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<document id="A14B511078C18292D0670F870F6E1990" ID-CLB-Dataset="46302" ID-DOI="10.5281/zenodo.207728" ID-GBIF-Dataset="98e2411c-6d02-4050-b16b-ab46286722a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="207728" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460392142752" checkinUser="plazi" docAuthor="Mariano, Rodolfo" docDate="2011" docId="03F687F2E1691151FF07FC135B7DF816" docLanguage="en" docName="zt03038p050.pdf" docOrigin="Zootaxa 3038" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paramaka pearljam Mariano, 2011, sp. n." docType="treatment" docVersion="10" lastPageNumber="49" masterDocId="FFCFFF8AE1681155FF90FFC55E17FFE3" masterDocTitle="A new species of Paramaka Savage &amp; Domínguez, 1992 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Brazil" masterLastPageNumber="50" masterPageNumber="45" pageNumber="46" updateTime="1698298332333" updateUser="ExternalLinkService">
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<mods:title id="1757E4B19184462D3C145AE307DD219E">A new species of Paramaka Savage &amp; Domínguez, 1992 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Brazil</mods:title>
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<mods:namePart id="24608F447BEAFCC95E158EEB1180502F">Mariano, Rodolfo</mods:namePart>
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<treatment id="03F687F2E1691151FF07FC135B7DF816" ID-DOI="http://doi.org/10.5281/zenodo.6186347" ID-GBIF-Taxon="119417027" ID-Zenodo-Dep="6186347" LSID="urn:lsid:plazi:treatment:03F687F2E1691151FF07FC135B7DF816" httpUri="http://treatment.plazi.org/id/03F687F2E1691151FF07FC135B7DF816" lastPageId="4" lastPageNumber="49" pageId="1" pageNumber="46">
<subSubSection id="C345656FE1691154FF07FC135F05FBF2" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="8BE036E4E1691154FF07FC135FCFFC13" blockId="1.[151,472,982,1041]" box="[151,472,982,1008]" pageId="1" pageNumber="46">
<heading id="D0A88188E1691154FF07FC135FCFFC13" bold="true" box="[151,472,982,1008]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
<emphasis id="B92BEAF6E1691154FF07FC135FCFFC13" bold="true" box="[151,472,982,1008]" pageId="1" pageNumber="46">
<taxonomicName id="4C5F4D67E1691154FF07FC135F90FC13" ID-CoL="4D298" box="[151,391,982,1008]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="pearljam" status="sp. nov.">
<emphasis id="B92BEAF6E1691154FF07FC135F90FC13" bold="true" box="[151,391,982,1008]" italics="true" pageId="1" pageNumber="46">Paramaka pearljam</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A218578DE1691154FE05FC135FCFFC13" box="[405,472,982,1008]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE036E4E1691154FF07FC3C5F05FBF2" blockId="1.[151,472,982,1041]" box="[151,274,1017,1041]" pageId="1" pageNumber="46">
(
<figureCitation id="13642A61E1691154FF0FFC3C5F1EFBF2" box="[159,265,1017,1041]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,249,1907,1930]" captionStartId-1="3.[151,250,1895,1918]" captionStartId-2="4.[151,250,976,999]" captionTargetBox-0="[201,1383,195,1856]" captionTargetBox-1="[151,1436,193,1863]" captionTargetBox-2="[211,1376,193,965]" captionTargetId-0="figure@2.[201,1384,195,1867]" captionTargetId-1="figure@3.[151,1436,193,1874]" captionTargetId-2="figure@4.[211,1376,193,965]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." captionText-1="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." captionText-2="FIGURE 3. Imago of Paramaka pearljam, sp. n. (lateral view)." httpUri-0="https://zenodo.org/record/207729/files/figure.png" httpUri-1="https://zenodo.org/record/207730/files/figure.png" httpUri-2="https://zenodo.org/record/207731/files/figure.png" pageId="1" pageNumber="46">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C345656FE1691154FF07FB845EE0FAEE" pageId="1" pageNumber="46" type="diagnosis">
<paragraph id="8BE036E4E1691154FF07FB845F38FB9E" blockId="1.[151,1437,1089,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF07FB845F1DFBB9" bold="true" box="[151,266,1089,1114]" pageId="1" pageNumber="46">Diagnosis</emphasis>
.
<taxonomicName id="4C5F4D67E1691154FE88FB845FEEFBBA" box="[280,505,1089,1113]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="pearljam">
<emphasis id="B92BEAF6E1691154FE88FB845FEEFBBA" box="[280,505,1089,1113]" italics="true" pageId="1" pageNumber="46">Paramaka pearljam</emphasis>
</taxonomicName>
can be separated from the other species of the genus by the following combination of characters:
</paragraph>
<paragraph id="8BE036E4E1691154FF55FB4F5EE0FAEE" blockId="1.[151,1437,1089,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF55FB4F5F32FB41" bold="true" box="[197,293,1162,1186]" pageId="1" pageNumber="46">Nymph.</emphasis>
1) gills on abdominal segments IVII and vestigial on abdominal segment VII, platelike, not ending in a apical process; 2) labrum expanded laterally, 1.3 maximum width of clypeus; 3) posterolateral projection weakly developed on abdominal segments VIII and more prominent on segment IX; 4) tarsal claw without an accessory denticle.
</paragraph>
</subSubSection>
<subSubSection id="C345656FE1691151FF55FADF5FFAF9FD" lastPageId="4" lastPageNumber="49" pageId="1" pageNumber="46" type="description">
<paragraph id="8BE036E4E1691154FF55FADF5A61FAB6" blockId="1.[151,1437,1089,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF55FADF5F4BFAD1" bold="true" box="[197,348,1306,1330]" pageId="1" pageNumber="46">Male Imago.</emphasis>
1) Coloration as in
<figureCitation id="13642A61E1691154FDAAFADC5C6CFAD2" box="[570,635,1305,1329]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="1" pageNumber="46">Fig. 2</figureCitation>
E and
<figureCitation id="13642A61E1691154FD53FADC5D11FAD2" box="[707,774,1305,1329]" captionStart="FIGURE 3" captionStartId="4.[151,250,976,999]" captionTargetBox="[211,1376,193,965]" captionTargetId="figure@4.[211,1376,193,965]" captionTargetPageId="4" captionText="FIGURE 3. Imago of Paramaka pearljam, sp. n. (lateral view)." httpUri="https://zenodo.org/record/207731/files/figure.png" pageId="1" pageNumber="46">Fig. 3</figureCitation>
; 2) styliger plate with a conspicuous and large median projection with apex curved ventrally; 3) each lobe of penes with a membranous projection.
</paragraph>
<paragraph id="8BE036E4E1691154FF57FAA75D9BFA9A" blockId="1.[151,1437,1089,2014]" box="[199,908,1377,1402]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF57FAA75F90FA99" bold="true" box="[199,391,1378,1402]" pageId="1" pageNumber="46">Mature Nymph.</emphasis>
(
<figureCitation id="13642A61E1691154FE06FAA45FC0FA9A" box="[406,471,1377,1401]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
) General coloration: brown yellowish.
</paragraph>
<paragraph id="8BE036E4E1691154FF55FA405EE1F9B2" blockId="1.[151,1437,1089,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF55FA405F12FA7E" bold="true" box="[197,261,1413,1437]" pageId="1" pageNumber="46">Head</emphasis>
. Yellowish; eyes of female black; antenna with pedicel yellow. Mouthparts: yellow; labrum (
<figureCitation id="13642A61E1691154FAABFA405B6AFA7E" box="[1339,1405,1413,1437]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
B) rounded with a median depression; labrum expanded laterally, 1.3 maximum width of clypeus, anterior margin almost straight; lateral margins strongly concave; mandibles (
<figureCitation id="13642A61E1691154FCC4FA085D83FA06" box="[852,916,1485,1509]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
C) yellowish, without bristles on outer dorsal margin; maxillae yellow (
<figureCitation id="13642A61E1691154FE2AFA345FECF9EA" box="[442,507,1521,1545]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
D); labium (
<figureCitation id="13642A61E1691154FD1AFA345CDEF9EA" box="[650,713,1521,1545]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
A) with segment III of palpi similar in length to segment I, and segment II slightly larger than I and III; paraglossa with numerous anterodorsal bristles; glossa with fine bristles on apex.
</paragraph>
<paragraph id="8BE036E4E1691154FF55F9985CBCF902" blockId="1.[151,1437,1089,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF55F9985F0AF996" bold="true" box="[197,285,1629,1653]" pageId="1" pageNumber="46">Thorax</emphasis>
. General color of pronotum, mesonotum and metanotum brown with suture brown yellowish. Legs yellowish; all femora with short spines and a few long setae near to apex on outer margin, and short spines along the inner margin (
<figureCitation id="13642A61E1691154FEF4F9605FB2F95E" box="[356,421,1701,1725]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
E); apex of claw without accessory denticle (
<figureCitation id="13642A61E1691154FC3EF9605DF8F95E" box="[942,1007,1701,1725]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
G); fore tarsi (
<figureCitation id="13642A61E1691154FB08F9605AC6F95E" box="[1176,1233,1701,1725]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig.1</figureCitation>
E) with spines on inner margin, progressively larger toward apex.
</paragraph>
<paragraph id="8BE036E4E1691154FF55F9285F21F85A" blockId="1.[151,1437,1089,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF55F9285F2EF8E5" bold="true" box="[197,313,1773,1798]" pageId="1" pageNumber="46">Abdomen</emphasis>
. (
<figureCitation id="13642A61E1691154FEC1F9285F87F8E6" box="[337,400,1773,1797]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
F, 1H) Tergites IIV yellow and tergites VX yellowbrownish with a medial black line on all; sternites IX yellow; gills presents in abdominal segments IVII, basally light gray, progressively darker distally to about 3/4 of distance from base, pale apically; median tracheal trunk gray; all gills on abdominal segments IVII and vestigial on abdominal segment VII, platelike, not ended in a thin apical process. Posterolateral projection weakly developed on abdominal segments VIII and more prominent on segment IX (
<figureCitation id="13642A61E1691154FB00F8B85AC6F876" box="[1168,1233,1917,1941]" captionStart="FIGURE 1" captionStartId="2.[151,249,1907,1930]" captionTargetBox="[201,1383,195,1856]" captionTargetId="figure@2.[201,1384,195,1867]" captionTargetPageId="2" captionText="FIGURE 1 (A G). Paramaka pearljam, sp. n., nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); H abdomen, lateral view (segments VII X)." httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="1" pageNumber="46">Fig. 1</figureCitation>
H). Terminal filaments broken.
</paragraph>
<paragraph id="8BE036E4E1691154FF57F8005B76F83E" blockId="1.[151,1437,1089,2014]" box="[199,1377,1989,2014]" pageId="1" pageNumber="46">
<emphasis id="B92BEAF6E1691154FF57F8005F41F83E" bold="true" box="[199,342,1989,2013]" pageId="1" pageNumber="46">Male Imago</emphasis>
(
<figureCitation id="13642A61E1691154FEF6F8005FA2F83E" box="[358,437,1989,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="1" pageNumber="46">Figs. 2</figureCitation>
,
<figureCitation id="13642A61E1691154FE51F8005FD8F83E" box="[449,463,1989,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,976,999]" captionTargetBox="[211,1376,193,965]" captionTargetId="figure@4.[211,1376,193,965]" captionTargetPageId="4" captionText="FIGURE 3. Imago of Paramaka pearljam, sp. n. (lateral view)." httpUri="https://zenodo.org/record/207731/files/figure.png" pageId="1" pageNumber="46">3</figureCitation>
) (in alcohol and slide). General coloration orange brown as in
<figureCitation id="13642A61E1691154FB00F8005AC7F83E" box="[1168,1232,1989,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="1" pageNumber="46">Fig. 2</figureCitation>
E and
<figureCitation id="13642A61E1691154FA88F8005B4BF83E" box="[1304,1372,1989,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,976,999]" captionTargetBox="[211,1376,193,965]" captionTargetId="figure@4.[211,1376,193,965]" captionTargetPageId="4" captionText="FIGURE 3. Imago of Paramaka pearljam, sp. n. (lateral view)." httpUri="https://zenodo.org/record/207731/files/figure.png" pageId="1" pageNumber="46">Fig. 3</figureCitation>
.
</paragraph>
<caption id="DF20666CE16A1157FF07F8B65CC9F824" httpUri="https://zenodo.org/record/207729/files/figure.png" pageId="2" pageNumber="47" targetBox="[201,1383,195,1856]" targetPageId="2">
<paragraph id="8BE036E4E16A1157FF07F8B65CC9F824" blockId="2.[151,1436,1907,1991]" pageId="2" pageNumber="47">
<emphasis id="B92BEAF6E16A1157FF07F8B65F48F869" bold="true" box="[151,351,1907,1930]" pageId="2" pageNumber="47">FIGURE 1 (AG).</emphasis>
<taxonomicName id="4C5F4D67E16A1157FEF7F8B15C24F869" box="[359,563,1908,1930]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="pearljam" status="sp. nov.">
<emphasis id="B92BEAF6E16A1157FEF7F8B15C24F869" box="[359,563,1908,1930]" italics="true" pageId="2" pageNumber="47">Paramaka pearljam</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAF6E16A1157FDD2F8B15C6BF86A" bold="true" box="[578,636,1908,1929]" pageId="2" pageNumber="47">
<taxonomicNameLabel id="A218578DE16A1157FDD2F8B15C6BF86A" box="[578,636,1908,1929]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, nymph. A, labium (ventral view); B, labrum (left: ventral view, right: dorsal view); C, right mandible (ventral view); D, maxilla (ventral view); E, foreleg; F, abdominal color pattern; G, foreleg (tibia and tarsal claw); Habdomen, lateral view (segments VIIX).
</paragraph>
</caption>
<caption id="DF20666CE16B1156FF07F8A25D19F858" httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="3" pageNumber="48" targetBox="[151,1436,193,1863]" targetPageId="3">
<paragraph id="8BE036E4E16B1156FF07F8A25D19F858" blockId="3.[151,1437,1895,1979]" pageId="3" pageNumber="48">
<emphasis id="B92BEAF6E16B1156FF07F8A25F4FF89D" bold="true" box="[151,344,1895,1918]" pageId="3" pageNumber="48">FIGURE 2 (AF).</emphasis>
<taxonomicName id="4C5F4D67E16B1156FECFF8AD5C3EF89D" box="[351,553,1896,1918]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="pearljam" status="sp. nov.">
<emphasis id="B92BEAF6E16B1156FECFF8AD5C3EF89D" box="[351,553,1896,1918]" italics="true" pageId="3" pageNumber="48">Paramaka pearljam</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAF6E16B1156FDA6F8AD5C78F89E" bold="true" box="[566,623,1896,1917]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="A218578DE16B1156FDA6F8AD5C78F89E" box="[566,623,1896,1917]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments IVII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection).
</paragraph>
</caption>
<caption id="DF20666CE16C1151FF07FC155D36FC04" box="[151,801,976,999]" httpUri="https://zenodo.org/record/207731/files/figure.png" pageId="4" pageNumber="49" targetBox="[211,1376,193,965]" targetPageId="4">
<paragraph id="8BE036E4E16C1151FF07FC155D36FC04" blockId="4.[151,801,976,999]" box="[151,801,976,999]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF07FC155F05FC05" bold="true" box="[151,274,976,999]" pageId="4" pageNumber="49">FIGURE 3.</emphasis>
Imago of
<taxonomicName id="4C5F4D67E16C1151FEEBFC145C50FC04" box="[379,583,977,999]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pearljam" status="sp. nov.">
<emphasis id="B92BEAF6E16C1151FEEBFC145C50FC04" box="[379,583,977,999]" italics="true" pageId="4" pageNumber="49">Paramaka pearljam</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAF6E16C1151FDC3FC145C9BFC05" bold="true" box="[595,652,977,998]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="A218578DE16C1151FDC3FC145C9BFC05" box="[595,652,977,998]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(lateral view).
</paragraph>
</caption>
<paragraph id="8BE036E4E16C1151FF55FBCA5DDCFBA9" blockId="4.[151,1436,1038,2037]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF55FBCA5F12FBC4" bold="true" box="[197,261,1039,1063]" pageId="4" pageNumber="49">Head</emphasis>
. Brown; eyes of male meeting on meson of head, upper portion of eyes brownish and lowers portion black; ocelli white; antennal scape brown, pedicel brown, flagellum pale.
</paragraph>
<paragraph id="8BE036E4E16C1151FF55FB925C03FB39" blockId="4.[151,1436,1038,2037]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF55FB925F0AFB8C" bold="true" box="[197,285,1111,1135]" pageId="4" pageNumber="49">Thorax</emphasis>
. Chestnut, with tergal suture yellowish brown, and sternal suture brown. Wings (
<figureCitation id="13642A61E16C1151FB52FB935B10FB8D" box="[1218,1287,1110,1134]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Fig. 2</figureCitation>
A, 2B): forewings hyaline with veins and crossveins yellowish.
<collectingCountry id="F3487674E16C1151FD79FBBF5D36FB71" box="[745,801,1146,1170]" name="India" pageId="4" pageNumber="49">Hind</collectingCountry>
wings: costal projection well developed. Legs: femora, tibia and tarsi yellowish, tarsal claws dissimilar, one hooked apically, other padlike. Except for apex of femur and base of tibia III tinged with black.
</paragraph>
<paragraph id="8BE036E4E16C1151FF55FB235C4FFA19" blockId="4.[151,1436,1038,2037]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF55FB235F2DFB1C" bold="true" box="[197,314,1254,1279]" pageId="4" pageNumber="49">Abdomen</emphasis>
(
<figureCitation id="13642A61E16C1151FED9FB235F80FB1D" box="[329,407,1254,1278]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Figs. 2</figureCitation>
E, 3). All tergites with a dorsal median black band; tergites I and II tinged with brown; III VI brown with a white anterior band, and IV and V with a white posterior band; VIIX dark brown; in lateral view, IIVI with a white anterior spot, and a small white spot near to spiracles; sternites III brown; IIIVI brown with two white anterior spots, III and IV with a white posterior band; VIIX dark brown;in some specimens general color of abdomen as in
<figureCitation id="13642A61E16C1151FE37FAB35FFBFA6D" box="[423,492,1398,1422]" captionStart="FIGURE 3" captionStartId="4.[151,250,976,999]" captionTargetBox="[211,1376,193,965]" captionTargetId="figure@4.[211,1376,193,965]" captionTargetPageId="4" captionText="FIGURE 3. Imago of Paramaka pearljam, sp. n. (lateral view)." httpUri="https://zenodo.org/record/207731/files/figure.png" pageId="4" pageNumber="49">Fig. 3</figureCitation>
. Genitalia (
<figureCitation id="13642A61E16C1151FDE2FAB35CD5FA6D" box="[626,706,1398,1422]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Figs. 2</figureCitation>
C, 2D, 2F): forceps yellowish; styliger plate with a conspicuous and large median projection with apex curved ventrally (indicated by an arrow in
<figureCitation id="13642A61E16C1151FBBFFA5F5A66FA51" box="[1071,1137,1434,1458]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Fig. 2</figureCitation>
D and
<figureCitation id="13642A61E16C1151FB2DFA5F5AE8FA51" box="[1213,1279,1434,1458]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Fig. 2</figureCitation>
F); inner margin of segment I of forceps with short spines; penis rounded on the apex and with a membranous inner projection (
<figureCitation id="13642A61E16C1151FF0FFA275EF4FA19" box="[159,227,1506,1530]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Fig. 2</figureCitation>
C). Terminal filament broken off.
</paragraph>
<paragraph id="8BE036E4E16C1151FF57F9C25FFAF9FD" blockId="4.[151,1436,1038,2037]" box="[199,493,1542,1567]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF57F9C25F6FF9FC" bold="true" box="[199,376,1543,1567]" pageId="4" pageNumber="49">Female Imago.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="C345656FE16C1151FF57F9EF5A91F985" pageId="4" pageNumber="49" type="biology_ecology">
<paragraph id="8BE036E4E16C1151FF57F9EF5A91F985" blockId="4.[151,1436,1038,2037]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF57F9EF5F30F9A0" bold="true" box="[199,295,1578,1603]" pageId="4" pageNumber="49">Biology.</emphasis>
The specimens of
<taxonomicName id="4C5F4D67E16C1151FD96F9EE5C9EF9A1" box="[518,649,1578,1602]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pearljam">
<emphasis id="B92BEAF6E16C1151FD96F9EE5C9EF9A1" box="[518,649,1578,1602]" italics="true" pageId="4" pageNumber="49">P. pearljam</emphasis>
</taxonomicName>
were collected in a low order permanent stony and sanding bottom stream. The collection of nymphs was made manually and the imagos caught at light trap.
</paragraph>
</subSubSection>
<subSubSection id="C345656FE16C1151FF57F9B65DA3F969" box="[199,948,1650,1675]" pageId="4" pageNumber="49" type="etymology">
<paragraph id="8BE036E4E16C1151FF57F9B65DA3F969" blockId="4.[151,1436,1038,2037]" box="[199,948,1650,1675]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF57F9B65F59F968" bold="true" box="[199,334,1651,1675]" pageId="4" pageNumber="49">Etymology.</emphasis>
Named to celebrate the 20th anniversary of Pearl Jam.
</paragraph>
</subSubSection>
<subSubSection id="C345656FE16C1151FF57F9535A31F91A" pageId="4" pageNumber="49" type="materials_examined">
<paragraph id="8BE036E4E16C1151FF57F9535A31F91A" blockId="4.[151,1436,1038,2037]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF57F9535F21F94C" bold="true" box="[199,310,1686,1711]" pageId="4" pageNumber="49">Material.</emphasis>
<typeStatus id="54E48846E16C1151FEAEF9535FB1F94D" box="[318,422,1686,1710]" pageId="4" pageNumber="49" type="holotype">Holotype</typeStatus>
: 3imago:
<collectingCountry id="F3487674E16C1151FDB2F9535C7EF94D" box="[546,617,1686,1710]" name="Brazil" pageId="4" pageNumber="49">Brazil</collectingCountry>
, Mato Grosso State, Ribeirão Cascalheira, Fazenda Manaus, tributary of ribeirão Bonito, 2º ponte,
<date id="FFE11024E16C1151FE2DF9785C27F936" box="[445,560,1725,1749]" pageId="4" pageNumber="49" value="2007-10-07">07.x.2007</date>
,
<geoCoordinate id="EE6B5023E16C1151FDADF9785CB1F936" box="[573,678,1722,1749]" direction="south" orientation="latitude" pageId="4" pageNumber="49" precision="925" value="-12.916667">S 12o 55</geoCoordinate>
599”
<geoCoordinate id="EE6B5023E16C1151FD7DF97B5D2EF936" box="[749,825,1725,1749]" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="925" value="-0.85">W 0 51</geoCoordinate>
o 53 474”,
<quantity id="4CA79B01E16C1151FC2CF9785DE8F936" box="[956,1023,1725,1749]" metricMagnitude="2" metricUnit="m" metricValue="3.29" pageId="4" pageNumber="49" unit="m" value="329.0">329m</quantity>
, light trap, Pinho, L. C.; Mateus, S.; Torati, L. &amp; Silva, F.R..
<typeStatus id="54E48846E16C1151FE37F9275C0FF919" box="[423,536,1762,1786]" pageId="4" pageNumber="49" type="paratype">Paratypes</typeStatus>
: 2 3imagos, 2 nymphs: same data as
<typeStatus id="54E48846E16C1151FC2CF9245A36F91A" box="[956,1057,1761,1785]" pageId="4" pageNumber="49" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C345656FE16C1151FF57F8C05B7DF816" pageId="4" pageNumber="49" type="discussion">
<paragraph id="8BE036E4E16C1151FF57F8C05B7DF816" blockId="4.[151,1436,1038,2037]" pageId="4" pageNumber="49">
<emphasis id="B92BEAF6E16C1151FF57F8C05F2CF8FE" bold="true" box="[199,315,1797,1821]" pageId="4" pageNumber="49">Remarks.</emphasis>
The nymph of this species has the typical gill colouration pattern of
<taxonomicName id="4C5F4D67E16C1151FBCCF8C05AC7F8FE" box="[1116,1232,1797,1821]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAF6E16C1151FBCCF8C05AC7F8FE" box="[1116,1232,1797,1821]" italics="true" pageId="4" pageNumber="49">Paramaka</emphasis>
</taxonomicName>
, but their apex is always blunt and not ending in an apical process, as it can be seen in
<taxonomicName id="4C5F4D67E16C1151FC0CF8EF5A04F8A2" box="[924,1043,1834,1857]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="B92BEAF6E16C1151FC0CF8EF5A04F8A2" box="[924,1043,1834,1857]" italics="true" pageId="4" pageNumber="49">P. convexa</emphasis>
</taxonomicName>
on gills IIIVI and in
<taxonomicName id="4C5F4D67E16C1151FA9EF8EF5B6DF8A2" box="[1294,1402,1834,1857]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="antonii">
<emphasis id="B92BEAF6E16C1151FA9EF8EF5B6DF8A2" box="[1294,1402,1834,1857]" italics="true" pageId="4" pageNumber="49">P. antonii</emphasis>
</taxonomicName>
on gills IVVI. The absence of an accessory denticle on tarsal claw is another character to differ
<taxonomicName id="4C5F4D67E16C1151FB24F88B5B23F886" box="[1204,1332,1869,1893]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pearljam">
<emphasis id="B92BEAF6E16C1151FB24F88B5B23F886" box="[1204,1332,1869,1893]" italics="true" pageId="4" pageNumber="49">P. pearljam</emphasis>
</taxonomicName>
from the other two
<taxonomicName id="4C5F4D67E16C1151FE9CF8B45F96F86A" box="[268,385,1905,1929]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAF6E16C1151FE9CF8B45F96F86A" box="[268,385,1905,1929]" italics="true" pageId="4" pageNumber="49">Paramaka</emphasis>
</taxonomicName>
species.
<bibRefCitation id="EFCE4B15E16C1151FE7AF8B45C9DF86A" author="Sartori" box="[490,650,1905,1929]" pageId="4" pageNumber="49" refString="Sartori M. (2005) A new species of the genus Paramaka Savage &amp; Dominguez, 1992 with some comments on related genera (Ephemeroptera, Leptophlebiidae, Atalophlebiinae). Studies on Neotropical Fauna and Environment 40 (3), 237 - 245." type="journal article" year="2005">Sartori (2005)</bibRefCitation>
mentioned as the main difference between
<taxonomicName id="4C5F4D67E16C1151FBECF8B75AFDF86A" box="[1148,1258,1906,1929]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="antonii">
<emphasis id="B92BEAF6E16C1151FBECF8B75AFDF86A" box="[1148,1258,1906,1929]" italics="true" pageId="4" pageNumber="49">P. antonii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D67E16C1151FAB3F8B75B8CF86A" box="[1315,1435,1906,1929]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="B92BEAF6E16C1151FAB3F8B75B8CF86A" box="[1315,1435,1906,1929]" italics="true" pageId="4" pageNumber="49">P. convexa</emphasis>
</taxonomicName>
the presence of a welldeveloped carina on the anterior margin of the clypeus. In imago the main difference is the form of the styliger plate, as in
<taxonomicName id="4C5F4D67E16C1151FD9CF87F5C98F832" box="[524,655,1977,2001]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pearljam">
<emphasis id="B92BEAF6E16C1151FD9CF87F5C98F832" box="[524,655,1977,2001]" italics="true" pageId="4" pageNumber="49">P. pearljam</emphasis>
</taxonomicName>
the projection on styliger plate is conspicuous and large with apex curved ventrally (
<figureCitation id="13642A61E16C1151FECDF8185FBCF816" box="[349,427,2013,2037]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1918]" captionTargetBox="[151,1436,193,1863]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURE 2 (A F). Paramaka pearljam, sp. n., imago. A, forewing; B, hind wing; C, detail of penis (ventral view); D, styliger plate (lateral view) (arrow indicating the median projection); E, abdominal color pattern (segments I VII) (dorsal view); F, styliger plate (ventral view) (arrow indicating median projection)." httpUri="https://zenodo.org/record/207730/files/figure.png" pageId="4" pageNumber="49">Figs. 2</figureCitation>
D, 2F), whereas
<taxonomicName id="4C5F4D67E16C1151FDF4F81B5CCCF816" box="[612,731,2014,2037]" class="Insecta" family="Leptophlebiidae" genus="Paramaka" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="B92BEAF6E16C1151FDF4F81B5CCCF816" box="[612,731,2014,2037]" italics="true" pageId="4" pageNumber="49">P. convexa</emphasis>
</taxonomicName>
has a median ridge projecting dorsally between the penes.
</paragraph>
</subSubSection>
</treatment>
</document>