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<document id="D7F2888FBA28CF2FDB5BB614D5F3992E" ID-CLB-Dataset="26683" ID-DOI="http://dx.doi.org/10.3897/evolsyst.1.14847" ID-GBIF-Dataset="ae0cbe4e-e7d0-477f-a4a8-1bc5d289c549" ID-Pensoft-Pub="2535-0730-1-77" ID-ZooBank="AC4ED29D692A4F16B30058F7AE2BF008" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="2535-0730-1-77" ModsDocOrigin="Evolutionary Systematics 1 (1)" ModsDocTitle="Portacosa, a new genus for the south-east Australian Grey Wolf Spider (Araneae, Lycosidae, Lycosinae)" checkinTime="1558621803442" checkinUser="pensoft" docAuthor="Framenau, Volker W." docDate="2017" docId="9E65FBC2795438944EB329191F6BFDBC" docLanguage="en" docName="EvolutSyst 1(1): 77-86" docOrigin="Evolutionary Systematics 1 (1)" docSource="http://dx.doi.org/10.3897/evolsyst.1.14847" docTitle="Portacosa Framenau, 2017, gen. n." docType="treatment" docUuid="8CDB0C42-1A51-455F-B368-5BD6ED3C094B" docUuidSource="ZooBank" docVersion="6" lastPageNumber="77" masterDocId="FFEBFFB2FFDFFFE5FFC8BD4FF16C1659" masterDocTitle="Portacosa, a new genus for the south-east Australian Grey Wolf Spider (Araneae, Lycosidae, Lycosinae)" masterLastPageNumber="86" masterPageNumber="77" pageNumber="77" updateTime="1701379450559" updateUser="ExternalLinkService">
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<mods:title id="522CCE78E4B89B68BB28704FFDEDCD25">Portacosa, a new genus for the south-east Australian Grey Wolf Spider (Araneae, Lycosidae, Lycosinae)</mods:title>
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<mods:namePart id="3415E8B818E3070F3A09E097510847F6">Framenau, Volker W.</mods:namePart>
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<treatment id="9E65FBC2795438944EB329191F6BFDBC" ID-GBIF-Taxon="157252056" LSID="urn:lsid:zoobank.org:act:8CDB0C42-1A51-455F-B368-5BD6ED3C094B" httpUri="http://treatment.plazi.org/id/9E65FBC2795438944EB329191F6BFDBC" lastPageId="1" lastPageNumber="77" pageId="0" pageNumber="77">
<subSubSection id="3BF372AC84C2098079C51FAB1C043A02" pageId="0" pageNumber="77" type="nomenclature">
<paragraph id="82E5D16C39F1DAC0A3FBFDA610289888" pageId="0" pageNumber="77">
<taxonomicName id="2BE5046585A9170CD6AF46C568A484C1" ID-CoL="63PNH" LSID="http://zoobank.org/8CDB0C42-1A51-455F-B368-5BD6ED3C094B" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
<taxonomicNameLabel id="7002EACA05E58C42CB2C8F9FF8AFA3D3" pageId="0" pageNumber="77">gen. n.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="EC93531C0113D8FF041BD517BA0E1D00" pageId="0" pageNumber="77" type="type species">
<paragraph id="5DCC0F26F63A9392CCF94F723633298B" pageId="0" pageNumber="77">Type species.</paragraph>
<paragraph id="0588E27F8B8D509B49027584DC8CA954" pageId="0" pageNumber="77">
<taxonomicName id="3421142F17F5921288FA65F2ABB42661" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa cinerea" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="cinerea">Portacosa cinerea</taxonomicName>
sp. n., designated here.
</paragraph>
</subSubSection>
<subSubSection id="27EA6ABA88B3CD150A29CC577F534A07" pageId="0" pageNumber="77" type="diagnosis">
<paragraph id="81D77FE87C5B01B43F92F757A394A0B9" pageId="0" pageNumber="77">Diagnosis.</paragraph>
<paragraph id="813559DA6A30D19652CE8908EEFB6170" pageId="0" pageNumber="77">
Somatic morphology, in particular the lack of a distinct colour pattern on carapace and abdomen (in particular in live spiders, Fig. 1
<normalizedToken id="EFEB82FBB9D1F2ADE7ED195FA2446065" originalValue="A–D">A-D</normalizedToken>
), places
<taxonomicName id="DF0E7512293180E3923E2339978E5B95" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. in close affinity with
<taxonomicName id="218B51138A14DBD10EDB63784B8EEB35" class="Arachnida" family="Lycosidae" genus="Hoggicosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoggicosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Hoggicosa</taxonomicName>
Roewer; however, males of
<taxonomicName id="8383BB4B52E014D19F579C7A95252557" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. lack the apical, dorsally bent setae on the cymbium tip and the median septum of the female epigyne is narrower anteriorly than the with of the posterior transverse part (see
<bibRefCitation id="BBC1D02212F0C239A383503B58D906B9" author="Langlands, PR" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="81" pagination="83 - 123" title="Systematic revision of Hoggicosa Roewer, 1960, the Australian ' bicolor' group of wolf spiders (Araneae: Lycosidae)." url="https://www.doi.org/10.1111/j.1096-3642.2009.00545.x" volume="158" year="2010">Langlands and Framenau 2010</bibRefCitation>
).
</paragraph>
<paragraph id="64E2B7D4F389C931ED304B23B1E072F7" pageId="0" pageNumber="77">
A putative apomorphy of the genus is the shape of the tegular apophysis. Unlike any other member of the family
<taxonomicName id="E3ADDF2231CC5423CE5845AAB20C360E" lsidName="" pageId="0" pageNumber="77" rank="subfamily" subfamily="Lycosinae">Lycosinae</taxonomicName>
in Australia, the ridge of the tegular apophysis (connecting its apical point and ventral process; see Fig. 3C, D) is very sharp and situated more towards the retrolateral edge of the tegular apophysis, whereas it reaches from the apical point to a more central ventral process in other
<taxonomicName id="2A5D3E1588D0BF207044E4FF57E98BFA" lsidName="" pageId="0" pageNumber="77" rank="subfamily" subfamily="Lycosinae">Lycosinae</taxonomicName>
in Australia. This corresponds to a continuous edge of the anterior hood of the epigyne with the median septum (Fig. 4A, C), whilst this edge is often interrupted in other
<taxonomicName id="D498B11A19D90BF8AE72C91ABF4C200D" lsidName="" pageId="0" pageNumber="77" rank="subfamily" subfamily="Lycosinae">Lycosinae</taxonomicName>
in Australia.
</paragraph>
<caption id="09E3E7C162EC4C60271805DA79AE5AB9" pageId="0" pageNumber="77">
<paragraph id="DECD6345A3DFC4F641B7CCB999B4A5E9" pageId="0" pageNumber="77">
Figure 1. Live images and habitat of
<taxonomicName id="EE8D8F25CEE074B582BEA378510A8C52" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa cinerea" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="cinerea">Portacosa cinerea</taxonomicName>
gen. n. and sp. n. A, male holotype (WAM T68032) from near Apex Park, Mildura, Victoria. B, male, same locality. C, female (WAM T56062) from Specimen Hill Bushland Reserve, Bendigo, Victoria, D, female from Red Hill, Canberra, Australian Capital Territory (WAM T67906); E, open
<taxonomicName id="8C5987A684F7C8CF426A301B01E64C32" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus camaldulensis" order="Myrtales" pageId="0" pageNumber="77" phylum="Tracheophyta" rank="species" species="camaldulensis">Eucalyptus camaldulensis</taxonomicName>
riparian woodland, near Apex Park, Mildura, Victoria, where the holotype male (WAM T68032) was found. F, road embankment in Red Hill, Canberra, habitat of female WAM T67906. G, closed burrow of holotype male (WAM T68032). H, open burrow of female WAM T67906 from Red Hill, Canberra.
</paragraph>
</caption>
<paragraph id="825790215ACD190AEEDD4F55AEF79273" pageId="0" pageNumber="77">
<taxonomicName id="C51E7C7D8C260163B84C97ACDA229C69" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. lacks any of the proposed synapomorphies of other Australian members of the
<taxonomicName id="43245F372303095A5076F2DAB0A01A80" lsidName="" pageId="0" pageNumber="77" rank="subfamily" subfamily="Lycosinae">Lycosinae</taxonomicName>
, i.e. the tegular apophysis is not retrolaterally incised as in
<taxonomicName id="D0C84D74FCB140F8C7E44191BAE9AB0A" class="Arachnida" family="Lycosidae" genus="Venator" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Venator" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Venator</taxonomicName>
Hogg, 1900 (
<bibRefCitation id="6CEF46E2D36687457E6EC251969949E2" author="Framenau, VW" journalOrPublisher="Zootaxa" pageId="4" pageNumber="81" pagination="541 - 555" title="Review of the Australian wolf spider genus Venator (Araneae, Lycosidae)." url="https://doi.org/10.11646/zootaxa.4013.4.5" volume="4013" year="2015">Framenau 2015</bibRefCitation>
), the carapace lacks a Union-Jack pattern as in
<taxonomicName id="8D29B7B2E645836D3BD19760914EFC77" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanicosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Tasmanicosa</taxonomicName>
Roewer, 1959 (
<bibRefCitation id="4E58639965D6A8FEBA86E7D7EA84F7CC" author="Framenau, VW" journalOrPublisher="Zootaxa" pageId="4" pageNumber="81" pagination="1 - 82" title="Revision of the Australian Union-Jack wolf spiders, genus Tasmanicosa (Araneae, Lycosidae, Lycosinae)." url="https://doi.org/10.11646/zootaxa.4213.1.1" volume="4213" year="2016">Framenau and Baehr 2016</bibRefCitation>
), the pedipalp lacks a large patch of apical setae as in
<taxonomicName id="AD23E4057EC73B3827E7D0153E403D2F" class="Arachnida" family="Lycosidae" genus="Knoelle" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Knoelle" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Knoelle</taxonomicName>
Framenau, 2006 (
<bibRefCitation id="8662C3E3486C37C0268654D13F59BE17" author="Framenau, VW" journalOrPublisher="Zootaxa" pageId="4" pageNumber="81" pagination="55 - 67" title="Knoelle, a new monotypic wolf spider genus from Australia (Araneae: Lycosidae)." volume="1281" year="2006 a">Framenau 2006a</bibRefCitation>
), the tegular apophysis is not elongated and spiders do not show turret-building behaviour as in
<taxonomicName id="CB13D947D321BA16426CF89FCD161B99" class="Arachnida" family="Lycosidae" genus="Dingosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dingosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Dingosa</taxonomicName>
(
<bibRefCitation id="0C5BD9F79639B1129369E44551002827" author="Framenau, VW" journalOrPublisher="Journal of Natural History" pageId="4" pageNumber="81" pagination="1603 - 1629" title="Revision of the Australian wolf spider genus Dingosa Roewer, 1955 (Araneae, Lycosidae, Lycosinae)." url="https://www.doi.org/10.1080/00222930701475717" volume="41" year="2007">Framenau and Baehr 2007</bibRefCitation>
), the carapace and abdomen are not dorsoventrally flattened as in
<taxonomicName id="EA54BBDA6B58B2C77A93DC2132FD51D0" class="Arachnida" family="Lycosidae" genus="Tapetosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tapetosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Tapetosa</taxonomicName>
Framenau, Main, Waldock &amp; Harvey, 2009 (
<bibRefCitation id="7D3CC1D3BE1FE3CEF6766B7A03DF1681" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="4" pageNumber="81" pagination="309 - 314" title="Tapetosa, a new monotypic wolf spider genus from Western Australia (Araneae: Lycosidae: Lycosinae)." url="https://www.doi.org/10.18195/issn.0312-3162.25(3).2009.309-314" volume="25" year="2009">Framenau et al. 2009</bibRefCitation>
), the tegular apophysis lacks apical serrations as in
<taxonomicName id="A369B7E385AD9943E277D3A396C60541" class="Arachnida" family="Lycosidae" genus="Costacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Costacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Costacosa</taxonomicName>
Framenau &amp; Leung, 2013 (
<bibRefCitation id="25894BE8BF28005D2FF041269CDFD55A" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum, Supplement" pageId="4" pageNumber="81" pagination="173 - 184" title="Costacosa, a new genus of wolf spider (Araneae, Lycosidae) from coastal north-west Western Australia." url="https://www.doi.org/10.18195/issn.0313-122x.83.2013.173-184" volume="83" year="2013">Framenau and Leung 2013</bibRefCitation>
), the abdomen is not dark with transverse light bands and spiders do not display turret-building behaviour as in
<taxonomicName id="3341D6B2D20DED764AA7337D4F826B3B" class="Arachnida" family="Lycosidae" genus="Mainosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mainosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Mainosa</taxonomicName>
Framenau, 2006 (
<bibRefCitation id="084B9D029F2FDD59DD5F4C4EAA450225" author="Framenau, VW" journalOrPublisher="Journal of Arachnology" pageId="4" pageNumber="81" pagination="206 - 213" title="Mainosa, a new genus for the Australian ' shuttlecock wolf spider' (Araneae, Lycosidae)." url="https://www.doi.org/10.1636/H04-90.1" volume="34" year="2006 b">Framenau 2006b</bibRefCitation>
), and male pedipalp cymbia do not have a compound apical hook as in
<taxonomicName id="F48F1E65A4D5BF62F64E957770F8E174" class="Arachnida" family="Lycosidae" genus="Venatrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Venatrix" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Venatrix</taxonomicName>
Roewer, 1960 and
<taxonomicName id="EA21425B92DB3FB51E6C59DCF459EF0D" class="Arachnida" family="Lycosidae" genus="Tuberculosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tuberculosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Tuberculosa</taxonomicName>
Framenau &amp; Yoo, 2006 (
<bibRefCitation id="17A36FAD0A139240B2AAACB374E23182" author="Framenau, VW" journalOrPublisher="Invertebrate Taxonomy" pageId="4" pageNumber="81" pagination="927 - 970" title="Revision of the wolf spider genus Venatrix Roewer (Araneae: Lycosidae)." url="https://www.doi.org/10.1071/IT01008" volume="15" year="2001">Framenau and Vink 2001</bibRefCitation>
;
<bibRefCitation id="A760D8EA8DF135467354B7EC0CE77364" author="Framenau, VW" journalOrPublisher="Invertebrate Systematics" pageId="4" pageNumber="81" pagination="185 - 202" title="Systematics of the new Australian wolf spider genus Tuberculosa (Araneae, Lycosidae)." url="https://doi.org/10.1071/IS05036" volume="20" year="2006">Framenau and Yoo 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="41A52E15D12F4FECBE9A1FA7102AB63C" pageId="0" pageNumber="77" type="etymology">
<paragraph id="A0CE31F1D4D083CCE06CC32F87DC28E7" pageId="0" pageNumber="77">Etymology.</paragraph>
<paragraph id="5C48A6E132FAD0D11BE755C25493580A" pageId="0" pageNumber="77">
The genus-group name is a composite noun derived from the Latin word portus - door, referring to the trapdoor-building behaviour of the type species and -cosa, a generic ending used for genera in the family
<taxonomicName id="74B549BF6532B4B117F62F70A29A3A38" family="Lycosidae" lsidName="" pageId="0" pageNumber="77" rank="family">Lycosidae</taxonomicName>
. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="2650C65E593BD6829E23E7848CFECEC5" pageId="0" pageNumber="77" type="description">
<paragraph id="F101EC518D56EDF2328E1DA7B0DBE5FC" pageId="0" pageNumber="77">Description.</paragraph>
<paragraph id="8C64EA8E99B3A32D8F54C5E79A5EBCB4" pageId="0" pageNumber="77">Large wolf spiders (TL 10.5-25.0 mm). Males slightly smaller than females. Carapace longer than wide, dorsal profile straight in lateral view. Carapace colouration brown with indistinct darker radial pattern, covered by grey pubescence that is denser between eyes, in particular in males. Abdomen dorsally with indistinct median chevron-pattern, which is less distinct in females and covered with dense grey pubescence, ventrally uniformly yellow-brown. AME larger than ALE, row of AE slightly procurved and narrower than row of PME. Chelicerae with three promarginal teeth with the median largest and three large, equally-sized retromarginal teeth. Leg formula IV&gt; I&gt; II&gt; III. Cymbium tip with approximately 20 straight spines (Fig. 3A, B). Tegulum of male pedipalp undivided (Fig, 3F); tegular apophysis with distinct ventral spur situated towards the retrolateral edge of the tegular apophysis, connected to apical point of tegular apophysis by sharp ridge (Fig. 3C, D). Embolus originating prolaterally on palea and curving ventrally around it, long and slender. Terminal apophysis broad and apically straight, pars pendula sickle-shaped (Figs 3E). Female epigyne with inverted T-shaped median septum, slightly longer than wide, anterior hoods distinct, connected continuously to medium septum (Figs 4A, C); spermathecal heads oval; spermathecal stalks S-shaped (Fig. 4B).</paragraph>
</subSubSection>
<subSubSection id="8582C059A00027DA5B5ECC994EC0A38B" lastPageId="1" lastPageNumber="78" pageId="0" pageNumber="77" type="composition">
<paragraph id="ACC8855A28318F3E22E91D8DE6F2855E" pageId="0" pageNumber="77">Composition.</paragraph>
<paragraph id="9690DCC0B5E41EC65A0F2ABC8CA6B9F7" pageId="0" pageNumber="77">
<taxonomicName id="A4F1B6B8C643FE135B0AB7C5FA3B16C6" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. currently includes only a single species,
<taxonomicName id="9F7A091E79991B1A616FC70AE66DD215" lsidName="P. cinerea" pageId="0" pageNumber="77" rank="species" species="cinerea">P. cinerea</taxonomicName>
gen. n. and sp. n.
</paragraph>
<paragraph id="A94BDE2A6AEC6078E520CB5338EB3859" lastPageId="1" lastPageNumber="78" pageId="0" pageNumber="77">
Systematics.
<taxonomicName id="A4183C04AF7F90242D7A80C46F6A94A9" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. is a member of the subfamily
<taxonomicName id="9D01D2674D94D05AC5838DD0D6E29E47" lsidName="" pageId="0" pageNumber="77" rank="subfamily" subfamily="Lycosinae">Lycosinae</taxonomicName>
Sundevall, 1833 based on the transverse orientation of the tegular apophysis that has a dorsal channel to guide the embolus (
<bibRefCitation id="BC057BC94B4EC4A0452A6DCE5673C676" pageId="0" pageNumber="77">Dondale 1986</bibRefCitation>
). However, sistergroup relationships remain unclear.
<taxonomicName id="0FCD568552B75524F5E26926F02299AF" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa cinerea" order="Araneae" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="cinerea">Portacosa cinerea</taxonomicName>
gen. n. and sp. n. was included in a world-wide multigene molecular analysis of the
<taxonomicName id="A592FCC92319A6469A35C2C3FDDB79B7" family="Lycosidae" lsidName="" pageId="0" pageNumber="77" rank="family">Lycosidae</taxonomicName>
as 'New Genus 6
<pageBreakToken id="8BB6B46AA651E36524176BE7B4969963" pageId="1" pageNumber="78" start="start">sp</pageBreakToken>
.' (
<bibRefCitation id="1771B4A23A92AFDEB77EF7F3D6D139D8" author="Murphy, NP" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="4" pageNumber="81" pagination="583 - 602" title="Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour." url="https://doi.org/10.1016/j.ympev.2005.09.004" volume="38" year="2006">Murphy et al. 2006</bibRefCitation>
). Maximum parsimony analyses placed
<taxonomicName id="7D406D44D6D3D185DB279109330218C4" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. as sister to the South American
<taxonomicName id="256A39734BE5ACCE9DBC30AE025281CA" class="Arachnida" family="Lycosidae" genus="Pavocosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pavocosa gallopavo" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="gallopavo">Pavocosa gallopavo</taxonomicName>
(
<normalizedToken id="C695D6AD69A4E5B718B7415B6BD6AD6E" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1941), both of which combined represented a sister taxon to a clade including
<taxonomicName id="FA161394A3E0423B726E570E210EE171" class="Arachnida" family="Lycosidae" genus="Hoggicosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoggicosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Hoggicosa</taxonomicName>
,
<taxonomicName id="9B39DCD6E8CF186593AAFAA83D8D1013" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Hogna</taxonomicName>
and
<taxonomicName id="0DFD95EDCB013E099A9726371AA0F852" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanicosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Tasmanicosa</taxonomicName>
. In contrast, Bayesian analyses placed
<taxonomicName id="A39E275DA37145E143C4E5D13F9B732C" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. in a poorly resolved polytomy including the above Australian genera as sister to
<taxonomicName id="5FB08F4536802E602DA343F62CE7DC8C" class="Arachnida" family="Lycosidae" genus="Pavocosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pavocosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Pavocosa</taxonomicName>
Roewer, 1960. Similarly, a morphological phylogenetic analysis of largely Australian
<taxonomicName id="E7295079499C686CDED2C1E93CEE7746" lsidName="" pageId="1" pageNumber="78" rank="subfamily" subfamily="Lycosinae">Lycosinae</taxonomicName>
did not resolve sistergroup relationships of
<taxonomicName id="BC6A769C55E9C314B965050A42035613" class="Arachnida" family="Lycosidae" genus="Portacosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Portacosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Portacosa</taxonomicName>
gen. n. (included in the analysis as 'Grey Wolf
<normalizedToken id="1283D2B656D4661949C93A8BC8CA37A8" originalValue="Spider’">Spider'</normalizedToken>
); however, that analysis was mainly focused on resolving relationships within
<taxonomicName id="484CAF38D7EDBC525B313932614D4986" class="Arachnida" family="Lycosidae" genus="Hoggicosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoggicosa" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Hoggicosa</taxonomicName>
(
<bibRefCitation id="3027F7DFBC3BB6BCEB15A80BAAB0530B" author="Langlands, PR" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="81" pagination="83 - 123" title="Systematic revision of Hoggicosa Roewer, 1960, the Australian ' bicolor' group of wolf spiders (Araneae: Lycosidae)." url="https://www.doi.org/10.1111/j.1096-3642.2009.00545.x" volume="158" year="2010">Langlands and Framenau 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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