treatments-xml/data/03/F6/87/03F687FE4347BC6D0CD7FBE4FD25741A.xml

<document id="C920AA930F2F826E9F029C98E872A7FF" ID-CLB-Dataset="2609" ID-DOI="10.11646/zootaxa.5091.2.3" ID-GBIF-Dataset="36ace73e-babc-4c20-a78f-fa772ce5f539" ID-ISSN="1175-5326" ID-Zenodo-Dep="5843837" ID-ZooBank="C8081B60-C872-4A19-9291-22A42DC8B763" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1642061981252" checkinUser="plazi" docAuthor="Mai, Zuqi, Hu, Jian, Minoshima, Yûsuke N., Jia, Fenglong &amp; Fikáček, Martin" docDate="2022" docId="03F687FE4347BC6D0CD7FBE4FD25741A" docLanguage="en" docName="zootaxa.5091.2.3.pdf" docOrigin="Zootaxa 5091 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dactylosternum nanlingensis Mai et Jia 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="281" masterDocId="FFCFFF86434EBC610C40FFFEFF88770C" masterDocTitle="Review of Dactylosternum Wollaston, 1854 from China and Japan (Coleoptera, Hydrophilidae, Sphaeridiinae)" masterLastPageNumber="300" masterPageNumber="269" pageNumber="278" updateTime="1699302224171" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="52A7FC5F79227FAD1CD3F4C946605A51">Review of Dactylosternum Wollaston, 1854 from China and Japan (Coleoptera, Hydrophilidae, Sphaeridiinae)</mods:title>
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<mods:namePart id="4A323B73A5AE999729275967211459EA">Mai, Zuqi</mods:namePart>
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<mods:affiliation id="365E0FF3DF2E229256420AC5C4905735">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; maizq 5 @ mail 2. sysu. edu. cn; http: // orcid. org / 0000 - 0003 - 3124 - 2021</mods:affiliation>
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<mods:namePart id="A6C9EBACD0519BDB3004563F34340569">Hu, Jian</mods:namePart>
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<mods:affiliation id="EF2EBC9E376A68EE5E97A6F74037AC2F">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; lsshj @ mail. sysu. edu. cn; http: // orcid. org / 0000 - 0002 - 4122 - 8533</mods:affiliation>
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<mods:namePart id="F8047B7930A2DEDCFB5B18F2FA2C2874">Minoshima, Yûsuke N.</mods:namePart>
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<mods:affiliation id="9F5298AC8F093A7F332683CBDD8AFE01">Natural History Division, Kitakyushu Museum of Natural History and Human History, 2 - 4 - 1 Higashida, Yahatahigashi-ku, Kitakyushushi, Fukuoka, 805 - 0071 Japan. minoshima @ kmnh. jp; http: // orcid. org / 0000 - 0002 - 2575 - 4082</mods:affiliation>
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<mods:namePart id="B1A2F2B9279832B3B548C9A6CEC4493A">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="9F4B9ABF89466FBA4C61397BF5DDB89C">School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, Guangdong, China. &amp; Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China. lssjfl @ mail. sysu. edu. cn; fenglongjia @ alilyun. com; http: // orcid. org / 0000 - 0003 - 2391 - 5038</mods:affiliation>
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<mods:namePart id="E3DB6C80CFD302FC0B7D56028FC8A30E">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="D71BC4C0AD9A1163D83ED2BE65DFC4B2">Department of Biological Sciences, National Sun Yat-sen University, No. 70 Lien-hai Rd., Kaohsiung City 80424. mfikacek @ gmail. com; http: // orcid. org / 0000 - 0002 - 2078 - 6798 &amp; National Museum, Department of Entomology, Cirkusová 1740, CZ- 19300 Praha 9, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 12843 Praha 2, Czech Republic Corresponding author</mods:affiliation>
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<taxonomicName id="4C5F4D6B4347BC680CD7FBE4FD087338" ID-CoL="9GCNT" authority="Mai et Jia" authorityName="Mai et Jia" authorityYear="2022" box="[151,640,1050,1077]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="278" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA4347BC680CD7FBE4FE727339" bold="true" box="[151,506,1050,1077]" italics="true" pageId="9" pageNumber="278">Dactylosternum nanlingensis</emphasis>
Mai et Jia
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(
<figureCitation id="13642A6D4347BC680CDFFBC1FF667355" box="[159,238,1087,1113]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1673,1697]" captionTargetBox="[236,1361,474,1632]" captionTargetId="figure-149@4.[151,1436,450,1647]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus of Dactylosternum spp. from China and Japan. 1: D. abdominale (Fabricius, 1792), 2: D. corbetti Balfour-Browne, 1942, 3: D. nanlingensis sp. nov., 4: D. pui Jia, 2002" figureDoi="http://doi.org/10.5281/zenodo.5843839" httpUri="https://zenodo.org/record/5843839/files/figure.png" pageId="9" pageNumber="278">Figs. 3</figureCitation>
, 
<figureCitation id="13642A6D4347BC680CBBFBC1FECA7355" box="[251,322,1087,1113]" captionStart="FIGURES 11–16" captionStartId="10.[151,264,1853,1877]" captionTargetBox="[255,1413,665,1819]" captionTargetId="figure-225@10.[151,1436,633,1830]" captionTargetPageId="10" captionText="FIGURES 11–16. Habitus of D. nanlingensis sp. nov. and D. songxiaobini sp. nov. 11–13: Habitus of D. nanlingensis sp. nov., holotype (11: ventral view, 12: lateral view, 13: frontal view), 14–16: Habitus of D. songxiaobini sp. nov., holotype (14: ventral view, 15: lateral view, 16: frontal view)" figureDoi="http://doi.org/10.5281/zenodo.5843843" httpUri="https://zenodo.org/record/5843843/files/figure.png" pageId="9" pageNumber="278">11–13</figureCitation>
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<figureCitation id="13642A6D4347BC680D0EFBC1FEE47355" box="[334,364,1087,1113]" captionStart="FIGURES 17-19" captionStartId="11.[151,264,1231,1255]" captionTargetBox="[151,1435,190,1207]" captionTargetId="figure-19@11.[151,1435,181,1207]" captionTargetPageId="11" captionText="FIGURES 17-19. Morphology of Dactylosternum spp. 17: hind wing of D. nanlingensis sp. nov., 18–19: Anterior margin of clypeus of D. hydrophiloides and D. pseudolatum sp. nov. (18: D. hydrophiloides (arrow indicates the marginal rim on anterior margin of clypeus), 19: D. pseudolatum sp. nov. (without marginal rim on anterior margin of clypeus))" figureDoi="http://doi.org/10.5281/zenodo.5843853" httpUri="https://zenodo.org/record/5843853/files/figure.png" pageId="9" pageNumber="278">17</figureCitation>
, 
<figureCitation id="13642A6D4347BC680D38FBC1FE1E7355" box="[376,406,1087,1113]" captionStart="FIGURES 20–24" captionStartId="13.[151,264,1946,1970]" captionTargetBox="[236,1409,756,1826]" captionTargetId="figure-217@13.[164,1423,741,1922]" captionTargetPageId="13" captionText="FIGURES 20–24. Aedeagus of Dactylosternum spp. 20: D. abdominale (specimen from Yonaguni-jima Island, Japan, dorsal view, showing variation of apex of paramera) 21: D. corbetti, 22: D. pui (phallobase missing), 23: D. nanlingensis sp. nov., paratype (23a: ventral view, 23b: dorsal view), 24: D. hydrophiloides" figureDoi="http://doi.org/10.5281/zenodo.5843857" httpUri="https://zenodo.org/record/5843857/files/figure.png" pageId="9" pageNumber="278">23</figureCitation>
, 
<figureCitation id="13642A6D4347BC680DE2FBC1FD9C7355" box="[418,532,1086,1113]" captionStart="FIGURES 39–47" captionStartId="19.[151,264,1466,1490]" captionTargetBox="[151,1431,183,1437]" captionTargetId="figure-19@19.[151,1436,181,1442]" captionTargetPageId="19" captionText="FIGURES 39–47. Morphology characters of Dactylosternum spp. SEM micrographs. 39–40: antennal club (39: D. nanlingensis sp. nov., 40: D. abdominale), 41–42: ventral surface of mesofemora (41: D. pui, 42: D. nanlingensis sp. nov.), 43–47: punctures on pronotum (43: D. pui, 44: D. songxiaobini sp. nov. (surface with two sizes of punctures, the blue arrow indicates the coarser puncture, the yellow arrow indicates the finer puncture), 45: D. nanlingensis sp. nov., 46: D. corbetti (arrow indicates two short divergent lines connected with a puncture), 47: D. abdominale (arrow indicates mesh-like microsculpture between punctures))" figureDoi="http://doi.org/10.5281/zenodo.5843879" httpUri="https://zenodo.org/record/5843879/files/figure.png" pageId="9" pageNumber="278">39, 42, 45</figureCitation>
, 
<figureCitation id="13642A6D4347BC680E60FBC1FDB77355" box="[544,575,1087,1113]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="9" pageNumber="278">49</figureCitation>
, 
<figureCitation id="13642A6D4347BC680E0AFBC1FD1C7356" box="[586,660,1087,1114]" captionStart="FIGURES 64–69" captionStartId="23.[151,264,1684,1708]" captionTargetBox="[158,1428,380,1658]" captionTargetId="figure-106@23.[158,1428,380,1660]" captionTargetPageId="23" captionText="FIGURES 64–69. Habitats of Dactylosternum spp. 64, 66: individual specimen of D. nanlingensis sp. nov. on the surface of a fallen wood in forest at night, 65: habitats of D. abdominale (Fabricius, 1792), D. corbetti Balfour-Browne, 1942 and D. hydrophiloides (MacLeay, 1825), specimens hiding in a decaying banana tree, 67: D. abdominale, 68: D. corbetti, 69: D. hydrophiloides. Figures 64, 66 photo by Mr. Bao-Ping Huang in Ruyuan County, Tianjingshan National Forest Park." figureDoi="http://doi.org/10.5281/zenodo.5843889" httpUri="https://zenodo.org/record/5843889/files/figure.png" pageId="9" pageNumber="278">64, 66</figureCitation>
).
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<emphasis id="B92BEAFA4347BC680CD7FB79FE4173AC" bold="true" box="[151,457,1158,1185]" pageId="9" pageNumber="278">Type material examined.</emphasis>
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:
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male (
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<collectingCounty id="62814E644347BC680F98FB79FB0773AD" box="[984,1167,1159,1185]" pageId="9" pageNumber="278">Ruyuan County</collectingCounty>
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, fallen rotten wood in forest, 
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<collectorName id="26AA533E4347BC680890FB55FA0873C9" box="[1232,1408,1195,1221]" pageId="9" pageNumber="278">Baoping Huang</collectorName>
&amp; 
<collectorName id="26AA533E4347BC680CD7FB31FECD73E5" box="[151,325,1231,1257]" pageId="9" pageNumber="278">Zhaoyang Tang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B373CB54347BC680D3BFB30FB5173E5" ID-GBIF-Occurrence="3436469337" box="[379,1241,1230,1257]" collectionCode="SYSU" pageId="9" pageNumber="278" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="B92BEAFA4347BC680D3BFB30FDA073E4" bold="true" box="[379,552,1230,1256]" pageId="9" pageNumber="278">
<typeStatus id="54E4884A4347BC680D3BFB30FDAB73E4" box="[379,547,1230,1256]" pageId="9" pageNumber="278" type="paratype">PARATYPES</typeStatus>
:
</emphasis>
<specimenCount id="9D59FD614347BC680E6FFB31FDF473E5" box="[559,636,1231,1257]" pageId="9" pageNumber="278" type="male">1 male</specimenCount>
, 
<specimenCount id="9D59FD614347BC680EC7FB30FD7B73E5" box="[647,755,1230,1257]" pageId="9" pageNumber="278" type="female">5 females</specimenCount>
(
<collectionCode id="ED4EAE2D4347BC680F41FB31FCC773E5" box="[769,847,1231,1257]" pageId="9" pageNumber="278">SYSU</collectionCode>
): same information as the 
<typeStatus id="54E4884A4347BC680833FB31FB5173E5" box="[1139,1241,1231,1257]" pageId="9" pageNumber="278" type="holotype">holotype</typeStatus>
</materialsCitation>
; 
<materialsCitation id="3B373CB54347BC6808A3FB31FC8C723D" ID-GBIF-Occurrence="3436469344" collectingDate="2020-08-18" collectionCode="SYSU" collectorName="Zhenming Yang" county="Ganzhou" elevation="526" latitude="24.541111" location="Xiasongtang" longLatPrecision="21" longitude="114.458885" municipality="Jiulianshan National Reserve" pageId="9" pageNumber="278" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="9D59FD614347BC6808A3FB31FAB973E5" box="[1251,1329,1231,1257]" pageId="9" pageNumber="278" type="male">1 male</specimenCount>
, 
<specimenCount id="9D59FD614347BC68097CFB31FA1473E5" box="[1340,1436,1231,1257]" pageId="9" pageNumber="278" type="female">1 female</specimenCount>
(
<collectionCode id="ED4EAE2D4347BC680CDFFB0DFF647201" box="[159,236,1267,1293]" pageId="9" pageNumber="278">SYSU</collectionCode>
), 
<emphasis id="B92BEAFA4347BC680D43FB0CFED27200" bold="true" box="[259,346,1266,1292]" pageId="9" pageNumber="278">
<collectingRegion id="499BF80A4347BC680D43FB0CFED27200" box="[259,346,1266,1292]" country="China" name="Jiangxi" pageId="9" pageNumber="278">Jiangxi</collectingRegion>
</emphasis>
, 
<collectingCounty id="62814E644347BC680D2CFB0DFE5C7201" box="[364,468,1267,1293]" pageId="9" pageNumber="278">Ganzhou</collectingCounty>
, 
<collectingMunicipality id="6B84AC924347BC680DA3FB0DFCBB7201" box="[483,819,1267,1293]" pageId="9" pageNumber="278">Jiulianshan National Reserve</collectingMunicipality>
, 
<location id="8E8060334347BC680F02FB0DFC5A7201" LSID="urn:lsid:plazi:treatment:03F687FE4347BC6D0CD7FBE4FD25741A:8E8060334347BC680F02FB0DFC5A7201" box="[834,978,1267,1293]" county="Ganzhou" latitude="24.541111" longLatPrecision="21" longitude="114.458885" municipality="Jiulianshan National Reserve" name="Xiasongtang" pageId="9" pageNumber="278" stateProvince="Jiangxi">Xiasongtang</location>
, 
<geoCoordinate id="EE6B502F4347BC680FA1FB0AFBEE7203" box="[993,1126,1268,1295]" degrees="24" direction="north" minutes="32" orientation="latitude" pageId="9" pageNumber="278" precision="15" seconds="28" value="24.541111">24°32′28″N</geoCoordinate>
<geoCoordinate id="EE6B502F4347BC680831FB0AFA8B7203" box="[1137,1283,1268,1295]" degrees="114" direction="east" minutes="27" orientation="longitude" pageId="9" pageNumber="278" precision="15" seconds="32" value="114.458885">114°27′32″E</geoCoordinate>
, under fallen rotten wood, 
<quantity id="4CA79B0D4347BC680D6CFAE8FEFD723D" box="[300,373,1302,1329]" metricMagnitude="2" metricUnit="m" metricValue="5.26" pageId="9" pageNumber="278" unit="m" value="526.0">
<elevation id="0072D1DB4347BC680D6CFAE8FEFD723D" box="[300,373,1302,1329]" metricMagnitude="2" metricUnit="m" metricValue="5.26" pageId="9" pageNumber="278" unit="m" value="526.0">526 m</elevation>
</quantity>
, 
<date id="FFE110284347BC680DC1FAE9FD9D723D" box="[385,533,1303,1329]" pageId="9" pageNumber="278" value="2020-08-18">
<collectingDate id="EFA5E9C04347BC680DC1FAE9FD9D723D" box="[385,533,1303,1329]" pageId="9" pageNumber="278" value="2020-08-18">18.VIII.2020</collectingDate>
</date>
, 
<collectorName id="26AA533E4347BC680E61FAE9FD5C723D" box="[545,724,1303,1329]" pageId="9" pageNumber="278">Zhenming Yang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34565634347BC680C87FAC4FD01715D" pageId="9" pageNumber="278" type="diagnosis">
<paragraph id="8BE036E84347BC680C87FAC4FD01715D" blockId="9.[151,1437,1158,2013]" pageId="9" pageNumber="278">
<emphasis id="B92BEAFA4347BC680C87FAC4FECA7258" bold="true" box="[199,322,1338,1364]" pageId="9" pageNumber="278">Diagnosis.</emphasis>
Length 
<quantity id="4CA79B0D4347BC680DE0FAC4FDA27259" box="[416,554,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="9" pageNumber="278" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
, ca. 1.4× as long as wide, widest at midlength. Body broadly oval, outline not parallel (
<figureCitation id="13642A6D4347BC680CBDFAA1FECA7275" box="[253,322,1375,1401]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1673,1697]" captionTargetBox="[236,1361,474,1632]" captionTargetId="figure-149@4.[151,1436,450,1647]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus of Dactylosternum spp. from China and Japan. 1: D. abdominale (Fabricius, 1792), 2: D. corbetti Balfour-Browne, 1942, 3: D. nanlingensis sp. nov., 4: D. pui Jia, 2002" figureDoi="http://doi.org/10.5281/zenodo.5843839" httpUri="https://zenodo.org/record/5843839/files/figure.png" pageId="9" pageNumber="278">Fig. 3</figureCitation>
), strongly convex (
<figureCitation id="13642A6D4347BC680E62FAA1FD267275" box="[546,686,1375,1401]" captionStart="FIGURES 11–16" captionStartId="10.[151,264,1853,1877]" captionTargetBox="[255,1413,665,1819]" captionTargetId="figure-225@10.[151,1436,633,1830]" captionTargetPageId="10" captionText="FIGURES 11–16. Habitus of D. nanlingensis sp. nov. and D. songxiaobini sp. nov. 11–13: Habitus of D. nanlingensis sp. nov., holotype (11: ventral view, 12: lateral view, 13: frontal view), 14–16: Habitus of D. songxiaobini sp. nov., holotype (14: ventral view, 15: lateral view, 16: frontal view)" figureDoi="http://doi.org/10.5281/zenodo.5843843" httpUri="https://zenodo.org/record/5843843/files/figure.png" pageId="9" pageNumber="278">Figs. 12–13</figureCitation>
). Antenna yellow-brown with club black, club loosely segmented (
<figureCitation id="13642A6D4347BC680CE0FA7DFF7B7291" box="[160,243,1411,1437]" captionStart="FIGURES 39–47" captionStartId="19.[151,264,1466,1490]" captionTargetBox="[151,1431,183,1437]" captionTargetId="figure-19@19.[151,1436,181,1442]" captionTargetPageId="19" captionText="FIGURES 39–47. Morphology characters of Dactylosternum spp. SEM micrographs. 39–40: antennal club (39: D. nanlingensis sp. nov., 40: D. abdominale), 41–42: ventral surface of mesofemora (41: D. pui, 42: D. nanlingensis sp. nov.), 43–47: punctures on pronotum (43: D. pui, 44: D. songxiaobini sp. nov. (surface with two sizes of punctures, the blue arrow indicates the coarser puncture, the yellow arrow indicates the finer puncture), 45: D. nanlingensis sp. nov., 46: D. corbetti (arrow indicates two short divergent lines connected with a puncture), 47: D. abdominale (arrow indicates mesh-like microsculpture between punctures))" figureDoi="http://doi.org/10.5281/zenodo.5843879" httpUri="https://zenodo.org/record/5843879/files/figure.png" pageId="9" pageNumber="278">Fig. 39</figureCitation>
). Surface of vertex, pronotum and elytron with fine punctures and microsculpture, each puncture connected with two short diverging lines forming a flying-bird-like impression (
<figureCitation id="13642A6D4347BC680FDDFA59FC7972CD" box="[925,1009,1446,1473]" captionStart="FIGURES 39–47" captionStartId="19.[151,264,1466,1490]" captionTargetBox="[151,1431,183,1437]" captionTargetId="figure-19@19.[151,1436,181,1442]" captionTargetPageId="19" captionText="FIGURES 39–47. Morphology characters of Dactylosternum spp. SEM micrographs. 39–40: antennal club (39: D. nanlingensis sp. nov., 40: D. abdominale), 41–42: ventral surface of mesofemora (41: D. pui, 42: D. nanlingensis sp. nov.), 43–47: punctures on pronotum (43: D. pui, 44: D. songxiaobini sp. nov. (surface with two sizes of punctures, the blue arrow indicates the coarser puncture, the yellow arrow indicates the finer puncture), 45: D. nanlingensis sp. nov., 46: D. corbetti (arrow indicates two short divergent lines connected with a puncture), 47: D. abdominale (arrow indicates mesh-like microsculpture between punctures))" figureDoi="http://doi.org/10.5281/zenodo.5843879" httpUri="https://zenodo.org/record/5843879/files/figure.png" pageId="9" pageNumber="278">Fig. 45</figureCitation>
). Elytra slightly explanate in posterior half, sutural stria reaching ca. one-thirds of elytra posteriorly, with 10 rows of serial punctures, all nearly reaching elytral base. Posterior margin of ventrite 5 with a broad marginal bead. 
<emphasis id="B92BEAFA4347BC680F99FA10FBC17105" bold="true" box="[985,1097,1518,1545]" italics="true" pageId="9" pageNumber="278">Aedeagus</emphasis>
(
<figureCitation id="13642A6D4347BC68081CFA11FB3B7105" box="[1116,1203,1519,1545]" captionStart="FIGURES 20–24" captionStartId="13.[151,264,1946,1970]" captionTargetBox="[236,1409,756,1826]" captionTargetId="figure-217@13.[164,1423,741,1922]" captionTargetPageId="13" captionText="FIGURES 20–24. Aedeagus of Dactylosternum spp. 20: D. abdominale (specimen from Yonaguni-jima Island, Japan, dorsal view, showing variation of apex of paramera) 21: D. corbetti, 22: D. pui (phallobase missing), 23: D. nanlingensis sp. nov., paratype (23a: ventral view, 23b: dorsal view), 24: D. hydrophiloides" figureDoi="http://doi.org/10.5281/zenodo.5843857" httpUri="https://zenodo.org/record/5843857/files/figure.png" pageId="9" pageNumber="278">Fig. 23</figureCitation>
): Parameres pointed at apex. Median lobe a little shorter than parameres, outer margin nearly parallel-sided in the medial, gradually narrowing apical, gonopore situated at apex.
</paragraph>
</subSubSection>
<subSubSection id="C34565634347BC6D0C87F9A4FC2C7669" lastPageId="12" lastPageNumber="281" pageId="9" pageNumber="278" type="description">
<paragraph id="8BE036E84347BC680C87F9A4FCAD7195" blockId="9.[151,1437,1158,2013]" pageId="9" pageNumber="278">
<emphasis id="B92BEAFA4347BC680C87F9A4FED17178" bold="true" box="[199,345,1626,1652]" pageId="9" pageNumber="278">Description.</emphasis>
Total length 
<quantity id="4CA79B0D4347BC680DACF9A4FDFB7179" box="[492,627,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="9" pageNumber="278" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
(
<typeStatus id="54E4884A4347BC680EC1F9A5FD617179" box="[641,745,1627,1653]" pageId="9" pageNumber="278" type="holotype">holotype</typeStatus>
: 
<quantity id="4CA79B0D4347BC680EB3F9A5FCC57179" box="[755,845,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="278" unit="mm" value="4.3">4.3 mm</quantity>
); maximum width 
<quantity id="4CA79B0D4347BC680860F9A5FB2F7179" box="[1056,1191,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="9" pageNumber="278" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 mm</quantity>
(
<typeStatus id="54E4884A4347BC6808F5F9A5FA957179" box="[1205,1309,1627,1653]" pageId="9" pageNumber="278" type="holotype">holotype</typeStatus>
: 
<quantity id="4CA79B0D4347BC680968F9A4FA187179" box="[1320,1424,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="9" pageNumber="278" unit="mm" value="2.75">2.75 mm</quantity>
); body broadly oval (
<figureCitation id="13642A6D4347BC680D32F981FE3E7195" box="[370,438,1663,1689]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1673,1697]" captionTargetBox="[236,1361,474,1632]" captionTargetId="figure-149@4.[151,1436,450,1647]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus of Dactylosternum spp. from China and Japan. 1: D. abdominale (Fabricius, 1792), 2: D. corbetti Balfour-Browne, 1942, 3: D. nanlingensis sp. nov., 4: D. pui Jia, 2002" figureDoi="http://doi.org/10.5281/zenodo.5843839" httpUri="https://zenodo.org/record/5843839/files/figure.png" pageId="9" pageNumber="278">Fig. 3</figureCitation>
), strongly convex (
<figureCitation id="13642A6D4347BC680ECFF981FC907195" box="[655,792,1663,1689]" captionStart="FIGURES 11–16" captionStartId="10.[151,264,1853,1877]" captionTargetBox="[255,1413,665,1819]" captionTargetId="figure-225@10.[151,1436,633,1830]" captionTargetPageId="10" captionText="FIGURES 11–16. Habitus of D. nanlingensis sp. nov. and D. songxiaobini sp. nov. 11–13: Habitus of D. nanlingensis sp. nov., holotype (11: ventral view, 12: lateral view, 13: frontal view), 14–16: Habitus of D. songxiaobini sp. nov., holotype (14: ventral view, 15: lateral view, 16: frontal view)" figureDoi="http://doi.org/10.5281/zenodo.5843843" httpUri="https://zenodo.org/record/5843843/files/figure.png" pageId="9" pageNumber="278">Figs. 12–13</figureCitation>
).
</paragraph>
<paragraph id="8BE036E84347BC680C87F95CFF777009" blockId="9.[151,1437,1158,2013]" pageId="9" pageNumber="278">
<emphasis id="B92BEAFA4347BC680C87F95CFEC271B1" bold="true" box="[199,330,1698,1725]" italics="true" pageId="9" pageNumber="278">Coloration.</emphasis>
Dorsal black, with lateral margin of elytron slightly paler (
<figureCitation id="13642A6D4347BC680FB8F95DFBB771B1" box="[1016,1087,1699,1725]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1673,1697]" captionTargetBox="[236,1361,474,1632]" captionTargetId="figure-149@4.[151,1436,450,1647]" captionTargetPageId="4" captionText="FIGURES 1–4. Habitus of Dactylosternum spp. from China and Japan. 1: D. abdominale (Fabricius, 1792), 2: D. corbetti Balfour-Browne, 1942, 3: D. nanlingensis sp. nov., 4: D. pui Jia, 2002" figureDoi="http://doi.org/10.5281/zenodo.5843839" httpUri="https://zenodo.org/record/5843839/files/figure.png" pageId="9" pageNumber="278">Fig. 3</figureCitation>
). Maxillary palps and antenna yellowish brown with club black. Ventral black, mouthparts and legs reddish brown, tarsomeres reddish yellow (
<figureCitation id="13642A6D4347BC680CE0F915FF797009" box="[160,241,1771,1797]" captionStart="FIGURES 11–16" captionStartId="10.[151,264,1853,1877]" captionTargetBox="[255,1413,665,1819]" captionTargetId="figure-225@10.[151,1436,633,1830]" captionTargetPageId="10" captionText="FIGURES 11–16. Habitus of D. nanlingensis sp. nov. and D. songxiaobini sp. nov. 11–13: Habitus of D. nanlingensis sp. nov., holotype (11: ventral view, 12: lateral view, 13: frontal view), 14–16: Habitus of D. songxiaobini sp. nov., holotype (14: ventral view, 15: lateral view, 16: frontal view)" figureDoi="http://doi.org/10.5281/zenodo.5843843" httpUri="https://zenodo.org/record/5843843/files/figure.png" pageId="9" pageNumber="278">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BE036E84347BC6B0C87F8F0FEC976DD" blockId="9.[151,1437,1158,2013]" lastBlockId="10.[151,1437,150,609]" lastPageId="10" lastPageNumber="279" pageId="9" pageNumber="278">
<emphasis id="B92BEAFA4347BC680C87F8F0FE847025" bold="true" box="[199,268,1806,1833]" italics="true" pageId="9" pageNumber="278">Head.</emphasis>
Clypeus weakly convex, surface with sparse, moderately coarse fine punctures, each puncture with a flying-bird-like microsculpture. Anterior margin of clypeus without marginal rim. Frontoclypeal suture weakly developed but distinct. Frons with punctures sparser than those on clypeus. Eyes of moderate size, nearly semicircular in dorsal view, separated by ca. 5.2× as wide as one eye, distinctly emarginate anteriorly in lateral view, smooth medially. Maxilla with basistipes bearing sparse long setae. Maxillary palps stout, palpomere 2 swollen, thicker than the following two palpomeres, palpomere 4 with obtuse apex, ca. 1.3× as long as penultimate. Mentum transverse, ca. 1.5× as wide as long, widest between anterolateral corners, slightly narrowing posteriorly, bearing sparse long setae, surface of mentum deeply concave anteromedially, lateral sides with series of fine setae. Submentum black, pubescent. Labial palps rather stout, palpomere 1 minute, palpomere 2 strongly widened distally, with dense brush of long yellow setae on its inner face and distal margin, palpomere 3 longer than other ones, a little narrower and and about 2× as long as penultimate. Antennal scapus longest, rather thick, ca. 3.5× as long as pedicellus, antennomere 3 thin and a little shorter than pedicellus, ca. 2× as long as antennomere 4, antennomeres 4 and 5 almost equal in length, slightly widening distally, without any seta, club densely pubescent, with few long setae, loosely segmented, ca. 6× as long as wide, antennomeres 7 and 8 almost equal in length, antennomere 9 symmetrical, ca. 1.8× as long as penultimate.
</paragraph>
<paragraph id="8BE036E84344BC6B0C87FE24FDFD756D" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="279">
<emphasis id="B92BEAFA4344BC6B0C87FE24FECC76F9" bold="true" box="[199,324,474,501]" italics="true" pageId="10" pageNumber="279">Prothorax.</emphasis>
Pronotum transverse, strongly convex, arcuately narrowing anteriorly, anterolateral angles obtusely rounded, posterolateral angles almost rectangular, anterior and lateral margins with narrow marginal bead, pronotum surface with punctures similar to those on vertex (
<figureCitation id="13642A6D4344BC6B0E95FDDDFCA27531" box="[725,810,546,573]" captionStart="FIGURES 39–47" captionStartId="19.[151,264,1466,1490]" captionTargetBox="[151,1431,183,1437]" captionTargetId="figure-19@19.[151,1436,181,1442]" captionTargetPageId="19" captionText="FIGURES 39–47. Morphology characters of Dactylosternum spp. SEM micrographs. 39–40: antennal club (39: D. nanlingensis sp. nov., 40: D. abdominale), 41–42: ventral surface of mesofemora (41: D. pui, 42: D. nanlingensis sp. nov.), 43–47: punctures on pronotum (43: D. pui, 44: D. songxiaobini sp. nov. (surface with two sizes of punctures, the blue arrow indicates the coarser puncture, the yellow arrow indicates the finer puncture), 45: D. nanlingensis sp. nov., 46: D. corbetti (arrow indicates two short divergent lines connected with a puncture), 47: D. abdominale (arrow indicates mesh-like microsculpture between punctures))" figureDoi="http://doi.org/10.5281/zenodo.5843879" httpUri="https://zenodo.org/record/5843879/files/figure.png" pageId="10" pageNumber="279">Fig. 45</figureCitation>
). Posterior margin slightly arcuate. Prosternum weakly tectiform medially, without median carina.
</paragraph>
<caption id="DF2066604344BC6B0CD7F8C3FDAE7091" ID-DOI="http://doi.org/10.5281/zenodo.5843843" ID-Zenodo-Dep="5843843" httpUri="https://zenodo.org/record/5843843/files/figure.png" pageId="10" pageNumber="279" startId="10.[151,264,1853,1877]" targetBox="[255,1413,665,1819]" targetPageId="10">
<paragraph id="8BE036E84344BC6B0CD7F8C3FDAE7091" blockId="10.[151,1436,1853,1949]" pageId="10" pageNumber="279">
<emphasis id="B92BEAFA4344BC6B0CD7F8C3FEDA7059" bold="true" box="[151,338,1853,1877]" pageId="10" pageNumber="279">FIGURES 11–16.</emphasis>
Habitus of 
<taxonomicName id="4C5F4D6B4344BC6B0D87F8C0FDED7059" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[455,613,1853,1877]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA4344BC6B0D87F8C0FDED7059" box="[455,613,1853,1877]" italics="true" pageId="10" pageNumber="279">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4344BC6B0E2BF8C3FD327059" bold="true" box="[619,698,1853,1877]" pageId="10" pageNumber="279">
<taxonomicNameLabel id="A21857814344BC6B0E2BF8C3FD327059" box="[619,698,1853,1877]" pageId="10" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C5F4D6B4344BC6B0EABF8C0FC047059" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[747,908,1853,1877]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="songxiaobini" status="sp. nov.">
<emphasis id="B92BEAFA4344BC6B0EABF8C0FC047059" box="[747,908,1853,1877]" italics="true" pageId="10" pageNumber="279">D. songxiaobini</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4344BC6B0FD1F8C3FC697059" bold="true" box="[913,993,1853,1877]" pageId="10" pageNumber="279">
<taxonomicNameLabel id="A21857814344BC6B0FD1F8C3FC697059" box="[913,993,1853,1877]" pageId="10" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
11–13: Habitus of 
<taxonomicName id="4C5F4D6B4344BC6B08E2F8C0FAC87059" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[1186,1344,1853,1877]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA4344BC6B08E2F8C0FAC87059" box="[1186,1344,1853,1877]" italics="true" pageId="10" pageNumber="279">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4344BC6B0906F8C3FA1D7059" bold="true" box="[1350,1429,1853,1877]" pageId="10" pageNumber="279">
<taxonomicNameLabel id="A21857814344BC6B0906F8C3FA1D7059" box="[1350,1429,1853,1877]" pageId="10" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (11: ventral view, 12: lateral view, 13: frontal view), 14–16: Habitus of 
<taxonomicName id="4C5F4D6B4344BC6B0F85F89CFBEE7075" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[965,1126,1889,1913]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="songxiaobini" status="sp. nov.">
<emphasis id="B92BEAFA4344BC6B0F85F89CFBEE7075" box="[965,1126,1889,1913]" italics="true" pageId="10" pageNumber="279">D. songxiaobini</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4344BC6B082CF89FFB347075" bold="true" box="[1132,1212,1889,1913]" pageId="10" pageNumber="279">
<taxonomicNameLabel id="A21857814344BC6B082CF89FFB347075" box="[1132,1212,1889,1913]" pageId="10" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (14: ventral view, 15: lateral view, 16: frontal view)
</paragraph>
</caption>
<caption id="DF2066604345BC6A0CD7FB31FB217223" ID-DOI="http://doi.org/10.5281/zenodo.5843853" ID-Zenodo-Dep="5843853" httpUri="https://zenodo.org/record/5843853/files/figure.png" pageId="11" pageNumber="280" startId="11.[151,264,1231,1255]" targetBox="[151,1435,190,1207]" targetPageId="11">
<paragraph id="8BE036E84345BC6A0CD7FB31FB217223" blockId="11.[151,1436,1231,1328]" pageId="11" pageNumber="280">
<emphasis id="B92BEAFA4345BC6A0CD7FB31FED973EB" bold="true" box="[151,337,1231,1255]" pageId="11" pageNumber="280">FIGURES 17-19.</emphasis>
Morphology of 
<taxonomicName id="4C5F4D6B4345BC6A0DBBFB31FD4673EB" box="[507,718,1231,1255]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="280" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92BEAFA4345BC6A0DBBFB31FD1573EB" box="[507,669,1231,1255]" italics="true" pageId="11" pageNumber="280">Dactylosternum</emphasis>
spp.
</taxonomicName>
17: hind wing of 
<taxonomicName id="4C5F4D6B4345BC6A0FC8FB2EFBA073EB" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[904,1064,1231,1255]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="280" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B92BEAFA4345BC6A0FC8FB2EFBA073EB" box="[904,1064,1231,1255]" italics="true" pageId="11" pageNumber="280">D. nanlingensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4345BC6A086FFB2EFB0973E4" bold="true" box="[1071,1153,1232,1256]" pageId="11" pageNumber="280">
<taxonomicNameLabel id="A21857814345BC6A086FFB2EFB0973E4" box="[1071,1153,1232,1256]" pageId="11" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 18–19: Anterior margin of clypeus of 
<taxonomicName id="4C5F4D6B4345BC6A0D46FB0AFE347207" baseAuthorityName="MacLeay" baseAuthorityYear="1825" box="[262,444,1267,1291]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Fungi" order="Coleoptera" pageId="11" pageNumber="280" phylum="Ascomycota" rank="species" species="hydrophiloides">
<emphasis id="B92BEAFA4345BC6A0D46FB0AFE347207" box="[262,444,1267,1291]" italics="true" pageId="11" pageNumber="280">D. hydrophiloides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D6B4345BC6A0DADFB0AFD057207" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[493,653,1267,1291]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="280" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA4345BC6A0DADFB0AFD057207" box="[493,653,1267,1291]" italics="true" pageId="11" pageNumber="280">D. pseudolatum</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4345BC6A0ED3FB0AFD6C7200" bold="true" box="[659,740,1268,1292]" pageId="11" pageNumber="280">
<taxonomicNameLabel id="A21857814345BC6A0ED3FB0AFD6C7200" box="[659,740,1268,1292]" pageId="11" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(18: 
<taxonomicName id="4C5F4D6B4345BC6A0F5AFB0AFC477207" baseAuthorityName="MacLeay" baseAuthorityYear="1825" box="[794,975,1267,1291]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Fungi" order="Coleoptera" pageId="11" pageNumber="280" phylum="Ascomycota" rank="species" species="hydrophiloides">
<emphasis id="B92BEAFA4345BC6A0F5AFB0AFC477207" box="[794,975,1267,1291]" italics="true" pageId="11" pageNumber="280">D. hydrophiloides</emphasis>
</taxonomicName>
(arrow indicates the marginal rim on anterior margin of clypeus), 19: 
<taxonomicName id="4C5F4D6B4345BC6A0DC9FAE6FDA17223" authority="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček, 2022" authorityName="Mai &amp; Hu &amp; Minoshima &amp; Jia &amp; Fikáček" authorityYear="2022" box="[393,553,1303,1327]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="280" phylum="Arthropoda" rank="species" species="pseudolatum" status="sp. nov.">
<emphasis id="B92BEAFA4345BC6A0DC9FAE6FDA17223" box="[393,553,1303,1327]" italics="true" pageId="11" pageNumber="280">D. pseudolatum</emphasis>
</taxonomicName>
<emphasis id="B92BEAFA4345BC6A0E70FAE6FD09723C" bold="true" box="[560,641,1304,1328]" pageId="11" pageNumber="280">
<taxonomicNameLabel id="A21857814345BC6A0E70FAE6FD09723C" box="[560,641,1304,1328]" pageId="11" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(without marginal rim on anterior margin of clypeus))
</paragraph>
</caption>
<paragraph id="8BE036E84345BC6A0C87FAA7FAB07067" blockId="11.[151,1436,1368,2043]" pageId="11" pageNumber="280">
<emphasis id="B92BEAFA4345BC6A0C87FAA7FE5A727F" bold="true" box="[199,466,1368,1395]" italics="true" pageId="11" pageNumber="280">Meso- and metathorax.</emphasis>
Scutellum slightly wider than long, in shape of equilateral triangle, with fine punctures similar to those on pronotum. Elytron strongly convex, sutural stria reaching ca. one-third of elytra posteriorly, with 10 longitudinal serial punctures, all nearly reaching elytral base, series 1–6 consisting of very fine and sparse punctures, lateral four series with punctures larger and deeper than in inner six series, series 5+6 linked together subposteriorly. Elytral intervals with fine punctures similar with those on pronotum (
<figureCitation id="13642A6D4345BC6A087DFA16FB18710E" box="[1085,1168,1512,1538]" captionStart="FIGURES 48–63" captionStartId="21.[151,264,1495,1519]" captionTargetBox="[153,1430,183,1463]" captionTargetId="figure-19@21.[151,1436,181,1471]" captionTargetPageId="21" captionText="FIGURES 48–63. Morphology characters of Dactylosternum spp. SEM micrographs. 48–50: mediobasal portion of elytron (48: D. songxiaobini sp. nov. (red arrows show primary serial punctures, green arrows show secondary serial punctures), 49: D. nanlingensis sp. nov., 50: D. corbetti), 51–55: punctures on posterior half of elytron (51: D. hydrophiloides (the red arrow indicates the ground puncture, the blue arrow indicates the micropuncture), 52: D. pseudolatum sp. nov., 53: D. salvazai, 54: D. latum, 55: D. frater sp. nov.), 56–59: anteromedial portion of metaventrite (56: D. pseudolatum sp. nov., 57: D. salvazai, 58: D. latum, 59: D. frater sp. nov.), 60–63: posteromedial portion of ventrite 5 (60: D. pseudolatum sp. nov., 61: D. salvazai, 62: D. latum, 63: D. frater sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.5843883" httpUri="https://zenodo.org/record/5843883/files/figure.png" pageId="11" pageNumber="280">Fig. 49</figureCitation>
). Elytral margin slightly explanate in posterior half. Hind wing (
<figureCitation id="13642A6D4345BC6A0E10F9F2FD2A712A" box="[592,674,1548,1574]" captionStart="FIGURES 17-19" captionStartId="11.[151,264,1231,1255]" captionTargetBox="[151,1435,190,1207]" captionTargetId="figure-19@11.[151,1435,181,1207]" captionTargetPageId="11" captionText="FIGURES 17-19. Morphology of Dactylosternum spp. 17: hind wing of D. nanlingensis sp. nov., 18–19: Anterior margin of clypeus of D. hydrophiloides and D. pseudolatum sp. nov. (18: D. hydrophiloides (arrow indicates the marginal rim on anterior margin of clypeus), 19: D. pseudolatum sp. nov. (without marginal rim on anterior margin of clypeus))" figureDoi="http://doi.org/10.5281/zenodo.5843853" httpUri="https://zenodo.org/record/5843853/files/figure.png" pageId="11" pageNumber="280">Fig. 17</figureCitation>
) well developed, with R-M loop widest at distal; RA 
<subScript id="17DB34AD4345BC6A08B4F9E1FA867121" attach="left" box="[1268,1294,1567,1581]" fontSize="6" pageId="11" pageNumber="280">3+4</subScript>
not attached to RA basally, forming a large opened radial cell; r3 absent; r4 present, connecting RA 
<subScript id="17DB34AD4345BC6A082FF9BDFB01715D" attach="left" box="[1135,1161,1603,1617]" fontSize="6" pageId="11" pageNumber="280">3+4</subScript>
to R-M loop; RA 
<subScript id="17DB34AD4345BC6A0910F9BDFAD0715D" attach="left" box="[1360,1368,1603,1617]" fontSize="6" pageId="11" pageNumber="280">3</subScript>
short, well defined at apical portion; RA 
<subScript id="17DB34AD4345BC6A0E60F999FDA07179" attach="left" box="[544,552,1639,1653]" fontSize="6" pageId="11" pageNumber="280">4</subScript>
fused with RP 
<subScript id="17DB34AD4345BC6A0E95F999FD557179" attach="left" box="[725,733,1639,1653]" fontSize="6" pageId="11" pageNumber="280">1</subScript>
forming a pigmented area, nearly reaching the margin of the wing apically, RP 
<subScript id="17DB34AD4345BC6A0D1CF975FEEC7195" attach="left" box="[348,356,1675,1689]" fontSize="6" pageId="11" pageNumber="280">1</subScript>
attached to R-M loop basally; RP 
<subScript id="17DB34AD4345BC6A0E9EF975FD6E7195" attach="left" box="[734,742,1675,1689]" fontSize="6" pageId="11" pageNumber="280">2</subScript>
forming a pigmented area wide on base, attached to R-M loop basally but not reaching the margin of the wing apically; RP 
<subScript id="17DB34AD4345BC6A0F14F951FCE671B1" attach="left" box="[852,878,1711,1725]" fontSize="6" pageId="11" pageNumber="280">3+4</subScript>
well developed, long and narrow, attached R-M loop basally, curved posteriorly and reaching wing margin apically; medial spur straight and sharp; MP 
<subScript id="17DB34AD4345BC6A0962F92DFAB471ED" attach="left" box="[1314,1340,1747,1761]" fontSize="6" pageId="11" pageNumber="280">3+4</subScript>
distinct, cambered, connected with MP 
<subScript id="17DB34AD4345BC6A0DB4F909FD867009" attach="left" box="[500,526,1783,1797]" fontSize="6" pageId="11" pageNumber="280">1+2</subScript>
, not attached to MP 
<subScript id="17DB34AD4345BC6A0EBCF909FC8C7009" attach="left" box="[764,772,1783,1797]" fontSize="6" pageId="11" pageNumber="280">3</subScript>
apically; MP 
<subScript id="17DB34AD4345BC6A0FE0F909FC207009" attach="left" box="[928,936,1783,1797]" fontSize="6" pageId="11" pageNumber="280">3</subScript>
and MP 
<subScript id="17DB34AD4345BC6A084DF909FB9D7009" attach="left" box="[1037,1045,1783,1797]" fontSize="6" pageId="11" pageNumber="280">4</subScript>
+CuA 
<subScript id="17DB34AD4345BC6A081AF909FBEA7009" attach="left" box="[1114,1122,1783,1797]" fontSize="6" pageId="11" pageNumber="280">1</subScript>
well defined, connected to basal portions of the veins proximally, reaching wing margin posteriorly; CuA 
<subScript id="17DB34AD4345BC6A0866F8E5FBA67025" attach="left" box="[1062,1070,1819,1833]" fontSize="6" pageId="11" pageNumber="280">1</subScript>
present, connect with MP 
<subScript id="17DB34AD4345BC6A0921F8E5FAE17025" attach="left" box="[1377,1385,1819,1833]" fontSize="6" pageId="11" pageNumber="280">3</subScript>
and MP 
<subScript id="17DB34AD4345BC6A0C80F8C1FF407041" attach="right" box="[192,200,1855,1869]" fontSize="6" pageId="11" pageNumber="280">4</subScript>
+CuA 
<subScript id="17DB34AD4345BC6A0D4CF8C1FE9C7041" attach="left" box="[268,276,1855,1869]" fontSize="6" pageId="11" pageNumber="280">1</subScript>
; CuA 
<subScript id="17DB34AD4345BC6A0D18F8C1FEE87041" attach="left" box="[344,352,1855,1869]" fontSize="6" pageId="11" pageNumber="280">2</subScript>
well defined, reaching the posterior edge of the wing apically; AA 
<subScript id="17DB34AD4345BC6A080CF8C1FBDC7041" attach="left" box="[1100,1108,1855,1869]" fontSize="6" pageId="11" pageNumber="280">3</subScript>
and AA 
<subScript id="17DB34AD4345BC6A08F1F8C1FB317041" attach="left" box="[1201,1209,1855,1869]" fontSize="6" pageId="11" pageNumber="280">4</subScript>
well defined, nearly reaching posterior margin of the wing; venation of anal field strongly reduced, composed of a short AP 
<subScript id="17DB34AD4345BC6A0958F89DFABA707D" attach="left" box="[1304,1330,1891,1905]" fontSize="6" pageId="11" pageNumber="280">3+4</subScript>
.
</paragraph>
<paragraph id="8BE036E84345BC6A0C87F88BFBC870F6" blockId="11.[151,1436,1368,2043]" pageId="11" pageNumber="280">
<emphasis id="B92BEAFA4345BC6A0C87F88BFE8D7083" bold="true" box="[199,261,1909,1935]" italics="true" pageId="11" pageNumber="280">Legs.</emphasis>
Reddish brown, rather short. Procoxae globular, pubescent; profemora bearing dense pubescence throughout; protarsomeres with dense pubescence ventrally.Ventral surface of mesofemora covered with pubescence on marginal portion (
<figureCitation id="13642A6D4345BC6A0DCCF842FE6A70DA" box="[396,482,1980,2006]" captionStart="FIGURES 39–47" captionStartId="19.[151,264,1466,1490]" captionTargetBox="[151,1431,183,1437]" captionTargetId="figure-19@19.[151,1436,181,1442]" captionTargetPageId="19" captionText="FIGURES 39–47. Morphology characters of Dactylosternum spp. SEM micrographs. 39–40: antennal club (39: D. nanlingensis sp. nov., 40: D. abdominale), 41–42: ventral surface of mesofemora (41: D. pui, 42: D. nanlingensis sp. nov.), 43–47: punctures on pronotum (43: D. pui, 44: D. songxiaobini sp. nov. (surface with two sizes of punctures, the blue arrow indicates the coarser puncture, the yellow arrow indicates the finer puncture), 45: D. nanlingensis sp. nov., 46: D. corbetti (arrow indicates two short divergent lines connected with a puncture), 47: D. abdominale (arrow indicates mesh-like microsculpture between punctures))" figureDoi="http://doi.org/10.5281/zenodo.5843879" httpUri="https://zenodo.org/record/5843879/files/figure.png" pageId="11" pageNumber="280">Fig. 42</figureCitation>
), metafemora without pubescence, with fine punctures and sparse microsculptures; meso- and metatarsomeres with dense pubescence ventrally, almost equal in length.
</paragraph>
<paragraph id="8BE036E84342BC6D0C87FF68FDF677D9" blockId="12.[151,1437,150,790]" pageId="12" pageNumber="281">
<emphasis id="B92BEAFA4342BC6D0C87FF68FEB477BD" bold="true" box="[199,316,150,177]" italics="true" pageId="12" pageNumber="281">Abdomen.</emphasis>
Pubescent, ventrite 1 with complete median carina. Posterior apex of ventrite 5 nearly truncate, with an incomplete marginal bead along margin.
</paragraph>
<paragraph id="8BE036E84342BC6D0C87FF20FC2C7669" blockId="12.[151,1437,150,790]" pageId="12" pageNumber="281">
<emphasis id="B92BEAFA4342BC6D0C87FF20FEFB77F5" bold="true" box="[199,371,222,249]" italics="true" pageId="12" pageNumber="281">Male genitalia</emphasis>
(
<figureCitation id="13642A6D4342BC6D0DC7FF21FE6877F5" box="[391,480,223,249]" captionStart="FIGURES 20–24" captionStartId="13.[151,264,1946,1970]" captionTargetBox="[236,1409,756,1826]" captionTargetId="figure-217@13.[164,1423,741,1922]" captionTargetPageId="13" captionText="FIGURES 20–24. Aedeagus of Dactylosternum spp. 20: D. abdominale (specimen from Yonaguni-jima Island, Japan, dorsal view, showing variation of apex of paramera) 21: D. corbetti, 22: D. pui (phallobase missing), 23: D. nanlingensis sp. nov., paratype (23a: ventral view, 23b: dorsal view), 24: D. hydrophiloides" figureDoi="http://doi.org/10.5281/zenodo.5843857" httpUri="https://zenodo.org/record/5843857/files/figure.png" pageId="12" pageNumber="281">Fig. 23</figureCitation>
). Aedeagus about 
<quantity id="4CA79B0D4342BC6D0EFAFF21FC9D77F5" box="[698,789,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="281" unit="mm" value="0.9">0.9 mm</quantity>
long, ca. 2.6× as long as wide. Phallobase short, with manubrium short and rounded posteriorly. Parameres widest in the middle, gradually narrowing towards apex, outside slightly and smoothly curved. Median lobe a little shorter than parameres, wide throughout, ca. 3× as long as phallobase, gradually narrowing apical, gonopore situated apically.
</paragraph>
</subSubSection>
<subSubSection id="C34565634342BC6D0C87FE91FBDD7685" box="[199,1109,367,393]" pageId="12" pageNumber="281" type="etymology">
<paragraph id="8BE036E84342BC6D0C87FE91FBDD7685" blockId="12.[151,1437,150,790]" box="[199,1109,367,393]" pageId="12" pageNumber="281">
<emphasis id="B92BEAFA4342BC6D0C87FE91FEC67685" bold="true" box="[199,334,367,393]" pageId="12" pageNumber="281">Etymology.</emphasis>
This species is named after the 
<typeStatus id="54E4884A4342BC6D0EF4FE91FD6C7685" box="[692,740,367,393]" pageId="12" pageNumber="281">type</typeStatus>
locality, the Nanling Mountains.
</paragraph>
</subSubSection>
<subSubSection id="C34565634342BC6D0C87FE6CFF5576F9" pageId="12" pageNumber="281" type="biology_ecology">
<paragraph id="8BE036E84342BC6D0C87FE6CFF5576F9" blockId="12.[151,1437,150,790]" pageId="12" pageNumber="281">
<emphasis id="B92BEAFA4342BC6D0C87FE6CFEA976A0" bold="true" box="[199,289,402,428]" pageId="12" pageNumber="281">Biology</emphasis>
(
<figureCitation id="13642A6D4342BC6D0D71FE6DFE3D76A2" box="[305,437,403,430]" captionStart="FIGURES 64–69" captionStartId="23.[151,264,1684,1708]" captionTargetBox="[158,1428,380,1658]" captionTargetId="figure-106@23.[158,1428,380,1660]" captionTargetPageId="23" captionText="FIGURES 64–69. Habitats of Dactylosternum spp. 64, 66: individual specimen of D. nanlingensis sp. nov. on the surface of a fallen wood in forest at night, 65: habitats of D. abdominale (Fabricius, 1792), D. corbetti Balfour-Browne, 1942 and D. hydrophiloides (MacLeay, 1825), specimens hiding in a decaying banana tree, 67: D. abdominale, 68: D. corbetti, 69: D. hydrophiloides. Figures 64, 66 photo by Mr. Bao-Ping Huang in Ruyuan County, Tianjingshan National Forest Park." figureDoi="http://doi.org/10.5281/zenodo.5843889" httpUri="https://zenodo.org/record/5843889/files/figure.png" pageId="12" pageNumber="281">Figs 64, 66</figureCitation>
). Specimens were collected from the surface of fallen wood in forest. The surface of the wood was wet and covered with thick layer of fulgal hyphae. Adults crawled quickly at the surface of the fallen wood.
</paragraph>
</subSubSection>
<subSubSection id="C34565634342BC6D0C87FE01FC1775FD" pageId="12" pageNumber="281" type="discussion">
<paragraph id="8BE036E84342BC6D0C87FE01FC1775FD" blockId="12.[151,1437,150,790]" pageId="12" pageNumber="281">
<emphasis id="B92BEAFA4342BC6D0C87FE01FEB37515" bold="true" box="[199,315,511,537]" pageId="12" pageNumber="281">Remarks.</emphasis>
This species is quite similar to 
<taxonomicName id="4C5F4D6B4342BC6D0EF6FE01FC6F7515" authority="Knisch, 1921" authorityName="Knisch" authorityYear="1921" box="[694,999,511,537]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="cycloides">
<emphasis id="B92BEAFA4342BC6D0EF6FE01FCCB7515" box="[694,835,511,537]" italics="true" pageId="12" pageNumber="281">D. cycloides</emphasis>
<bibRefCitation id="EFCE4B194342BC6D0F0EFE01FC6F7515" author="Knisch, A." box="[846,999,511,537]" pageId="12" pageNumber="281" pagination="55 - 88" refId="ref22834" refString="Knisch, A. (1921) Die exotischen Hydrophiliden des Deutschen Entomologischen Museums (Col.) Archiv fur Naturgeschichte, 8, 55 - 88. [85 A (1919)]" type="journal article" year="1921">Knisch, 1921</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F34876784342BC6D0872FE01FB177515" box="[1074,1183,511,537]" name="Indonesia" pageId="12" pageNumber="281">Indonesia</collectingCountry>
and 
<collectingCountry id="F34876784342BC6D089DFE01FACC7515" box="[1245,1348,511,537]" name="Malaysia" pageId="12" pageNumber="281">Malaysia</collectingCountry>
and 
<taxonomicName id="4C5F4D6B4342BC6D09C1FE01FE0D7531" authority="Fauvel, 1883" authorityName="Fauvel" authorityYear="1883" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="auripes">
<emphasis id="B92BEAFA4342BC6D09C1FE01FF637530" italics="true" pageId="12" pageNumber="281">D. auripes</emphasis>
Fauvel, 1883
</taxonomicName>
(
<figureCitation id="13642A6D4342BC6D0DD4FDDDFE6E7531" box="[404,486,547,573]" captionStart="FIGURES 70–71" captionStartId="24.[151,264,1327,1351]" captionTargetBox="[244,1400,309,1243]" captionTargetId="figure-30@24.[151,1436,211,1282]" captionTargetPageId="24" captionText="FIGURES 70–71. Habitus of Dactylosternum spp. 70: Dactylosternum auripes Fauvel, 1883, 71: Dactylosternum dytiscoides (Fabricius, 1775)" figureDoi="http://doi.org/10.5281/zenodo.5843895" httpUri="https://zenodo.org/record/5843895/files/figure.png" pageId="12" pageNumber="281">Fig. 70</figureCitation>
) from 
<collectingCountry id="F34876784342BC6D0E73FDDDFD6C7531" box="[563,740,547,573]" name="New Caledonia" pageId="12" pageNumber="281">New Caledonia</collectingCountry>
. These two species also have a loose antennal club and highly convex body. This new species can be distinguished from 
<taxonomicName id="4C5F4D6B4342BC6D0F7AFDB9FC4F756D" authorityName="Knisch" authorityYear="1921" box="[826,967,583,609]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="cycloides">
<emphasis id="B92BEAFA4342BC6D0F7AFDB9FC4F756D" box="[826,967,583,609]" italics="true" pageId="12" pageNumber="281">D. cycloides</emphasis>
</taxonomicName>
by posterior margin of ventrite 5 with a broad marginal bead (marginal bead not distinct in 
<taxonomicName id="4C5F4D6B4342BC6D0E97FD95FCE97589" authorityName="Knisch" authorityYear="1921" box="[727,865,619,645]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="cycloides">
<emphasis id="B92BEAFA4342BC6D0E97FD95FCE97589" box="[727,865,619,645]" italics="true" pageId="12" pageNumber="281">D. cycloides</emphasis>
</taxonomicName>
), apex of median lobe rounded but sharper (with a duller apex in 
<taxonomicName id="4C5F4D6B4342BC6D0D79FD71FE4C75A5" authorityName="Knisch" authorityYear="1921" box="[313,452,655,681]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="cycloides">
<emphasis id="B92BEAFA4342BC6D0D79FD71FE4C75A5" box="[313,452,655,681]" italics="true" pageId="12" pageNumber="281">D. cycloides</emphasis>
</taxonomicName>
). It differs from 
<taxonomicName id="4C5F4D6B4342BC6D0EC3FD71FD7175A4" authorityName="Fauvel" authorityYear="1883" box="[643,761,655,680]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="auripes">
<emphasis id="B92BEAFA4342BC6D0EC3FD71FD7175A4" box="[643,761,655,680]" italics="true" pageId="12" pageNumber="281">D. auripes</emphasis>
</taxonomicName>
by shallower and finer serial punctures on elytron (strongly impressed punctures in 
<taxonomicName id="4C5F4D6B4342BC6D0DE5FD4DFD9475C0" authorityName="Fauvel" authorityYear="1883" box="[421,540,691,716]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="auripes">
<emphasis id="B92BEAFA4342BC6D0DE5FD4DFD9475C0" box="[421,540,691,716]" italics="true" pageId="12" pageNumber="281">D. auripes</emphasis>
</taxonomicName>
), inner 5 punctural series reaching elytral base (not reaching elytral base in 
<taxonomicName id="4C5F4D6B4342BC6D09C1FD4DFF6375FC" authorityName="Fauvel" authorityYear="1883" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="auripes">
<emphasis id="B92BEAFA4342BC6D09C1FD4DFF6375FC" italics="true" pageId="12" pageNumber="281">D. auripes</emphasis>
</taxonomicName>
), median lobe rounded apically (apex truncate in 
<taxonomicName id="4C5F4D6B4342BC6D0F59FD29FC0775FC" authorityName="Fauvel" authorityYear="1883" box="[793,911,727,752]" class="Insecta" family="Hydrophilidae" genus="Dactylosternum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="auripes">
<emphasis id="B92BEAFA4342BC6D0F59FD29FC0775FC" box="[793,911,727,752]" italics="true" pageId="12" pageNumber="281">D. auripes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34565634342BC6D0C87FD04FD25741A" box="[199,685,762,790]" pageId="12" pageNumber="281" type="distribution">
<paragraph id="8BE036E84342BC6D0C87FD04FD25741A" blockId="12.[151,1437,150,790]" box="[199,685,762,790]" pageId="12" pageNumber="281">
<emphasis id="B92BEAFA4342BC6D0C87FD04FEE97418" bold="true" box="[199,353,762,788]" pageId="12" pageNumber="281">Distribution.</emphasis>
<collectingCountry id="F34876784342BC6D0D27FD05FE227419" box="[359,426,763,789]" name="China" pageId="12" pageNumber="281">China</collectingCountry>
(
<collectingRegion id="499BF80A4342BC6D0DF9FD05FDC87419" box="[441,576,763,789]" country="China" name="Guangdong" pageId="12" pageNumber="281">Guangdong</collectingRegion>
, 
<collectingRegion id="499BF80A4342BC6D0E0BFD05FD29741A" box="[587,673,763,790]" country="China" name="Jiangxi" pageId="12" pageNumber="281">Jiangxi</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>