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<mods:title id="948D997AC19A88EE1FDCD030D5BE5258">Review of the genus Singilis Rambur, 1837 of North Africa and Iberian Peninsula (Coleoptera: Carabidae: Lebiini)</mods:title>
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<treatment id="A55B87E1913FFFE625FDFE2B4E209C7C" ID-DOI="http://doi.org/10.5281/zenodo.6044469" ID-GBIF-Taxon="127694671" ID-Zenodo-Dep="6044469" LSID="urn:lsid:plazi:treatment:A55B87E1913FFFE625FDFE2B4E209C7C" httpUri="http://treatment.plazi.org/id/A55B87E1913FFFE625FDFE2B4E209C7C" lastPageNumber="271" pageId="7" pageNumber="271">
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<paragraph id="2D4D36F7913FFFE625FDFE2B4F9C9EDA" blockId="7.[151,896,439,466]" box="[151,896,439,466]" pageId="7" pageNumber="271">
<heading id="7605819B913FFFE625FDFE2B4F9C9EDA" bold="true" box="[151,896,439,466]" fontSize="11" level="1" pageId="7" pageNumber="271" reason="1">
<emphasis id="1F86EAE5913FFFE625FDFE2B4F9C9EDA" bold="true" box="[151,896,439,466]" pageId="7" pageNumber="271">
<taxonomicName id="EAF24D74913FFFE625FDFE2B4EEF9EDA" ID-CoL="8LY8N" authority="Escalera, 1914" authorityName="Escalera" authorityYear="1914" box="[151,755,439,466]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="subSpecies" species="soror" status="comb. nov." subGenus="Singilis" subSpecies="melillensis">
<emphasis id="1F86EAE5913FFFE625FDFE2B4E2B9ED9" bold="true" box="[151,567,439,465]" italics="true" pageId="7" pageNumber="271">Singilis (Singilis) soror melillensis</emphasis>
<bibRefCitation id="49634B06913FFFE62754FE244EEF9EDA" author="Escalera" box="[574,755,440,466]" pageId="7" pageNumber="271" refString="Escalera, M. M. de la (1914) Los Coleopteros de Marruecos. Trabajos del Museo de Ciencias Naturales, Zoology, 11, 1 - 553." type="journal article" year="1914">Escalera, 1914</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="04B5579E913FFFE62790FE244F9C9EDA" box="[762,896,440,466]" pageId="7" pageNumber="271">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="65E8657C913FFFE625FDFE624E899D3B" pageId="7" pageNumber="271" type="reference_group">
<paragraph id="2D4D36F7913FFFE625FDFE624E0B9D1C" blockId="7.[151,661,510,563]" box="[151,535,510,532]" pageId="7" pageNumber="271">
<treatmentCitationGroup id="0DE211D9913FFFE625FDFE624E0B9D1C" box="[151,535,510,532]" pageId="7" pageNumber="271">
<taxonomicName id="EAF24D74913FFFE625FDFE624E0B9D1C" authority="Escalera, 1914: 47" authorityName="Escalera" authorityPageNumber="47" authorityYear="1914" box="[151,535,510,532]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="species" species="melillensis">
<emphasis id="1F86EAE5913FFFE625FDFE624D499D1C" box="[151,341,510,532]" italics="true" pageId="7" pageNumber="271">Singilis melillensis</emphasis>
<treatmentCitation id="AC5310E6913FFFE62436FE624E0B9D1C" author="Escalera" box="[348,535,510,532]" page="47" pageId="7" pageNumber="271" year="1914">
<bibRefCitation id="49634B06913FFFE62436FE624DE89D1C" author="Escalera" box="[348,500,510,532]" pageId="7" pageNumber="271" refString="Escalera, M. M. de la (1914) Los Coleopteros de Marruecos. Trabajos del Museo de Ciencias Naturales, Zoology, 11, 1 - 553." type="journal article" year="1914">Escalera, 1914</bibRefCitation>
: 47
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D4D36F7913FFFE625FDFD814E899D3B" blockId="7.[151,661,510,563]" box="[151,661,541,563]" pageId="7" pageNumber="271">
<emphasis id="1F86EAE5913FFFE625FDFD814D609D3B" box="[151,380,541,563]" italics="true" pageId="7" pageNumber="271">
<taxonomicName id="EAF24D74913FFFE625FDFD814D499D3B" box="[151,341,541,563]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="species" species="melillensis">Singilis melillensis</taxonomicName>
ab.
</emphasis>
<taxonomicName id="EAF24D74913FFFE624E8FD814E039D3B" authority="Mateu" authorityName="Mateu" box="[386,543,541,563]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="variety" species="alternans" variety="unicolor">
<emphasis id="1F86EAE5913FFFE624E8FD814DC49D3B" box="[386,472,541,563]" italics="true" pageId="7" pageNumber="271">unicolor</emphasis>
Mateu
</taxonomicName>
[nom.nud.]
</paragraph>
</subSubSection>
<subSubSection id="65E8657C913FFFE625FDFDC348D19C58" pageId="7" pageNumber="271" type="materials_examined">
<paragraph id="2D4D36F7913FFFE625FDFDC348D19C58" blockId="7.[151,1437,607,885]" pageId="7" pageNumber="271">
<materialsCitation id="9D9A3CAA913FFFE625FDFDC34CE49D94" ID-GBIF-Occurrence="1427604093" collectingDate="1909-12" collectionCode="MNCN" collectorName="Arias" country="Morocco" location="Material" pageId="7" pageNumber="271" specimenCount="1" stateProvince="Melilla" typeStatus="holotype">
<emphasis id="1F86EAE5913FFFE625FDFDC34D9D9D70" bold="true" box="[151,385,607,632]" pageId="7" pageNumber="271">
<location id="282D602C913FFFE625FDFDC34CE39D70" LSID="urn:lsid:plazi:treatment:A55B87E1913FFFE625FDFE2B4E209C7C:282D602C913FFFE625FDFDC34CE39D70" box="[151,255,607,632]" country="Morocco" name="Material" pageId="7" pageNumber="271" stateProvince="Melilla">Material</location>
examined.
</emphasis>
<collectingCountry id="55E57667913FFFE624E3FDFC4E079D70" box="[393,539,608,632]" name="Morocco" pageId="7" pageNumber="271">MOROCCO</collectingCountry>
: “
<taxonomicName id="EAF24D74913FFFE6275AFDC34F9D9D70" authority="Esc. Tipo" authorityName="Esc. Tipo" box="[560,897,607,632]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="species" species="melillensis">Singilis melillensis Esc. Tipo</taxonomicName>
”, “
<collectingRegion id="EF36F815913FFFE626CBFDC34FEB9D70" box="[929,1015,607,632]" country="Spain" name="Melilla" pageId="7" pageNumber="271">Melilla</collectingRegion>
,
<collectingDate id="4908E9DF913FFFE62169FDFC48779D70" box="[1027,1131,608,632]" pageId="7" pageNumber="271" value="1909-12">
<date id="594C1037913FFFE62169FDFC48779D70" box="[1027,1131,608,632]" pageId="7" pageNumber="271" value="1909-12">XII.1909</date>
</collectingDate>
,
<collectorName id="80075321913FFFE62112FDC348AB9D70" box="[1144,1207,607,632]" pageId="7" pageNumber="271">Arias</collectorName>
” (
<typeStatus id="F2498855913FFFE621BAFDC349219D70" box="[1232,1341,607,632]" pageId="7" pageNumber="271" type="holotype">Holotype</typeStatus>
, female,
<collectionCode id="4BE3AE32913FFFE625FDFD184CEC9D94" box="[151,240,644,668]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="7" pageNumber="271">MNCN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9D9A3CAA913FFFE62469FD184EA59D94" ID-GBIF-Occurrence="1427604092" box="[259,697,644,669]" collectingDate="1909-12" collectionCode="MNCN" collectorName="Arias" country="Spain" location="Melilla" pageId="7" pageNumber="271" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="55E57667913FFFE62469FD184D4B9D95" box="[259,343,644,669]" name="Spain" pageId="7" pageNumber="271">Melilla</collectingCountry>
,
<date id="594C1037913FFFE6240BFD184DD59D94" box="[353,457,644,668]" pageId="7" pageNumber="271" value="1909-12">
<collectingDate id="4908E9DF913FFFE6240BFD184DD59D94" box="[353,457,644,668]" pageId="7" pageNumber="271" value="1909-12">XII.1909</collectingDate>
</date>
,
<collectorName id="80075321913FFFE624BEFD184E0D9D95" box="[468,529,644,669]" pageId="7" pageNumber="271">Arias</collectorName>
(
<specimenCount id="3BF4FD7E913FFFE62749FD184E509D94" box="[547,588,644,668]" pageId="7" pageNumber="271" type="male">1 ♂</specimenCount>
,
<collectionCode id="4BE3AE32913FFFE62732FD184EAD9D94" box="[600,689,644,668]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="7" pageNumber="271">MNCN</collectionCode>
)
</materialsCitation>
; Taurirt, B Sicar, Marruecos, A. Cobos leg. (14 ex., ISNB);
<materialsCitation id="9D9A3CAA913FFFE62039FD194F329DC8" ID-GBIF-Occurrence="1427604096" collectionCode="ISNB" country="Melilla, Marruecos" location="Pardo Alcaide" municipality="Monte Arruit" pageId="7" pageNumber="271" specimenCount="1" typeStatus="holotype">
<collectingMunicipality id="CD29AC8D913FFFE62039FD194CC29DC8" pageId="7" pageNumber="271">Monte Arruit</collectingMunicipality>
,
<collectingCountry id="55E57667913FFFE62584FD3B4DD09DB7" box="[238,460,679,704]" isEnumeration="true" pageId="7" pageNumber="271">Melilla, Marruecos</collectingCountry>
,
<location id="282D602C913FFFE624B6FD3B4E639DC8" LSID="urn:lsid:plazi:treatment:A55B87E1913FFFE625FDFE2B4E209C7C:282D602C913FFFE624B6FD3B4E639DC8" box="[476,639,679,704]" country="Melilla, Marruecos" municipality="Monte Arruit" name="Pardo Alcaide" pageId="7" pageNumber="271">Pardo Alcaide</location>
(1 ex.,
<collectionCode id="4BE3AE32913FFFE62788FD344F3A9DC8" box="[738,806,680,704]" pageId="7" pageNumber="271">ISNB</collectionCode>
)
</materialsCitation>
; “
<emphasis id="1F86EAE5913FFFE62621FD3448A99DC8" box="[843,1205,679,704]" italics="true" pageId="7" pageNumber="271">
<taxonomicName id="EAF24D74913FFFE62621FD344FFF9DC8" box="[843,995,679,704]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="species" species="melillensis">S. melillensis</taxonomicName>
ab.
<taxonomicName id="EAF24D74913FFFE62177FD3B48609DC8" box="[1053,1148,679,704]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="variety" species="alternans" variety="unicolor">unicolor</taxonomicName>
nov.
</emphasis>
”, “Taurirt, B Sicar, Marruecos, A. Cobos leg.” (Holotype, male, ISNB);
<materialsCitation id="9D9A3CAA913FFFE6278CFD5048D19C58" ID-GBIF-Occurrence="1427604097" collectingDate="1930-06" collectionCode="MNCN" collectorName="Axdir, B. &amp; C. Bolivar &amp; Therefore I" country="Spain" location="Rif" pageId="7" pageNumber="271" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectorName id="80075321913FFFE6278CFD504F519DEC" box="[742,845,716,741]" pageId="7" pageNumber="271">Axdir, B.</collectorName>
Urriaguel,
<location id="282D602C913FFFE626A4FD504FE99DED" LSID="urn:lsid:plazi:treatment:A55B87E1913FFFE625FDFE2B4E209C7C:282D602C913FFFE626A4FD504FE99DED" box="[974,1013,716,741]" country="Spain" name="Rif" pageId="7" pageNumber="271">Rif</location>
,
<collectingDate id="4908E9DF913FFFE6216AFD50487C9DEC" box="[1024,1120,716,740]" pageId="7" pageNumber="271" value="1930-06">
<date id="594C1037913FFFE6216AFD50487C9DEC" box="[1024,1120,716,740]" pageId="7" pageNumber="271" value="1930-06">VI.1930</date>
</collectingDate>
.
<collectorName id="80075321913FFFE62100FD5048C49DED" box="[1130,1240,716,741]" pageId="7" pageNumber="271">C.Bolivar</collectorName>
(
<specimenCount id="3BF4FD7E913FFFE62180FD50490E9DEC" box="[1258,1298,716,740]" pageId="7" pageNumber="271" type="female">1 ♀</specimenCount>
,
<collectionCode id="4BE3AE32913FFFE6204AFD5049659DEC" box="[1312,1401,716,740]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="7" pageNumber="271">MNCN</collectionCode>
).
<emphasis id="1F86EAE5913FFFE625FDFD734D039C00" bold="true" box="[151,287,751,776]" pageId="7" pageNumber="271">Comments.</emphasis>
This taxon was described as a distinct species from
<collectingCountry id="55E57667913FFFE62619FD734FD89C00" box="[883,964,751,776]" name="Spain" pageId="7" pageNumber="271">Melilla</collectingCountry>
on the basis of specimens with very long pubescence (
<figureCitation id="B5C92A72913FFFE62444FC884D6F9C24" box="[302,371,788,813]" captionStart="FIGURES 1 11" captionStartId="4.[151,264,1821,1844]" captionTargetBox="[167,1431,212,1789]" captionTargetId="figure@4.[151,1436,195,1797]" captionTargetPageId="4" captionText="FIGURES 1 11. Habitus of Singilis bicolor - group. 1 — S. bicolor; 2 — S. alternans; 3 — S. soror melillensis (Taurirt); 4 — S. soror riffensis syn. n., Paratype (Isaguen); 5 6 — S. soror riffensis syn. n. (Ketama); 7 — S. soror s. str. (15 km SE Berkane); 8 — S. soror s. str. = S. pardoi syn. n., Paratype (El Haraig); 9 — S. soror s. str. (Oudjda); 10 — S. soror oranensis ssp. n. Holotype; 11 — S. soror oranensis Paratype." httpUri="https://zenodo.org/record/291795/files/figure.png" pageId="7" pageNumber="271">Fig 3</figureCitation>
). No other differential characters were found between this taxon and other populations.
<collectorName id="80075321913FFFE625FDFCAB4D099C47" box="[151,277,823,848]" pageId="7" pageNumber="271">Therefore I</collectorName>
propose to consider
<taxonomicName id="EAF24D74913FFFE62495FCA44E8F9C58" box="[511,659,823,848]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="species" species="melillensis">
<emphasis id="1F86EAE5913FFFE62495FCA44E8F9C58" box="[511,659,823,848]" italics="true" pageId="7" pageNumber="271">S. melillensis</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="EAF24D74913FFFE62602FCA44FA29C58" box="[872,958,824,848]" class="Insecta" family="Carabidae" genus="Singilis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="271" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="1F86EAE5913FFFE62602FCA44FA29C58" box="[872,958,824,848]" italics="true" pageId="7" pageNumber="271">S. soror</emphasis>
</taxonomicName>
. Aedeagus as in fig. 18.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="65E8657C913FFFE625ADFCC04E209C7C" box="[199,572,860,885]" pageId="7" pageNumber="271" type="distribution">
<paragraph id="2D4D36F7913FFFE625ADFCC04E209C7C" blockId="7.[151,1437,607,885]" box="[199,572,860,885]" pageId="7" pageNumber="271">
<emphasis id="1F86EAE5913FFFE625ADFCC04D7C9C7D" bold="true" box="[199,352,860,885]" pageId="7" pageNumber="271">Distribution.</emphasis>
<collectingCountry id="55E57667913FFFE6240DFCC14DD09C7C" box="[359,460,861,884]" name="Morocco" pageId="7" pageNumber="271">Morocco</collectingCountry>
(
<figureCitation id="B5C92A72913FFFE624B6FCC04E339C7C" box="[476,559,860,885]" captionStart="FIGURE 12" captionStartId="5.[151,250,1108,1130]" captionTargetBox="[151,1436,193,1087]" captionTargetId="figure@5.[151,1436,193,1087]" captionTargetPageId="5" captionText="FIGURE 12. Distribution map of Singilis bicolor - group." httpUri="https://zenodo.org/record/291384/files/figure.png" pageId="7" pageNumber="271">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>