368 lines
52 KiB
XML
368 lines
52 KiB
XML
<document id="1B49DEB9BA027C2830370F6E15A5FB2A" ID-DOI="10.11646/zootaxa.3972.2.6" ID-GBIF-Dataset="eee2db07-ee84-4161-87af-54c590a748cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="233469" ID-ZooBank="8D47A1B5-5AB3-4570-BC70-4CD81B340A87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461187497085" checkinUser="plazi" docAuthor="Golovatch, Sergei, Evsyukov, Aleksandr & Reip, Hans S." docDate="2015" docId="A5747B12121912685DEEFF57FA2FFA64" docLanguage="en" docName="zt03972p266.pdf" docOrigin="Zootaxa 3972 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hirudicryptus abchasicus Golovatch, Evsyukov & Reip, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="262" masterDocId="594D036A121312645D79FFC0FFB3FFCE" masterDocTitle="Colobognatha millipedes in the Caucasus (Diplopoda: Polyzoniida, Platydesmida, Siphonocryptida)" masterLastPageNumber="266" masterPageNumber="250" pageNumber="260" updateTime="1698599875139" updateUser="plazi">
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<mods:mods id="97E416C2AB324D803703A5CE9960887F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="38EDE6D5938EB514DCCB1288A0687AF0">
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<mods:title id="72E068C9E85B0271468641D52EF3CE49">Colobognatha millipedes in the Caucasus (Diplopoda: Polyzoniida, Platydesmida, Siphonocryptida)</mods:title>
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<mods:name id="A2AD9218A71C5F8E98EEE7AE1BA4ACDA" type="personal">
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<mods:namePart id="F6A46C4B44E2D4DD0CADFDB0DE0EBE37">Golovatch, Sergei</mods:namePart>
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<mods:roleTerm id="315D6F8A5BCB44CC9086DD07A9927DA0">Author</mods:roleTerm>
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<mods:namePart id="00BAC1979706AD4038A905A613483634">Evsyukov, Aleksandr</mods:namePart>
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</mods:name>
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<mods:name id="13F6DE8A715FF7F25D6B95D284D4C989" type="personal">
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<mods:roleTerm id="7B4BF975FB034374FFAE7E7DBE98BC86">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9B142C4843E8A2AD92DEA27CA9E2BC02">Reip, Hans S.</mods:namePart>
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<mods:typeOfResource id="A8CD97B0FE355117DAD776EE32FB2941">text</mods:typeOfResource>
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<mods:title id="5BF5C68E093A8395648F480ABCEEA300">Zootaxa</mods:title>
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<mods:part id="EE0C396918D3D14C2D68CEBF33856912">
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<mods:date id="0CBD9480C08133B3D0155CE4A789B51D">2015</mods:date>
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<mods:detail id="F34330E00F6F1E894844A904885D737F" type="volume">
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<mods:number id="4820090A6478E56F12AFF936082D2505">3972</mods:number>
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<mods:identifier id="FD961B92FE2006F1863292C3AA6C0109" type="DOI">10.11646/zootaxa.3972.2.6</mods:identifier>
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<treatment id="A5747B12121912685DEEFF57FA2FFA64" ID-DOI="http://doi.org/10.5281/zenodo.6107430" ID-GBIF-Taxon="119628871" ID-Zenodo-Dep="6107430" LSID="urn:lsid:plazi:treatment:A5747B12121912685DEEFF57FA2FFA64" httpUri="http://treatment.plazi.org/id/A5747B12121912685DEEFF57FA2FFA64" lastPageId="12" lastPageNumber="262" pageId="10" pageNumber="260">
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<subSubSection id="65C7998F1219126E5DEEFF57FEB5FF1D" pageId="10" pageNumber="260" type="nomenclature">
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<paragraph id="2D62CA041219126E5DEEFF57FD81FF7C" blockId="10.[151,562,151,211]" box="[151,562,151,178]" pageId="10" pageNumber="260">
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<heading id="762A7D681219126E5DEEFF57FD81FF7C" bold="true" box="[151,562,151,178]" fontSize="11" level="1" pageId="10" pageNumber="260" reason="1">
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<emphasis id="1FA916161219126E5DEEFF57FD81FF7C" bold="true" box="[151,562,151,178]" pageId="10" pageNumber="260">
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<taxonomicName id="EADDB1871219126E5DEEFF57FE7FFF7F" ID-CoL="849D6" box="[151,460,151,177]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="species" species="abchasicus" status="sp. nov.">
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<emphasis id="1FA916161219126E5DEEFF57FE7FFF7F" bold="true" box="[151,460,151,177]" italics="true" pageId="10" pageNumber="260">Hirudicryptus abchasicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="049AAB6D1219126E5CADFF58FD81FF7C" box="[468,562,152,178]" pageId="10" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2D62CA041219126E5DEEFF7AFEB5FF1D" blockId="10.[151,562,151,211]" box="[151,262,186,211]" pageId="10" pageNumber="260">
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<figureCitation id="B5E6D6811219126E5DEEFF7AFEB5FF1D" box="[151,262,186,211]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="11.[151,250,1918,1940]" captionStartId-1="12.[151,250,658,680]" captionStartId-2="13.[151,250,1956,1978]" captionStartId-3="14.[151,250,1151,1173]" captionStartId-4="14.[151,250,1961,1983]" captionTargetBox-0="[175,1420,210,1871]" captionTargetBox-1="[161,1424,203,628]" captionTargetBox-2="[160,1424,201,1907]" captionTargetBox-3="[150,1433,193,1128]" captionTargetBox-4="[151,1436,1284,1940]" captionTargetId-0="figure@11.[151,1436,193,1897]" captionTargetId-1="figure@12.[151,1435,193,637]" captionTargetId-2="figure@13.[159,1428,195,1934]" captionTargetId-3="figure@14.[150,1436,193,1129]" captionTargetId-4="figure@14.[151,1436,1284,1940]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="14" captionText-0="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." captionText-1="FIGURE 7. Hirudicryptus abchasicus sp. nov., male holotype. A: head, collum and segment 2, oral view. B: midbody paraterga, lateral view. C: caudal end of body, dorsal view. Scale bar: 0.5 mm." captionText-2="FIGURE 8. Hirudicryptus abchasicus sp. nov., male holotype. A: leg 11, caudal view. B: anterior gonopods, anterior view; C: tip of right anterior gonopod, anterodorsal view. D: right posterior gonopod, ventrolateral view. Scale bar: 0.1 mm." captionText-3="FIGURE 9. Distribution of Colobognatha in the Caucasus and adjacent parts of Turkey and Iran. Blue square— Hirudisoma roseum (Victor, 1839); red asterisk— Fioria hyrcana Golovatch, 1980; green triangle— Hirudicryptus abchasicus sp. nov." captionText-4="FIGURE 10. Distribution of the genus Hirudicryptus." httpUri-0="https://zenodo.org/record/233475/files/figure.png" httpUri-1="https://zenodo.org/record/233476/files/figure.png" httpUri-2="https://zenodo.org/record/233477/files/figure.png" httpUri-3="https://zenodo.org/record/233478/files/figure.png" httpUri-4="https://zenodo.org/record/233479/files/figure.png" pageId="10" pageNumber="260">Figs 6–10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="65C7998F1219126E5DEEFEC2FCD3FEAD" pageId="10" pageNumber="260" type="materials_examined">
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<paragraph id="2D62CA041219126E5DEEFEC2FB0BFEF1" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
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<emphasis id="1FA916161219126E5DEEFEC2FE3BFED5" bold="true" box="[151,392,258,283]" pageId="10" pageNumber="260">Material examined.</emphasis>
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<typeStatus id="F26674A61219126E5CEEFEC2FDB3FED5" box="[407,512,258,283]" pageId="10" pageNumber="260" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="4BCC52C11219126E5F25FEC3FD06FED5" box="[604,693,259,283]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="260">ZMUM</collectionCode>
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ρ2583),
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<emphasis id="1FA916161219126E5E53FEC2FC71FED5" bold="true" box="[810,962,258,283]" pageId="10" pageNumber="260">ABKHAZIA</emphasis>
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, Gumista Nature Reserve, forest litter,
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<date id="5963ECC41219126E5DEEFEE7FEB8FEF1" box="[151,267,295,319]" pageId="10" pageNumber="260" value="1982-06-04">4.VI.1982</date>
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, leg. J. Boháč.
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<typeStatus id="F26674A61219126E5CCCFEE7FDA4FEF1" box="[437,535,295,319]" pageId="10" pageNumber="260" type="paratype">Paratype</typeStatus>
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female (
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<collectionCode id="4BCC52C11219126E5F00FEE7FD61FEF1" box="[633,722,295,319]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="260">ZMUM</collectionCode>
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ρ2584), same data, together with
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<typeStatus id="F26674A61219126E5936FEE6FB06FEF1" box="[1103,1205,294,319]" pageId="10" pageNumber="260" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="2D62CA041219126E5DBEFE8AFCD3FEAD" blockId="10.[151,1437,258,2014]" box="[199,864,330,355]" pageId="10" pageNumber="260">
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<emphasis id="1FA916161219126E5DBEFE8AFEA5FEAD" bold="true" box="[199,278,330,355]" pageId="10" pageNumber="260">Name:</emphasis>
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To emphasize Abkhazia, the
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<emphasis id="1FA916161219126E5F18FE8CFD57FEAD" box="[609,740,332,355]" italics="true" pageId="10" pageNumber="260">terra typica</emphasis>
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; adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="65C7998F1219126E5DBEFEAEFF52FDD9" pageId="10" pageNumber="260" type="diagnosis">
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<paragraph id="2D62CA041219126E5DBEFEAEFF52FDD9" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
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<emphasis id="1FA916161219126E5DBEFEAEFEF0FE49" bold="true" box="[199,323,366,391]" pageId="10" pageNumber="260">Diagnosis:</emphasis>
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Distinguished from all congeners but
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<taxonomicName id="EADDB1871219126E5F80FEAFFBF7FE49" authority="Loksa, 1967" authorityName="Loksa" authorityYear="1967" box="[761,1092,366,391]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="species" species="canariensis">
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<emphasis id="1FA916161219126E5F80FEAFFC2EFE48" box="[761,925,367,390]" italics="true" pageId="10" pageNumber="260">H. canariensis</emphasis>
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(Loksa, 1967)
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</taxonomicName>
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by the much shorter antennae which are only about as long as collum width, as well as by the smaller and much shorter collum which only forms a quite inconspicuous casque-like shield to cover the head. Differs from
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<taxonomicName id="EADDB1871219126E5EBDFE77FBDBFE00" box="[964,1128,439,462]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="species" species="canariensis">
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<emphasis id="1FA916161219126E5EBDFE77FBDBFE00" box="[964,1128,439,462]" italics="true" pageId="10" pageNumber="260">H. canariensis</emphasis>
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</taxonomicName>
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by the concave last tergum and less numerous, 9–11 + 9–11 vs 14–15 + 14–15, tubercles at the caudal margin of midbody terga. See also Key below.
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</paragraph>
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</subSubSection>
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<subSubSection id="65C7998F1219126E5DBEFDE2FB12F970" pageId="10" pageNumber="260" type="description">
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<paragraph id="2D62CA041219126E5DBEFDE2FC2EFD4D" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
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<emphasis id="1FA916161219126E5DBEFDE2FEE8FDF5" bold="true" box="[199,347,546,571]" pageId="10" pageNumber="260">Description:</emphasis>
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Length about 8.5 (
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<typeStatus id="F26674A61219126E5F4AFDE2FD2AFDF5" box="[563,665,546,571]" pageId="10" pageNumber="260" type="holotype">holotype</typeStatus>
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) or
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<quantity id="EA2567E11219126E5FBEFDE3FCBAFDF4" box="[711,777,547,570]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="260" unit="mm" value="7.0">7 mm</quantity>
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(
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<typeStatus id="F26674A61219126E5E61FDE3FCCDFDF5" box="[792,894,547,571]" pageId="10" pageNumber="260" type="paratype">paratype</typeStatus>
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), width ca 0.9 and 1.0 (
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<typeStatus id="F26674A61219126E59F0FDE2FB5DFDF5" box="[1161,1262,546,571]" pageId="10" pageNumber="260" type="holotype">holotype</typeStatus>
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) or 0.8 and
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<quantity id="EA2567E11219126E5800FDE3FF70FD90" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="260" unit="mm" value="0.9">0.9 mm</quantity>
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(
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<typeStatus id="F26674A61219126E5DAAFD87FE84FD91" box="[211,311,583,607]" pageId="10" pageNumber="260" type="paratype">paratype</typeStatus>
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) on midbody pro- and metazonae, respectively.
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<typeStatus id="F26674A61219126E5E1BFD86FC79FD91" box="[866,970,582,607]" pageId="10" pageNumber="260" type="holotype">Holotype</typeStatus>
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with 48 podous + 1 apodous segment + telson,
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<typeStatus id="F26674A61219126E5D91FDABFEFBFD4D" box="[232,328,619,643]" pageId="10" pageNumber="260" type="paratype">paratype</typeStatus>
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with 43p + 1a + telson. Length to width ratio> 10:1.
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</paragraph>
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<paragraph id="2D62CA041219126E5DBEFD4EFCC1FC09" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
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Coloration in alcohol rather uniformly red- or grey-brown to yellowish with a narrow light axial line (alcohol material), anterior part of body in
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<typeStatus id="F26674A61219126E5F6EFD73FDC4FD05" box="[535,631,691,715]" pageId="10" pageNumber="260" type="paratype">paratype</typeStatus>
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darker, marbled grey; metatergal bosses/tubercles at caudal margin and antennae mostly dark grey-brown while a very wide mid-dorsal stripe lighter both in holo- and
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<typeStatus id="F26674A61219126E59B4FD17FA9EFD21" box="[1229,1325,727,751]" pageId="10" pageNumber="260" type="paratype">paratype</typeStatus>
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(
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<figureCitation id="B5E6D6811219126E5847FD16FA32FD21" box="[1342,1409,726,751]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Fig. 6</figureCitation>
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A, C). Ocelli inside a black ocular field. Body very clearly flattened dorsoventrally, hirudiform, regularly, but strongly broadened until segment 6 or 7, similarly regularly and clearly narrowing towards telson on last 4 or 5 segments. Head very small and rather narrow, coniform, about 1/3 as broad as collum and about twice as long as broad; rostrum narrowly rounded (
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<figureCitation id="B5E6D6811219126E5CAFFCA6FD92FCB1" box="[470,545,870,895]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Figs 6</figureCitation>
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A, 7A). Ocelli large, convex, 2+2 on each side of head arranged in a nearly longitudinal line, each ocellus with a long seta at base. Antennae short, stout, erect, rod-shaped, not clavate, about as long as collum width; antennomere 6 largest (
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<figureCitation id="B5E6D6811219126E5FCCFC6EFD4FFC09" box="[693,764,942,967]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Figs 6</figureCitation>
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A, B, 7A).
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</paragraph>
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<paragraph id="2D62CA041219126E5DBEFC12FD9DFA28" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
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Collum biconvex, short and broad (
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<figureCitation id="B5E6D6811219126E5F27FC12FD11FC25" box="[606,674,978,1003]" captionStart="FIGURE 7" captionStartId="12.[151,250,658,680]" captionTargetBox="[161,1424,203,628]" captionTargetId="figure@12.[151,1435,193,637]" captionTargetPageId="12" captionText="FIGURE 7. Hirudicryptus abchasicus sp. nov., male holotype. A: head, collum and segment 2, oral view. B: midbody paraterga, lateral view. C: caudal end of body, dorsal view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/233476/files/figure.png" pageId="10" pageNumber="260">Fig. 7</figureCitation>
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A), like head characteristically strongly inclined forward to form a small casque-like shield, also distinctly separated from next tergum (
|
||
<figureCitation id="B5E6D6811219126E5EBCFC36FBBEFBC1" box="[965,1037,1014,1039]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Fig. 6</figureCitation>
|
||
B, D); paraterga on collum small, strongly declined, subvertical, each delimited by an oblique sharp bend; dorsal surface rather flat, only slightly impressed medially; caudal margin with a transverse row of several small grains/tubercles (
|
||
<figureCitation id="B5E6D6811219126E599FFBFEFA9CFB99" box="[1254,1327,1086,1111]" captionStart="FIGURE 7" captionStartId="12.[151,250,658,680]" captionTargetBox="[161,1424,203,628]" captionTargetId="figure@12.[151,1435,193,637]" captionTargetPageId="12" captionText="FIGURE 7. Hirudicryptus abchasicus sp. nov., male holotype. A: head, collum and segment 2, oral view. B: midbody paraterga, lateral view. C: caudal end of body, dorsal view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/233476/files/figure.png" pageId="10" pageNumber="260">Fig. 7</figureCitation>
|
||
A) and a considerable lighter gap between paramedian pair (
|
||
<figureCitation id="B5E6D6811219126E5F9DFBA2FC98FBB5" box="[740,811,1122,1147]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Fig. 6</figureCitation>
|
||
). Axial suture inconspicuous, but visible starting with collum. Following paraterga nearly acute at caudal corner, especially clearly drawn behind tergal margin only in 2nd and a few caudalmost segments, paraterga of penultimate segment fused medially into a broad, terminally slightly concave plate with two caudolateral denticles and a vestigial axial suture (
|
||
<figureCitation id="B5E6D6811219126E5E85FB11FBF4FB24" box="[1020,1095,1233,1258]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Figs 6</figureCitation>
|
||
C, 7C). Lateral peritremes on paraterga evident, delimited by a sulcus both dorsally and, to lesser degree, ventrally. Metaterga slightly, but clearly elevated caudally, each with 9–11 + 9–11 small bosses/tubercles at caudal margin, like collum with a considerable lighter gap between paramedian pair. Surface of collum and following metaterga faintly rugulose and microtuberculate, dull all over. Ozopores lateral, starting with segment 5, small, inconspicuous, each lying at bottom of a small oblong groove in front of caudolateral corner of a clearly thickened, nearly porostele-shaped peritreme (
|
||
<figureCitation id="B5E6D6811219126E5C6AFA69FEEBFA0C" box="[275,344,1449,1474]" captionStart="FIGURE 7" captionStartId="12.[151,250,658,680]" captionTargetBox="[161,1424,203,628]" captionTargetId="figure@12.[151,1435,193,637]" captionTargetPageId="12" captionText="FIGURE 7. Hirudicryptus abchasicus sp. nov., male holotype. A: head, collum and segment 2, oral view. B: midbody paraterga, lateral view. C: caudal end of body, dorsal view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/233476/files/figure.png" pageId="10" pageNumber="260">Fig. 7</figureCitation>
|
||
B). Telson small, completely concealed in dorsal view by caudal plate of penultimate segment’s fused paraterga (
|
||
<figureCitation id="B5E6D6811219126E5C28FA0DFE2BFA28" box="[337,408,1485,1510]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Figs 6</figureCitation>
|
||
A, C, D, 7C).
|
||
</paragraph>
|
||
<paragraph id="2D62CA041219126E5DBEFA32FE1EF99C" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
|
||
Sterna about 2/3 as wide as coxae, the latter almost in touch medially (
|
||
<figureCitation id="B5E6D6811219126E5EA2FA31FBADF9C4" box="[987,1054,1521,1546]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="10" pageNumber="260">Fig. 6</figureCitation>
|
||
A, C, E). Legs 6-segmented. Male coxa 2 with a short, tube-shaped, simple, basal gonapophysis directed caudomesad. Claws simple (
|
||
<figureCitation id="B5E6D6811219126E5992F9D5FA9DF9E0" box="[1259,1326,1557,1582]" captionStart="FIGURE 8" captionStartId="13.[151,250,1956,1978]" captionTargetBox="[160,1424,201,1907]" captionTargetId="figure@13.[159,1428,195,1934]" captionTargetPageId="13" captionText="FIGURE 8. Hirudicryptus abchasicus sp. nov., male holotype. A: leg 11, caudal view. B: anterior gonopods, anterior view; C: tip of right anterior gonopod, anterodorsal view. D: right posterior gonopod, ventrolateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/233477/files/figure.png" pageId="10" pageNumber="260">Fig. 8</figureCitation>
|
||
A). Coxal sacks starting with leg 3.
|
||
</paragraph>
|
||
<paragraph id="2D62CA041219126E5DBEF99DFB12F970" blockId="10.[151,1437,258,2014]" pageId="10" pageNumber="260">
|
||
Anterior gonopods (
|
||
<figureCitation id="B5E6D6811219126E5CD4F99DFE42F9B8" box="[429,497,1629,1654]" captionStart="FIGURE 8" captionStartId="13.[151,250,1956,1978]" captionTargetBox="[160,1424,201,1907]" captionTargetId="figure@13.[159,1428,195,1934]" captionTargetPageId="13" captionText="FIGURE 8. Hirudicryptus abchasicus sp. nov., male holotype. A: leg 11, caudal view. B: anterior gonopods, anterior view; C: tip of right anterior gonopod, anterodorsal view. D: right posterior gonopod, ventrolateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/233477/files/figure.png" pageId="10" pageNumber="260">Fig. 8</figureCitation>
|
||
B) apparently 5-segmented, C-shaped, strongly incrassate and stout. Tarsus with a strong apical stylet and a gutter for accommodation of posterior gonopod tarsus (
|
||
<figureCitation id="B5E6D6811219126E5948F941FBC7F954" box="[1073,1140,1665,1690]" captionStart="FIGURE 8" captionStartId="13.[151,250,1956,1978]" captionTargetBox="[160,1424,201,1907]" captionTargetId="figure@13.[159,1428,195,1934]" captionTargetPageId="13" captionText="FIGURE 8. Hirudicryptus abchasicus sp. nov., male holotype. A: leg 11, caudal view. B: anterior gonopods, anterior view; C: tip of right anterior gonopod, anterodorsal view. D: right posterior gonopod, ventrolateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/233477/files/figure.png" pageId="10" pageNumber="260">Fig. 8</figureCitation>
|
||
B, C). Posterior gonopods (
|
||
<figureCitation id="B5E6D6811219126E5DE6F965FF52F970" box="[159,225,1701,1726]" captionStart="FIGURE 8" captionStartId="13.[151,250,1956,1978]" captionTargetBox="[160,1424,201,1907]" captionTargetId="figure@13.[159,1428,195,1934]" captionTargetPageId="13" captionText="FIGURE 8. Hirudicryptus abchasicus sp. nov., male holotype. A: leg 11, caudal view. B: anterior gonopods, anterior view; C: tip of right anterior gonopod, anterodorsal view. D: right posterior gonopod, ventrolateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/233477/files/figure.png" pageId="10" pageNumber="260">Fig. 8</figureCitation>
|
||
D) apparently 6-segmented, much slenderer; tarsus longest, flagelliform, very simple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65C7998F121912685DBEF909FB75FB59" lastPageId="12" lastPageNumber="262" pageId="10" pageNumber="260" type="discussion">
|
||
<paragraph id="2D62CA04121912685DBEF909FE39FC2A" blockId="10.[151,1437,258,2014]" lastBlockId="12.[151,1437,755,1104]" lastPageId="12" lastPageNumber="262" pageId="10" pageNumber="260">
|
||
<emphasis id="1FA916161219126E5DBEF909FE87F92C" bold="true" box="[199,308,1737,1762]" pageId="10" pageNumber="260">Remarks</emphasis>
|
||
. The order
|
||
<taxonomicName id="EADDB1871219126E5CCEF909FDC3F92C" box="[439,624,1737,1762]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="order">Siphonocryptida</taxonomicName>
|
||
has hitherto been known to contain only six species in two genera and a single family (
|
||
<bibRefCitation id="494CB7F51219126E5C40F92DFD37F8C8" author="Enghoff" box="[313,644,1773,1798]" pageId="10" pageNumber="260" refString="Enghoff, H. & Golovatch, S. I. (1995) A revision of the Siphonocryptidae (Diplopoda, Polyzoniida). Zoologica Scripta, 24 (1), 29 - 41. Available from: http: // www. readcube. com / articles / 10.1111 % 2 Fj. 1463 - 6409.1995. tb 00473. x" type="journal article" year="1995">Enghoff and Golovatch 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="494CB7F51219126E5FE9F92EFCD3F8C8" author="Korsos" box="[656,864,1773,1798]" pageId="10" pageNumber="260" refString="Korsos, Z., Enghoff, H., Chang, H. W. (2008) A most unusual anomal distribution pattern: A new siphonocryptid millipede from Taiwan (Diplopoda, Sipjonocryptida). Acta Zoologica Academiae Scientarum Hungaricae, 54, 151 - 157." type="journal article" year="2008">
|
||
Korsós
|
||
<emphasis id="1FA916161219126E5F9FF92FFCABF8C8" box="[742,792,1773,1798]" italics="true" pageId="10" pageNumber="260">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="494CB7F51219126E5E15F92EFC14F8C8" author="Korsos" box="[876,935,1774,1798]" pageId="10" pageNumber="260" refString="Korsos, Z., Geoffroy, J. - J., Mauries, J. - P. (2009) The fifth element: Reconnection of the disjunct distribution of the members of Siphonocryptida (Diplopoda) with the description of a new species from Nepal. Journal of Natural History, 43, 435 - 445. doi: 10.1080 / 00222930802610428" type="journal article" year="2009">2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="494CB7F51219126E5ECAF92DFBE1F8C8" author="Enghoff" box="[947,1106,1773,1798]" pageId="10" pageNumber="260" refString="Enghoff, H. (2010) A new strikingly coloured species of Siphonocryptus, sixth of its order (Diplopoda: Siphonocryptida). Zootaxa 2681, 66 - 68." type="journal article" year="2010">Enghoff 2010</bibRefCitation>
|
||
). The group’s highly peculiar distribution only emphasizes its relictual status (
|
||
<bibRefCitation id="494CB7F51219126E5FB1F8D1FBA6F8E4" author="Shelley" box="[712,1045,1809,1834]" pageId="10" pageNumber="260" refString="Shelley, R. M. & Golovatch, S. I. (2011) Atlas of myriapod biogeography. I. Indigenous ordinal and supra-ordinal distributions in the Diplopoda: Perspectives on taxon origins and ages, and a hypothesis on the origin and early evolution of the class. Insecta Mundi 0 158, 1 - 134." type="journal article" year="2011">Shelley and Golovatch 2011</bibRefCitation>
|
||
). Thus, the genus
|
||
<taxonomicName id="EADDB1871219126E5997F8D1FE80F880" authority="Pocock, 1894" authorityName="Pocock" authorityYear="1894" class="Diplopoda" family="Siphonocryptidae" genus="Siphonocryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FA916161219126E5997F8D1FA2FF8E4" box="[1262,1436,1809,1834]" italics="true" pageId="10" pageNumber="260">Siphonocryptus</emphasis>
|
||
Pocock, 1894
|
||
</taxonomicName>
|
||
, comprises three species, one in Sumatra,
|
||
<collectingCountry id="55CA8A941219126E5E6CF8F5FC36F880" box="[789,901,1845,1870]" name="Indonesia" pageId="10" pageNumber="260">Indonesia</collectingCountry>
|
||
, the other two in continental Western
|
||
<collectingCountry id="55CA8A941219126E584CF8F5FA2FF880" box="[1333,1436,1845,1870]" name="Malaysia" pageId="10" pageNumber="260">Malaysia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="494CB7F51219126E5DE7F899FE40F8BC" author="Enghoff" box="[158,499,1881,1906]" pageId="10" pageNumber="260" refString="Enghoff, H. & Golovatch, S. I. (1995) A revision of the Siphonocryptidae (Diplopoda, Polyzoniida). Zoologica Scripta, 24 (1), 29 - 41. Available from: http: // www. readcube. com / articles / 10.1111 % 2 Fj. 1463 - 6409.1995. tb 00473. x" type="journal article" year="1995">Enghoff and Golovatch 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="494CB7F51219126E5F7BF899FD16F8BC" author="Enghoff" box="[514,677,1881,1906]" pageId="10" pageNumber="260" refString="Enghoff, H. (2010) A new strikingly coloured species of Siphonocryptus, sixth of its order (Diplopoda: Siphonocryptida). Zootaxa 2681, 66 - 68." type="journal article" year="2010">Enghoff 2010</bibRefCitation>
|
||
). In contrast, the distribution pattern of
|
||
<taxonomicName id="EADDB1871219126E5900F899FEFDF858" authority="Enghoff & Golovatch 1995" authorityName="Enghoff & Golovatch" authorityYear="1995" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FA916161219126E5900F899FAA5F8BC" box="[1145,1302,1881,1906]" italics="true" pageId="10" pageNumber="260">Hirudicryptus</emphasis>
|
||
<bibRefCitation id="494CB7F51219126E5859F899FEFDF858" author="Enghoff" pageId="10" pageNumber="260" refString="Enghoff, H. & Golovatch, S. I. (1995) A revision of the Siphonocryptidae (Diplopoda, Polyzoniida). Zoologica Scripta, 24 (1), 29 - 41. Available from: http: // www. readcube. com / articles / 10.1111 % 2 Fj. 1463 - 6409.1995. tb 00473. x" type="journal article" year="1995">Enghoff & Golovatch 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
is trans-Palaearctic (
|
||
<figureCitation id="B5E6D6811219126E5F3CF8BDFD2EF858" box="[581,669,1917,1942]" captionStart="FIGURE 10" captionStartId="14.[151,250,1961,1983]" captionTargetBox="[151,1436,1284,1940]" captionTargetId="figure@14.[151,1436,1284,1940]" captionTargetPageId="14" captionText="FIGURE 10. Distribution of the genus Hirudicryptus." httpUri="https://zenodo.org/record/233479/files/figure.png" pageId="10" pageNumber="260">Fig. 10</figureCitation>
|
||
). The
|
||
<typeStatus id="F26674A61219126E5F95F8BEFCAFF858" box="[748,796,1918,1942]" pageId="10" pageNumber="260">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="EADDB1871219126E5EF3F8BFFB6CF858" authority="Loksa, 1967" authorityName="Loksa" authorityYear="1967" box="[906,1247,1917,1942]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis id="1FA916161219126E5EF3F8BFFB82F85B" box="[906,1073,1918,1942]" italics="true" pageId="10" pageNumber="260">H. canariensis</emphasis>
|
||
(Loksa, 1967)
|
||
</taxonomicName>
|
||
, occurs only on Madeira and the Canaries, where it is largely confined to the relict laurisilva biome. The second species,
|
||
<taxonomicName id="EADDB1871219126E58F8F862FD16F810" authority="Korsos, Enghoff & Chang, 2008" authorityName="Korsos, Enghoff & Chang" authorityYear="2008" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="species" species="taiwanensis">
|
||
<emphasis id="1FA916161219126E58F8F862FEAFF813" italics="true" pageId="10" pageNumber="260">H. taiwanensis</emphasis>
|
||
<bibRefCitation id="494CB7F51219126E5C5BF806FD16F810" author="Korsos" box="[290,677,1989,2014]" pageId="10" pageNumber="260" refString="Korsos, Z., Enghoff, H., Chang, H. W. (2008) A most unusual anomal distribution pattern: A new siphonocryptid millipede from Taiwan (Diplopoda, Sipjonocryptida). Acta Zoologica Academiae Scientarum Hungaricae, 54, 151 - 157." type="journal article" year="2008">Korsós, Enghoff & Chang, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, is endemic to
|
||
<collectingCountry id="55CA8A941219126E5E22F805FC03F810" box="[859,944,1989,2014]" name="Taiwan" pageId="10" pageNumber="260">Taiwan</collectingCountry>
|
||
, whereas the third,
|
||
<taxonomicName id="EADDB1871219126859F3F807FD9AFCC2" authority="Korsos, Geoffroy & Mauries, 2009" authorityName="Korsos, Geoffroy & Mauries" authorityYear="2009" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" lastPageId="12" lastPageNumber="262" order="Siphonocryptida" pageId="10" pageNumber="260" phylum="Arthropoda" rank="species" species="quintumelementum">
|
||
<emphasis id="1FA916161219126E59F3F807FA32F810" box="[1162,1409,1989,2014]" italics="true" pageId="10" pageNumber="260">H. quintumelementum</emphasis>
|
||
<bibRefCitation id="494CB7F5121F12685DEEFD34FD9AFCC2" author="Korsos" box="[151,553,755,780]" pageId="12" pageNumber="262" refString="Korsos, Z., Geoffroy, J. - J., Mauries, J. - P. (2009) The fifth element: Reconnection of the disjunct distribution of the members of Siphonocryptida (Diplopoda) with the description of a new species from Nepal. Journal of Natural History, 43, 435 - 445. doi: 10.1080 / 00222930802610428" type="journal article" year="2009">Korsós, Geoffroy & Mauriès, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, is only known from a few females collected at a
|
||
<quantity id="EA2567E1121F1268591FFD33FB0FFCC5" box="[1126,1212,755,780]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="262" unit="m" value="2500.0">2500 m</quantity>
|
||
elevation in
|
||
<collectingCountry id="55CA8A94121F1268582BFD33FA2BFCC2" box="[1362,1432,755,780]" name="Nepal" pageId="12" pageNumber="262">Nepal</collectingCountry>
|
||
, Himalayas (
|
||
<bibRefCitation id="494CB7F5121F12685C66FCD7FDC3FCFE" author="Enghoff" box="[287,624,791,816]" pageId="12" pageNumber="262" refString="Enghoff, H. & Golovatch, S. I. (1995) A revision of the Siphonocryptidae (Diplopoda, Polyzoniida). Zoologica Scripta, 24 (1), 29 - 41. Available from: http: // www. readcube. com / articles / 10.1111 % 2 Fj. 1463 - 6409.1995. tb 00473. x" type="journal article" year="1995">Enghoff and Golovatch 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="494CB7F5121F12685F07FCD8FCE6FCFE" author="Korsos" box="[638,853,791,816]" pageId="12" pageNumber="262" refString="Korsos, Z., Enghoff, H., Chang, H. W. (2008) A most unusual anomal distribution pattern: A new siphonocryptid millipede from Taiwan (Diplopoda, Sipjonocryptida). Acta Zoologica Academiae Scientarum Hungaricae, 54, 151 - 157." type="journal article" year="2008">
|
||
Korsós
|
||
<emphasis id="1FA91616121F12685FAEFCD9FCB9FCFE" box="[727,778,791,816]" italics="true" pageId="12" pageNumber="262">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="494CB7F5121F12685E1AFCD8FC13FCFE" author="Korsos" box="[867,928,792,816]" pageId="12" pageNumber="262" refString="Korsos, Z., Geoffroy, J. - J., Mauries, J. - P. (2009) The fifth element: Reconnection of the disjunct distribution of the members of Siphonocryptida (Diplopoda) with the description of a new species from Nepal. Journal of Natural History, 43, 435 - 445. doi: 10.1080 / 00222930802610428" type="journal article" year="2009">2009</bibRefCitation>
|
||
). It may well be that the distribution pattern under consideration dates back at least to the Oligocene times of the so-called “Warm Earth” to have highly probable explanations rooted in palaeobotanical evidence. These imply a gradual shrinkage and disruption ever since of the previously dominating and continuous subtropical biome (
|
||
<bibRefCitation id="494CB7F5121F12685ED6FC43FBDBFC52" author="Golovatch" box="[943,1128,899,924]" pageId="12" pageNumber="262" refString="Golovatch, S. I. (1997) On the main patterns of millipede distribution and faunogenesis in Eurasia (Diplopoda). Entomologica Scandinavica, Supplement 51, 199 - 208." type="book chapter" year="1997">Golovatch 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="494CB7F5121F1268590DFC43FA94FC52" author="Zherikhin" box="[1140,1319,899,924]" pageId="12" pageNumber="262" refString="Zherikhin, V. V. (2003) [A history of biomes]. In: KMK Scientific Press, Moscow, 98 - 188. [in Russian]" type="book chapter" year="2003">Zherikhin 2003</bibRefCitation>
|
||
). Being so vastly disjunct, the present-day distribution of
|
||
<taxonomicName id="EADDB187121F12685FDDFC67FCEEFC0E" box="[676,861,935,960]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="order">Siphonocryptida</taxonomicName>
|
||
is best accounted for by extinction events (
|
||
<bibRefCitation id="494CB7F5121F1268583EFC67FECDFC2A" author="Shelley" pageId="12" pageNumber="262" refString="Shelley, R. M. & Golovatch, S. I. (2011) Atlas of myriapod biogeography. I. Indigenous ordinal and supra-ordinal distributions in the Diplopoda: Perspectives on taxon origins and ages, and a hypothesis on the origin and early evolution of the class. Insecta Mundi 0 158, 1 - 134." type="journal article" year="2011">Shelley and Golovatch 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="79A29A8C1218126F5DEEF8BEFCEFF81F" httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="11" pageNumber="261" targetBox="[175,1420,210,1871]" targetPageId="11">
|
||
<paragraph id="2D62CA041218126F5DEEF8BEFCEFF81F" blockId="11.[151,1436,1918,2001]" pageId="11" pageNumber="261">
|
||
<emphasis id="1FA916161218126F5DEEF8BEFEA6F85D" bold="true" box="[151,277,1918,1940]" pageId="11" pageNumber="261">FIGURE 6.</emphasis>
|
||
<taxonomicName id="EADDB1871218126F5C64F8BEFD96F85A" box="[285,549,1918,1940]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="11" pageNumber="261" phylum="Arthropoda" rank="species" species="abchasicus" status="sp. nov.">
|
||
<emphasis id="1FA916161218126F5C64F8BEFD96F85A" box="[285,549,1918,1940]" italics="true" pageId="11" pageNumber="261">Hirudicryptus abchasicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FA916161218126F5F55F8BFFDCCF85A" bold="true" box="[556,639,1919,1940]" pageId="11" pageNumber="261">
|
||
<taxonomicNameLabel id="049AAB6D1218126F5F55F8BFFDCCF85A" box="[556,639,1919,1940]" pageId="11" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype (A–C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="79A29A8C121F12685DEEFD52FC1EFD09" httpUri="https://zenodo.org/record/233476/files/figure.png" pageId="12" pageNumber="262" targetBox="[161,1424,203,628]" targetPageId="12">
|
||
<paragraph id="2D62CA04121F12685DEEFD52FC1EFD09" blockId="12.[151,1436,658,711]" pageId="12" pageNumber="262">
|
||
<emphasis id="1FA91616121F12685DEEFD52FEABFD66" bold="true" box="[151,280,658,680]" pageId="12" pageNumber="262">FIGURE 7.</emphasis>
|
||
<taxonomicName id="EADDB187121F12685C5AFD52FD9EFD66" box="[291,557,658,680]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="species" species="abchasicus" status="sp. nov.">
|
||
<emphasis id="1FA91616121F12685C5AFD52FD9EFD66" box="[291,557,658,680]" italics="true" pageId="12" pageNumber="262">Hirudicryptus abchasicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="049AAB6D121F12685F41FD53FD3EFD66" box="[568,653,659,680]" pageId="12" pageNumber="262" rank="species">
|
||
<emphasis id="1FA91616121F12685F41FD53FD3EFD66" bold="true" box="[568,653,659,680]" pageId="12" pageNumber="262">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype. A: head, collum and segment 2, oral view. B: midbody paraterga, lateral view. C: caudal end of body, dorsal view. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D62CA04121F12685DBEFC2FFB7FFB9E" blockId="12.[151,1437,755,1104]" pageId="12" pageNumber="262">
|
||
The discovery of
|
||
<taxonomicName id="EADDB187121F12685CDDFC31FDF8FBC6" box="[420,587,1007,1032]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="species" species="abchasicus" status="sp. nov.">
|
||
<emphasis id="1FA91616121F12685CDDFC31FDF8FBC6" box="[420,587,1007,1032]" italics="true" pageId="12" pageNumber="262">H. abchasicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="049AAB6D121F12685F23FC2FFD0FFBC6" box="[602,700,1007,1032]" pageId="12" pageNumber="262" rank="species">
|
||
<emphasis id="1FA91616121F12685F23FC2FFD0FFBC6" bold="true" box="[602,700,1007,1032]" pageId="12" pageNumber="262">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the Caucasus very nicely bridges the huge gap between Macaronesia and the Himalayas (Fig, 10). This species is clearly a highly relictual element in the Caucasian fauna. Luckily, it stems from a nature reserve, thus demanding no other special measures of protection.
|
||
</paragraph>
|
||
<paragraph id="2D62CA04121F12685DEEFBBFFB75FB59" blockId="12.[151,1222,1151,1176]" box="[151,1222,1151,1176]" pageId="12" pageNumber="262">
|
||
The following key can be offered to separate all four presently known species of
|
||
<taxonomicName id="EADDB187121F1268595BFBBFFB0CFB56" box="[1058,1215,1151,1176]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FA91616121F1268595BFBBFFB0CFB56" box="[1058,1215,1151,1176]" italics="true" pageId="12" pageNumber="262">Hirudicryptus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65C7998F121F12685DEEFB07FA2FFA64" pageId="12" pageNumber="262" type="key">
|
||
<key id="594C9D35121F12685DEEFB07FA2FFA64" pageId="12" pageNumber="262">
|
||
<keyStep id="9629D2A1121F12685DEEFB07FA2FFB37" pageId="12" pageNumber="262">
|
||
<paragraph id="2D62CA04121F12685DEEFB07FA28FB12" blockId="12.[151,1436,1223,1450]" box="[151,1435,1223,1244]" pageId="12" pageNumber="262">
|
||
<keyLead id="962C6931121F12685DEEFB07FA28FB12" box="[151,1435,1223,1244]" pageId="12" pageNumber="262">1. Caudal margin of last tergum straight, devoid of lateral prongs.................................................. 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="2D62CA04121F12685DEEFB24FA2FFB37" blockId="12.[151,1436,1223,1450]" box="[151,1436,1252,1273]" pageId="12" pageNumber="262">
|
||
<keyLead id="962C6931121F12685DEEFB24FA2FFB37" box="[151,1436,1252,1273]" pageId="12" pageNumber="262">
|
||
- Caudal margin of last tergum clearly concave, flanked by small lateral prongs (
|
||
<figureCitation id="B5E6D681121F12685EAFFB24FBA1FB37" box="[982,1042,1252,1273]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="12" pageNumber="262">Figs 6</figureCitation>
|
||
C, 7C).......................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="9629D2A1121F12685DEEFAC2FA28FAFB" pageId="12" pageNumber="262">
|
||
<paragraph id="2D62CA04121F12685DEEFAC2FA29FAD9" blockId="12.[151,1436,1223,1450]" box="[151,1434,1282,1303]" pageId="12" pageNumber="262">
|
||
<keyLead id="962C6931121F12685DEEFAC2FA29FAD9" box="[151,1434,1282,1303]" pageId="12" pageNumber="262">
|
||
2. Midbody terga with 14–15 + 14–15 tubercles at caudal margin. Macaronesia...........................
|
||
<taxonomicName id="EADDB187121F12685876FAC2FA29FAD9" ID-CoL="3M4P2" box="[1295,1434,1282,1303]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis id="1FA91616121F12685876FAC2FA29FAD9" box="[1295,1434,1282,1303]" italics="true" pageId="12" pageNumber="262">H. canariensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="2D62CA04121F12685DEEFADFFA28FAFB" blockId="12.[151,1436,1223,1450]" box="[151,1435,1311,1333]" pageId="12" pageNumber="262">
|
||
<keyLead id="962C6931121F12685DEEFADFFA28FAFB" box="[151,1435,1311,1333]" pageId="12" pageNumber="262">
|
||
- Midbody terga with 9–11 + 9–11 tubercles at caudal margin.
|
||
<collectingCountry id="55CA8A94121F12685E53FADFFCD0FAFA" box="[810,867,1311,1332]" name="Nepal" pageId="12" pageNumber="262">Nepal</collectingCountry>
|
||
.............................
|
||
<taxonomicName id="EADDB187121F126859BEFAE0FA28FAFB" ID-CoL="3M4P3" box="[1223,1435,1311,1333]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="species" species="quintumelementum">
|
||
<emphasis id="1FA91616121F126859BEFAE0FA28FAFB" box="[1223,1435,1311,1333]" italics="true" pageId="12" pageNumber="262">H. quintumelementum</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="9629D2A1121F12685DEEFAFCFA2FFA64" pageId="12" pageNumber="262">
|
||
<paragraph id="2D62CA04121F12685DEEFAFCFA2FFAA1" blockId="12.[151,1436,1223,1450]" pageId="12" pageNumber="262">
|
||
<keyLead id="962C6931121F12685DEEFAFCFA2FFAA1" pageId="12" pageNumber="262">
|
||
3. Midbody terga with 6–8 + 6–8 tubercles at caudal margin Adults larger: length
|
||
<quantity id="EA2567E1121F12685968FAFCFB13FA9F" box="[1041,1184,1340,1361]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" metricValueMax="1.68" metricValueMin="1.05" pageId="12" pageNumber="262" unit="mm" value="13.65" valueMax="16.8" valueMin="10.5">10.5–16.8 mm</quantity>
|
||
(males), up to
|
||
<quantity id="EA2567E1121F1268583BFAFCFA28FA9F" box="[1346,1435,1340,1361]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="12" pageNumber="262" unit="mm" value="19.4">19.4 mm</quantity>
|
||
(females), width 1.2–2.0 or
|
||
<quantity id="EA2567E1121F12685C95FA9AFDD2FAA1" box="[492,609,1370,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="12" pageNumber="262" unit="mm" value="1.9" valueMax="2.5" valueMin="1.3">1.3–2.5 mm</quantity>
|
||
, respectively.
|
||
<collectingCountry id="55CA8A94121F12685F90FA9AFC81FAA1" box="[745,818,1370,1391]" name="Taiwan" pageId="12" pageNumber="262">Taiwan</collectingCountry>
|
||
.......................................
|
||
<taxonomicName id="EADDB187121F12685874FA9AFA2FFAA1" ID-CoL="3M4P4" box="[1293,1436,1370,1391]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="species" species="taiwanensis">
|
||
<emphasis id="1FA91616121F12685874FA9AFA2FFAA1" box="[1293,1436,1370,1391]" italics="true" pageId="12" pageNumber="262">H. taiwanensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="2D62CA04121F12685DEEFAB7FA2FFA64" blockId="12.[151,1436,1223,1450]" pageId="12" pageNumber="262">
|
||
<keyLead id="962C6931121F12685DEEFAB7FA2FFA64" pageId="12" pageNumber="262">
|
||
- Midbody terga with 9–11 + 9–11 tubercles at caudal margin (
|
||
<figureCitation id="B5E6D681121F12685E50FAB7FCD0FA42" box="[809,867,1399,1420]" captionStart="FIGURE 6" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[175,1420,210,1871]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 6. Hirudicryptus abchasicus sp. nov., male holotype (A – C) and female paratype (D, E). A, D, E: habitus, mainly sublateral, lateral and mainly ventral views, respectively. B: anterior part of body, subventral view. C: anterior and posterior parts of body, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/233475/files/figure.png" pageId="12" pageNumber="262">Fig. 6</figureCitation>
|
||
C). Adults smaller,
|
||
<quantity id="EA2567E1121F12685965FAB7FBCEFA42" box="[1052,1149,1399,1420]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.5" metricValueMin="7.0" pageId="12" pageNumber="262" unit="mm" value="7.75" valueMax="8.5" valueMin="7.0">7–8.5 mm</quantity>
|
||
long, 0.9–1.0 mm wide. Cau- casus..............................................................................
|
||
<taxonomicName id="EADDB187121F126859B8FA55FAFBFA64" ID-CoL="849D6" box="[1217,1352,1428,1450]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="12" pageNumber="262" phylum="Arthropoda" rank="species" species="abchasicus" status="sp. nov.">
|
||
<emphasis id="1FA91616121F126859B8FA55FAFBFA64" box="[1217,1352,1428,1450]" italics="true" pageId="12" pageNumber="262">H. abchasicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="049AAB6D121F12685836FA54FA2FFA64" box="[1359,1436,1428,1450]" pageId="12" pageNumber="262" rank="species">
|
||
<emphasis id="1FA91616121F12685836FA54FA2FFA64" bold="true" box="[1359,1436,1428,1450]" pageId="12" pageNumber="262">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |