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<mods:title id="A8005BB7E4B950B6EA093E952BA21FF8">A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea</mods:title>
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<mods:namePart id="534C0C2489BEA5D75A411A95C51A2010">Phan, Quoc Toan</mods:namePart>
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<taxonomicName id="4FC64D46FFE8EC15B594FF10FD73D3C4" ID-CoL="L4P2" authority="Davies &amp; Yang, 1996" authorityName=": Davies &amp; Yang" authorityYear="1996" box="[151,716,151,178]" class="Insecta" family="Euphaeidae" genus="Bayadera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nephelopennis">
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<emphasis id="BAB2EAD7FFE8EC15B594FF10FE7DD3C7" bold="true" box="[151,450,151,177]" italics="true" pageId="13" pageNumber="164">Bayadera nephelopennis</emphasis>
<bibRefCitation id="EC574B34FFE8EC15B4CAFF1FFD73D3C4" author="Davies" box="[457,716,152,178]" pageId="13" pageNumber="164" refString="Davies, D. A. L. &amp; Yang, B. (1996) New species of Bayadera Selys and Schmidtiphaea Asahina from China (Odonata, Euphaeidae). Tijdschrift voor Entomologie, 139, 145 - 155." type="journal article" year="1996">Davies &amp; Yang, 1996</bibRefCitation>
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(
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)
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<taxonomicName id="4FC64D46FFE8EC15B594FE85FD38D26E" ID-CoL="L4P2" authority=": Davies &amp; Yang (1996)" authorityName=": Davies &amp; Yang" authorityYear="1996" box="[151,647,258,280]" class="Insecta" family="Euphaeidae" genus="Bayadera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nephelopennis">
<emphasis id="BAB2EAD7FFE8EC15B594FE85FE2DD26E" box="[151,402,258,280]" italics="true" pageId="13" pageNumber="164">Bayadera nephelopennis</emphasis>
:
<bibRefCitation id="EC574B34FFE8EC15B49CFE85FD38D26E" author="Davies" box="[415,647,258,280]" pageId="13" pageNumber="164" refString="Davies, D. A. L. &amp; Yang, B. (1996) New species of Bayadera Selys and Schmidtiphaea Asahina from China (Odonata, Euphaeidae). Tijdschrift voor Entomologie, 139, 145 - 155." type="journal article" year="1996">Davies &amp; Yang (1996)</bibRefCitation>
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, pp. 148149,
<figureCitation id="10FD2A40FFE8EC15B61FFE85FC2FD26E" box="[796,912,258,280]" captionStart-0="FIGURE 12" captionStart-1="FIGURE13" captionStart-2="FIGURE 14" captionStart-3="FIGURE15" captionStart-4="FIGURE 16" captionStart-5="FIGURE 17" captionStartId-0="18.[151,250,1124,1146]" captionStartId-1="20.[151,250,1462,1484]" captionStartId-2="23.[151,250,1889,1911]" captionStartId-3="24.[151,250,929,951]" captionStartId-4="24.[151,250,1970,1992]" captionStartId-5="25.[151,250,1366,1388]" captionTargetBox-0="[285,1295,203,1100]" captionTargetBox-1="[411,1176,447,1440]" captionTargetBox-2="[220,1341,852,1868]" captionTargetBox-3="[186,1410,221,879]" captionTargetBox-4="[286,1291,1018,1921]" captionTargetBox-5="[248,1338,210,1339]" captionTargetId-0="figure@18.[281,1305,195,1102]" captionTargetId-1="figure@20.[411,1176,447,1440]" captionTargetId-2="figure@23.[184,1403,844,1868]" captionTargetId-3="figure@24.[153,1433,193,907]" captionTargetId-4="figure@24.[281,1305,1015,1949]" captionTargetId-5="figure@25.[217,1369,193,1345]" captionTargetPageId-0="18" captionTargetPageId-1="20" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="24" captionTargetPageId-5="25" captionText-0="FIGURE 12. Dysphaea spp. [12ab] D. basitincta. (12a) ♂, Cao Bang and (12b) ♀, Hainan (rearranged from Fig. 7, p. 307, Hämäläinen 2012). [12cd] D. gloriosa; (12c) ♂ and (12d) ♀, Quang Nam. [12ef] D. haomiao; (12e) ♂, Cao Bang and (12f) ♀, Guizhou (rearranged from Fig.5, p. 307, Hämäläinen 2012)." captionText-1="FIGURE13.Euphaea decorata.(13a)♂,Pia Oac Nature Reserve,Cao Bang Province; (13b)♀, Mau Son mountain,Lang Son Province (photographed by Q.T.Phan)." captionText-2="FIGURE 14. Dorsal view of wings of male Euphaea spp. (14a) E. hirta; (14b) E. masoni; (14c) E. guerini; (14d) E. saola sp. nov." captionText-3="FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. IX show abdominal segments S110." captionText-4="FIGURE 16. Euphaea spp. all ♀. (16ab) body and head of E. hirta, Nam Nung; (16c) E. masoni, Bao Loc; (16d) E. guerini, Tay Giang; (16e) E. saola sp. nov., A Luoi." captionText-5="FIGURE 17. Euphaea spp. [17ad] E. ochracea, Mang Den, Kon Tum &amp; [17e], Penang, Malaysia; [17fj] holotype (♂) and paratype (♀) of E. sanguinea sp. nov. (17a, f) male habitus; (17b, g) female habitus; (17c) genital ligula, ventral view; (17e, j) left cercus, interolateral view; (17h) male head in antero-dorsal view; (17d, i) female heads in antero-dorsal view. Scale bars in mm." httpUri-0="https://zenodo.org/record/3096091/files/figure.png" httpUri-1="https://zenodo.org/record/3096093/files/figure.png" httpUri-2="https://zenodo.org/record/3096097/files/figure.png" httpUri-3="https://zenodo.org/record/3096107/files/figure.png" httpUri-4="https://zenodo.org/record/3096109/files/figure.png" httpUri-5="https://zenodo.org/record/3096113/files/figure.png" pageId="13" pageNumber="164">Figs 1217</figureCitation>
.
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<collectorName id="25335313FFE8EC15B7B0FE09FC87D2D0" box="[691,824,397,422]" pageId="13" pageNumber="164">T. Kompier</collectorName>
leg. (TKC). [
<collectingCountry id="F0D17655FFE8EC15B6DBFE0AFB9FD2D0" box="[984,1056,397,422]" name="China" pageId="13" pageNumber="164">China</collectingCountry>
]
<specimenCount id="9EC0FD4CFFE8EC15B131FE09FBE8D2D0" box="[1074,1111,398,422]" pageId="13" pageNumber="164" type="male">3♂</specimenCount>
,
<location id="8D19601EFFE8EC15B165FE0AFB68D2D0" LSID="urn:lsid:plazi:treatment:006F87D3FFE8EC15B594FF10FA38D078:8D19601EFFE8EC15B165FE0AFB68D2D0" box="[1126,1239,397,422]" country="China" county="Sa Pa" municipality="Lao Cai Prov." name="Emeishan" pageId="13" pageNumber="164" stateProvince="Sichuan">Emeishan</location>
,
<elevation id="03EBD1F6FFE8EC15B1E5FE09FA8CD2D3" box="[1254,1331,398,422]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="164" unit="m" value="800.0">
<quantity id="4F3E9B20FFE8EC15B1E5FE09FA8CD2D3" box="[1254,1331,398,422]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="164" unit="m" value="800.0">800 m</quantity>
</elevation>
,
<collectingRegion id="4A02F827FFE8EC15B041FE0AFA23D2D0" box="[1346,1436,397,422]" country="China" name="Sichuan" pageId="13" pageNumber="164">Sichuan</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFE8EC15B5DCFE36FED5D2BC" box="[223,362,433,458]" pageId="13" pageNumber="164" value="2017-07-25">
<date id="FC781005FFE8EC15B5DCFE36FED5D2BC" box="[223,362,433,458]" pageId="13" pageNumber="164" value="2017-07-25">25.VII.2017</date>
</collectingDate>
,
<collectorName id="25335313FFE8EC15B475FE35FE00D2BC" box="[374,447,433,458]" pageId="13" pageNumber="164">X. Liu</collectorName>
leg. (FHC).
</materialsCitation>
</paragraph>
<paragraph id="887936C5FFE8EC15B5C4FE52FEBED164" blockId="13.[151,1437,325,782]" pageId="13" pageNumber="164">
<materialsCitation id="38AE3C98FFE8EC15B5C4FE52FEBED164" ID-GBIF-Occurrence="1805097974" collectingDate="2017-05-28" collectingDateMax="2017-05-29" collectingDateMin="2017-05-28" collectorName="T. Kompier." country="Vietnam" location="Other" pageId="13" pageNumber="164" specimenCount="1" stateProvince="Lai Chau">
<emphasis id="BAB2EAD7FFE8EC15B5C4FE52FC99D298" bold="true" box="[199,806,469,494]" pageId="13" pageNumber="164">
<location id="8D19601EFFE8EC15B5C4FE52FEB0D298" LSID="urn:lsid:plazi:treatment:006F87D3FFE8EC15B594FF10FA38D078:8D19601EFFE8EC15B5C4FE52FEB0D298" box="[199,271,469,494]" country="Vietnam" name="Other" pageId="13" pageNumber="164" stateProvince="Lai Chau">Other</location>
materials confirmed by field observations.
</emphasis>
[
<collectingCountry id="F0D17655FFE8EC15B63AFE52FC21D298" box="[825,926,469,494]" name="Vietnam" pageId="13" pageNumber="164">Vietnam</collectingCountry>
] 17 exs,
<collectingRegion id="4A02F827FFE8EC15B113FE52FBC2D298" box="[1040,1149,469,494]" country="Vietnam" name="Lai Chau" pageId="13" pageNumber="164">Lai Chau</collectingRegion>
Prov.,
<collectingDate id="EC3CE9EDFFE8EC15B1D5FE51FACAD298" box="[1238,1397,470,494]" pageId="13" pageNumber="164" value="2017-05-28" valueMax="2017-05-29" valueMin="2017-05-28">
<date id="FC781005FFE8EC15B1D5FE51FACAD298" box="[1238,1397,470,494]" pageId="13" pageNumber="164" value="2017-05-28" valueMax="2017-05-29" valueMin="2017-05-28">2829.V.2017</date>
</collectingDate>
,
<collectorName id="25335313FFE8EC15B085FE51FEBED164" pageId="13" pageNumber="164">T. Kompier.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="887936C5FFE8EC15B5C4FD9AFDFFD19C" blockId="13.[151,1437,325,782]" pageId="13" pageNumber="164">
<emphasis id="BAB2EAD7FFE8EC15B5C4FD9AFEAED140" bold="true" box="[199,273,541,566]" pageId="13" pageNumber="164">Notes.</emphasis>
This is a new record for
<collectingCountry id="F0D17655FFE8EC15B72FFD9AFD30D140" box="[556,655,541,566]" name="Vietnam" pageId="13" pageNumber="164">Vietnam</collectingCountry>
. Male
<taxonomicName id="4FC64D46FFE8EC15B7D8FD9AFC4ED140" authorityName=": Davies &amp; Yang" authorityYear="1996" box="[731,1009,541,566]" class="Insecta" family="Euphaeidae" genus="Bayadera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nephelopennis">
<emphasis id="BAB2EAD7FFE8EC15B7D8FD9AFC4ED140" box="[731,1009,541,566]" italics="true" pageId="13" pageNumber="164">Bayadera nephelopennis</emphasis>
</taxonomicName>
is characterized by a black body with pruinosity on legs, basal abdomen, and ventral sides of synthorax (
<figureCitation id="10FD2A40FFE8EC15B684FDC6FC66D12F" box="[903,985,577,602]" captionStart="FIGURE 9" captionStartId="13.[151,250,1717,1739]" captionTargetBox="[289,1295,838,1682]" captionTargetId="figure@13.[281,1305,834,1711]" captionTargetPageId="13" captionText="FIGURE 9. Bayadera nephelopennis. [9ae] ♂ and [9f] ♀. (9a) head and thorax; (9b) appendages, dorsal view; (9c) appendages, lateral view; (9d) appendages, ventral view; (9e) wings; (9f) head and thorax." httpUri="https://zenodo.org/record/3096075/files/figure.png" pageId="13" pageNumber="164">Fig. 9a</figureCitation>
). Wings are cloudy light brown in basal two-thirds with hyaline apex in Chinese males, but hyaline with slightly darkened apex in Vietnamese ones (
<figureCitation id="10FD2A40FFE8EC15B06CFDE2FF0BD1D4" captionStart="FIGURE 9" captionStartId="13.[151,250,1717,1739]" captionTargetBox="[289,1295,838,1682]" captionTargetId="figure@13.[281,1305,834,1711]" captionTargetPageId="13" captionText="FIGURE 9. Bayadera nephelopennis. [9ae] ♂ and [9f] ♀. (9a) head and thorax; (9b) appendages, dorsal view; (9c) appendages, lateral view; (9d) appendages, ventral view; (9e) wings; (9f) head and thorax." httpUri="https://zenodo.org/record/3096075/files/figure.png" pageId="13" pageNumber="164">Fig. 9e</figureCitation>
). Anal appendages are long in lateral view and cerci are characterized by two well defined closely spaced tubercles (
<figureCitation id="10FD2A40FFE8EC15B415FD2AFE2ED1B0" box="[278,401,685,710]" captionStart="FIGURE 9" captionStartId="13.[151,250,1717,1739]" captionTargetBox="[289,1295,838,1682]" captionTargetId="figure@13.[281,1305,834,1711]" captionTargetPageId="13" captionText="FIGURE 9. Bayadera nephelopennis. [9ae] ♂ and [9f] ♀. (9a) head and thorax; (9b) appendages, dorsal view; (9c) appendages, lateral view; (9d) appendages, ventral view; (9e) wings; (9f) head and thorax." httpUri="https://zenodo.org/record/3096075/files/figure.png" pageId="13" pageNumber="164">Fig. 9bd</figureCitation>
). Female is separated from other
<taxonomicName id="4FC64D46FFE8EC15B645FD2AFC0CD1B0" authorityName="Selys" authorityYear="1853" box="[838,947,685,710]" class="Insecta" family="Euphaeidae" genus="Bayadera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="BAB2EAD7FFE8EC15B645FD2AFC0CD1B0" box="[838,947,685,710]" italics="true" pageId="13" pageNumber="164">Bayadera</emphasis>
</taxonomicName>
species by a broad dorsal stripe over mesepisternum of synthorax (
<figureCitation id="10FD2A40FFE8EC15B4E0FD56FD8DD19C" box="[483,562,721,746]" captionStart="FIGURE 9" captionStartId="13.[151,250,1717,1739]" captionTargetBox="[289,1295,838,1682]" captionTargetId="figure@13.[281,1305,834,1711]" captionTargetPageId="13" captionText="FIGURE 9. Bayadera nephelopennis. [9ae] ♂ and [9f] ♀. (9a) head and thorax; (9b) appendages, dorsal view; (9c) appendages, lateral view; (9d) appendages, ventral view; (9e) wings; (9f) head and thorax." httpUri="https://zenodo.org/record/3096075/files/figure.png" pageId="13" pageNumber="164">Fig. 9f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0DC654EFFE8EC15B5C4FD72FA38D078" box="[199,1415,757,782]" pageId="13" pageNumber="164" type="distribution">
<paragraph id="887936C5FFE8EC15B5C4FD72FA38D078" blockId="13.[151,1437,325,782]" box="[199,1415,757,782]" pageId="13" pageNumber="164">
<emphasis id="BAB2EAD7FFE8EC15B5C4FD72FEDFD078" bold="true" box="[199,352,757,782]" pageId="13" pageNumber="164">Distribution.</emphasis>
<collectingCountry id="F0D17655FFE8EC15B464FD72FE79D078" box="[359,454,757,782]" name="Vietnam" pageId="13" pageNumber="164">Vietnam</collectingCountry>
(
<collectingRegion id="4A02F827FFE8EC15B4D6FD71FD8FD078" box="[469,560,757,782]" country="Vietnam" name="Lao Cai" pageId="13" pageNumber="164">Lao Cai</collectingRegion>
and
<collectingRegion id="4A02F827FFE8EC15B765FD72FD72D078" box="[614,717,757,782]" country="Vietnam" name="Lai Chau" pageId="13" pageNumber="164">Lai Chau</collectingRegion>
Provinces),
<collectingCountry id="F0D17655FFE8EC15B659FD72FC22D078" box="[858,925,757,782]" name="China" pageId="13" pageNumber="164">China</collectingCountry>
(
<collectingRegion id="4A02F827FFE8EC15B6A8FD72FBCBD078" box="[939,1140,757,782]" country="China" name="Sichuan" pageId="13" pageNumber="164">Sichuan Province</collectingRegion>
) (
<bibRefCitation id="EC574B34FFE8EC15B189FD72FAC5D078" author="Davies" box="[1162,1402,757,782]" pageId="13" pageNumber="164" refString="Davies, D. A. L. &amp; Yang, B. (1996) New species of Bayadera Selys and Schmidtiphaea Asahina from China (Odonata, Euphaeidae). Tijdschrift voor Entomologie, 139, 145 - 155." type="journal article" year="1996">Davies &amp; Yang 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>