDF9566DFDB7" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[852,1107,518,545]" class="Insecta" family="Dynastidae" genus="Homophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
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||
<emphasis id="4992894A1F36FFEDCE5CFDF9566DFDB7" bold="true" box="[852,1107,518,545]" pageId="17" pageNumber="170">
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||
<emphasis id="4992894A1F36FFEDCE5CFDF95639FDB7" bold="true" box="[852,1031,518,545]" italics="true" pageId="17" pageNumber="170">Homophileurus</emphasis>
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Kolbe
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</emphasis>
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||
</taxonomicName>
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="C0124DFD1F36FFEDCD84FDD5566DFD65" pageId="17" pageNumber="170">
|
||
<paragraph id="7B5955581F36FFEDCD84FDD5566AFD1C" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
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||
<keyLead id="C017F66D1F36FFEDCD84FDD5566AFD1C" pageId="17" pageNumber="170">
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||
7. Pronotum convex, without longitudinal furrow (
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<figureCitation id="E3DD49DD1F36FFEDCE3EFDD551AEFDD2" box="[822,912,554,580]" captionStart="Figure 29" captionStartId="18.[140,212,706,728]" captionTargetBox="[405,849,207,626]" captionTargetId="figure@18.[371,876,162,686]" captionTargetPageId="18" captionText="Figure 29. Pronotum of Palaeophileurus sclateri (Bates) (scale bar: 5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770656" httpUri="https://zenodo.org/record/3770656/files/figure.png" pageId="17" pageNumber="170">Fig. 29</figureCitation>
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). Elytra opaque, with rows of punctures slightly impressed or absent. First tarsomere of metatibia without apical, spiniform process (
|
||
<figureCitation id="E3DD49DD1F36FFEDCFA5FD8F50C1FD1C" box="[685,767,624,650]" captionStart="Figure 30" captionStartId="18.[140,212,1155,1177]" captionTargetBox="[272,992,882,1069]" captionTargetId="figure@18.[225,1023,829,1125]" captionTargetPageId="18" captionText="Figure 30. Metatarsus of Palaeophileurus sclateri (Bates) (scale bar: 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770658" httpUri="https://zenodo.org/record/3770658/files/figure.png" pageId="17" pageNumber="170">Fig. 30</figureCitation>
|
||
)........
|
||
<taxonomicName id="BCE62EDB1F36FFEDCE4EFD8F566AFD1C" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[838,1108,624,651]" class="Insecta" family="Dynastidae" genus="Palaeophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F36FFEDCE4EFD8F566AFD1C" bold="true" box="[838,1108,624,651]" pageId="17" pageNumber="170">
|
||
<emphasis id="4992894A1F36FFEDCE4EFD8F563BFD1D" bold="true" box="[838,1029,624,651]" italics="true" pageId="17" pageNumber="170">Palaeophileurus</emphasis>
|
||
Kolbe
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F36FFEDCD84FD6B566DFD65" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FD6B566DFD65" pageId="17" pageNumber="170">
|
||
– Pronotum with longitudinal furrow, with or without tubercles or fovea. Elytra shiny, with impressed rows of punctures. First tarsomere of metatibia with apical, spiniform process (Fig. 31)......................................................
|
||
<emphasis id="4992894A1F36FFEDC94CFD26566DFD65" bold="true" box="[1092,1107,729,755]" pageId="17" pageNumber="170">8</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F36FFEDCD84FD02566DFC51" pageId="17" pageNumber="170">
|
||
<paragraph id="7B5955581F36FFEDCD84FD02566AFCCB" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FD02566AFCCB" pageId="17" pageNumber="170">
|
||
8. Tubercles or horns of frons placed near center of head (
|
||
<figureCitation id="E3DD49DD1F36FFEDCE7CFD0251F4FC81" box="[884,970,765,791]" captionStart="Figure 32" captionStartId="19.[140,212,618,640]" captionTargetBox="[481,783,185,554]" captionTargetId="figure@19.[443,804,165,598]" captionTargetPageId="19" captionText="Figure 32. Head and pronotum of Hemiphileurus sp. (scale bar: 5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770660" httpUri="https://zenodo.org/record/3770660/files/figure.png" pageId="17" pageNumber="170">Fig. 32</figureCitation>
|
||
). Pronotum without fovea, longitudinal furrow usually incomplete (not reaching anterior margin).....................................................................
|
||
<taxonomicName id="BCE62EDB1F36FFEDCE5BFCBC566AFCCB" authority="Kolbe" authorityName="Kolbe" authorityYear="1910" box="[851,1108,835,862]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F36FFEDCE5BFCBC566AFCCB" bold="true" box="[851,1108,835,862]" pageId="17" pageNumber="170">
|
||
<emphasis id="4992894A1F36FFEDCE5BFCBC563BFCC8" bold="true" box="[851,1029,835,862]" italics="true" pageId="17" pageNumber="170">Hemiphileurus</emphasis>
|
||
Kolbe
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F36FFEDCD84FC98566DFC51" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FC98566DFC51" pageId="17" pageNumber="170">
|
||
– Tubercles or horns of frons placed near lateral margin of head. Pronotum with anterior, wide fovea, longitudinal furrow usually complete, anterior margin with tubercle at center (
|
||
<figureCitation id="E3DD49DD1F36FFEDCF52FC52509BFC51" box="[602,677,941,967]" captionStart="Figure 33" captionStartId="19.[140,212,1069,1091]" captionTargetBox="[426,826,686,1006]" captionTargetId="figure@19.[402,846,650,1049]" captionTargetPageId="19" captionText="Figure 33. Pronotum of Phileurus didymus (Linnaeus) (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770662" httpUri="https://zenodo.org/record/3770662/files/figure.png" pageId="17" pageNumber="170">Fig 33</figureCitation>
|
||
) ......................................................
|
||
<emphasis id="4992894A1F36FFEDC94CFC52566DFC51" bold="true" box="[1092,1107,941,967]" pageId="17" pageNumber="170">9</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F36FFEDCD84FC2F566DFBC5" pageId="17" pageNumber="170">
|
||
<paragraph id="7B5955581F36FFEDCD84FC2F566DFB9B" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FC2F566DFB9B" pageId="17" pageNumber="170">
|
||
9. Prosternal process triangular, with angulate projection or large conical knob of posterior surface..................................................
|
||
<taxonomicName id="BCE62EDB1F36FFEDCE48FC0C566DFB9B" authority="Endrodi" authorityName="Endrodi" authorityYear="1978" box="[832,1107,1010,1038]" class="Insecta" family="Dynastidae" genus="Paraphileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F36FFEDCE48FC0C566DFB9B" bold="true" box="[832,1107,1010,1038]" pageId="17" pageNumber="170">
|
||
<emphasis id="4992894A1F36FFEDCE48FC0C51D7FB98" bold="true" box="[832,1001,1011,1038]" italics="true" pageId="17" pageNumber="170">Paraphileurus</emphasis>
|
||
Endrődi
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F36FFEDCD84FBE8566DFBC5" blockId="17.[140,1108,167,1108]" pageId="17" pageNumber="170">
|
||
<keyLead id="C017F66D1F36FFEDCD84FBE8566DFBC5" pageId="17" pageNumber="170">
|
||
– Prosternal process variable, never with angulate projection........................... ......................................................................................
|
||
<taxonomicName id="BCE62EDB1F36FFEDCE7EFBC6566DFBC5" authority="Latreille" authorityName="Latreille" authorityYear="1807" box="[886,1107,1081,1108]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F36FFEDCE7EFBC6566DFBC5" bold="true" box="[886,1107,1081,1108]" pageId="17" pageNumber="170">
|
||
<emphasis id="4992894A1F36FFEDCE7EFBC651D7FBC2" bold="true" box="[886,1001,1081,1108]" italics="true" pageId="17" pageNumber="170">Phileurus</emphasis>
|
||
Latreille
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
<caption id="2F9905D01F36FFEDCD84F989509AF91A" box="[140,676,1654,1676]" pageId="17" pageNumber="170">
|
||
<paragraph id="7B5955581F36FFEDCD84F989509AF91A" blockId="17.[140,676,1654,1676]" box="[140,676,1654,1676]" pageId="17" pageNumber="170">
|
||
<emphasis id="4992894A1F36FFEDCD84F98952C5F91A" bold="true" box="[140,251,1654,1676]" pageId="17" pageNumber="170">Figure 28.</emphasis>
|
||
Head of
|
||
<taxonomicName id="BCE62EDB1F36FFEDCC5AF98953E9F91D" authorityName="Kolbe" authorityYear="1910" box="[338,471,1654,1675]" class="Insecta" family="Dynastidae" genus="Archophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F36FFEDCC5AF98953E9F91D" box="[338,471,1654,1675]" italics="true" pageId="17" pageNumber="170">Archophileurus</emphasis>
|
||
</taxonomicName>
|
||
sp. (scale bar: 2 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2F9905D01F35FFEECD84FD3D517EFD41" ID-DOI="http://doi.org/10.5281/zenodo.3770656" ID-Zenodo-Dep="3770656" box="[140,832,706,728]" httpUri="https://zenodo.org/record/3770656/files/figure.png" pageId="18" pageNumber="171" startId="18.[140,212,706,728]" targetBox="[405,849,207,626]" targetPageId="18">
|
||
<paragraph id="7B5955581F35FFEECD84FD3D517EFD41" blockId="18.[140,832,706,728]" box="[140,832,706,728]" pageId="18" pageNumber="171">
|
||
<emphasis id="4992894A1F35FFEECD84FD3D52C5FD4E" bold="true" box="[140,251,706,728]" pageId="18" pageNumber="171">Figure 29.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName id="BCE62EDB1F35FFEECC76FD3D50ACFD4E" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[382,658,706,728]" class="Insecta" family="Dynastidae" genus="Palaeophileurus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="171" phylum="Arthropoda" rank="species" species="sclateri">
|
||
<emphasis id="4992894A1F35FFEECC76FD3D5072FD41" box="[382,588,706,727]" italics="true" pageId="18" pageNumber="171">Palaeophileurus sclateri</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
(scale bar: 5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2F9905D01F35FFEECD84FB7C5178FB0E" ID-DOI="http://doi.org/10.5281/zenodo.3770658" ID-Zenodo-Dep="3770658" box="[140,838,1155,1177]" httpUri="https://zenodo.org/record/3770658/files/figure.png" pageId="18" pageNumber="171" startId="18.[140,212,1155,1177]" targetBox="[272,992,882,1069]" targetPageId="18">
|
||
<paragraph id="7B5955581F35FFEECD84FB7C5178FB0E" blockId="18.[140,838,1155,1177]" box="[140,838,1155,1177]" pageId="18" pageNumber="171">
|
||
<emphasis id="4992894A1F35FFEECD84FB7C52C5FB0F" bold="true" box="[140,251,1155,1177]" pageId="18" pageNumber="171">Figure 30.</emphasis>
|
||
Metatarsus of
|
||
<taxonomicName id="BCE62EDB1F35FFEECC8CFB7C50A6FB0F" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[388,664,1155,1177]" class="Insecta" family="Dynastidae" genus="Palaeophileurus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="171" phylum="Arthropoda" rank="species" species="sclateri">
|
||
<emphasis id="4992894A1F35FFEECC8CFB7C506CFB0E" box="[388,594,1155,1176]" italics="true" pageId="18" pageNumber="171">Palaeophileurus sclateri</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
(scale bar: 2 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7B5955581F35FFEECD84F989517EF91A" blockId="18.[140,832,1654,1676]" box="[140,832,1654,1676]" pageId="18" pageNumber="171">
|
||
<emphasis id="4992894A1F35FFEECD84F98952C5F91A" bold="true" box="[140,251,1654,1676]" pageId="18" pageNumber="171">
|
||
<figureCitation id="E3DD49DD1F35FFEECD84F98952CEF91A" box="[140,240,1654,1676]" captionStart="Figure 3" captionStartId="5.[140,212,965,987]" captionTargetBox="[499,717,649,885]" captionTargetId="figure@5.[475,772,625,945]" captionTargetPageId="5" captionText="Figure 3. Right mandible of Lycomedes sp. (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770633" httpUri="https://zenodo.org/record/3770633/files/figure.png" pageId="18" pageNumber="171">Figure 3</figureCitation>
|
||
|.
|
||
</emphasis>
|
||
Metatarsus of
|
||
<taxonomicName id="BCE62EDB1F35FFEECC8CF98950AFF91A" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[388,657,1654,1676]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="171" phylum="Arthropoda" rank="species" species="didymus">
|
||
<emphasis id="4992894A1F35FFEECC8CF9895016F91D" box="[388,552,1654,1675]" italics="true" pageId="18" pageNumber="171">Phileurus didymus</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
(scale bar:
|
||
<quantity id="BC1EF8BD1F35FFEECFF1F988510BF91A" box="[761,821,1655,1676]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="171" unit="mm" value="2.0">2 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<caption id="2F9905D01F34FFEFCD84FD95510EFDE9" ID-DOI="http://doi.org/10.5281/zenodo.3770660" ID-Zenodo-Dep="3770660" box="[140,816,618,640]" httpUri="https://zenodo.org/record/3770660/files/figure.png" pageId="19" pageNumber="172" startId="19.[140,212,618,640]" targetBox="[481,783,185,554]" targetPageId="19">
|
||
<paragraph id="7B5955581F34FFEFCD84FD95510EFDE9" blockId="19.[140,816,618,640]" box="[140,816,618,640]" pageId="19" pageNumber="172">
|
||
<emphasis id="4992894A1F34FFEFCD84FD9552C5FD16" bold="true" box="[140,251,618,640]" pageId="19" pageNumber="172">Figure 32.</emphasis>
|
||
Head and pronotum of
|
||
<taxonomicName id="BCE62EDB1F34FFEFCCD7FD95505DFDE9" authorityName="Kolbe" authorityYear="1910" box="[479,611,618,639]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F34FFEFCCD7FD95505DFDE9" box="[479,611,618,639]" italics="true" pageId="19" pageNumber="172">Hemiphileurus</emphasis>
|
||
</taxonomicName>
|
||
sp. (scale bar: 5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2F9905D01F34FFEFCD84FBD2510EFBD5" ID-DOI="http://doi.org/10.5281/zenodo.3770662" ID-Zenodo-Dep="3770662" box="[140,816,1069,1091]" httpUri="https://zenodo.org/record/3770662/files/figure.png" pageId="19" pageNumber="172" startId="19.[140,212,1069,1091]" targetBox="[426,826,686,1006]" targetPageId="19">
|
||
<paragraph id="7B5955581F34FFEFCD84FBD2510EFBD5" blockId="19.[140,816,1069,1091]" box="[140,816,1069,1091]" pageId="19" pageNumber="172">
|
||
<emphasis id="4992894A1F34FFEFCD84FBD252C5FBD5" bold="true" box="[140,251,1069,1091]" pageId="19" pageNumber="172">Figure 33.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName id="BCE62EDB1F34FFEFCC76FBD250B5FBD5" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[382,651,1069,1091]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="172" phylum="Arthropoda" rank="species" species="didymus">
|
||
<emphasis id="4992894A1F34FFEFCC76FBD253ECFBD4" box="[382,466,1069,1090]" italics="true" pageId="19" pageNumber="172">Phileurus</emphasis>
|
||
<emphasis id="4992894A1F34FFEFCCD0FBD2501CFBD4" box="[472,546,1069,1090]" italics="true" pageId="19" pageNumber="172">didymus</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
(scale bar: 1 cm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7B5955581F34FFEFCD84FB6251D8FB2E" blockId="19.[140,998,1181,1208]" box="[140,998,1181,1208]" pageId="19" pageNumber="172">
|
||
<emphasis id="4992894A1F34FFEFCD84FB6251D8FB2E" bold="true" box="[140,998,1181,1208]" pageId="19" pageNumber="172">
|
||
Clave para los adultos de las tribus de
|
||
<taxonomicName id="BCE62EDB1F34FFEFCF9EFB625110FB2E" box="[662,814,1181,1208]" class="Insecta" family="Dynastinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="172" phylum="Arthropoda" rank="family">Dynastinae</taxonomicName>
|
||
de
|
||
<collectingCountry id="03F115C81F34FFEFCE68FB6251D8FB2E" box="[864,998,1181,1208]" name="Colombia" pageId="19" pageNumber="172">Colombia</collectingCountry>
|
||
</emphasis>
|
||
</paragraph>
|
||
<subSubSection id="33FC06D31F34FFE8CD84FB15566DFD46" lastPageId="20" lastPageNumber="173" pageId="19" pageNumber="172" type="key">
|
||
<keyStep id="C0124DFD1F34FFEFCD84FB15566DFAFB" pageId="19" pageNumber="172">
|
||
<paragraph id="7B5955581F34FFEFCD84FB15566DFAB1" blockId="19.[140,1108,1258,1636]" pageId="19" pageNumber="172">
|
||
<keyLead id="C017F66D1F34FFEFCD84FB15566DFAB1" pageId="19" pageNumber="172">
|
||
1.
|
||
<collectingRegion id="B9229BBA1F34FFEFCDFEFB15536CFA92" box="[246,338,1258,1284]" country="Myanmar" name="Chin" pageId="19" pageNumber="172">Mentón</collectingRegion>
|
||
ensanchado o expandido, cubriendo la base de los palpos labiales. Cuerpo mas o menos dorsoventralmente aplanado.......................
|
||
<taxonomicName id="BCE62EDB1F34FFEFCED1FAF2566DFAB1" ID-CoL="8RYTV" authorityName="Burmeister" authorityYear="1847" box="[985,1107,1293,1319]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="172" phylum="Arthropoda" rank="tribe" tribe="Phileurini">
|
||
<emphasis id="4992894A1F34FFEFCED1FAF2566DFAB1" bold="true" box="[985,1107,1293,1319]" pageId="19" pageNumber="172">Phileurini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F34FFEFCD84FACE566DFAFB" blockId="19.[140,1108,1258,1636]" pageId="19" pageNumber="172">
|
||
<keyLead id="C017F66D1F34FFEFCD84FACE566DFAFB" pageId="19" pageNumber="172">
|
||
–
|
||
<collectingRegion id="B9229BBA1F34FFEFCDFEFACF536CFADC" box="[246,338,1328,1354]" country="Myanmar" name="Chin" pageId="19" pageNumber="172">Mentón</collectingRegion>
|
||
estrecho o angosto, sin cubrir la base de los palpos labiales. Cuerpo no dorsoventralmente aplanado .................................................................
|
||
<emphasis id="4992894A1F34FFEFC94CFAAC566DFAFB" bold="true" box="[1092,1107,1363,1389]" pageId="19" pageNumber="172">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F34FFEFCD84FA88566DF9F5" pageId="19" pageNumber="172">
|
||
<paragraph id="7B5955581F34FFEFCD84FA88566AFA6C" blockId="19.[140,1108,1258,1636]" pageId="19" pageNumber="172">
|
||
<keyLead id="C017F66D1F34FFEFCD84FA88566AFA6C" pageId="19" pageNumber="172">
|
||
2. Cabeza y pronoto convexos, sin cuernos, tubérculos o fóveas, en ambos sexos, o con tubérculos inconspicuos detrás de la sutura fronto-clipeal. Machos con las uñas anteriores más desarrolladas en muchas especies. Tarsos de todas las piernas generalmente largos y cilíndricos..............................
|
||
<taxonomicName id="BCE62EDB1F34FFEFCEA9FA1F566AFA6C" ID-CoL="8RYSP" box="[929,1108,1504,1530]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="172" phylum="Arthropoda" rank="tribe" tribe="Cyclocephalini">
|
||
<emphasis id="4992894A1F34FFEFCEA9FA1F566AFA6C" bold="true" box="[929,1108,1504,1530]" pageId="19" pageNumber="172">Cyclocephalini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F34FFEFCD84F9FB566DF9F5" blockId="19.[140,1108,1258,1636]" pageId="19" pageNumber="172">
|
||
<keyLead id="C017F66D1F34FFEFCD84F9FB566DF9F5" pageId="19" pageNumber="172">
|
||
– Cabeza y/o pronoto con cuernos, tubérculos o fóveas en ambos sexos. Machos con uñas anteriores de tamaño variable, desarrollados o no. Protarsos y mesotarsos generalmente cortos y triangulares............................................
|
||
<emphasis id="4992894A1F34FFEFC94CF9B6566DF9F5" bold="true" box="[1092,1107,1609,1635]" pageId="19" pageNumber="172">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F33FFE8CD84FF58566DFEE6" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FF58566DFE91" blockId="20.[140,1108,167,721]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FF58566DFE91" pageId="20" pageNumber="173">
|
||
3. Tarsos anteriores de los machos más largos que las tibias respectivas. Segmentos tarsales posteriores generalmente cilíndricos, segmento basal con fuerte espina apical.................................................................................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CED4FF12566DFE91" ID-CoL="628CX" authorityName="MacLeay" authorityYear="1819" box="[988,1107,237,263]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="tribe" tribe="Dynastini">
|
||
<emphasis id="4992894A1F33FFE8CED4FF12566DFE91" bold="true" box="[988,1107,237,263]" pageId="20" pageNumber="173">Dynastini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84FEEE566DFEE6" blockId="20.[140,1108,167,721]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FEEE566DFEE6" pageId="20" pageNumber="173">
|
||
– Tarsos anteriores de los machos más cortos o de igual tamaño que las tibias respectivas. Segmentos tarsales expandidos en forma de triángulo, segmento basal de forma variable...............................................................................
|
||
<emphasis id="4992894A1F33FFE8C94CFEA9566DFEE6" bold="true" box="[1092,1107,342,368]" pageId="20" pageNumber="173">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F33FFE8CD84FE85566DFDB7" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FE85566DFE21" blockId="20.[140,1108,167,721]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FE85566DFE21" pageId="20" pageNumber="173">
|
||
4. Élitros tomentosos o irregularmente punteados. Puntuaciones ligeramente marcadas. Cuerpo ligeramente deprimido.............................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CEA0FE62566DFE21" authorityName="Burmeister" authorityYear="1847" box="[936,1107,413,439]" class="Insecta" family="Dynastidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="178" phylum="Arthropoda" rank="tribe" tribe="Agaocephalini">
|
||
<emphasis id="4992894A1F33FFE8CEA0FE62566DFE21" bold="true" box="[936,1107,413,439]" pageId="20" pageNumber="173">Agaocephalini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84FE3E566DFDB7" blockId="20.[140,1108,167,721]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FE3E566DFDB7" pageId="20" pageNumber="173">
|
||
– Élitros lisos o con puntuaciones dispuestas en hileras distinguibles (excepto en algunas especies de
|
||
<taxonomicName id="BCE62EDB1F33FFE8CF01FE1C5091FE6B" authorityName="Burmeister" authorityYear="1847" box="[521,687,483,509]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F33FFE8CF01FE1C5091FE6B" box="[521,687,483,509]" italics="true" pageId="20" pageNumber="173">Heterogomphus</emphasis>
|
||
</taxonomicName>
|
||
). Puntuaciones fuerte o moderadamente marcadas. Cuerpo no deprimido, robusto........................................
|
||
<emphasis id="4992894A1F33FFE8C94CFDF8566DFDB7" bold="true" box="[1092,1107,519,545]" pageId="20" pageNumber="173">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F33FFE8CD84FDD5566DFD46" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FDD5566DFDF1" blockId="20.[140,1108,167,721]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FDD5566DFDF1" pageId="20" pageNumber="173">
|
||
5. Ápice de las metatibias generalmente truncado, liso o ligeramente prolongado (Fig 1). Dimorfismo sexual poco acentuado....................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CEBDFDB2566DFDF1" ID-CoL="8RYTT" authorityName="Mulsant" authorityYear="1842" box="[949,1107,589,615]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="tribe" tribe="Pentodontini">
|
||
<emphasis id="4992894A1F33FFE8CEBDFDB2566DFDF1" bold="true" box="[949,1107,589,615]" pageId="20" pageNumber="173">Pentodontini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84FD8E566DFD46" blockId="20.[140,1108,167,721]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FD8E566DFD46" pageId="20" pageNumber="173">
|
||
– Ápice de las metatibias generalmente crenulado o com dientes agudos notórios (excepto para las espécies de
|
||
<taxonomicName id="BCE62EDB1F33FFE8CF6EFD6C5084FD3B" authorityName="Hope" authorityYear="1837" box="[614,698,659,685]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F33FFE8CF6EFD6C5084FD3B" box="[614,698,659,685]" italics="true" pageId="20" pageNumber="173">Coelosis</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="E3DD49DD1F33FFE8CFD9FD6C5130FD38" box="[721,782,659,686]" captionStart="Figure 2" captionStartId="5.[140,212,580,602]" captionTargetBox="[321,958,196,485]" captionTargetId="figure@5.[245,1002,165,551]" captionTargetPageId="5" captionText="Figure 2. Metatibia of Heterogomphus schoenherri Burmeister (scale bar: 5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770631" httpUri="https://zenodo.org/record/3770631/files/figure.png" pageId="20" pageNumber="173">Fig 2</figureCitation>
|
||
). Dimorfismo sexual evidente ....................................................................................................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CEE5FD49566DFD46" ID-CoL="8RYTF" authorityName="Mulsant & Rey" authorityYear="1842" box="[1005,1107,694,720]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="tribe" tribe="Oryctini">
|
||
<emphasis id="4992894A1F33FFE8CEE5FD49566DFD46" bold="true" box="[1005,1107,694,720]" pageId="20" pageNumber="173">Oryctini</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
<paragraph id="7B5955581F33FFE8CD84FCD051ECFCDC" blockId="20.[140,978,815,842]" box="[140,978,815,842]" pageId="20" pageNumber="173">
|
||
<emphasis id="4992894A1F33FFE8CD84FCD051ECFCDC" bold="true" box="[140,978,815,842]" pageId="20" pageNumber="173">
|
||
Clave para los géneros de
|
||
<taxonomicName id="BCE62EDB1F33FFE8CCE6FCD05091FCDC" authorityName="Burmeister" authorityYear="1847" box="[494,687,815,842]" class="Insecta" family="Dynastidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="178" phylum="Arthropoda" rank="tribe" tribe="Agaocephalini">Agaocephalini</taxonomicName>
|
||
adultos de
|
||
<collectingCountry id="03F115C81F33FFE8CE44FCD051ECFCDC" box="[844,978,815,842]" name="Colombia" pageId="20" pageNumber="173">Colombia</collectingCountry>
|
||
</emphasis>
|
||
</paragraph>
|
||
<subSubSection id="33FC06D31F33FFE9CD84FC98566AFE4C" lastPageId="21" lastPageNumber="174" pageId="20" pageNumber="173" type="key">
|
||
<keyStep id="C0124DFD1F33FFE8CD84FC98566DFC35" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FC98566DFC16" blockId="20.[140,1108,870,1636]" box="[140,1107,870,897]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FC98566DFC16" box="[140,1107,870,897]" pageId="20" pageNumber="173">
|
||
1. Superficie de los élitros tomentosa, de color pardo grisáceo ........................
|
||
<emphasis id="4992894A1F33FFE8C94CFC99566DFC16" bold="true" box="[1092,1107,870,896]" pageId="20" pageNumber="173">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84FC75566DFC35" blockId="20.[140,1108,870,1636]" box="[140,1107,905,932]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FC75566DFC35" box="[140,1107,905,932]" pageId="20" pageNumber="173">
|
||
– Superficie de los élitros glabra, irregularmente punteada.............................
|
||
<emphasis id="4992894A1F33FFE8C94CFC76566DFC35" bold="true" box="[1092,1107,905,931]" pageId="20" pageNumber="173">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F33FFE8CD84FC52566DFB0C" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FC52566DFBA6" blockId="20.[140,1108,870,1636]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FC52566DFBA6" pageId="20" pageNumber="173">
|
||
2. Lado exterior de las mandíbulas con 3 dientes (
|
||
<figureCitation id="E3DD49DD1F33FFE8CE2AFC525156FC51" box="[802,872,941,967]" captionStart="Figure 3" captionStartId="5.[140,212,965,987]" captionTargetBox="[499,717,649,885]" captionTargetId="figure@5.[475,772,625,945]" captionTargetPageId="5" captionText="Figure 3. Right mandible of Lycomedes sp. (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770633" httpUri="https://zenodo.org/record/3770633/files/figure.png" pageId="20" pageNumber="173">Fig. 3</figureCitation>
|
||
). Proceso prosternal corto o moderadamente largo. Pronoto con cuerno fuerte, corto a largo, plano, estrecho, generalmente dirigido hacia arriba, recto (
|
||
<figureCitation id="E3DD49DD1F33FFE8CE8FFC0C51F4FB98" box="[903,970,1011,1038]" captionStart="Figure 4" captionStartId="5.[140,212,1654,1676]" captionTargetBox="[371,827,1029,1631]" captionTargetId="figure@5.[345,903,1015,1634]" captionTargetPageId="5" captionText="Figure 4. Male head and pronotum (lateral view) of Lycomedes sp. (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770635" httpUri="https://zenodo.org/record/3770635/files/figure.png" pageId="20" pageNumber="173">Fig. 4</figureCitation>
|
||
) ................ ........................................................................................
|
||
<emphasis id="4992894A1F33FFE8CE8BFBE9566DFBA6" bold="true" box="[899,1107,1045,1073]" pageId="20" pageNumber="173">
|
||
<taxonomicName id="BCE62EDB1F33FFE8CE8BFBE951C1FBA7" box="[899,1023,1046,1073]" class="Insecta" family="Dynastidae" genus="Lycomedes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F33FFE8CE8BFBE951C1FBA7" bold="true" box="[899,1023,1046,1073]" italics="true" pageId="20" pageNumber="173">Lycomedes</emphasis>
|
||
</taxonomicName>
|
||
<collectingRegion id="B9229BBA1F33FFE8C90EFBEA566DFBA6" box="[1030,1107,1045,1072]" country="Germany" name="Bremen" pageId="20" pageNumber="173">Bréme</collectingRegion>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84FBC5566DFB0C" blockId="20.[140,1108,870,1636]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FBC5566DFB0C" pageId="20" pageNumber="173">
|
||
– Lado exterior de las mandíbulas con 2 dientes (
|
||
<figureCitation id="E3DD49DD1F33FFE8CE2AFBC55156FBC3" box="[802,872,1082,1109]" captionStart="Figure 5" captionStartId="6.[140,212,464,486]" captionTargetBox="[515,729,195,424]" captionTargetId="figure@6.[496,751,165,444]" captionTargetPageId="6" captionText="Figure 5. Right mandible of Spodistes sp. (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770637" httpUri="https://zenodo.org/record/3770637/files/figure.png" pageId="20" pageNumber="173">Fig. 5</figureCitation>
|
||
). Proceso prosternal ausente. Pronoto con cuerno largo, dirigido hacia delante, delgado y curvo (
|
||
<figureCitation id="E3DD49DD1F33FFE8CDF5FB7F5307FB0D" box="[253,313,1152,1179]" captionStart="Figure 6" captionStartId="6.[140,212,1043,1065]" captionTargetBox="[322,937,544,1011]" captionTargetId="figure@6.[305,943,514,1023]" captionTargetPageId="6" captionText="Figure 6. Male head and pronotum (lateral view) of Spodistes sp. (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770639" httpUri="https://zenodo.org/record/3770639/files/figure.png" pageId="20" pageNumber="173">Fig 6</figureCitation>
|
||
).........................................................................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CE54FB7F566DFB0C" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[860,1107,1152,1179]" class="Insecta" family="Dynastidae" genus="Spodistes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F33FFE8CE54FB7F566DFB0C" bold="true" box="[860,1107,1152,1179]" pageId="20" pageNumber="173">
|
||
<emphasis id="4992894A1F33FFE8CE54FB7F51F7FB0D" bold="true" box="[860,969,1152,1179]" italics="true" pageId="20" pageNumber="173">Spodistes</emphasis>
|
||
Burmeister
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F33FFE8CD84FB5B566DFAFB" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FB5B566AFAB1" blockId="20.[140,1108,870,1636]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FB5B566AFAB1" pageId="20" pageNumber="173">
|
||
3. Tibia anterior con 4 dientes, diente basal muy corto en los machos braquíceros. Cabeza con dos cuernos largos, divergentes, proyectados hacia delante; curvados hacia arriba, ápices redondeados (
|
||
<figureCitation id="E3DD49DD1F33FFE8CE4BFB1551B8FA92" box="[835,902,1258,1284]" captionStart="Figure 7" captionStartId="6.[150,222,1653,1675]" captionTargetBox="[382,861,1196,1590]" captionTargetId="figure@6.[375,873,1102,1632]" captionTargetPageId="6" captionText="Figure 7. Male head and pronotum (lateral view) of Aegopsis curvicornis Burmeister (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770641" httpUri="https://zenodo.org/record/3770641/files/figure.png" pageId="20" pageNumber="173">Fig. 7</figureCitation>
|
||
) ......................... ..................................................................................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CE6CFAF2566AFAB1" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[868,1108,1293,1319]" class="Insecta" family="Dynastidae" genus="Aegopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F33FFE8CE6CFAF2566AFAB1" bold="true" box="[868,1108,1293,1319]" pageId="20" pageNumber="173">
|
||
<emphasis id="4992894A1F33FFE8CE6CFAF251F9FAB1" bold="true" box="[868,967,1293,1319]" italics="true" pageId="20" pageNumber="173">Aegopsis</emphasis>
|
||
Burmeister
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84FACE566DFAFB" blockId="20.[140,1108,870,1636]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FACE566DFAFB" pageId="20" pageNumber="173">
|
||
– Tibia anterior con 3 dientes bien definidos. Cabeza con cuerno o grueso, curvado hacia arriba, ápice bifurcado .........................................................
|
||
<emphasis id="4992894A1F33FFE8C94CFAAC566DFAFB" bold="true" box="[1092,1107,1363,1389]" pageId="20" pageNumber="173">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F33FFE8CD84FA88566DF9F2" pageId="20" pageNumber="173">
|
||
<paragraph id="7B5955581F33FFE8CD84FA88566DFA6C" blockId="20.[140,1108,870,1636]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84FA88566DFA6C" pageId="20" pageNumber="173">
|
||
4. Machos con cuerno cefálico corto y grueso, ligeramente curvo. Pronoto con prominencia dirigida hacia delante de ápice agudo, superficie lisa. Machos sin cuernos y tubérculos en cabeza y pronoto. Cabeza y pronoto de color negro, élitros marrón rojizo...............................................
|
||
<taxonomicName id="BCE62EDB1F33FFE8CE33FA1F566DFA6C" authority="Thomson" authorityName="Thomson" authorityYear="1859" box="[827,1107,1504,1531]" class="Insecta" family="Dynastidae" genus="Mitracephala" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F33FFE8CE33FA1F566DFA6C" bold="true" box="[827,1107,1504,1531]" pageId="20" pageNumber="173">
|
||
<emphasis id="4992894A1F33FFE8CE33FA1F51E2FA6D" bold="true" box="[827,988,1504,1531]" italics="true" pageId="20" pageNumber="173">Mitracephala</emphasis>
|
||
Thomson
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F33FFE8CD84F9FB566DF9F2" blockId="20.[140,1108,870,1636]" pageId="20" pageNumber="173">
|
||
<keyLead id="C017F66D1F33FFE8CD84F9FB566DF9F2" pageId="20" pageNumber="173">
|
||
– Machos con cuerno cefálico largo, curvado hacia arriba, ápice bifurcado. Pronoto convexo o con prominencia alta. Cuerpo de color negro, marrón o con manchas oscuras.........................................................................................
|
||
<emphasis id="4992894A1F33FFE8C94CF9B5566DF9F2" bold="true" box="[1092,1107,1610,1636]" pageId="20" pageNumber="173">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F32FFE9CD84FF59566AFE4C" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FF59566DFEDB" blockId="21.[140,1108,166,475]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FF59566DFEDB" pageId="21" pageNumber="174">
|
||
5. Machos con cuerno cefálico bifurcado en el ápice. Pronoto con prominencia ancha y alta de ápice redondeado, superficie densamente punteada. Machos sin cuerno frontal. Elitros con hileras de puntuaciones fuertemente marcadas. Proceso prosternal alto, ápice dilatado. Cuerpo de color negro o marrón...... .................................................................................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE46FECC566DFEDB" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[846,1107,306,334]" class="Insecta" family="Dynastidae" genus="Horridocalia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE46FECC566DFEDB" bold="true" box="[846,1107,306,334]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE46FECC51D7FED8" bold="true" box="[846,1001,307,334]" italics="true" pageId="21" pageNumber="174">Horridocalia</emphasis>
|
||
Endrődi
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84FEA8566AFE4C" blockId="21.[140,1108,166,475]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FEA8566AFE4C" pageId="21" pageNumber="174">
|
||
– Machos con cuerno cefálico emarginado o bifurcado. Hembras con dos tubérculos transversales en la cabeza. Pronoto convexo y liso. Élitros fuertemente reticulados y finamente punteados. Proceso prosternal ausente. Cuerpo de color pardo amarillo con manchas oscuras.....................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE66FE3F566AFE4C" authority="Prell" box="[878,1108,448,475]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE66FE3F566AFE4C" bold="true" box="[878,1108,448,475]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE66FE3F562AFE4D" bold="true" box="[878,1044,448,475]" italics="true" pageId="21" pageNumber="174">Brachysiderus</emphasis>
|
||
Prell
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
<paragraph id="7B5955581F32FFE9CD84FDC651E5FDC2" blockId="21.[140,987,569,596]" box="[140,987,569,596]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CD84FDC651E5FDC2" bold="true" box="[140,987,569,596]" pageId="21" pageNumber="174">
|
||
Clave para los géneros de
|
||
<taxonomicName id="BCE62EDB1F32FFE9CCF9FDC65086FDC2" box="[497,696,569,596]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
|
||
adultos de
|
||
<collectingCountry id="03F115C81F32FFE9CE5DFDC651E5FDC2" box="[853,987,569,596]" name="Colombia" pageId="21" pageNumber="174">Colombia</collectingCountry>
|
||
</emphasis>
|
||
</paragraph>
|
||
<subSubSection id="33FC06D31F32FFEACD84FD8F566DFEE6" lastPageId="22" lastPageNumber="175" pageId="21" pageNumber="174" type="key">
|
||
<keyStep id="C0124DFD1F32FFE9CD84FD8F566DFC35" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FD8F566DFC81" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FD8F566DFC81" pageId="21" pageNumber="174">
|
||
1. Tarsos anteriores de los machos engrosados; uña externa más grande que la interna. Hembras con tarsos anteriores y uñas simples. Longitud del clípeo subigual o mayor que la longitud de la frente; apice parabólico, cuadrado, ligeramente emarginado, o redondeado. Tibia posterior semicircular en sección transvesal ...................................................................................................
|
||
<emphasis id="4992894A1F32FFE9C94CFD02566DFC81" bold="true" box="[1092,1107,765,791]" pageId="21" pageNumber="174">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84FCDE566DFC35" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FCDE566DFC35" pageId="21" pageNumber="174">
|
||
– Tarsos anteriores de los machos y hembras sin engrosamiento. Uñas tarsales simples. Longitud del clípeo mas corta que la longitud de la frente; apice trapezoidal (
|
||
<figureCitation id="E3DD49DD1F32FFE9CC89FC985381FC17" box="[385,447,871,897]" captionStart="Figure 8" captionStartId="7.[140,212,1654,1676]" captionText="Figure 8. Head of Stenocrates sp. (scale bar: 2 mm)." pageId="21" pageNumber="174">Fig 8</figureCitation>
|
||
). Tibia posterior aplanada dorsoventralmente, no semicircular en sección transversal......................................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE4AFC75566DFC35" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[834,1107,905,932]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE4AFC75566DFC35" bold="true" box="[834,1107,905,932]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE4AFC7551F6FC32" bold="true" box="[834,968,906,932]" italics="true" pageId="21" pageNumber="174">Stenocrates</emphasis>
|
||
Burmeister
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F32FFE9CD84FC52566DFC7C" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FC52566DFC51" blockId="21.[140,1108,624,1601]" box="[140,1107,940,968]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FC52566DFC51" box="[140,1107,940,968]" pageId="21" pageNumber="174">
|
||
2. Propigidio con numerosas sedas largas, visibles.................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE8DFC53566DFC51" authority="Casey" authorityName="Casey" authorityYear="1915" box="[901,1107,940,967]" class="Insecta" family="Dynastidae" genus="Chalepides" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE8DFC53566DFC51" bold="true" box="[901,1107,940,967]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE8DFC535639FC51" bold="true" box="[901,1031,940,967]" italics="true" pageId="21" pageNumber="174">Chalepides</emphasis>
|
||
Casey
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84FC2E566DFC7C" blockId="21.[140,1108,624,1601]" box="[140,1107,976,1003]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FC2E566DFC7C" box="[140,1107,976,1003]" pageId="21" pageNumber="174">
|
||
– Propigidio sin sedas largas y abundantes.....................................................
|
||
<emphasis id="4992894A1F32FFE9C94CFC2F566DFC7C" bold="true" box="[1092,1107,976,1002]" pageId="21" pageNumber="174">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F32FFE9CD84FC0B566DFBA7" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FC0B566DFB9B" blockId="21.[140,1108,624,1601]" box="[140,1107,1011,1038]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FC0B566DFB9B" box="[140,1107,1011,1038]" pageId="21" pageNumber="174">
|
||
3. Clípeo con ápice puntiagudo o estrechamente parabólico ..........................
|
||
<emphasis id="4992894A1F32FFE9C94CFC0C566DFB9B" bold="true" box="[1092,1107,1011,1037]" pageId="21" pageNumber="174">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84FBE8566DFBA7" blockId="21.[140,1108,624,1601]" box="[140,1107,1047,1073]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FBE8566DFBA7" box="[140,1107,1047,1073]" pageId="21" pageNumber="174">
|
||
– Clípeo con ápice redondeado, truncado, trapezoidal o emarginado ............
|
||
<emphasis id="4992894A1F32FFE9C94CFBE8566DFBA7" bold="true" box="[1092,1107,1047,1073]" pageId="21" pageNumber="174">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F32FFE9CD84FBC5566AFA95" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FBC5566DFB0C" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FBC5566DFB0C" pageId="21" pageNumber="174">
|
||
4. Clípeo con ápice acumindo (
|
||
<figureCitation id="E3DD49DD1F32FFE9CF34FBC550BEFBC2" box="[572,640,1082,1108]" captionStart="Figure 9" captionStartId="8.[139,211,547,569]" captionTargetBox="[429,829,209,446]" captionTargetId="figure@8.[396,849,161,529]" captionTargetPageId="8" captionText="Figure 9. Head of Mimeoma sp. (scale bar: 2 mm). (Figure taken from Ratcliffe 2003)." figureDoi="http://doi.org/10.5281/zenodo.3770643" httpUri="https://zenodo.org/record/3770643/files/figure.png" pageId="21" pageNumber="174">Fig. 9</figureCitation>
|
||
).
|
||
<collectingRegion id="B9229BBA1F32FFE9CF9EFBC550CCFBC2" box="[662,754,1082,1108]" country="Myanmar" name="Chin" pageId="21" pageNumber="174">Mentón</collectingRegion>
|
||
con ápice emarginado, superficie sin surco en el tercio apical. Base del pronoto con línea marginal............ ..........................................................................................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE87FB7F566DFB0C" authority="Casey" authorityName="Casey" authorityYear="1915" box="[911,1107,1152,1178]" class="Insecta" family="Dynastidae" genus="Mimeoma" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE87FB7F566DFB0C" bold="true" box="[911,1107,1152,1178]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE87FB7F5639FB0C" bold="true" box="[911,1031,1152,1178]" italics="true" pageId="21" pageNumber="174">Mimeoma</emphasis>
|
||
Casey
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84FB5B566AFA95" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FB5B566AFA95" pageId="21" pageNumber="174">
|
||
– Clípeo con ápice estrechamente parabólico (Fig 10).
|
||
<collectingRegion id="B9229BBA1F32FFE9CE45FB5B5197FB28" box="[845,937,1188,1214]" country="Myanmar" name="Chin" pageId="21" pageNumber="174">Mentón</collectingRegion>
|
||
con ápice profundamente emarginado, superficie con surco en el tercio apical. Base del pronoto sin línea marginal .........................................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE5EFB16566AFA95" authority="Erichson" authorityName="Erichson" authorityYear="1847" box="[854,1108,1257,1284]" class="Insecta" family="Dynastidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE5EFB16566AFA95" bold="true" box="[854,1108,1257,1284]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE5EFB1651DCFA92" bold="true" box="[854,994,1257,1284]" italics="true" pageId="21" pageNumber="174">Ancognatha</emphasis>
|
||
Erichson
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F32FFE9CD84FAF2566DFA06" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FAF2566DFADC" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FAF2566DFADC" pageId="21" pageNumber="174">
|
||
5. Clípeo trapezoidal o casi trapezoidal. Margen anterior del pronoto con una proyección central (Fig. 11) ...........................................
|
||
<taxonomicName id="BCE62EDB1F32FFE9CE7EFACF566DFADC" authority="Harold" authorityName="Harold" authorityYear="1869" box="[886,1107,1328,1354]" class="Insecta" family="Dynastidae" genus="Dyscinetus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F32FFE9CE7EFACF566DFADC" bold="true" box="[886,1107,1328,1354]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE7EFACF51C8FADC" bold="true" box="[886,1014,1328,1354]" italics="true" pageId="21" pageNumber="174">Dyscinetus</emphasis>
|
||
Harold
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84FAAB566DFA06" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FAAB566DFA06" pageId="21" pageNumber="174">
|
||
– Clípeo cuadrado, cuadrangular o parabólico. Margen anterior del pronoto simple........................................................................................................
|
||
<emphasis id="4992894A1F32FFE9C94CFA89566DFA06" bold="true" box="[1092,1107,1398,1424]" pageId="21" pageNumber="174">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F32FFE9CD84FA65566DF9D7" pageId="21" pageNumber="174">
|
||
<paragraph id="7B5955581F32FFE9CD84FA65566AFA6C" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84FA65566AFA6C" pageId="21" pageNumber="174">
|
||
6. Clípeo con los lados generalmente divergentes desde la base, ápice ampliamente redondeado (Fig. 12). Maxila sin dientes (excepto en Aspilodea fuliginea).....................................................................................
|
||
<emphasis id="4992894A1F32FFE9CE92FA1F566AFA6C" bold="true" box="[922,1108,1504,1531]" pageId="21" pageNumber="174">
|
||
<emphasis id="4992894A1F32FFE9CE92FA1F5633FA6D" bold="true" box="[922,1037,1504,1531]" italics="true" pageId="21" pageNumber="174">Aspilodea</emphasis>
|
||
Bates
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F32FFE9CD84F9FB566DF9D7" blockId="21.[140,1108,624,1601]" pageId="21" pageNumber="174">
|
||
<keyLead id="C017F66D1F32FFE9CD84F9FB566DF9D7" pageId="21" pageNumber="174">
|
||
– Clípeo con lados convergentes desde la base, ápice redondeado, truncado o emarginado. Maxila con dientes evidentes..................................................
|
||
<emphasis id="4992894A1F32FFE9C94CF9D8566DF9D7" bold="true" box="[1092,1107,1575,1601]" pageId="21" pageNumber="174">7</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F31FFEACD84FF58566DFEE6" pageId="22" pageNumber="175">
|
||
<paragraph id="7B5955581F31FFEACD84FF58566DFE91" blockId="22.[140,1108,167,369]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FF58566DFE91" pageId="22" pageNumber="175">
|
||
7. Clípeo con los lados convergentes, ápice redondeado, parabólico o emarginado (Fig. 13). Machos con los tarsos anteriores engrosados ........................ ...................................................................................
|
||
<taxonomicName id="BCE62EDB1F31FFEACE55FF13566DFE91" authority="Dejean" authorityName="Dejean" authorityYear="1821" box="[861,1107,236,263]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACE55FF13566DFE91" bold="true" box="[861,1107,236,263]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACE55FF1351C9FE91" bold="true" box="[861,1015,236,263]" italics="true" pageId="22" pageNumber="175">Cyclocephala</emphasis>
|
||
Dejean
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F31FFEACD84FEEE566DFEE6" blockId="22.[140,1108,167,369]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FEEE566DFEE6" pageId="22" pageNumber="175">
|
||
– Clípeo subcuadrado, lados ligeramente convergentes, ápice truncado o emarginado. Machos con los tarsos anteriores simples no engrosados................... ..................................................................................
|
||
<taxonomicName id="BCE62EDB1F31FFEACE52FEA9566DFEE6" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[858,1107,342,369]" class="Insecta" family="Dynastidae" genus="Erioscelis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACE52FEA9566DFEE6" bold="true" box="[858,1107,342,369]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACE52FEA951F6FEE7" bold="true" box="[858,968,342,369]" italics="true" pageId="22" pageNumber="175">Erioscelis</emphasis>
|
||
Burmeister
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
<paragraph id="7B5955581F31FFEACD84FE3051AAFE7C" blockId="22.[140,916,463,490]" box="[140,916,463,490]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACD84FE3051AAFE7C" bold="true" box="[140,916,463,490]" pageId="22" pageNumber="175">
|
||
Clave para los géneros de
|
||
<taxonomicName id="BCE62EDB1F31FFEACCF9FE30504CFE7C" authorityName="MacLeay" authorityYear="1819" box="[497,626,463,490]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="tribe" tribe="Dynastini">Dynastini</taxonomicName>
|
||
adultos de
|
||
<collectingCountry id="03F115C81F31FFEACE06FE3051AAFE7C" box="[782,916,463,490]" name="Colombia" pageId="22" pageNumber="175">Colombia</collectingCountry>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="33FC06D31F31FFEACD84FDF8566AFB9B" pageId="22" pageNumber="175" type="key">
|
||
<keyStep id="C0124DFD1F31FFEACD84FDF8566DFCAC" pageId="22" pageNumber="175">
|
||
<paragraph id="7B5955581F31FFEACD84FDF8566AFD3B" blockId="22.[140,1108,519,1038]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FDF8566AFD3B" pageId="22" pageNumber="175">
|
||
1. Clípeo emarginado o con 2 dientes espiniformes ampliamente separados. Machos con cuerno cefálico largo y ligeramente aplanado dorso ventralmente, bifurcado en el ápice. Pronoto de los machos con 2 cuernos agudos, 1 en cada ángulo exterior (Fig. 14). Hembras con tubérculo frontal doble o simple, pronoto convexo ...................................................
|
||
<taxonomicName id="BCE62EDB1F31FFEACE82FD6B566AFD3B" authority="Kirby" authorityName="Kirby" authorityYear="1825" box="[906,1108,659,686]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACE82FD6B566AFD3B" bold="true" box="[906,1108,659,686]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACE82FD6B5639FD38" bold="true" box="[906,1031,660,686]" italics="true" pageId="22" pageNumber="175">Megasoma</emphasis>
|
||
Kirby
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F31FFEACD84FD48566DFCAC" blockId="22.[140,1108,519,1038]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FD48566DFCAC" pageId="22" pageNumber="175">
|
||
– Clípeo truncado o con 2 dientes apicales próximos. Machos con cuerno cefálico largo, con dientes en la superficie dorsal, ápice acuminado o redondeado. Pronoto de los machos con cuerno central. Hembras con tubérculo frontal, pronoto convexo............................................................................
|
||
<emphasis id="4992894A1F31FFEAC94CFCDF566DFCAC" bold="true" box="[1092,1107,800,826]" pageId="22" pageNumber="175">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F31FFEACD84FCBB566AFB9B" pageId="22" pageNumber="175">
|
||
<paragraph id="7B5955581F31FFEACD84FCBB566DFC32" blockId="22.[140,1108,519,1038]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FCBB566DFC32" pageId="22" pageNumber="175">
|
||
2. Mandíbulas con ápice entero, o con una muesca. Pronoto de los machos con cuerno dirigido hacia arriba de ápice acuminado, o con tubérculo erecto y ancho (Fig. 15). Pronoto de las hembras generalmente punteado ...........
|
||
<taxonomicName id="BCE62EDB1F31FFEACEB6FC76566DFC32" authority="Hope" authorityName="Hope" box="[958,1107,905,932]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACEB6FC76566DFC32" bold="true" box="[958,1107,905,932]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACEB6FC765632FC32" bold="true" box="[958,1036,905,932]" italics="true" pageId="22" pageNumber="175">Golofa</emphasis>
|
||
Hope
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F31FFEACD84FC51566AFB9B" blockId="22.[140,1108,519,1038]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FC51566AFB9B" pageId="22" pageNumber="175">
|
||
– Mandíbulas con 2 dientes apicales. Pronoto de los machos con cuerno dirigido hacia delante, ápice agudo (Fig. 16). Pronoto de las hembras generalmente rugoso............................................................................
|
||
<taxonomicName id="BCE62EDB1F31FFEACE7EFC0B566AFB9B" authority="MacLeay" authorityName="MacLeay" authorityYear="1819" box="[886,1108,1011,1038]" class="Insecta" family="Dynastidae" genus="Dynastes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACE7EFC0B566AFB9B" bold="true" box="[886,1108,1011,1038]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACE7EFC0B51DFFB98" bold="true" box="[886,993,1012,1038]" italics="true" pageId="22" pageNumber="175">Dynastes</emphasis>
|
||
MacLeay
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
<paragraph id="7B5955581F31FFEACD84FB9351B8FB11" blockId="22.[140,902,1132,1159]" box="[140,902,1132,1159]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACD84FB9351B8FB11" bold="true" box="[140,902,1132,1159]" pageId="22" pageNumber="175">
|
||
Clave para los géneros de
|
||
<taxonomicName id="BCE62EDB1F31FFEACCF9FB93505DFB11" authorityName="Mulsant & Rey" authorityYear="1842" box="[497,611,1132,1159]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="tribe" tribe="Oryctini">Oryctini</taxonomicName>
|
||
adultos de
|
||
<collectingCountry id="03F115C81F31FFEACE08FB9351B8FB11" box="[768,902,1132,1159]" name="Colombia" pageId="22" pageNumber="175">Colombia</collectingCountry>
|
||
</emphasis>
|
||
</paragraph>
|
||
<subSubSection id="33FC06D31F31FFEBCD84FB5B566DFB0C" lastPageId="23" lastPageNumber="176" pageId="22" pageNumber="175" type="key">
|
||
<keyStep id="C0124DFD1F31FFEACD84FB5B566DFB76" pageId="22" pageNumber="175">
|
||
<paragraph id="7B5955581F31FFEACD84FB5B566DFB2B" blockId="22.[140,1108,1187,1601]" box="[140,1107,1187,1214]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FB5B566DFB2B" box="[140,1107,1187,1214]" pageId="22" pageNumber="175">
|
||
1. Tibia anterior tridentada ............................................................................
|
||
<emphasis id="4992894A1F31FFEAC94CFB5C566DFB2B" bold="true" box="[1092,1107,1187,1213]" pageId="22" pageNumber="175">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F31FFEACD84FB38566DFB76" blockId="22.[140,1108,1187,1601]" box="[140,1107,1222,1249]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FB38566DFB76" box="[140,1107,1222,1249]" pageId="22" pageNumber="175">
|
||
– Tibia anterior cuadridentada ......................................................................
|
||
<emphasis id="4992894A1F31FFEAC94CFB39566DFB76" bold="true" box="[1092,1107,1222,1248]" pageId="22" pageNumber="175">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F31FFEACD84FB15566DFA07" pageId="22" pageNumber="175">
|
||
<paragraph id="7B5955581F31FFEACD84FB15566DFAB1" blockId="22.[140,1108,1187,1601]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FB15566DFAB1" pageId="22" pageNumber="175">
|
||
2. Élitros lisos, brillantes, de color negro. Clípeo emarginado o truncado (Bidentado en
|
||
<taxonomicName id="BCE62EDB1F31FFEACC47FAF153DEFAB1" box="[335,480,1293,1319]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="species" species="philoctetes">
|
||
<emphasis id="4992894A1F31FFEACC47FAF153DEFAB1" box="[335,480,1293,1319]" italics="true" pageId="22" pageNumber="175">M. philoctetes</emphasis>
|
||
</taxonomicName>
|
||
). Mandíbulas bidentadas (Fig. 17).......
|
||
<taxonomicName id="BCE62EDB1F31FFEACE9BFAF2566DFAB1" authority="Hope" authorityName="Hope" authorityYear="1837" box="[915,1107,1293,1319]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACE9BFAF2566DFAB1" bold="true" box="[915,1107,1293,1319]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACE9BFAF25635FAB1" bold="true" box="[915,1035,1293,1319]" italics="true" pageId="22" pageNumber="175">Megaceras</emphasis>
|
||
Hope
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F31FFEACD84FACE566DFA07" blockId="22.[140,1108,1187,1601]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FACE566DFA07" pageId="22" pageNumber="175">
|
||
– Élitros con 5 filas de puntuaciones sobre una superficie opaca, de color pardorojo. Clípeo estrecho con ápice bidentado. Mandíbulas tridentazas (Fig. 18) ..............................................................................................
|
||
<taxonomicName id="BCE62EDB1F31FFEACEA3FA89566DFA07" authority="Hope" authorityName="Hope" authorityYear="1837" box="[939,1107,1398,1425]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACEA3FA89566DFA07" bold="true" box="[939,1107,1398,1425]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACEA3FA895637FA07" bold="true" box="[939,1033,1398,1425]" italics="true" pageId="22" pageNumber="175">Coelosis</emphasis>
|
||
Hope
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F31FFEBCD84FA65566DFEBC" lastPageId="23" lastPageNumber="176" pageId="22" pageNumber="175">
|
||
<paragraph id="7B5955581F31FFEACD84FA65566DF9D6" blockId="22.[140,1108,1187,1601]" pageId="22" pageNumber="175">
|
||
<keyLead id="C017F66D1F31FFEACD84FA65566DF9D6" pageId="22" pageNumber="175">
|
||
3. Élitros con filas de puntuaciones en surcos profundos. Mandíbulas con 2 lóbulos arredondados, similarmente grandes, proyectados desde la parte inferior del clipeo. Cabeza con tubérculo cónico cerca de la margen anterior del clipeo. Machos con prominencia pronotal formando uma cavidad triangular y alcanzando la margen posterior del pronoto .............
|
||
<taxonomicName id="BCE62EDB1F31FFEACE54F9D9566DF9D6" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[860,1107,1573,1601]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F31FFEACE54F9D9566DF9D6" bold="true" box="[860,1107,1573,1601]" pageId="22" pageNumber="175">
|
||
<emphasis id="4992894A1F31FFEACE54F9D951D7F9D7" bold="true" box="[860,1001,1574,1601]" italics="true" pageId="22" pageNumber="175">Gibboryctes</emphasis>
|
||
Endrődi
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F30FFEBCD84FF58566DFEBC" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FF58566DFEBC" pageId="23" pageNumber="176">
|
||
– Élitros lisos, rugosos o cubiertos densamente de puntuaciones, nunca en filas. Mandíbulas variablemente dentadas, nunca con 2 lóbulos grandes arredondeados. Cabeza con 1–2 tubérculos cónicos, nunca cerca del clípeo. Machos con prominencia pronotal de forma variable ..............................................
|
||
<emphasis id="4992894A1F30FFEBC94CFEEF566DFEBC" bold="true" box="[1092,1107,272,298]" pageId="23" pageNumber="176">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F30FFEBCD84FECB566DFDF1" pageId="23" pageNumber="176">
|
||
<paragraph id="7B5955581F30FFEBCD84FECB566DFE4C" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FECB566DFE4C" pageId="23" pageNumber="176">
|
||
4. Cuerpo alongado y subparalelo. Tibia anterior con dientes proyectados casi en angulos rectos. Ápice de la tibia posterior con dos dientes fuertes. Machos con la mitad anterior del pronoto casi liso, con un cuerno (o tubérculo) medio (Fig. 19). Hembras sin cavidad pronotal en el pronoto........................... ...............................................................................
|
||
<taxonomicName id="BCE62EDB1F30FFEBCE4CFE3F566DFE4C" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[836,1107,448,475]" class="Insecta" family="Dynastidae" genus="Podischnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F30FFEBCE4CFE3F566DFE4C" bold="true" box="[836,1107,448,475]" pageId="23" pageNumber="176">
|
||
<emphasis id="4992894A1F30FFEBCE4CFE3F51F6FE4D" bold="true" box="[836,968,448,475]" italics="true" pageId="23" pageNumber="176">Podischnus</emphasis>
|
||
Burmeister
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F30FFEBCD84FE1B566DFDF1" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FE1B566DFDF1" pageId="23" pageNumber="176">
|
||
– Cuerpo ancho, lados arredondeados. Tibia anterior con dientes proyectados oblicuamente. Ápice da tibia posterior crenulado o con
|
||
<geoCoordinate id="1ED2339F1F30FFEBCE64FDF85199FDB7" box="[876,935,519,545]" degrees="1, 3" direction="west" orientation="longitude" pageId="23" pageNumber="176" precision="5555" value="-1.3">1,3 o</geoCoordinate>
|
||
4 dientes. Machos con la mitad anterior del pronoto densamente cubierto de puntuaciones o rugoso. Hembras con o sin cavidad pronotal en el pronoto .....................
|
||
<emphasis id="4992894A1F30FFEBC94CFDB2566DFDF1" bold="true" box="[1092,1107,589,615]" pageId="23" pageNumber="176">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F30FFEBCD84FD8F566DFCAC" pageId="23" pageNumber="176">
|
||
<paragraph id="7B5955581F30FFEBCD84FD8F566DFD47" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FD8F566DFD47" pageId="23" pageNumber="176">
|
||
5. Machos y hembras con cuerno em la cabeza. Proceso prosternal corto, no producido después de la mitad de los protrocanteres. Pronoto con margen anterior emarginada hacia el centro (
|
||
<figureCitation id="E3DD49DD1F30FFEBCF9BFD4850DAFD47" box="[659,740,695,721]" captionStart="Figure 20" captionStartId="13.[140,212,713,735]" captionText="Figure 20. Pronotum of Enema pan (Fabricius) (scale bar: 1 cm)." pageId="23" pageNumber="176">Fig. 20</figureCitation>
|
||
) ..........................
|
||
<taxonomicName id="BCE62EDB1F30FFEBCEBFFD48566DFD47" authority="Hope" authorityName="Hope" authorityYear="1837" box="[951,1107,695,721]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F30FFEBCEBFFD48566DFD47" bold="true" box="[951,1107,695,721]" pageId="23" pageNumber="176">
|
||
<emphasis id="4992894A1F30FFEBCEBFFD485637FD47" bold="true" box="[951,1033,695,721]" italics="true" pageId="23" pageNumber="176">Enema</emphasis>
|
||
Hope
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F30FFEBCD84FD25566DFCAC" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FD25566DFCAC" pageId="23" pageNumber="176">
|
||
– Machos con o sin cuerno en la cabeza. Hembras nunca con cuerno. Proceso prosternal largo, producido después de la mitad de los protrocanteres. Pronoto con margen anterior no emarginada hacia el centro ...........................
|
||
<emphasis id="4992894A1F30FFEBC94CFCDF566DFCAC" bold="true" box="[1092,1107,800,826]" pageId="23" pageNumber="176">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="C0124DFD1F30FFEBCD84FCBC566DFB0C" pageId="23" pageNumber="176">
|
||
<paragraph id="7B5955581F30FFEBCD84FCBC566DFC7C" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FCBC566DFC7C" pageId="23" pageNumber="176">
|
||
6. Frente de los machos e hembras con dos tubérculos cónicos, nunca con cuerno. Mandíbulas con tres dientes. Pronoto en los machos con cuerno subapical e generalmente con cuernos laterales o una crista triangular elevada a cada lado (Fig. 21); hembras con cavidad pronotal en el tercio anterior del pronoto................................................................................
|
||
<taxonomicName id="BCE62EDB1F30FFEBCE95FC2F566DFC7C" authority="Hope" authorityName="Hope" authorityYear="1837" box="[925,1107,976,1002]" class="Insecta" family="Dynastidae" genus="Strategus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F30FFEBCE95FC2F566DFC7C" bold="true" box="[925,1107,976,1002]" pageId="23" pageNumber="176">
|
||
<emphasis id="4992894A1F30FFEBCE95FC2F5634FC7C" bold="true" box="[925,1034,976,1002]" italics="true" pageId="23" pageNumber="176">Strategus</emphasis>
|
||
Hope
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="7B5955581F30FFEBCD84FC0B566DFB0C" blockId="23.[140,1108,164,1179]" pageId="23" pageNumber="176">
|
||
<keyLead id="C017F66D1F30FFEBCD84FC0B566DFB0C" pageId="23" pageNumber="176">
|
||
– Frente de los machos con cuerno. Frente de las hembras con dos pequeños tubérculos. Mandíbulas simples o con dos dientes arredondeados. Pronoto en los machos con cuerno o protuberancia originándose desde la mitad posterior y con o sin cuernos laterales; hembras generalmente sin cavidad pronotal .... ........................................................................
|
||
<taxonomicName id="BCE62EDB1F30FFEBCE18FB7F566DFB0C" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[784,1107,1152,1179]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4992894A1F30FFEBCE18FB7F566DFB0C" bold="true" box="[784,1107,1152,1179]" pageId="23" pageNumber="176">
|
||
<emphasis id="4992894A1F30FFEBCE18FB7F51F6FB0D" bold="true" box="[784,968,1152,1179]" italics="true" pageId="23" pageNumber="176">Heterogomphus</emphasis>
|
||
Burmeister
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |