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<document id="E0348F7B07EA46E3EA5DC2843507C709" ID-CLB-Dataset="138993" ID-DOI="10.11646/zootaxa.5182.3.4" ID-GBIF-Dataset="93b0e11c-0298-431a-af60-3a9ec2aadc1c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7053043" ID-ZooBank="C6D187FF-BCFD-4249-9322-05E99AB898AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1662450029242" checkinUser="plazi" docAuthor="Kóbor, Péter &amp; Kondorosy, Előd" docDate="2022" docId="DC1787D591406E6AFF20FA7FFE653105" docLanguage="en" docName="zootaxa.5182.3.4.pdf" docOrigin="Zootaxa 5182 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Germalus pulchellus Kóbor &amp; Kondorosy 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="270" masterDocId="202EFFAD91426E6FFFB7FFE4FFB73754" masterDocTitle="The big-eyed bug genus Germalus Stål, 1862 in New Guinea (Heteroptera: Lygaeoidea: Geocoridae)" masterLastPageNumber="278" masterPageNumber="265" pageNumber="267" updateTime="1699377966083" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C306AC34F3495F5754F8F901932FCD28">The big-eyed bug genus Germalus Stål, 1862 in New Guinea (Heteroptera: Lygaeoidea: Geocoridae)</mods:title>
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<mods:affiliation id="A81D27903CF2E5D29CC9584EE9FD8A16">Hungarian University of Agriculture and Life Sciences, Georgikon Campus, Department of Conservation Biology, 16 Deák Ferenc Street, H- 8360 Keszthely, Hungary</mods:affiliation>
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<heading id="0F4981AF91406E6DFF20FA7FFE4F32E1" bold="true" box="[151,504,1435,1462]" fontSize="11" level="1" pageId="2" pageNumber="267" reason="1">
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<taxonomicName id="93BE4D4091406E6DFF20FA7FFE2432E2" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[151,403,1435,1462]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="267" phylum="Arthropoda" rank="species" species="pulchellus" status="sp. nov.">
<emphasis id="66CAEAD191406E6DFF20FA7FFE2432E2" bold="true" box="[151,403,1435,1462]" italics="true" pageId="2" pageNumber="267">Germalus pulchellus</emphasis>
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<taxonomicNameLabel id="7DF957AA91406E6DFE2DFA7FFE4F32E1" box="[410,504,1435,1461]" pageId="2" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="540136C391406E6DFF20FA5BFEE4328D" blockId="2.[151,504,1435,1497]" box="[151,339,1471,1497]" pageId="2" pageNumber="267">
(
<figureCitation id="CC852A4691406E6DFF28FA5BFEB6328D" box="[159,257,1471,1497]" captionStart="FIGURE 1" captionStartId="3.[151,250,1877,1901]" captionTargetBox="[164,1435,345,1854]" captionTargetId="figure-93@3.[151,1435,345,1854]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of New Guinean Germalus species. A. G. fuscovittatus Malipatil, 2013 (BMNH), B. G. pulchellus sp. nov. (Holotype, NHMW), C. G. kontschani sp. nov. (Holotype, ZSM), D G. cucullatus sp. nov. (Holotype, RBINS), E. G. pappi sp. nov. (holotype, ZSM), F. G. sellatus sp. nov. (Holotype, MFNB). (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7053047" httpUri="https://zenodo.org/record/7053047/files/figure.png" pageId="2" pageNumber="267">Figs. 1B</figureCitation>
,
<figureCitation id="CC852A4691406E6DFEBAFA5BFE87328D" box="[269,304,1471,1497]" captionStart="FIGURE 3" captionStartId="9.[151,250,1721,1745]" captionTargetBox="[151,1436,181,1697]" captionTargetId="figure-17@9.[151,1436,181,1697]" captionTargetPageId="9" captionText="FIGURE 3. Peritreme of metathoracic scent efferent apparatus (indicated by arrows) of New Guinean Germalus species. A. G. fuscovittatus Malipatil, 2013, B. G. pulchellus sp. nov., C. G. kontschani sp. nov., D. G. cucullatus sp. nov., E. G. pappi sp. nov., F. G. sellatus sp. nov. (images not to scale)." figureDoi="http://doi.org/10.5281/zenodo.7053053" httpUri="https://zenodo.org/record/7053053/files/figure.png" pageId="2" pageNumber="267">3B</figureCitation>
,
<figureCitation id="CC852A4691406E6DFE8BFA5BFEFC328D" box="[316,331,1471,1497]" captionStart="FIGURE 4" captionStartId="10.[151,250,1507,1531]" captionTargetBox="[151,1428,605,1484]" captionTargetId="figure-211@10.[151,1436,597,1484]" captionTargetPageId="10" captionText="FIGURE 4. Distribution map of New Guinean Germalus species." figureDoi="http://doi.org/10.5281/zenodo.7053055" httpUri="https://zenodo.org/record/7053055/files/figure.png" pageId="2" pageNumber="267">4</figureCitation>
)
</paragraph>
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<materialsCitation id="E4D63C9E91406E6DFF20F9E3FF543111" ID-GBIF-Occurrence="3898573301" collectionCode="NHMW" collectorName="A. Riedel" elevation="2000" location="IRIAN JAYA" pageId="2" pageNumber="267" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="8B05886191406E6DFF20F9E3FEB33175" box="[151,260,1543,1569]" pageId="2" pageNumber="267" type="holotype">Holotype</typeStatus>
:
<specimenCount id="42B8FD4A91406E6DFEB8F9ECFE933176" box="[271,292,1544,1570]" pageId="2" pageNumber="267" type="male"></specimenCount>
(
<collectionCode id="32AFAE0691406E6DFE83F9E3FE203175" box="[308,407,1543,1569]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="267" type="Museum">NHMW</collectionCode>
): “Diuremna /
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/ 911. IX. //
<collectingRegion id="967AF82191406E6DFCC3F9E3FBBD3175" box="[884,1034,1543,1569]" country="Indonesia" name="Papua Barat" pageId="2" pageNumber="267">IRIAN JAYA</collectingRegion>
/ Jayawijaya-Prov. / leg.
<collectorName id="F94B531591406E6DFA9EF9E3FA2F3175" box="[1321,1432,1543,1569]" pageId="2" pageNumber="267">A. Riedel</collectorName>
, 1992.”
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<typeStatus id="8B05886191406E6DFF70F9B4FE9A313E" box="[199,301,1616,1642]" pageId="2" pageNumber="267" type="paratype">Paratype</typeStatus>
:
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(
<collectionCode id="32AFAE0691406E6DFEEAF9ABFE77313D" box="[349,448,1615,1641]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="267" type="Museum">NHMW</collectionCode>
): “Diuremna /
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/ 911. IX. //
<collectingRegion id="967AF82191406E6DFC2FF9ABFB9A313D" box="[920,1069,1615,1641]" country="Indonesia" name="Papua Barat" pageId="2" pageNumber="267">IRIAN JAYA</collectingRegion>
/ Jayawijaya-Prov. / leg.
<collectorName id="F94B531591406E6DFAFEF9ABFF0B31D9" pageId="2" pageNumber="267">A. Riedel</collectorName>
, 1992.”
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</paragraph>
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<emphasis id="66CAEAD191406E6DFF70F973FEE631E5" bold="true" box="[199,337,1687,1713]" pageId="2" pageNumber="267">Description</emphasis>
.
</paragraph>
<paragraph id="540136C391406E6CFF70F95FFB1B3615" blockId="2.[151,1437,1543,2038]" lastBlockId="3.[151,1437,151,321]" lastPageId="3" lastPageNumber="268" pageId="2" pageNumber="267">
<emphasis id="66CAEAD191406E6DFF70F95FFEE23181" box="[199,341,1723,1749]" italics="true" pageId="2" pageNumber="267">Colouration.</emphasis>
Head generally ochraceous, vertex and clypeus with blackish decoration of variable extent (
<figureCitation id="CC852A4691406E6DFA8EF95FFA273181" box="[1337,1424,1723,1749]" captionStart="FIGURE 2" captionStartId="4.[151,250,1863,1887]" captionTargetBox="[151,1437,181,1840]" captionTargetId="figure-36@4.[151,1437,181,1840]" captionTargetPageId="4" captionText="FIGURE 2. Detail pictures of New Guinean Germalus species: A. variability of vertex pattern in G. pulchellus (holotype and paratype, NHMW); B. labial trough (indicated with arrow) of G. fuscovittatus, C. sutures of abdominal tergites 4/55/6 (indicated with arrow) of G. fuscovittatus, D. venation of metathoracic wing in G. fuscovittatus (abbreviations: h—hamus, iv—intervannals, M—media, R—radius) (images not to scale)." figureDoi="http://doi.org/10.5281/zenodo.7053049" httpUri="https://zenodo.org/record/7053049/files/figure.png" pageId="2" pageNumber="267">Fig. 2A</figureCitation>
). Eyestalk with blackish irregular spots posteriad reaching nearly the midline of eyestalk dorsally. Ocelli hyaline. Antennomere I ochraceous with apical half strongly infuscate dorsally; antennomeres IIIII narrowly infuscate at base and apex; antennomere IV mostly dark fuscous with narrow ochraceous annulation basally. Mandibular plates with three irregular brownish spots between apex and ocular sulcus. Labiomeres ochraceous, extreme apex of labiomere IV black. Thorax. Pronotum ochraceous with blackish punctuation and black longitudinal band medially. Humeral angles with a blackish ovoid spot and a tooth-shaped irregular brownish spots on posterior margin. Scutellum blackish with small ovoid ochraceous spots at basal and apical edges of median trifurcate carina. Clavus of hemelytron blackish with an irregular, elongate ochraceous spots at base of claval furrow and corial margin of clavus. Corium mostly ochraceous, punctuation along veins blackish; apical third of Cu and adjoining part of margin of corium with irregular infuscate area. Exocorium narrowly marginated with dark brown. Thoracic pleurites and sternites ochraceous with blackish punctuation; meso- and metathorax with an extended irregular blackish spot interrupted by the peritreme. Femora and tibiae with small brownish maculae. Abdominal tergites blackish; abdominal sternites ochraceous with dorsal margins brownish posteriad; sternites IV and VI with irregular brownish spots of variable extent laterad.
</paragraph>
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<paragraph id="540136C391416E6CFF20F8B1FBE830E1" blockId="3.[151,1436,1877,1973]" pageId="3" pageNumber="268">
<emphasis id="66CAEAD191416E6CFF20F8B1FEA33039" bold="true" box="[151,276,1877,1901]" pageId="3" pageNumber="268">FIGURE 1.</emphasis>
Dorsal habitus of New Guinean
<taxonomicName id="93BE4D4091416E6CFDD3F8B1FC8F3039" box="[612,824,1877,1901]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="66CAEAD191416E6CFDD3F8B1FD7F3039" box="[612,712,1877,1901]" italics="true" pageId="3" pageNumber="268">Germalus</emphasis>
species. A
</taxonomicName>
.
<taxonomicName id="93BE4D4091416E6CFCF4F8B1FB313039" authority="Malipatil, 2013" authorityName="Malipatil" authorityYear="2013" box="[835,1158,1877,1901]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="fuscovittatus">
<emphasis id="66CAEAD191416E6CFCF4F8B1FCE23038" box="[835,853,1877,1900]" italics="true" pageId="3" pageNumber="268">G</emphasis>
.
<emphasis id="66CAEAD191416E6CFCD5F8B1FC553039" box="[866,994,1877,1901]" italics="true" pageId="3" pageNumber="268">fuscovittatus</emphasis>
<bibRefCitation id="302F4B3291416E6CFC5EF8B1FB313039" author="Malipatil, M. B. &amp; Blackett, M. J." box="[1001,1158,1877,1901]" pageId="3" pageNumber="268" pagination="257 - 300" refId="ref8783" refString="Malipatil, M. B. &amp; Blackett, M. J. (2013) Review and revision of Australian Germalus Stal, with new genera and further new species of Australian Geocorinae (Hemiptera: Heteroptera: Geocoridae). Zootaxa, 3746 (2), 257 - 300. https: // doi. org / 10.11646 / zootaxa. 3746.2.3" type="journal article" year="2013">Malipatil, 2013</bibRefCitation>
</taxonomicName>
(BMNH), B.
<taxonomicName id="93BE4D4091416E6CFAA2F8B1FA2B3039" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[1301,1436,1877,1901]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="pulchellus" status="sp. nov.">
<emphasis id="66CAEAD191416E6CFAA2F8B1FA903038" box="[1301,1319,1877,1900]" italics="true" pageId="3" pageNumber="268">G</emphasis>
.
<emphasis id="66CAEAD191416E6CFA83F8B1FA2B3039" box="[1332,1436,1877,1901]" italics="true" pageId="3" pageNumber="268">pulchellus</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191416E6CFF20F89DFF5F30C5" bold="true" box="[151,232,1913,1937]" pageId="3" pageNumber="268">
<taxonomicNameLabel id="7DF957AA91416E6CFF20F89DFF5F30C5" box="[151,232,1913,1937]" pageId="3" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holotype, NHMW), C.
<taxonomicName id="93BE4D4091416E6CFE5DF89DFDC030C5" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[490,631,1913,1937]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="kontschani" status="sp. nov.">
<emphasis id="66CAEAD191416E6CFE5DF89DFE4B30C4" box="[490,508,1913,1936]" italics="true" pageId="3" pageNumber="268">G</emphasis>
.
<emphasis id="66CAEAD191416E6CFDBEF89DFDC030C5" box="[521,631,1913,1937]" italics="true" pageId="3" pageNumber="268">kontschani</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191416E6CFDC9F89DFD7830C5" bold="true" box="[638,719,1913,1937]" pageId="3" pageNumber="268">
<taxonomicNameLabel id="7DF957AA91416E6CFDC9F89DFD7830C5" box="[638,719,1913,1937]" pageId="3" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holotype, ZSM), D
<taxonomicName id="93BE4D4091416E6CFC1BF89DFB8330C5" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[940,1076,1913,1937]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="cucullatus" status="sp. nov.">
<emphasis id="66CAEAD191416E6CFC1BF89DFC0930C4" box="[940,958,1913,1936]" italics="true" pageId="3" pageNumber="268">G</emphasis>
.
<emphasis id="66CAEAD191416E6CFC7BF89DFB8330C5" box="[972,1076,1913,1937]" italics="true" pageId="3" pageNumber="268">cucullatus</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191416E6CFB8CF89DFB3B30C5" bold="true" box="[1083,1164,1913,1937]" pageId="3" pageNumber="268">
<taxonomicNameLabel id="7DF957AA91416E6CFB8CF89DFB3B30C5" box="[1083,1164,1913,1937]" pageId="3" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holotype, RBINS), E.
<taxonomicName id="93BE4D4091416E6CFA33F89DFF6630E1" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="pappi" status="sp. nov.">
<emphasis id="66CAEAD191416E6CFA33F89DFA2130C4" box="[1412,1430,1913,1936]" italics="true" pageId="3" pageNumber="268">G</emphasis>
.
<emphasis id="66CAEAD191416E6CFF20F87AFF6630E1" box="[151,209,1950,1973]" italics="true" pageId="3" pageNumber="268">pappi</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191416E6CFF60F879FE9F30E1" bold="true" box="[215,296,1949,1973]" pageId="3" pageNumber="268">
<taxonomicNameLabel id="7DF957AA91416E6CFF60F879FE9F30E1" box="[215,296,1949,1973]" pageId="3" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, ZSM), F.
<taxonomicName id="93BE4D4091416E6CFE49F879FDDD30E1" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[510,618,1949,1973]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="sellatus" status="sp. nov.">
<emphasis id="66CAEAD191416E6CFE49F879FDA730E0" box="[510,528,1949,1972]" italics="true" pageId="3" pageNumber="268">G</emphasis>
.
<emphasis id="66CAEAD191416E6CFDAAF879FDDD30E1" box="[541,618,1949,1973]" italics="true" pageId="3" pageNumber="268">sellatus</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191416E6CFDC6F879FD7630E1" bold="true" box="[625,705,1949,1973]" pageId="3" pageNumber="268">
<taxonomicNameLabel id="7DF957AA91416E6CFDC6F879FD7630E1" box="[625,705,1949,1973]" pageId="3" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holotype, MFNB). (scale bar = 1 mm).
</paragraph>
</caption>
<caption id="00C1664B91466E6BFF20F8A3FD603098" ID-DOI="http://doi.org/10.5281/zenodo.7053049" ID-Zenodo-Dep="7053049" httpUri="https://zenodo.org/record/7053049/files/figure.png" pageId="4" pageNumber="269" startId="4.[151,250,1863,1887]" targetBox="[151,1437,181,1840]" targetPageId="4">
<paragraph id="540136C391466E6BFF20F8A3FD603098" blockId="4.[151,1437,1863,1996]" pageId="4" pageNumber="269">
<emphasis id="66CAEAD191466E6BFF20F8A3FEA3300B" bold="true" box="[151,276,1863,1887]" pageId="4" pageNumber="269">FIGURE 2.</emphasis>
Detail pictures of New Guinean
<taxonomicName id="93BE4D4091466E6BFDD0F8ACFCA83034" box="[615,799,1864,1888]" pageId="4" pageNumber="269">
<emphasis id="66CAEAD191466E6BFDD0F8ACFD7C3034" box="[615,715,1864,1888]" italics="true" pageId="4" pageNumber="269">Germalus</emphasis>
species
</taxonomicName>
: A. variability of vertex pattern in
<taxonomicName id="93BE4D4091466E6BFB36F8ACFABF3034" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[1153,1288,1864,1888]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191466E6BFB36F8ACFB24300B" box="[1153,1171,1864,1887]" italics="true" pageId="4" pageNumber="269">G</emphasis>
.
<emphasis id="66CAEAD191466E6BFB17F8ACFABF3034" box="[1184,1288,1864,1888]" italics="true" pageId="4" pageNumber="269">pulchellus</emphasis>
</taxonomicName>
(holotype and paratype, NHMW); B. labial trough (indicated with arrow) of
<taxonomicName id="93BE4D4091466E6BFCA2F888FC0230D0" authorityName="Malipatil" authorityYear="2013" box="[789,949,1900,1924]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="276" phylum="Arthropoda" rank="species" species="fuscovittatus">
G.
<emphasis id="66CAEAD191466E6BFC82F888FC0230D0" box="[821,949,1900,1924]" italics="true" pageId="4" pageNumber="269">fuscovittatus</emphasis>
</taxonomicName>
, C. sutures of abdominal tergites 4/55/6 (indicated with arrow) of
<taxonomicName id="93BE4D4091466E6BFED8F874FDB830FC" authorityName="Malipatil" authorityYear="2013" box="[367,527,1936,1960]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="276" phylum="Arthropoda" rank="species" species="fuscovittatus">
G.
<emphasis id="66CAEAD191466E6BFE38F874FDB830FC" box="[399,527,1936,1960]" italics="true" pageId="4" pageNumber="269">fuscovittatus</emphasis>
</taxonomicName>
, D. venation of metathoracic wing in
<taxonomicName id="93BE4D4091466E6BFC20F874FB8030FC" authorityName="Malipatil" authorityYear="2013" box="[919,1079,1936,1960]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="fuscovittatus">
<emphasis id="66CAEAD191466E6BFC20F874FC1E30F3" box="[919,937,1936,1959]" italics="true" pageId="4" pageNumber="269">G</emphasis>
.
<emphasis id="66CAEAD191466E6BFC00F874FB8030FC" box="[951,1079,1936,1960]" italics="true" pageId="4" pageNumber="269">fuscovittatus</emphasis>
</taxonomicName>
(abbreviations: h—hamus, iv—intervannals, M—media, R—radius) (images not to scale).
</paragraph>
</caption>
<paragraph id="540136C391476E6AFF70FF73FA2A34B9" blockId="5.[151,1437,151,1617]" pageId="5" pageNumber="270">
<emphasis id="66CAEAD191476E6AFF70FF73FE8137E5" box="[199,310,151,177]" italics="true" pageId="5" pageNumber="270">Structure.</emphasis>
Integument of head dull, impunctate. Eyes moderately stylate; eye stalks moderately projected; ocular sulcus complete, distinct. Ocelli situated near ocular sulcus at basal part of vertex; ocellus-eye distance to interocellar distance: 1: 1.39. Clypeus rounded exceeding the mandibular plates, margins of clypeus converging towards base, constricted medially. Antenniferous tubercles well-developed, rounded, but not visible in dorsal view. Antennomere I shortest, granuliform with two short, erect setae basally; antennomere II longest, cylindrical; antennomere III somewhat shorter than IV subequal in length; antennomere III cylindrical, antennomere IV fusiform. Ratio of antennomeres: 1: 2.41: 1.69: 2.00. Mandibular and maxillar plates clearly separated; ventral margin of bucculae forming a U-shaped labial trough. Labiomere I surpassing base of head; labiomeres IIII subequal; labiomere IV somewhat shorter, apex of labiomere IV reaching line of metacoxae. Ratio of labiomeres: 1: 1.0: 1.0: 0.75. Thorax. Pronotum trapeziform with lateral margin slightly carinate and constricted medially; pronotum ratio: 1: 1.55. Integument of pronotum deeply punctate except collar-like spot at anterior margin, pronotal callosities and humeral angles. Scutellum triangular, subequilateral; scutellum length to width: 1: 1.21. Scutellum strongly punctate except well-developed, elevated, median trifurcate carina. Hemelytron macropterous. Clavus well-developed, margins parallel; claval commissure complete with a line of punctures at full length. Margins densely punctate at almost full length of both scutellar and corial margins; punctuation at corial margin continues at basal part of posterior edge of corium. Length of claval commissure to median length of scutellum: 1: 1.74. Corium punctate along venation as follows: Cu punctate at apical half of length, M-R punctate at full length. Exocorium widening towards apex. Membrane well developed, surpassing apex of abdomen, without conspicuous venation. Thoracic pleurites and sternites densely punctate except collar of prosternite, supracoxal lobes, peritreme and posterior lobe of metapleurite. Peritreme auricular with an elongate, crescent-shaped dorsal supporting process (
<figureCitation id="CC852A4691476E6AFBB7FCA7FBEF3409" box="[1024,1112,835,861]" captionStart="FIGURE 3" captionStartId="9.[151,250,1721,1745]" captionTargetBox="[151,1436,181,1697]" captionTargetId="figure-17@9.[151,1436,181,1697]" captionTargetPageId="9" captionText="FIGURE 3. Peritreme of metathoracic scent efferent apparatus (indicated by arrows) of New Guinean Germalus species. A. G. fuscovittatus Malipatil, 2013, B. G. pulchellus sp. nov., C. G. kontschani sp. nov., D. G. cucullatus sp. nov., E. G. pappi sp. nov., F. G. sellatus sp. nov. (images not to scale)." figureDoi="http://doi.org/10.5281/zenodo.7053053" httpUri="https://zenodo.org/record/7053053/files/figure.png" pageId="5" pageNumber="270">Fig. 3B</figureCitation>
); peritremal surface distinct. Femora fusiform, fore femora with minute tooth-like processes apically on dorsal and ventral side. Fore tibiae clubshaped, middle and hind tibiae rather cylindrical. Tarsal claws evenly curved with well-defined blunt spur basally. Unguitractor plate with conspicuous, ovoid parempodia-like pads. Abdomen impunctate, integument covered with fine silvery pubescence, genital segments setose. Female ovipositor elongate, bisecting abdominal sternites VVII.
</paragraph>
<paragraph id="540136C391476E6AFF70FC13FA34330D" blockId="5.[151,1437,151,1617]" pageId="5" pageNumber="270">
<emphasis id="66CAEAD191476E6AFF70FC13FEC13345" bold="true" box="[199,374,1015,1041]" pageId="5" pageNumber="270">Measurements</emphasis>
(
<typeStatus id="8B05886191476E6AFE33FC13FE5A3345" box="[388,493,1015,1041]" pageId="5" pageNumber="270" type="holotype">holotype</typeStatus>
). Total body length: 5.81; length of head: 0.76, width of head: 1.68; distance of ocelli: 0.43; labiomeres: 0.670.670.670.50; length of antennomeres: 0.390.940.660.78; length of pronotum: 1.20, width of pronotum: 1.86; length of scutellum: 0.82; width of scutellum: 0.99; length of claval commissure: 0.47.
</paragraph>
</subSubSection>
<subSubSection id="1CA4654891476E6AFF70FB86FB343295" pageId="5" pageNumber="270" type="diagnosis">
<paragraph id="540136C391476E6AFF70FB86FB343295" blockId="5.[151,1437,151,1617]" pageId="5" pageNumber="270">
<emphasis id="66CAEAD191476E6AFF70FB86FE8D3328" bold="true" box="[199,314,1122,1148]" pageId="5" pageNumber="270">Diagnosis</emphasis>
.
<taxonomicName id="93BE4D4091476E6AFEFFFB87FE6A3329" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[328,477,1123,1149]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus" status="sp. nov.">
<emphasis id="66CAEAD191476E6AFEFFFB87FEEB3329" box="[328,348,1123,1149]" italics="true" pageId="5" pageNumber="270">G</emphasis>
.
<emphasis id="66CAEAD191476E6AFEDDFB87FE6A3329" box="[362,477,1123,1149]" italics="true" pageId="5" pageNumber="270">pulchellus</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191476E6AFE53FB87FD893329" bold="true" box="[484,574,1123,1149]" pageId="5" pageNumber="270">
<taxonomicNameLabel id="7DF957AA91476E6AFE53FB87FD893329" box="[484,574,1123,1149]" pageId="5" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="93BE4D4091476E6AFD0CFB87FCAA3328" authority="Kóbor &amp; Kondorosy, 2022" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[699,797,1123,1149]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="pappi" status="sp. nov.">
G.
<emphasis id="66CAEAD191476E6AFD6AFB87FCAA3328" box="[733,797,1123,1148]" italics="true" pageId="5" pageNumber="270">pappi</emphasis>
</taxonomicName>
<emphasis id="66CAEAD191476E6AFC93FB87FCCA3329" bold="true" box="[804,893,1123,1149]" pageId="5" pageNumber="270">
<taxonomicNameLabel id="7DF957AA91476E6AFC93FB87FCCA3329" box="[804,893,1123,1149]" pageId="5" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from New
<collectingCountry id="2CA9765391476E6AFC4AFB87FBE63329" box="[1021,1105,1123,1149]" name="Guinea-Bissau" pageId="5" pageNumber="270">Guinea</collectingCountry>
,
<taxonomicName id="93BE4D4091476E6AFBEBFB87FEBB33F5" authority="Kobor &amp; Kondorosy, 2022" authorityName="Kobor &amp; Kondorosy" authorityYear="2022" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="66CAEAD191476E6AFBEBFB87FBC73329" box="[1116,1136,1123,1149]" italics="true" pageId="5" pageNumber="270">G</emphasis>
.
<emphasis id="66CAEAD191476E6AFBC9FB87FB623329" box="[1150,1237,1123,1149]" italics="true" pageId="5" pageNumber="270">scabrus</emphasis>
<bibRefCitation id="302F4B3291476E6AFB6BFB87FEBB33F5" author="Kobor, P. &amp; Kondorosy, E." pageId="5" pageNumber="270" pagination="2199 - 2212" refId="ref8724" refString="Kobor, P. &amp; Kondorosy, E. (2022) Big-eyed bugs of the Solomon Islands: taxonomy, ecological and evolutionary significance (Heteroptera: Lygaeoidea: Geocoridae). Journal of Natural History, 55 (35 - 36), 2199 - 2212. https: // doi. org / 10.1080 / 00222933.2021.2003460" type="journal article" year="2022">Kóbor &amp; Kondorosy, 2022</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="2CA9765391476E6AFECBFB63FD8033F5" box="[380,567,1159,1185]" name="Solomon Islands" pageId="5" pageNumber="270">Solomon Islands</collectingCountry>
and the Australian species,
<taxonomicName id="93BE4D4091476E6AFCC4FB63FB2833F5" authority="Distant, 1918" authorityName="Distant" authorityYear="1918" box="[883,1183,1159,1185]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="coloratus">
<emphasis id="66CAEAD191476E6AFCC4FB63FC3033F5" box="[883,903,1159,1185]" italics="true" pageId="5" pageNumber="270">G</emphasis>
.
<emphasis id="66CAEAD191476E6AFC21FB63FBB733F5" box="[918,1024,1159,1185]" italics="true" pageId="5" pageNumber="270">coloratus</emphasis>
Distant,
<quantity id="93469B2691476E6AFBD0FB63FB2833F5" box="[1127,1183,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="4.87172" pageId="5" pageNumber="270" unit="in" value="1918.0">1918</quantity>
</taxonomicName>
in general features. In
<taxonomicName id="93BE4D4091476E6AFF20FB4FFF4E3390" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[151,249,1195,1221]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pappi">
<emphasis id="66CAEAD191476E6AFF20FB4FFF4E3390" box="[151,249,1195,1221]" italics="true" pageId="5" pageNumber="270">G. pappi</emphasis>
</taxonomicName>
pronotum with broad blackish transverse band across pronotal callosities (in
<taxonomicName id="93BE4D4091476E6AFBECFB4FFB473391" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[1115,1264,1195,1221]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191476E6AFBECFB4FFB473391" box="[1115,1264,1195,1221]" italics="true" pageId="5" pageNumber="270">G. pulchellus</emphasis>
</taxonomicName>
pronotum with narrow longitudinal band medially) and the supportive process of peritreme almost straight (in
<taxonomicName id="93BE4D4091476E6AFB6DFB2BFAC633BD" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[1242,1393,1231,1257]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191476E6AFB6DFB2BFAC633BD" box="[1242,1393,1231,1257]" italics="true" pageId="5" pageNumber="270">G. pulchellus</emphasis>
</taxonomicName>
the supportive process strongly curved, crescent-shaped). In
<taxonomicName id="93BE4D4091476E6AFCA2FB17FC383259" authorityName="Kobor &amp; Kondorosy" authorityYear="2022" box="[789,911,1267,1293]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="66CAEAD191476E6AFCA2FB17FC383259" box="[789,911,1267,1293]" italics="true" pageId="5" pageNumber="270">G. scabrus</emphasis>
</taxonomicName>
pronotum with both longitudinal and transversal bands (in
<taxonomicName id="93BE4D4091476E6AFE9BFAF3FE753265" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[300,450,1303,1329]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191476E6AFE9BFAF3FE753265" box="[300,450,1303,1329]" italics="true" pageId="5" pageNumber="270">G. pulchellus</emphasis>
</taxonomicName>
pronotum with only median longitudinal band) and no collar-like impunctate spot present at anterior margin of pronotum (in
<taxonomicName id="93BE4D4091476E6AFDF9FADFFD533201" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[590,740,1339,1365]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191476E6AFDF9FADFFD533201" box="[590,740,1339,1365]" italics="true" pageId="5" pageNumber="270">G. pulchellus</emphasis>
</taxonomicName>
pronotum with distinct collar-like impunctate spot at anterior margin). In
<taxonomicName id="93BE4D4091476E6AFEACFABBFE10322D" authorityName="Distant" authorityYear="1918" box="[283,423,1375,1401]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="coloratus">
<emphasis id="66CAEAD191476E6AFEACFABBFE10322D" box="[283,423,1375,1401]" italics="true" pageId="5" pageNumber="270">G. coloratus</emphasis>
</taxonomicName>
decoration of pronotum consists of spots at humeral angles (in
<taxonomicName id="93BE4D4091476E6AFBDBFABBFAB6322D" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[1132,1281,1375,1401]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191476E6AFBDBFABBFAB6322D" box="[1132,1281,1375,1401]" italics="true" pageId="5" pageNumber="270">G. pulchellus</emphasis>
</taxonomicName>
decoration of pronotum consists of median longitudinal band and spots at humeral angles) and median trifurcate carina of scutellum entirely ochraceous (in
<taxonomicName id="93BE4D4091476E6AFE66FA43FDD03295" authorityName="Kóbor &amp; Kondorosy" authorityYear="2022" box="[465,615,1447,1473]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="66CAEAD191476E6AFE66FA43FDD03295" box="[465,615,1447,1473]" italics="true" pageId="5" pageNumber="270">G. pulchellus</emphasis>
</taxonomicName>
only apices of trifurcate carina are ochraceous).
</paragraph>
</subSubSection>
<subSubSection id="1CA4654891476E6AFF70FA2FFDF0315D" pageId="5" pageNumber="270" type="etymology">
<paragraph id="540136C391476E6AFF70FA2FFDF0315D" blockId="5.[151,1437,151,1617]" pageId="5" pageNumber="270">
<emphasis id="66CAEAD191476E6AFF70FA2FFEFF32B1" bold="true" box="[199,328,1483,1509]" pageId="5" pageNumber="270">Etymology</emphasis>
. Species epithet Latin, meaning “beautiful, pretty” and refers to the decorative appearance of this insect, especially the pattern of vertex.
</paragraph>
</subSubSection>
<subSubSection id="1CA4654891476E6AFF70F9F6FE653105" pageId="5" pageNumber="270" type="distribution">
<paragraph id="540136C391476E6AFF70F9F6FE653105" blockId="5.[151,1437,151,1617]" pageId="5" pageNumber="270">
<emphasis id="66CAEAD191476E6AFF70F9F6FEEE3178" bold="true" box="[199,345,1554,1580]" pageId="5" pageNumber="270">Distribution</emphasis>
.
<typeStatus id="8B05886191476E6AFEDCF9F7FE643179" box="[363,467,1555,1581]" pageId="5" pageNumber="270" type="holotype">Holotype</typeStatus>
and
<typeStatus id="8B05886191476E6AFDA5F9F7FDC53179" box="[530,626,1555,1581]" pageId="5" pageNumber="270" type="paratype">paratype</typeStatus>
were collected near Irian Jaya, New
<collectingCountry id="2CA9765391476E6AFB9BF9F7FBCA3179" box="[1068,1149,1555,1581]" name="Guinea-Bissau" pageId="5" pageNumber="270">Guinea</collectingCountry>
(fig. 4), habitat: Central Range montane rain forests.
</paragraph>
</subSubSection>
</treatment>
</document>