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<mods:title id="616182E4DE23BD279D78C39BD4165DD0">New species of spongillafly genus Sisyra Burmeister, 1839 (Neuroptera: Sisyridae) from China and Laos</mods:title>
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<mods:namePart id="65701A204BB13E33FA971B7236C48DB1">Yang, Ying</mods:namePart>
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<treatment id="A75A87D45C29FFEC70CE2D9CFA4B8C0B" ID-DOI="http://doi.org/10.5281/zenodo.12173787" ID-Zenodo-Dep="12173787" LSID="urn:lsid:plazi:treatment:A75A87D45C29FFEC70CE2D9CFA4B8C0B" httpUri="http://treatment.plazi.org/id/A75A87D45C29FFEC70CE2D9CFA4B8C0B" lastPageNumber="139" pageId="10" pageNumber="139">
<subSubSection id="67E965495C29FFEC70CE2D9CFDDF88DB" box="[136,526,971,995]" pageId="10" pageNumber="139" type="nomenclature">
<paragraph id="2F4C36C25C29FFEC70CE2D9CFDDF88DB" blockId="10.[136,1452,971,1060]" box="[136,526,971,995]" pageId="10" pageNumber="139">
<heading id="740481AE5C29FFEC70CE2D9CFDDF88DB" box="[136,526,971,995]" fontSize="10" level="2" pageId="10" pageNumber="139" reason="2">
<taxonomicName id="E8F34D415C29FFEC70CE2D9CFE7888DB" ID-CoL="6YSLL" authority="Yang, 1986" authorityName="Yang" authorityYear="1986" box="[136,425,971,995]" class="Insecta" family="Sisyridae" genus="Sisyra" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="yunana">
<emphasis id="1D87EAD05C29FFEC70CE2D9CFE7888DB" bold="true" box="[136,425,971,995]" pageId="10" pageNumber="139">
<emphasis id="1D87EAD05C29FFEC70CE2D9CFEF788DA" bold="true" box="[136,294,971,995]" italics="true" pageId="10" pageNumber="139">Sisyra yunana</emphasis>
<bibRefCitation id="4B624B335C29FFEC716A2D9CFE7888DB" author="Yang" box="[300,425,971,995]" pageId="10" pageNumber="139" pagination="423 - 434" refId="ref6714" refString="Yang, C. K. 1986. Thirty new species and four new genera of Neuroptera from Yunnan, and the family Nemopteridae new to China. Acta Agriculturae Universitatis Pekinensis, 12: 153 - 166, 423 - 434." type="journal article" year="1986">Yang, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation id="B7C82A475C29FFEC71802D9CFDD488DB" box="[454,517,971,995]" captionStart="Figure 8" captionStartId="11.[155,218,2057,2079]" captionTargetBox="[203,1396,422,2006]" captionTargetId="figure-105@11.[189,1399,421,2009]" captionTargetPageId="11" captionText="Figure 8. Sisyra yunana Yang, 1986, male genitalia. AB. lateral view. CD. dorsal view; EF. caudal view. Scale bars=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12172264" httpUri="https://zenodo.org/record/12172264/files/figure.png" pageId="10" pageNumber="139">Fig. 8</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67E965495C29FFEC70CE2DB9FE7C8F1B" pageId="10" pageNumber="139" type="reference_group">
<paragraph id="2F4C36C25C29FFEC70CE2DB9FE7C8F1B" blockId="10.[136,1452,971,1060]" pageId="10" pageNumber="139">
<treatmentCitationGroup id="0FE311EC5C29FFEC70CE2DB9FE7C8F1B" pageId="10" pageNumber="139">
<treatmentCitation id="AE5210D35C29FFEC70CE2DB9FE678F3C" author="Yang" box="[136,438,1006,1028]" page="155" pageId="10" pageNumber="139" year="1986">
<taxonomicName id="E8F34D415C29FFEC70CE2DB9FE678F3C" ID-CoL="6YSLL" authority="Yang, 1986: 155" authorityName="Yang" authorityPageNumber="155" authorityYear="1986" box="[136,438,1006,1028]" class="Insecta" family="Sisyridae" genus="Sisyra" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="yunana">
<emphasis id="1D87EAD05C29FFEC70CE2DB9FEC38F3B" box="[136,274,1006,1027]" italics="true" pageId="10" pageNumber="139">Sisyra yunana</emphasis>
<bibRefCitation id="4B624B335C29FFEC71512DB9FE678F3C" author="Yang" box="[279,438,1006,1028]" pageId="10" pageNumber="139" pagination="423 - 434" refId="ref6714" refString="Yang, C. K. 1986. Thirty new species and four new genera of Neuroptera from Yunnan, and the family Nemopteridae new to China. Acta Agriculturae Universitatis Pekinensis, 12: 153 - 166, 423 - 434." type="journal article" year="1986">Yang, 1986: 155</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="AE5210D35C29FFEC72D52DB9FCB88F3B" author="Yang" box="[659,873,1006,1027]" page="103" pageId="10" pageNumber="139" year="1999">
<bibRefCitation id="4B624B335C29FFEC72D52DB9FCB88F3B" author="Yang" box="[659,873,1006,1027]" pageId="10" pageNumber="139" pagination="102 - 115" refId="ref6757" refString="Yang, C. K., Yang, X. K., Liu, Z. Q. 1999. Neuroptera. In: Chen, S. C. (ed.). Pictorial Handbook of Rare and Precious Insects in China. Chinese Edition. China Forestry Publishing House, Beijing. pp. 102 - 115." type="book chapter" year="1999">
Yang
<emphasis id="1D87EAD05C29FFEC728D2DB9FD278F3B" box="[715,758,1006,1027]" italics="true" pageId="10" pageNumber="139">et al</emphasis>
., 1999: 103
</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="AE5210D35C29FFEC744F2DB9FB0D8F3B" author="Yang &amp; Liu &amp; A revision of the spongillafly genus &amp; Sisyra Burmeister" box="[1033,1244,1006,1028]" page="29" pageId="10" pageNumber="139" year="2023">
<bibRefCitation id="4B624B335C29FFEC744F2DB9FB0D8F3B" author="Yang &amp; Liu &amp; A revision of the spongillafly genus &amp; Sisyra Burmeister" box="[1033,1244,1006,1028]" pageId="10" pageNumber="139" pagination="1 - 37" refId="ref6868" refString="Yang, Y., Liu, X. Y. 2023. A revision of the spongillafly genus Sisyra Burmeister, 1839 (Neuroptera: Sisyridae) from China. Aquatic Insects, 1 - 37. doi: 10.1080 / 01650424.2023.2251660" type="book chapter" year="2023">Yang &amp; Liu, 2023: 29</bibRefCitation>
</treatmentCitation>
(redescription).
<typeStatus id="F04888605C29FFEC753A2DB9FA7D8F3B" box="[1404,1452,1006,1027]" pageId="10" pageNumber="139">Type</typeStatus>
locality:
<collectingCountry id="57E476525C29FFEC714B2A59FE978F1C" box="[269,326,1038,1060]" name="China" pageId="10" pageNumber="139">China</collectingCountry>
(
<collectingRegion id="ED37F8205C29FFEC71152A59FE738F1B" box="[339,418,1038,1059]" country="China" name="Yunnan" pageId="10" pageNumber="139">Yunnan</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="67E965495C29FFEC70F82A14FA4B8C0B" pageId="10" pageNumber="139" type="diagnosis">
<paragraph id="2F4C36C25C29FFEC70F82A14FD308FFB" blockId="10.[136,1458,1091,1843]" pageId="10" pageNumber="139">Diagnosis. Head brown; antenna with basal two thirds brown (same color as head), distal one third of flagellum yellow (progressively lighter but darkened on most distal five flagellomeres). Forewing costal space weakly broadened basally. Male sternum 9 short, and subquadrate, strongly protruding anteroventrally. Gonocoxites 9 strongly swollen basally, with distal half slender, bearing a rounded tubercle at middle.</paragraph>
<paragraph id="2F4C36C25C29FFEC70FB2A9AFEC88D25" blockId="10.[136,1458,1091,1843]" pageId="10" pageNumber="139">Description of male genitalia. Sternum 9 well developed, strongly protruding anteroventrally, subquadrate in caudal view, caudal portion with many transverse wrinkles and protruding trichobotheria, each bearing extremely long bristles. Gonocoxites 9 long and directed posteroventrally; strongly swollen basally, with distal half strongly narrowed; in dorsal view swollen base with a tridactylous-like projection (each tip with a long bristle) and an additional thick bristle on inner margin; a rounded tubercle present on dorsal portion at middle, with many short setae; distal half curved medially in dorsal view, with apex modified into a narrow subtriangular lobe, densely covered by short setae; a long bristle present at tip. Gonocoxites 11 arcuate, ventral margin slightly concaved dorsally in lateral view, without any projection; gonostyli 11 paired, subtriangular, dorsal margin concaved ventrally and lateral margin concaved medially in caudal view. Ectoproct subtrapezoidal, covered by densely long setae; callus cerci with three to five trichobothria. Hypandrium internum indiscernible.</paragraph>
<paragraph id="2F4C36C25C29FFEC70FB287FFD298D90" blockId="10.[136,1458,1091,1843]" pageId="10" pageNumber="139">
Material examined.
<materialsCitation id="9F9B3C9F5C29FFEC71DA287FFD918D5B" collectingDate="1981-03-12" collectionCode="CAU" collectorName="Chikun Yang" country="China" county="Xishuangbanna" latitude="22.166666" location="Sancha River" longLatPrecision="1261" longitude="100.85" municipality="Jinghong" pageId="10" pageNumber="139" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="F04888605C29FFEC71DA287FFE2E8D78" box="[412,511,1576,1600]" pageId="10" pageNumber="139" type="holotype">Holotype</typeStatus>
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,
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:
<collectingRegion id="ED37F8205C29FFEC72C6287FFD058D07" box="[640,724,1576,1599]" country="China" name="Yunnan" pageId="10" pageNumber="139">Yunnan</collectingRegion>
,
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,
<collectingMunicipality id="CF28ACB85C29FFEC73DC287FFC2F8D78" box="[922,1022,1576,1600]" pageId="10" pageNumber="139">Jinghong</collectingMunicipality>
,
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,
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,
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,
<date id="5B4D10025C29FFEC75D7287FFEC38D5B" pageId="10" pageNumber="139" value="1981-03-12">
<collectingDate id="4B09E9EA5C29FFEC75D7287FFEC38D5B" pageId="10" pageNumber="139" value="1981-03-12">12 March 1981</collectingDate>
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,
<collectorName id="820653145C29FFEC7164281CFE638D5B" box="[290,434,1611,1635]" pageId="10" pageNumber="139">Chikun Yang</collectorName>
leg. (
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)
</materialsCitation>
.
<materialsCitation id="9F9B3C9F5C29FFEC720B281CFCE68DBD" collectingDate="1982-05-17" collectionCode="CAU" collectorName="Chikun Yang" country="China" latitude="22.333334" location="Longzhou" longLatPrecision="1260" longitude="106.85" municipality="Chongzuo" pageId="10" pageNumber="139" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Guangxi">
<typeStatus id="F04888605C29FFEC720B281CFCF78D5B" box="[589,806,1611,1635]" isOtherTypeStatus="true" pageId="10" pageNumber="139">Additional material</typeStatus>
.
<specimenCount id="39F5FD4B5C29FFEC7373281CFC878D5A" box="[821,854,1607,1635]" count="2" pageId="10" pageNumber="139" type="male">2♂</specimenCount>
<specimenCount id="39F5FD4B5C29FFEC7310281CFCA98D5A" box="[854,888,1607,1635]" count="2" pageId="10" pageNumber="139" type="female">2♀</specimenCount>
,
<collectingCountry id="57E476525C29FFEC73CD281CFC1E8D5B" box="[907,975,1611,1635]" name="China" pageId="10" pageNumber="139">China</collectingCountry>
:
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,
<collectingMunicipality id="CF28ACB85C29FFEC740A281CFB6D8D5B" box="[1100,1212,1611,1635]" pageId="10" pageNumber="139">Chongzuo</collectingMunicipality>
,
<location id="2A2C60195C29FFEC748B281CFAED8D5B" LSID="urn:lsid:plazi:treatment:A75A87D45C29FFEC70CE2D9CFA4B8C0B:2A2C60195C29FFEC748B281CFAED8D5B" box="[1229,1340,1611,1635]" country="China" latitude="22.333334" longLatPrecision="1260" longitude="106.85" municipality="Chongzuo" name="Longzhou" pageId="10" pageNumber="139" stateProvince="Guangxi">Longzhou</location>
,
<geoCoordinate id="4AC750055C29FFEC750A281CFA768D5A" box="[1356,1447,1609,1635]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="10" pageNumber="139" precision="925" value="22.333334">22°20N</geoCoordinate>
,
<geoCoordinate id="4AC750055C29FFEC70CE283AFF3F8DBD" box="[136,238,1644,1669]" degrees="106" direction="east" minutes="51" orientation="longitude" pageId="10" pageNumber="139" precision="925" value="106.85">106°51E</geoCoordinate>
,
<date id="5B4D10025C29FFEC70BB283AFE468DBD" box="[253,407,1645,1670]" pageId="10" pageNumber="139" value="1982-05-17">
<collectingDate id="4B09E9EA5C29FFEC70BB283AFE468DBD" box="[253,407,1645,1670]" pageId="10" pageNumber="139" value="1982-05-17">17 May 1982</collectingDate>
</date>
,
<collectingMethod id="F6B24ED55C29FFEC71E0283AFDDF8DBD" box="[422,526,1645,1669]" pageId="10" pageNumber="139">light trap</collectingMethod>
,
<collectorName id="820653145C29FFEC725B283AFD7D8DBE" box="[541,684,1645,1670]" pageId="10" pageNumber="139">Chikun Yang</collectorName>
leg. (
<collectionCode id="49E2AE075C29FFEC72B7283AFCFE8DBD" box="[753,815,1645,1669]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="10" pageNumber="139">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9F9B3C9F5C29FFEC7300283AFD248D90" collectingDate="1984-05-20" collectionCode="CAU" collectorName="Fasheng Li" country="China" county="Chongzuo" latitude="22.466667" location="Nonggang" longLatPrecision="1260" longitude="106.95" municipality="Longzhou" pageId="10" pageNumber="139" specimenCount="3" specimenCount-male="3" stateProvince="Guangxi">
<specimenCount id="39F5FD4B5C29FFEC7300283AFCB98DBD" box="[838,872,1642,1669]" count="3" pageId="10" pageNumber="139" type="male">3♂</specimenCount>
,
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:
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,
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,
<collectingMunicipality id="CF28ACB85C29FFEC74FE283AFAF68DBD" box="[1208,1319,1645,1669]" pageId="10" pageNumber="139">Longzhou</collectingMunicipality>
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<location id="2A2C60195C29FFEC75702839FA798DBE" LSID="urn:lsid:plazi:treatment:A75A87D45C29FFEC70CE2D9CFA4B8C0B:2A2C60195C29FFEC75702839FA798DBE" box="[1334,1448,1646,1670]" country="China" county="Chongzuo" latitude="22.466667" longLatPrecision="1260" longitude="106.95" municipality="Longzhou" name="Nonggang" pageId="10" pageNumber="139" stateProvince="Guangxi">Nonggang</location>
,
<geoCoordinate id="4AC750055C29FFEC70CE28C7FF358D9F" box="[136,228,1679,1704]" degrees="22" direction="north" minutes="28" orientation="latitude" pageId="10" pageNumber="139" precision="925" value="22.466667">22°28N</geoCoordinate>
,
<geoCoordinate id="4AC750055C29FFEC70A928C7FE848D9F" box="[239,341,1679,1704]" degrees="106" direction="east" minutes="57" orientation="longitude" pageId="10" pageNumber="139" precision="925" value="106.95">106°57E</geoCoordinate>
,
<date id="5B4D10025C29FFEC712628C7FE218D90" box="[352,496,1680,1704]" pageId="10" pageNumber="139" value="1984-05-20">
<collectingDate id="4B09E9EA5C29FFEC712628C7FE218D90" box="[352,496,1680,1704]" pageId="10" pageNumber="139" value="1984-05-20">20 May 1984</collectingDate>
</date>
,
<collectorName id="820653145C29FFEC71BD28C7FDA28D90" box="[507,627,1680,1704]" pageId="10" pageNumber="139">Fasheng Li</collectorName>
leg. (
<collectionCode id="49E2AE075C29FFEC72E928C7FD3C8D90" box="[687,749,1680,1704]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="10" pageNumber="139">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2F4C36C25C29FFEC70F828E4FDB48DF3" blockId="10.[136,1458,1091,1843]" box="[190,613,1715,1739]" pageId="10" pageNumber="139">
Distribution.
<collectingCountry id="57E476525C29FFEC710A28E4FE5D8DF3" box="[332,396,1715,1739]" name="China" pageId="10" pageNumber="139">China</collectingCountry>
(
<collectingRegion id="ED37F8205C29FFEC71DC28E4FE2B8DF3" box="[410,506,1715,1739]" country="China" name="Guangxi" pageId="10" pageNumber="139">Guangxi</collectingRegion>
,
<collectingRegion id="ED37F8205C29FFEC724528E4FD8B8DF3" box="[515,602,1715,1739]" country="China" name="Yunnan" pageId="10" pageNumber="139">Yunnan</collectingRegion>
).
</paragraph>
<paragraph id="2F4C36C25C29FFEC70F82882FA4B8C0B" blockId="10.[136,1458,1091,1843]" pageId="10" pageNumber="139">
Remarks. The pairing of the male to the female of
<taxonomicName id="E8F34D415C29FFEC728C2881FCE08DD5" authorityName="Yang" authorityYear="1986" box="[714,817,1750,1773]" class="Insecta" family="Sisyridae" genus="Sisyra" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="yunana">
<emphasis id="1D87EAD05C29FFEC728C2881FCE08DD5" box="[714,817,1750,1773]" italics="true" pageId="10" pageNumber="139">S. yunana</emphasis>
</taxonomicName>
is primarily based on the same color pattern of the antennae, which is a very important character for distinguishing the species of
<taxonomicName id="E8F34D415C29FFEC733228AFFC668C37" authorityName="Burmeister" authorityYear="1839" box="[884,951,1784,1807]" class="Insecta" family="Sisyridae" genus="Sisyra" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="1D87EAD05C29FFEC733228AFFC668C37" box="[884,951,1784,1807]" italics="true" pageId="10" pageNumber="139">Sisyra</emphasis>
</taxonomicName>
and stable within same species. Nevertheless, further male specimens of this species from the
<typeStatus id="F04888605C29FFEC72CE294CFD678C0B" box="[648,694,1819,1843]" pageId="10" pageNumber="139">type</typeStatus>
locality in
<collectingRegion id="ED37F8205C29FFEC736B294CFCAE8C0A" box="[813,895,1819,1842]" country="China" name="Yunnan" pageId="10" pageNumber="139">Yunnan</collectingRegion>
are needed for confirming the present association.
</paragraph>
</subSubSection>
</treatment>
</document>