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<document id="9DC52C8A4BB3C4B471E662A2143CE5B0" ID-DOI="10.5281/zenodo.188586" ID-GBIF-Dataset="ddfbe6a3-4ba0-477c-82ba-0190603b6c15" ID-ISSN="1175-5326" ID-Zenodo-Dep="188586" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461095488965" checkinUser="plazi" docAuthor="Bosselaers, Jan" docDate="2009" docId="03B9291AFFD8F16EFF64F592FD55D157" docLanguage="en" docName="Bosselaers_2009.imd" docOrigin="Zootaxa 2141" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cybaeodes liocraninus Simon 1913, new combination" docType="treatment" docVersion="13" lastPageNumber="49" masterDocId="FF805162FFD1F162FFF3F327FF8FD505" masterDocTitle="Studies in Liocranidae (Araneae): redescriptions and transfers in Apostenus Westring and Brachyanillus Simon, as well as description of a new genus" masterLastPageNumber="55" masterPageNumber="37" pageNumber="46" updateTime="1698589017302" updateUser="plazi">
<mods:mods id="0FF44A230526155F95967D77DD77335D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="94994A973B7BA715EE777124011DA01F">Studies in Liocranidae (Araneae): redescriptions and transfers in Apostenus Westring and Brachyanillus Simon, as well as description of a new genus</mods:title>
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<mods:namePart id="C9E556A0D11496FDEED1F6FB7E518956">Bosselaers, Jan</mods:namePart>
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<mods:date id="4F4E56DB5F058532B852D296E643628F">2009</mods:date>
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<mods:classification id="34887FF8C54AEF2FAE917790210A53EA">journal article</mods:classification>
<mods:identifier id="883015E332D06F9F09BB84AB53ABB40F" type="DOI">10.5281/zenodo.188586</mods:identifier>
<mods:identifier id="E35F368C58376719C6C23E82A71E8E9E" type="GBIF-Dataset">ddfbe6a3-4ba0-477c-82ba-0190603b6c15</mods:identifier>
<mods:identifier id="190907FAB36E0A49F0CA01FFF7AC289C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9280EAEEE18E7C458694ED3F8593B2D4" type="Zenodo-Dep">188586</mods:identifier>
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<treatment id="03B9291AFFD8F16EFF64F592FD55D157" ID-DOI="http://doi.org/10.5281/zenodo.5576009" ID-GBIF-Taxon="119620239" ID-Zenodo-Dep="5576009" LSID="urn:lsid:plazi:treatment:03B9291AFFD8F16EFF64F592FD55D157" httpUri="http://treatment.plazi.org/id/03B9291AFFD8F16EFF64F592FD55D157" lastPageId="12" lastPageNumber="49" pageId="9" pageNumber="46">
<subSubSection id="C30ACB87FFD8F16BFF64F592FE56D3F7" pageId="9" pageNumber="46" type="nomenclature">
<paragraph id="8BAF980CFFD8F16BFF64F592FCB5D3CA" blockId="9.[151,826,1717,1778]" box="[151,826,1717,1743]" pageId="9" pageNumber="46">
<heading id="D0E72F60FFD8F16BFF64F592FCB5D3CA" bold="true" box="[151,826,1717,1743]" fontSize="11" level="1" pageId="9" pageNumber="46" reason="1">
<emphasis id="B964441EFFD8F16BFF64F592FCB5D3CA" bold="true" box="[151,826,1717,1743]" pageId="9" pageNumber="46">
<taxonomicName id="4C10E38FFFD8F16BFF64F592FDD2D3CA" ID-CoL="32RC8" authority="Simon, 1913" authorityName="Simon" authorityYear="1913" box="[151,605,1717,1743]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="liocraninus" status="comb. nov.">
<emphasis id="B964441EFFD8F16BFF64F592FE25D3CA" bold="true" box="[151,426,1717,1743]" italics="true" pageId="9" pageNumber="46">Cybaeodes liocraninus</emphasis>
(
<bibRefCitation id="EF81E5FDFFD8F16BFE4BF592FDDAD3CA" author="Simon" box="[440,597,1717,1743]" pageId="9" pageNumber="54" refString="Simon, E. (1913) Araneae et Opiliones (Quatrieme serie). In Biospeologica, XXX. Archives de zoologie experimentale et generale 52 (2), 359 - 386." type="journal article" year="1913">Simon, 1913</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A257F965FFD8F16BFD97F592FCB5D3CA" box="[612,826,1717,1743]" pageId="9" pageNumber="46">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF980CFFD8F16BFF64F5FFFE56D3F7" blockId="9.[151,826,1717,1778]" box="[151,473,1752,1778]" pageId="9" pageNumber="46">
<figureCitation id="132B8489FFD8F16BFF64F5FFFE89D3F7" box="[151,262,1752,1778]" captionStart="FIGURE 2 A – K" captionStartId="4.[151,257,1819,1843]" captionTargetBox="[168,1425,219,1763]" captionTargetId="figure@4.[151,1436,194,1795]" captionTargetPageId="4" captionText="FIGURE 2 A – K. Leg spination schematics. A, legend; B, Apostenus humilis, male holotype; C, Apostenus humilis, female, MNHN 25; D, Apostenus ochraceus, female; E, Agraecina rutilia, female holotype; F, Agraecina lineata, female, MNHN 5653; G, Cybaeodes liocraninus, holotype; H, Cybaeodes testaceus, female holotype, MNHN 3107; I, Cybaeodes madidus, female holotype, MNHN 1677; J, Cybaeodes marinae, female, Italy, Castelporziano, AMNH; K, Arabelia pheidoleicomes, female holotype." httpUri="https://zenodo.org/record/188588/files/figure.png" pageId="9" pageNumber="46">Figs. 2 G</figureCitation>
; 3 C–G; 6 A; 8 E.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFD8F16BFF64F402FDF6D239" box="[151,633,1829,1852]" pageId="9" pageNumber="46" type="reference_group">
<paragraph id="8BAF980CFFD8F16BFF64F402FDF6D239" blockId="9.[151,633,1829,1852]" box="[151,633,1829,1852]" pageId="9" pageNumber="46">
<treatmentCitationGroup id="AB00BF22FFD8F16BFF64F402FDF6D239" box="[151,633,1829,1852]" pageId="9" pageNumber="46">
<taxonomicName id="4C10E38FFFD8F16BFF64F402FDFBD239" authority="Simon, 1913: 378" authorityName="Simon" authorityPageNumber="378" authorityYear="1913" box="[151,628,1829,1852]" class="Arachnida" family="Liocranidae" genus="Brachyanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFD8F16BFF64F402FE23D239" box="[151,428,1829,1852]" italics="true" pageId="9" pageNumber="46">Brachyanillus liocraninus</emphasis>
<treatmentCitation id="0AB1BE1DFFD8F16BFE40F402FDFBD239" author="Simon" box="[435,628,1829,1852]" page="378" pageId="9" pageNumber="46" year="1913">
<bibRefCitation id="EF81E5FDFFD8F16BFE40F402FDB0D239" author="Simon" box="[435,575,1829,1852]" pageId="9" pageNumber="54" refString="Simon, E. (1913) Araneae et Opiliones (Quatrieme serie). In Biospeologica, XXX. Archives de zoologie experimentale et generale 52 (2), 359 - 386." type="journal article" year="1913">Simon, 1913</bibRefCitation>
: 378
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFD8F168FF64F449FDF1D228" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46" type="materials_examined">
<paragraph id="8BAF980CFFD8F16BFF64F449FEB7D2AA" blockId="9.[151,1437,1899,1967]" pageId="9" pageNumber="46">
<materialsCitation id="3B789251FFD8F16BFF64F449FEBBD2AA" ID-GBIF-Occurrence="1503373083" collectionCode="MNHN" country="Algeria" determinerName="E. S. G tte Misserghin" location="Algeria" pageId="9" pageNumber="46" specimenCode="499" specimenCount="1" specimenCount-juvenile="1" typeStatus="holotype">
<emphasis id="B964441EFFD8F16BFF64F449FEDBD28D" bold="true" box="[151,340,1902,1928]" pageId="9" pageNumber="46">
<typeStatus id="54AB26AEFFD8F16BFF64F449FF59D28D" box="[151,214,1902,1928]" pageId="9" pageNumber="46">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AB26AEFFD8F16BFEAEF448FE41D28C" box="[349,462,1903,1929]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
juvenile,
<collectingCountry id="F307D89CFFD8F16BFDB9F448FD26D28C" box="[586,681,1903,1929]" name="Algeria" pageId="9" pageNumber="46">Algeria</collectingCountry>
, &quot;
<taxonomicName id="4C10E38FFFD8F16BFD31F448FB8ED28D" box="[706,1025,1903,1928]" class="Arachnida" family="Liocranidae" genus="Brachyanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFD8F16BFD31F448FB8ED28D" box="[706,1025,1903,1928]" italics="true" pageId="9" pageNumber="46">Brachyanillus liocraninus</emphasis>
</taxonomicName>
E.S. G tte Misserghin (
<collectingCountry id="F307D89CFFD8F16BFAFCF448FAC0D28C" box="[1295,1359,1903,1929]" name="Algeria" pageId="9" pageNumber="46">Oran</collectingCountry>
) R.J.&quot; [
<collectionCode id="ED0100C9FFD8F16BFF6EF4B2FF71D2AA" LSID="urn:lsid:biocol.org:col:34988" box="[157,254,1941,1967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="46">MNHN</collectionCode>
499]
</materialsCitation>
.
</paragraph>
<caption id="DF6FC884FFDBF168FF64F5A6FB1FD3BE" httpUri="https://zenodo.org/record/188591/files/figure.png" pageId="10" pageNumber="47" targetBox="[222,1344,231,1608]" targetPageId="10">
<paragraph id="8BAF980CFFDBF168FF64F5A6FB1FD3BE" blockId="10.[151,1435,1665,1723]" pageId="10" pageNumber="47">
<emphasis id="B964441EFFDBF168FF64F5A6FEDED39C" bold="true" box="[151,337,1665,1689]" pageId="10" pageNumber="47">FIGURE 5 A–G.</emphasis>
<taxonomicName id="4C10E38FFFDBF168FEAFF5A5FDBAD39C" box="[348,565,1666,1689]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="madidus">
<emphasis id="B964441EFFDBF168FEAFF5A5FDBAD39C" box="[348,565,1666,1689]" italics="true" pageId="10" pageNumber="47">Cybaeodes madidus</emphasis>
</taxonomicName>
, holotype. A, carapace, do; B, carapace, lat; C, sternum and mouth parts, ve; D, carapace, fr; E, epigyne; F, leg II, tarsal tip; G, leg IV, tarsal tip. Scale bars: A–E: 0.5; F–G: 0.1.
</paragraph>
</caption>
<paragraph id="8BAF980CFFDBF168FF36F5CBFDF1D228" blockId="10.[151,1436,1770,2030]" pageId="10" pageNumber="47">
<materialsCitation id="3B789251FFDBF168FF36F5CBFDF5D228" ID-GBIF-Occurrence="1503373075" collectingDate="2013-03-02" collectionCode="MNHN" collectorName="Gr. te Misserghin" country="Algeria" determinerName=". S. biosp" location="Algeria" pageId="10" pageNumber="47" specimenCode="621" specimenCount="1" specimenCount-juvenile="1">
<emphasis id="B964441EFFDBF168FF36F5CBFD95D203" bold="true" box="[197,538,1772,1798]" pageId="10" pageNumber="47">Other material examined.</emphasis>
<specimenCount id="9D165385FFDBF168FDD0F5CBFD2DD203" box="[547,674,1772,1798]" pageId="10" pageNumber="47" type="juvenile">1 juvenile</specimenCount>
,
<collectingCountry id="F307D89CFFDBF168FD5CF5CBFC80D203" box="[687,783,1772,1798]" name="Algeria" pageId="10" pageNumber="47">Algeria</collectingCountry>
, &quot;
<taxonomicName id="4C10E38FFFDBF168FCD9F5CAFBE2D203" box="[810,1133,1773,1798]" class="Arachnida" family="Liocranidae" genus="Brachyanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFDBF168FCD9F5CAFBE2D203" box="[810,1133,1773,1798]" italics="true" pageId="10" pageNumber="47">Brachyanillus liocraninus</emphasis>
</taxonomicName>
E.S. biosp. n° 621. Gr.te Misserghin.
<collectingCountry id="F307D89CFFDBF168FEDBF434FEE9D228" box="[296,358,1811,1837]" name="Algeria" pageId="10" pageNumber="47">Oran</collectingCountry>
.
<date id="FFAEBECCFFDBF168FE81F434FE44D228" box="[370,459,1811,1837]" pageId="10" pageNumber="47" value="1913-03-02">2 III 13</date>
&quot; [
<collectionCode id="ED0100C9FFDBF168FE17F434FDC8D228" LSID="urn:lsid:biocol.org:col:34988" box="[484,583,1811,1837]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="47">MNHN</collectionCode>
621]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFDBF168FF36F41EFF5CD27E" pageId="10" pageNumber="47" type="diagnosis">
<paragraph id="8BAF980CFFDBF168FF36F41EFF5CD27E" blockId="10.[151,1436,1770,2030]" pageId="10" pageNumber="47">
<emphasis id="B964441EFFDBF168FF36F41EFECBD256" bold="true" box="[197,324,1849,1875]" pageId="10" pageNumber="47">Diagnosis.</emphasis>
<taxonomicName id="4C10E38FFFDBF168FEBFF41DFE7BD256" box="[332,500,1850,1875]" class="Arachnida" family="Liocranidae" genus="Brachyanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFDBF168FEBFF41DFE7BD256" box="[332,500,1850,1875]" italics="true" pageId="10" pageNumber="47">C. liocraninus</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="4C10E38FFFDBF168FC84F41DFC77D256" box="[887,1016,1850,1875]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFDBF168FC84F41DFC77D256" box="[887,1016,1850,1875]" italics="true" pageId="10" pageNumber="47">Cybaeodes</emphasis>
</taxonomicName>
species by the complete absence of eyes.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFDBF16EFF36F4A0FC36D49F" lastPageId="12" lastPageNumber="49" pageId="10" pageNumber="47" type="description">
<paragraph id="8BAF980CFFDBF16EFF36F4A0FC36D49F" blockId="10.[151,1436,1770,2030]" lastBlockId="12.[151,1437,152,1106]" lastPageId="12" lastPageNumber="49" pageId="10" pageNumber="47">
<emphasis id="B964441EFFDBF168FF36F4A0FED2D2A4" bold="true" box="[197,349,1927,1953]" pageId="10" pageNumber="47">Description.</emphasis>
Juvenile (
<typeStatus id="54AB26AEFFDBF168FE24F4A0FDCBD2A4" box="[471,580,1927,1953]" pageId="10" pageNumber="47" type="holotype">holotype</typeStatus>
). Total length 4.10. Carapace length 1.74,
<geoCoordinate id="EE24FECBFFDBF168FBCBF4A0FB04D2A4" box="[1080,1163,1927,1953]" direction="west" orientation="longitude" pageId="10" pageNumber="47" precision="555" value="-1.32">w 1.32</geoCoordinate>
, flattened (
<figureCitation id="132B8489FFDBF168FAFCF4A0FAEFD2A4" box="[1295,1376,1927,1953]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="10" pageNumber="47">Figs. 3</figureCitation>
D, 6 A), pale yellowish brown. Fovea dark brown, deep, length 0.16, anterior end 1.16 from front end of carapace. No eyes (
<figureCitation id="132B8489FFDBF168FEF5F4F3FEC3D2EB" box="[262,332,2004,2030]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="10" pageNumber="47">Fig. 3</figureCitation>
C, F). Chilum absent. Chelicerae yellow, with a row of thick, stout hairs along promarginal cheliceral rim. Three teeth along promarginal rim, halfway between fang base and base of paturon, largest tooth in the middle. Two small teeth on retromarginal rim, close to fang base. Sternum yellow, smooth, shieldshaped, length 1.05,
<geoCoordinate id="EE24FECBFFDAF169FE78F417FE50D24F" box="[395,479,1840,1866]" direction="west" orientation="longitude" pageId="11" pageNumber="48" precision="555" value="-0.84">w 0.84</geoCoordinate>
. No PCT, no ICS (
<figureCitation id="132B8489FFDAF169FD4EF417FC8CD24F" box="[701,771,1840,1866]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="11" pageNumber="48">Fig. 3</figureCitation>
E), PLB inconspicuous and isolated. Labium as wide as long. Endites subrectangular, frontally rounded, with oblique depression, apical hair tuft and serrula. Abdomen greyish white. ALS subconical, almost contiguous at their bases, with small apical segment. PMS small and slender, not flattened. PLS subconical, separated by slightly less than their length, terminal segment small, conical. Trochanters deeply notched, no rh, feathery hairs present on legs. Legs long, unicolorous yellow, leg formula 4123, leg length I 6.10 II 5.79 III 4.97 IV 7.47. Patellar indentation 2/3 of pa length, mt and ta with erectile bristles, vt preening brush on mt III, ta IV with pl basal spine, ta IV with a row of three do trichobothria in basal half. Tarsal claws with 5–7 teeth and five pairs of tenent hairs (
<figureCitation id="132B8489FFDDF16EFB8CF3C2FB49D5FA" box="[1151,1222,229,255]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="12" pageNumber="49">Fig. 3</figureCitation>
G). Leg spination (
<figureCitation id="132B8489FFDDF16EFF6CF22BFE8DD423" box="[159,258,268,294]" captionStart="FIGURE 2 A – K" captionStartId="4.[151,257,1819,1843]" captionTargetBox="[168,1425,219,1763]" captionTargetId="figure@4.[151,1436,194,1795]" captionTargetPageId="4" captionText="FIGURE 2 A – K. Leg spination schematics. A, legend; B, Apostenus humilis, male holotype; C, Apostenus humilis, female, MNHN 25; D, Apostenus ochraceus, female; E, Agraecina rutilia, female holotype; F, Agraecina lineata, female, MNHN 5653; G, Cybaeodes liocraninus, holotype; H, Cybaeodes testaceus, female holotype, MNHN 3107; I, Cybaeodes madidus, female holotype, MNHN 1677; J, Cybaeodes marinae, female, Italy, Castelporziano, AMNH; K, Arabelia pheidoleicomes, female holotype." httpUri="https://zenodo.org/record/188588/files/figure.png" pageId="12" pageNumber="49">Fig. 2 G</figureCitation>
) fe: I pl 0-0-1 do 1-1-0; II pl 0-0-1 do 1-1-0; III do 1-3-2; IV do 1-2-2; ti: I plv 1-1-1-1-1 rlv 1-1-1-1- 1; II plv 0-0-0-1 rlv 1-1-1-1; III pl 1-0-1 do 1-1-0 rl 1-0-1 plv 1-1-1 rlv 1-1-1; IV pl 1-0-1 do 1-1-0 rl 1-0-1 plv 1-1-1 rlv 1-1-1; mt: I plv 1-0-0 rlv 1-0-0; II plv 1-0-0 rlv 1-0-0; III pl 2-1-1 do 0-1-0 rl 1-1-1 plv 1-1-1 rlv 1-0- 1 vts 1; IV pl 1-1-1 rl 1-1-1 plv 1-1-1 rlv 1-1-1 vts 1; ta: IV pl 1-0-0.
</paragraph>
</subSubSection>
<caption id="DF6FC884FFDAF169FF64F55DFF54D3B1" httpUri="https://zenodo.org/record/188592/files/figure.png" pageId="11" pageNumber="48" targetBox="[193,1359,232,1595]" targetPageId="11">
<paragraph id="8BAF980CFFDAF169FF64F55DFF54D3B1" blockId="11.[151,1436,1658,1716]" pageId="11" pageNumber="48">
<emphasis id="B964441EFFDAF169FF64F55DFEBBD397" bold="true" box="[151,308,1658,1682]" pageId="11" pageNumber="48">FIGURE 6 A.</emphasis>
<taxonomicName id="4C10E38FFFDAF169FECFF55CFDBED397" box="[316,561,1659,1682]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFDAF169FECFF55CFDBED397" box="[316,561,1659,1682]" italics="true" pageId="11" pageNumber="48">Cybaeodes liocraninus</emphasis>
</taxonomicName>
, holotype.
<emphasis id="B964441EFFDAF169FD59F55DFD4DD397" bold="true" box="[682,706,1658,1682]" pageId="11" pageNumber="48">B.</emphasis>
<taxonomicName id="4C10E38FFFDAF169FD39F55CFC2FD397" box="[714,928,1659,1682]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="madidus">
<emphasis id="B964441EFFDAF169FD39F55CFC2FD397" box="[714,928,1659,1682]" italics="true" pageId="11" pageNumber="48">Cybaeodes madidus</emphasis>
</taxonomicName>
, holotype. A, habitus, lat; B, habitus, lat. Scale bar: 1.
</paragraph>
</caption>
<subSubSection id="C30ACB87FFDDF16EFF36F281FB44D12E" pageId="12" pageNumber="49" type="discussion">
<paragraph id="8BAF980CFFDDF16EFF36F281FD8ED787" blockId="12.[151,1437,152,1106]" pageId="12" pageNumber="49">
<emphasis id="B964441EFFDDF16EFF36F281FEDCD4C5" bold="true" box="[197,339,422,448]" pageId="12" pageNumber="49">Discussion.</emphasis>
Although Simon refers to the
<typeStatus id="54AB26AEFFDDF16EFD30F281FD79D4C5" box="[707,758,422,448]" pageId="12" pageNumber="49">type</typeStatus>
specimen as a female (without any comments about the genital morphology), study of the
<typeStatus id="54AB26AEFFDDF16EFDDBF2EAFDD5D4E2" box="[552,602,461,487]" pageId="12" pageNumber="49">type</typeStatus>
specimen as well as an additional specimen from his collection proves that he only saw juveniles. Nevertheless, the species is redescribed here and not treated as a
<emphasis id="B964441EFFDDF16EFB16F2D3FA1BD708" box="[1253,1428,500,525]" italics="true" pageId="12" pageNumber="49">nomen dubium</emphasis>
, because the completely anophthalmic specimens clearly belong to a separate species not described elsewhere in
<taxonomicName id="4C10E38FFFDDF16EFF44F166FEC2D75E" box="[183,333,577,603]" class="Arachnida" family="Liocranidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="family">Liocranidae</taxonomicName>
, and they present enough distinctive characters to allow attributing newly collected adult specimens to the same species.
</paragraph>
<paragraph id="8BAF980CFFDDF16EFF36F1A9FB44D12E" blockId="12.[151,1437,152,1106]" pageId="12" pageNumber="49">
Apart from the absence of eyes, which is often considered a typical energy-saving adaptation of cave animals (
<bibRefCitation id="EF81E5FDFFDDF16EFEF6F192FE0AD7CA" author="Barr" box="[261,389,693,719]" pageId="12" pageNumber="53" refString="Barr, T. (1968) Cave ecology and the evolution of troglobites. Evolutionary Biology 2, 35 - 102. Bernard, F. (1968) Les fourmis (Hymenoptera Formicidae) d' Europe occidentale et septentrionale. Masson et Cie., Paris, 411 pp." type="book chapter" year="1968">Barr 1968</bibRefCitation>
;
<bibRefCitation id="EF81E5FDFFDDF16EFE61F192FDC5D7CA" author="Diamond" box="[402,586,693,719]" pageId="12" pageNumber="54" refString="Diamond, J. (1996) Competition for brain space. Nature, London 382, 756 - 757." type="journal article" year="1996">Diamond 1996</bibRefCitation>
;
<bibRefCitation id="EF81E5FDFFDDF16EFDA5F192FCAFD7CA" author="Bosselaers" box="[598,800,693,719]" pageId="12" pageNumber="54" refString="Bosselaers, J. (1998) Nesticus henderickxi (Araneae, Nesticidae), a new blind troglobitic spider from Crete. Bulletin of the British arachnological Society 11, 9 - 14." type="journal article" year="1998">Bosselaers 1998</bibRefCitation>
), the general morphology of
<taxonomicName id="4C10E38FFFDDF16EFB72F192FAA1D7CB" box="[1153,1326,693,718]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFDDF16EFB72F192FAA1D7CB" box="[1153,1326,693,718]" italics="true" pageId="12" pageNumber="49">C. liocraninus</emphasis>
</taxonomicName>
does not justify a place in a separate genus and advocates the inclusion of the species in the genus
<taxonomicName id="4C10E38FFFDDF16EFB55F1FBFAA8D7F0" box="[1190,1319,732,757]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFDDF16EFB55F1FBFAA8D7F0" box="[1190,1319,732,757]" italics="true" pageId="12" pageNumber="49">Cybaeodes</emphasis>
</taxonomicName>
(compare
<figureCitation id="132B8489FFDDF16EFF64F025FE8BD619" box="[151,260,770,796]" captionStart="FIGURE 6 A" captionStartId="11.[151,255,1658,1682]" captionTargetBox="[193,1359,232,1595]" captionTargetId="figure@11.[185,1402,197,1633]" captionTargetPageId="11" captionText="FIGURE 6 A. Cybaeodes liocraninus, holotype. B. Cybaeodes madidus, holotype. A, habitus, lat; B, habitus, lat. Scale bar: 1." httpUri="https://zenodo.org/record/188592/files/figure.png" pageId="12" pageNumber="49">Figs. 6 A</figureCitation>
and B,
<figureCitation id="132B8489FFDDF16EFE93F025FE3FD619" box="[352,432,770,796]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="12" pageNumber="49">Figs. 3</figureCitation>
C–G with
<figureCitation id="132B8489FFDDF16EFDC0F025FD4BD619" box="[563,708,770,796]" captionStart="FIGURE 4 A – G" captionStartId="8.[151,255,1826,1850]" captionTargetBox="[155,1429,217,1755]" captionTargetId="figure@8.[151,1436,194,1802]" captionTargetPageId="8" captionText="FIGURE 4 A – G. Cybaeodes testaceus, holotype. A, carapace, do; B, carapace, lat; C, sternum and mouth parts, ve; D, carapace, fr; E, epigyne; F, leg I, tarsal tip; G, leg III, tarsal tip. Scale bars: A – D: 0.5; E: 0.25; F – G: 0.1." httpUri="https://zenodo.org/record/188590/files/figure.png" pageId="12" pageNumber="49">Figs. 4 A–G</figureCitation>
and
<figureCitation id="132B8489FFDDF16EFD0DF025FCC2D619" box="[766,845,770,796]" captionStart="FIGURE 5 A – G" captionStartId="10.[151,255,1665,1689]" captionTargetBox="[222,1344,231,1608]" captionTargetId="figure@10.[152,1435,197,1640]" captionTargetPageId="10" captionText="FIGURE 5 A – G. Cybaeodes madidus, holotype. A, carapace, do; B, carapace, lat; C, sternum and mouth parts, ve; D, carapace, fr; E, epigyne; F, leg II, tarsal tip; G, leg IV, tarsal tip. Scale bars: A – E: 0.5; F – G: 0.1." httpUri="https://zenodo.org/record/188591/files/figure.png" pageId="12" pageNumber="49">5 A–G</figureCitation>
, and
<figureCitation id="132B8489FFDDF16EFC78F025FC66D619" box="[907,1001,770,796]" captionStart="FIGURE 8 A – H" captionStartId="15.[151,259,1348,1372]" captionTargetBox="[151,1435,313,1323]" captionTargetId="figure@15.[151,1436,313,1324]" captionTargetPageId="15" captionText="FIGURE 8 A – H. Dorsal views of specimens. A, Apostenus humilis, holotype; B, Apostenus ochraceus, Lesbos, Kratigos; C, Agraecina rutilia, holotype; D, Agraecina lineata, female, MNHN 5653; E, Cybaeodes liocraninus, holotype; F, Cybaeodes testaceus, holotype; G, Cybaeodes madidus, holotype; H, Arabelia pheidoleicomes sp. n., Lesbos, Agiasos, Megali Limni. Scale bars: 0.5." httpUri="https://zenodo.org/record/188593/files/figure.png" pageId="12" pageNumber="49">Fig. 8 E</figureCitation>
with
<figureCitation id="132B8489FFDDF16EFBDEF025FB34D619" box="[1069,1211,770,796]" captionStart="FIGURE 8 A – H" captionStartId="15.[151,259,1348,1372]" captionTargetBox="[151,1435,313,1323]" captionTargetId="figure@15.[151,1436,313,1324]" captionTargetPageId="15" captionText="FIGURE 8 A – H. Dorsal views of specimens. A, Apostenus humilis, holotype; B, Apostenus ochraceus, Lesbos, Kratigos; C, Agraecina rutilia, holotype; D, Agraecina lineata, female, MNHN 5653; E, Cybaeodes liocraninus, holotype; F, Cybaeodes testaceus, holotype; G, Cybaeodes madidus, holotype; H, Arabelia pheidoleicomes sp. n., Lesbos, Agiasos, Megali Limni. Scale bars: 0.5." httpUri="https://zenodo.org/record/188593/files/figure.png" pageId="12" pageNumber="49">Figs. 8 F, G</figureCitation>
). The leg spination of
<taxonomicName id="4C10E38FFFDDF16EFF44F00EFED1D647" box="[183,350,809,834]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFDDF16EFF44F00EFED1D647" box="[183,350,809,834]" italics="true" pageId="12" pageNumber="49">C. liocraninus</emphasis>
</taxonomicName>
(compare
<figureCitation id="132B8489FFDDF16EFE2FF00EFDB1D646" box="[476,574,809,835]" captionStart="FIGURE 2 A – K" captionStartId="4.[151,257,1819,1843]" captionTargetBox="[168,1425,219,1763]" captionTargetId="figure@4.[151,1436,194,1795]" captionTargetPageId="4" captionText="FIGURE 2 A – K. Leg spination schematics. A, legend; B, Apostenus humilis, male holotype; C, Apostenus humilis, female, MNHN 25; D, Apostenus ochraceus, female; E, Agraecina rutilia, female holotype; F, Agraecina lineata, female, MNHN 5653; G, Cybaeodes liocraninus, holotype; H, Cybaeodes testaceus, female holotype, MNHN 3107; I, Cybaeodes madidus, female holotype, MNHN 1677; J, Cybaeodes marinae, female, Italy, Castelporziano, AMNH; K, Arabelia pheidoleicomes, female holotype." httpUri="https://zenodo.org/record/188588/files/figure.png" pageId="12" pageNumber="49">Fig. 2 G</figureCitation>
with
<figureCitation id="132B8489FFDDF16EFD72F00EFC86D646" box="[641,777,809,835]" captionStart="FIGURE 2 A – K" captionStartId="4.[151,257,1819,1843]" captionTargetBox="[168,1425,219,1763]" captionTargetId="figure@4.[151,1436,194,1795]" captionTargetPageId="4" captionText="FIGURE 2 A – K. Leg spination schematics. A, legend; B, Apostenus humilis, male holotype; C, Apostenus humilis, female, MNHN 25; D, Apostenus ochraceus, female; E, Agraecina rutilia, female holotype; F, Agraecina lineata, female, MNHN 5653; G, Cybaeodes liocraninus, holotype; H, Cybaeodes testaceus, female holotype, MNHN 3107; I, Cybaeodes madidus, female holotype, MNHN 1677; J, Cybaeodes marinae, female, Italy, Castelporziano, AMNH; K, Arabelia pheidoleicomes, female holotype." httpUri="https://zenodo.org/record/188588/files/figure.png" pageId="12" pageNumber="49">Figs. 2 H–J</figureCitation>
) as well as the presence of five pairs of tenent hairs and 5–7 toothed claws on the tarsal tips (compare
<figureCitation id="132B8489FFDDF16EFD3FF077FC9BD66F" box="[716,788,848,874]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="12" pageNumber="49">Fig. 3</figureCitation>
G with
<figureCitation id="132B8489FFDDF16EFC8AF077FB85D66F" box="[889,1034,848,874]" captionStart="FIGURE 4 A – G" captionStartId="8.[151,255,1826,1850]" captionTargetBox="[155,1429,217,1755]" captionTargetId="figure@8.[151,1436,194,1802]" captionTargetPageId="8" captionText="FIGURE 4 A – G. Cybaeodes testaceus, holotype. A, carapace, do; B, carapace, lat; C, sternum and mouth parts, ve; D, carapace, fr; E, epigyne; F, leg I, tarsal tip; G, leg III, tarsal tip. Scale bars: A – D: 0.5; E: 0.25; F – G: 0.1." httpUri="https://zenodo.org/record/188590/files/figure.png" pageId="12" pageNumber="49">Figs. 4 F, G</figureCitation>
and
<figureCitation id="132B8489FFDDF16EFBB5F077FB1CD66F" box="[1094,1171,848,874]" captionStart="FIGURE 5 A – G" captionStartId="10.[151,255,1665,1689]" captionTargetBox="[222,1344,231,1608]" captionTargetId="figure@10.[152,1435,197,1640]" captionTargetPageId="10" captionText="FIGURE 5 A – G. Cybaeodes madidus, holotype. A, carapace, do; B, carapace, lat; C, sternum and mouth parts, ve; D, carapace, fr; E, epigyne; F, leg II, tarsal tip; G, leg IV, tarsal tip. Scale bars: A – E: 0.5; F – G: 0.1." httpUri="https://zenodo.org/record/188591/files/figure.png" pageId="12" pageNumber="49">5 F, G</figureCitation>
) further confirm this.
<bibRefCitation id="EF81E5FDFFDDF16EFF64F051FE7BD695" box="[151,500,886,912]" pageId="12" pageNumber="54" refString="Platnick, N. &amp; Di Franco, F. (1992) On the relationships of the spider genus Cybaeodes (Araneae, Dionycha). American Museum Novitates 3053, 1 - 9." type="journal article">
Platnick &amp;
<collectingCountry id="F307D89CFFDDF16EFED1F051FECFD695" box="[290,320,886,912]" name="Iceland" pageId="12" pageNumber="49">Di</collectingCountry>
Franco (1992)
</bibRefCitation>
already stated &quot;...we would not be surprised to find that blind, cavernicolous members of
<taxonomicName id="4C10E38FFFDDF16EFED9F0BAFE25D6B3" box="[298,426,925,950]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFDDF16EFED9F0BAFE25D6B3" box="[298,426,925,950]" italics="true" pageId="12" pageNumber="49">Cybaeodes</emphasis>
</taxonomicName>
occur in Mediterranean caves&quot; and
<bibRefCitation id="EF81E5FDFFDDF16EFCA2F0BAFBBFD6B2" author="Ribera" box="[849,1072,925,951]" pageId="12" pageNumber="54" refString="Ribera, C. (2004) Dysdera valentina (Araneae, Dysderidae), una nueva especie de la provincia de Valencia, con algunas adiciones a la fauna cavernicola iberica. Revista Iberica de Aracnologia 9, 211 - 215." type="journal article" year="2004">Ribera (2004: 215)</bibRefCitation>
reports that material identified as
<taxonomicName id="4C10E38FFFDDF16EFF48F0E3FD83D6D8" box="[187,524,964,989]" class="Arachnida" family="Liocranidae" genus="Brachyanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFDDF16EFF48F0E3FD83D6D8" box="[187,524,964,989]" italics="true" pageId="12" pageNumber="49">Brachyanillus liocraninus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF81E5FDFFDDF16EFDCCF0E3FC90D6DB" author="Fage" box="[575,799,964,990]" pageId="12" pageNumber="54" refString="Fage, L. (1931) Araneae, 5 me serie, precedee d' un essai sur l'evolution souterraine et son determinisme. In Biospeologica, LV. Archives de zoologie experimentale et generale 71, 91 - 291." type="journal article" year="1931">Fage (1931: 209)</bibRefCitation>
actually belongs to
<taxonomicName id="4C10E38FFFDDF16EFBDBF0E3FB3BD6D8" box="[1064,1204,964,989]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFDDF16EFBDBF0E3FB3BD6D8" box="[1064,1204,964,989]" italics="true" pageId="12" pageNumber="49">Cybaeodes</emphasis>
</taxonomicName>
. Ribera (personal communication) captured several species of blind
<taxonomicName id="4C10E38FFFDDF16EFD03F0CDFCFDD106" box="[752,882,1002,1027]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFDDF16EFD03F0CDFCFDD106" box="[752,882,1002,1027]" italics="true" pageId="12" pageNumber="49">Cybaeodes</emphasis>
</taxonomicName>
, with a somatic morphology indistinguishable from Simon's
<taxonomicName id="4C10E38FFFDDF16EFECFF736FE6FD12F" box="[316,480,1041,1066]" class="Arachnida" family="Liocranidae" genus="Brachyanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFDDF16EFECFF736FE6FD12F" box="[316,480,1041,1066]" italics="true" pageId="12" pageNumber="49">Brachyanillus</emphasis>
</taxonomicName>
, in Spanish caves. Description of these new species is ongoing.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFDDF16EFF36F710FD55D157" box="[197,730,1079,1106]" pageId="12" pageNumber="49" type="distribution">
<paragraph id="8BAF980CFFDDF16EFF36F710FD55D157" blockId="12.[151,1437,152,1106]" box="[197,730,1079,1106]" pageId="12" pageNumber="49">
<emphasis id="B964441EFFDDF16EFF36F710FEEAD154" bold="true" box="[197,357,1079,1105]" pageId="12" pageNumber="49">Distribution.</emphasis>
<collectingCountry id="F307D89CFFDDF16EFE9FF71FFE48D157" box="[364,455,1080,1106]" name="Algeria" pageId="12" pageNumber="49">Algeria</collectingCountry>
,
<collectingCountry id="F307D89CFFDDF16EFE20F71FFD99D157" box="[467,534,1080,1106]" name="Spain" pageId="12" pageNumber="49">Spain</collectingCountry>
(
<bibRefCitation id="EF81E5FDFFDDF16EFDD6F71FFD41D157" author="Platnick" box="[549,718,1080,1106]" pageId="12" pageNumber="54" refString="Platnick, N. (2009) The World Spider Catalog, version 9.5. The American Museum of Natural History. Available from: http: // research. amnh. org / entomology / spiders / catalog /. html (18 May 2009)" type="journal article" year="2009">Platnick 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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