treatments-xml/data/98/D7/EA/98D7EA1C817A580F9B767EC3E73A8F9A.xml

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<document ID-CLB-Dataset="89309" ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.82461" ID-GBIF-Dataset="92e148c9-6ca3-489d-8b0b-7bca978ee901" ID-PMC="PMC9848772" ID-Pensoft-Pub="1313-2970-1110-1" ID-Pensoft-UUID="88D66028C7A752528DD9138A2105F895" ID-PubMed="36761453" ID-ZooBank="01302157EE404B9999DD3EC3377D929C" ModsDocID="1313-2970-1110-1" checkinTime="1657006639549" checkinUser="pensoft" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Lin, Aung &amp; Panha, Somsak" docDate="2022" docId="98D7EA1C817A580F9B767EC3E73A8F9A" docLanguage="en" docName="ZooKeys 1110: 1-37" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.82461" docTitle="Burmochlamys Pholyotha &amp; Panha 2022, gen. nov." docType="treatment" docUuid="060B5C90-76D8-44D5-96CC-924066E94F59" docUuidSource="ZooBank" docVersion="5" id="88D66028C7A752528DD9138A2105F895" lastPageNumber="1" masterDocId="88D66028C7A752528DD9138A2105F895" masterDocTitle="Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="1" updateTime="1732937909010" updateUser="ExternalLinkService">
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<mods:titleInfo id="E0AEE1AF6B669C525B4F7D4D2E563C21">
<mods:title id="EAD3D9CDB3368EC348CC260E5BE72BEA">Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae)</mods:title>
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<mods:name id="60EF97C383DB7ACA5385EAC2D8BEA3E6" type="personal">
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<mods:namePart id="BD0116EE9B254263D2221C535D19E83C">Pholyotha, Arthit</mods:namePart>
<mods:affiliation id="126974EFE0B57C96B5882F69B207A1F6">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
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<mods:namePart id="B17FBAF3F18BDC631645FBD189FB8D5F">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="B9D411591403897B0906BA136A458F22">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
<mods:name id="989C06F66B301D9E421F2088F077F3EB" type="personal">
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<mods:namePart id="E3642C2F6D7B527958FA1C8572339321">Lin, Aung</mods:namePart>
<mods:affiliation id="E072028FB22B1A46FB13B331CAD0EA4C">Fauna and Flora International, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
</mods:name>
<mods:name id="4887A0CA4BB0928E91FDF627C2E9EC09" type="personal">
<mods:role id="8F9C19DD7AA846A529095818F799521C">
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<mods:namePart id="56471CC473F72DEB157355A1E670BBF7">Panha, Somsak</mods:namePart>
<mods:affiliation id="BBE625CD97DAF45548784D134298F4BC">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok, Thailand</mods:affiliation>
<mods:nameIdentifier id="0083EC1C75522E9B924B8B7E16264859" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="2C4C95BB6CC966CBA2EAF0EA92175747">text</mods:typeOfResource>
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<mods:title id="6F5E0CE6B96D8432CA8EACF7AD15E0A4">ZooKeys</mods:title>
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<mods:part id="F7E7D8EA5DE3E5A04324134D278F6179">
<mods:date id="1BB53C2EDAC45B8C6F5DE4AE07D44FBA">2022</mods:date>
<mods:detail id="42F744043BB2A1EB13AFE3C50F404BBD" type="pubDate">
<mods:number id="6E7DB097D722D78570F805CECB84B1AC">2022-07-04</mods:number>
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<mods:detail id="74DD7B42FC3E333319E4D0090465CB34" type="volume">
<mods:number id="C7A3C01F714AA0F1DD8E096649EF7929">1110</mods:number>
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<mods:start id="1FF798D88D1C1418E6D90C583A48CAB4">1</mods:start>
<mods:end id="4A7BCBBAAF50BA547F111249ED785070">37</mods:end>
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<mods:url id="E62931AF7F1A243BBDD72DFF7475EF75">http://dx.doi.org/10.3897/zookeys.1110.82461</mods:url>
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<mods:classification id="E9731D9D6A6646AE3612BB1C216DC48C">journal article</mods:classification>
<mods:identifier id="B2995F40DCAAA82BA7487B1BC1FF7D11" type="DOI">http://dx.doi.org/10.3897/zookeys.1110.82461</mods:identifier>
<mods:identifier id="AF6B37840AFC045F69EDE50CAB32AB93" type="Pensoft-Pub">1313-2970-1110-1</mods:identifier>
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<treatment id="98D7EA1C817A580F9B767EC3E73A8F9A" ID-GBIF-Taxon="196387952" LSID="urn:lsid:zoobank.org:act:060B5C90-76D8-44D5-96CC-924066E94F59" httpUri="http://treatment.plazi.org/id/98D7EA1C817A580F9B767EC3E73A8F9A" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Helicarionidae" scope_subFamily="Durgellinae" scope_superOrder="Eupulmonata">
<subSubSection id="A6823BE1A618BD67FCA1E68D11D3F333" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="04B606B16FF91C68CC287FD012C7746D" pageId="0" pageNumber="1">
Genus
<taxonomicName id="8641A899A359809E3A09FE5A33000625" LSID="https://zoobank.org/060B5C90-76D8-44D5-96CC-924066E94F59" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus" status="gen. nov.">Burmochlamys Pholyotha &amp; Panha</taxonomicName>
<taxonomicNameLabel id="D8660E8F82C03BEF6A240971F1D145DA" pageId="0" pageNumber="1">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F906FA37B88EE838875AB6A020399CB0" pageId="0" pageNumber="1" type="type species">
<paragraph id="889FFDC59945C057BC92802E7A493462" pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph id="10A65BB8F1C212A63AEC062D848699FA" pageId="0" pageNumber="1">
<taxonomicName id="DB52A5E48A42B73629642A17C811E457" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys fasciola" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="150DCF0FF32CCD7E1B82F826721B5431" italics="true" pageId="0" pageNumber="1">Burmochlamys fasciola</emphasis>
</taxonomicName>
sp. nov., by original designation.
</paragraph>
</subSubSection>
<subSubSection id="5989ED5FEB6EB9B821E1C69140D3F775" pageId="0" pageNumber="1" type="etymology">
<paragraph id="7EB72EE1C26A71271B46F8EF7FA7D147" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="C543978A3A3A2F87AD41A316B871BC04" pageId="0" pageNumber="1">
The name combines
<taxonomicName id="01D4B8D3C779A53C36686E35D8FCE9C4" genus="Burmo" lsidName="Burmo" pageId="0" pageNumber="1" rank="genus">
<emphasis id="02D672D5BE2EE6B6B4DDEAED67164036" italics="true" pageId="0" pageNumber="1">Burmo</emphasis>
</taxonomicName>
in reference to Burma, the historical name of Myanmar, and the Greek word
<emphasis id="8758AB378170876B7C15A56FC79A3188" italics="true" pageId="0" pageNumber="1">chlamys</emphasis>
meaning mantle or cloak in reference to land snail with well-developed mantle extensions. Therefore, the generic name means the Burmese land snail with the well-developed mantle extensions. The gender of the new generic name is feminine.
</paragraph>
</subSubSection>
<subSubSection id="1619C222EE53D73FCE3301F89A396D50" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="0B48A6FE4812DB5330294B2F95A2F2A4" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="06AE7FD8A6E5E31CBDF5D05111A16FA3" pageId="0" pageNumber="1">
Shell subglobose to globose, small size, little high spire, and sculptured with spiral furrows and undulating radial lines. Snail with five well-developed mantle extensions; caudal horn raised. Genitalia with penial retractor muscle attached at tip of epiphallic caecum; gametolytic organ with rather short to moderate cylindrical duct and bulbous sac; well-developed dart apparatus; flagellum absent.
<taxonomicName id="FB92BDC3ADC378C8CAB7D9717D3E263B" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">Radula</taxonomicName>
with large monocuspid central tooth and attached by two smaller teeth; laterals and marginals undifferentiated, large monocuspid, and at base of each tooth on outer side attached by a smaller tooth.
</paragraph>
</subSubSection>
<subSubSection id="41E1E4CCB6E6D595837D64F7C50AFF4A" pageId="0" pageNumber="1" type="description">
<paragraph id="77CE749265926AC9C71D306A009A738A" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="A0058BBED128A74ED2BB3E1413C1D457" pageId="0" pageNumber="1">
<emphasis id="6A5716EA4229C1FD97CD5677BFA6F7F7" bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
subglobose to globose, small-sized, thin, whitish to brownish, with or without yellowish brown band on the periphery. Shell surface with distinct to faintly spiral furrows, crossed by distinct to faintly undulating radial lines. Whorls
<normalizedToken id="26C5ACF4F15AE39CB9CF999D56C63465" originalValue="5½">51/2</normalizedToken>
-7, regularly increasing; spire rather elevated; body whorl rounded. Aperture oblique and crescentic with simple lip. Umbilicus open, narrow to moderate, and deep.
</paragraph>
<paragraph id="532789D2E9095F8DDD18EBF8D85E7DC7" pageId="0" pageNumber="1">
<emphasis id="0628021B5873CEDF58F6201BCF6EE494" bold="true" italics="true" pageId="0" pageNumber="1">Animal</emphasis>
reticulated skin with pale to dark greyish or with a brown or yellow tinge. Mantle lobes or mantle extension well developed, divided into two shell lobes and three dorsal lobes, and somewhat thickened near their margins (Fig.
<figureCitation id="7E1C9157A1D05D9206143CCEEF8121ED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">3A-C</figureCitation>
). Shell lobes short to moderate, slender, finger-like, and same colour as body. Right shell lobe (rsl) rather longer and larger than left shell lobe (lsl). Dorsal lobes large, broad, normally larger than shell lobes, crescent-shaped, and same colour as body. Right dorsal lobe (rdl) larger than left dorsal lobe. Left dorsal lobe divided into anterior (ant-ldl) and posterior lobes (post-ldl). Sole tripartite and lateral foot margin present. Caudal foss (cf) and caudal horn (ch) present (Fig.
<figureCitation id="4863D9CA2BDB23624C16116F237F3DB0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">3B</figureCitation>
).
</paragraph>
<caption id="C456A3A9A2B1DBEBC69CA66BA9811ACB" doi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="669A73C739EDC3B77B5BCF51442406F5" pageId="0" pageNumber="1">
<emphasis id="BA2A2E94D05B210E88F7770301FD9F63" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
Synoptic illustration of mantle extensions, body terminology and radular morphology of
<taxonomicName id="3DE0A441D4E2CF9517E078C25C0DB891" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="7C10259F521EAAC6393A578258162F05" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov.
<emphasis id="E9B92DA5312D6E8DFA723AC1E3791B22" bold="true" pageId="0" pageNumber="1">A</emphasis>
right and left shell lobes of
<taxonomicName id="722A27390235EBE84C61A5F69F4DB65A" lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="FF1A83C9C1C683A0D03A1715E332E57B" italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="2F514368CBEF9931EA02CA3CD549E541" bold="true" pageId="0" pageNumber="1">B</emphasis>
mantle extensions (shell lobes and dorsal lobes) and posterior body of
<taxonomicName id="F69498F33ECCA86F187455317A58D9DC" lsidName="B. poongee" pageId="0" pageNumber="1" rank="species" species="poongee">
<emphasis id="82149524D5F258F58F7DE12A3CB39CEC" italics="true" pageId="0" pageNumber="1">B. poongee</emphasis>
</taxonomicName>
<emphasis id="3C6714DB91DF1205E27CFE8B536F3553" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of mantle extensions (shell lobes and dorsal lobes) of
<taxonomicName id="4773188F3A0587B9EB7C8ADF1263C36E" lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis id="D544C3E8DED403F370065F988FD01CFA" italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
<emphasis id="A35887C436182D3C7B5434D9FA4624DB" bold="true" pageId="0" pageNumber="1">D, E</emphasis>
representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth.
</paragraph>
</caption>
<paragraph id="A0F619DFB98D0C0BAFD81FCFB31A7BEF" pageId="0" pageNumber="1">
<emphasis id="6126A6913BE9ACC9CEA946390E8A0CF9" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
possess penis with thin penial sheath; penial retractor muscle varying in size and attached at tip of short and straight epiphallic caecum. Flagellum absent. Dart apparatus present. Gametolytic organ with rather short to moderate cylindrical duct and bulbous sac.
</paragraph>
<paragraph id="42AC848E72A57CB84FB32A4FDB905354" pageId="0" pageNumber="1">
<taxonomicName id="3F4D709FB8667EB13B0B503CE4ABD73C" genus="Radular" lsidName="Radular teeth" pageId="0" pageNumber="1" rank="species" species="teeth">
<emphasis id="7841DA868D4842C1BC5FFDBD166219CF" bold="true" italics="true" pageId="0" pageNumber="1">Radular teeth</emphasis>
</taxonomicName>
arranged in wide U-shaped rows. Central tooth monocuspid, large, narrow to broad spatulate shape, and with two smaller triangular-shaped teeth located at base (Fig.
<figureCitation id="C1B16761B9C4B9C25F39EC55794D427D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">3D</figureCitation>
). Lateral and marginal teeth not differentiated, monocuspid, large spatulate shape, and with only one smaller triangular-shaped tooth located at base on outer side. From inner to outer, lateromarginal teeth gradually narrower, smaller, and rather pointed cusp (Fig.
<figureCitation id="123E43B41B560E8CF97821AB2B57E1EF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">3D, E</figureCitation>
), and tiny inner cusp sometimes present on outermost teeth (yellow arrow in Fig.
<figureCitation id="C3452DF6581774838736F1F626D57256" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">3E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="806E33426303AD2183CDFD0E561A7891" pageId="0" pageNumber="1" type="constituent species">
<paragraph id="60C964C08E616B9A587B05EC49EFCD58" pageId="0" pageNumber="1">Constituent species.</paragraph>
<paragraph id="E1BC509CED3717B495F37019762AF3F9" pageId="0" pageNumber="1">
<taxonomicName id="A8DB8ED17919EEE9EA4CECBFFF64E258" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="4E07B921D6CF5971BCB3FA0408306AA3" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. currently contains:
<taxonomicName id="9C0A8709471C5BA56B3371F49606A01F" lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis id="6890F61F2029D6F0E141E50FCA2505A2" italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
(Benson, 1859), comb. nov.,
<taxonomicName id="51FABC5172B3D8CD7186E626FA8DCDF5" lsidName="B. cauisa" pageId="0" pageNumber="1" rank="species" species="cauisa">
<emphasis id="B3FC309DD2029D2F4FFA809E3C874127" italics="true" pageId="0" pageNumber="1">B. cauisa</emphasis>
</taxonomicName>
(Benson, 1859), comb. nov.,
<taxonomicName id="131D317CEEDB3B7C95959BD09F80594B" lsidName="B. perpaula" pageId="0" pageNumber="1" rank="species" species="perpaula">
<emphasis id="2380CBA6C6E62BB5F4700A94447DB789" italics="true" pageId="0" pageNumber="1">B. perpaula</emphasis>
</taxonomicName>
(Benson, 1859), comb. nov.,
<taxonomicName id="204EF2C86847EF94D184F4772510E24C" lsidName="B. poongee" pageId="0" pageNumber="1" rank="species" species="poongee">
<emphasis id="719D689794D36D3F118FA8BACDD74254" italics="true" pageId="0" pageNumber="1">B. poongee</emphasis>
</taxonomicName>
(Theobald, 1859), comb. nov.,
<taxonomicName id="A006B5093FE6D3B0D9D3166CB9D42235" lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis id="4595DA3400370C32C6BF1A0C8CB41267" italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="0F57BF80F8B2A78C5C825A4053BD965B" lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="CE2F5541097D65C254C616F1123B6BC5" italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="5BF4CAB7F683F774792FFD474A077838" lsidName="B. moulmeinica" pageId="0" pageNumber="1" rank="species" species="moulmeinica">
<emphasis id="E575F61A2BA85A1F7BAB567C22470E56" italics="true" pageId="0" pageNumber="1">B. moulmeinica</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="272F545B0DE604A7F23D9908D02C8274" lsidName="B. versicolor" pageId="0" pageNumber="1" rank="species" species="versicolor">
<emphasis id="3D7D1B514BDC799A7195B89DCA1FCBB8" italics="true" pageId="0" pageNumber="1">B. versicolor</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName id="B1FBE81564E82AC796F3DFBED1D302EE" lsidName="B. whitteni" pageId="0" pageNumber="1" rank="species" species="whitteni">
<emphasis id="16251546D9F30ED7F2B11561B822CA06" italics="true" pageId="0" pageNumber="1">B. whitteni</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="FBD5AAC1A21285EEEA12312B547EF3A2" pageId="0" pageNumber="1" type="distribution">
<paragraph id="F1F4DD6424EB4241669E1DD1AFC7B9C9" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="DEAA66CFC7FCFF88CD09A6B36E164B11" pageId="0" pageNumber="1">
<taxonomicName id="0F46F28CCF37B09B339355EC1ADD13FF" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="62C09F26695EEE1B25EACCC3D935BBD5" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. shows a remarkable degree of endemism and localisation being restricted to the limestone karsts in the south of Salween River basin, Myanmar (Fig.
<figureCitation id="21CE9518260B97CF0D553D48DE55CD42" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the Salween River basin (known as Thanlwin River) in the southeastern Myanmar showing the sampling sites. Coloured triangles indicate the localities recorded for each Burmochlamys species. White circles indicate the localities recorded for Sophina species. White squares indicate the sampling localities with no records of both Sophina and Burmochlamys species." figureDoi="10.3897/zookeys.1110.82461.figure1" httpUri="https://binary.pensoft.net/fig/709567" pageId="0" pageNumber="1">1</figureCitation>
). The limestone formations in the region are notable for their fragmented, island-like nature, with hills, caves, and towers forming archipelagos of habitat islands and some areas are temporarily flooded during the monsoon season (Fig.
<figureCitation id="9C30179B8386B452C968592B67BF5423" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat of some species of Burmochlamys gen. nov. in the karst basin of Hpa-An, Kayin State, Myanmar. A type locality of B. fasciola sp. nov. at Bardai Mountain and microhabitat structure of the karst wall B newly discovered locality of B. poongee at Kaw Ka Thaung Cave C type locality of B. albida sp. nov. at Waiponla Monastery D type locality of B. moulmeinica sp. nov. at Lun Nya Pagoda E type locality of B. versicolor sp. nov. at Bayin Nyi Cave. Photos by Ruttapon Srisonchai." figureDoi="10.3897/zookeys.1110.82461.figure2" httpUri="https://binary.pensoft.net/fig/709568" pageId="0" pageNumber="1">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5640020C9ECD6765F6B8115FB51D216D" pageId="0" pageNumber="1" type="remarks">
<paragraph id="8C0DEC117815506155F7205D620EDBAB" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="FA787B93CC2C8E50FD2433E8A152E09B" pageId="0" pageNumber="1">
<taxonomicName id="955BF0DD453D07CB1A93984356FB33E1" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="386DF75437124F2D479F1740B8BAF605" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. possesses a similar radular morphology (monocuspid and spatulate shape) to those of
<taxonomicName id="E5AECE86DDD41E90168FD5D01A4644A3" authorityName="C. Tumpeesuwan &amp; S. Tumpeesuwan" authorityYear="2017" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" lsidName="Aenigmatoconcha" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="6C21D5DF4A4E76630992229A1171DF7E" italics="true" pageId="0" pageNumber="1">Aenigmatoconcha</emphasis>
</taxonomicName>
Tumpeesuwan &amp; Tumpeesuwan, 2017,
<taxonomicName id="F89A9E9BD21B1BF6B3FB2A9CCE623484" authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepotaxis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="F467227E694B90FB3945C26A7ED564CA" italics="true" pageId="0" pageNumber="1">Chalepotaxis</emphasis>
</taxonomicName>
Ancey, 1887, and
<emphasis id="A2023E8D6DB3142F44B6F4C31F2DF3A1" italics="true" pageId="0" pageNumber="1">Sophina</emphasis>
. However, the new genus is easy to distinguish from these three genera by having a microscopic shell sculpture, slender mantle extensions (left and right shell lobes) and genitalia with a well-developed dart apparatus and without a flagellum. In contrast, those three genera have a smooth shell surface and well-developed left and right shell lobes that can be enlarged and cover most of the shell. The genitalia of
<emphasis id="C1367E2A95288C3A0F7AC0A9C1D3EF3F" italics="true" pageId="0" pageNumber="1">Sophina</emphasis>
is more similar to that of
<taxonomicName id="E3648FB4C913C11130BF461718CF1544" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="804AD77BC6079A38F45BE624EB56F1C2" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov., while
<taxonomicName id="6B70666539640F3E74B0BD151E30D517" authorityName="C. Tumpeesuwan &amp; S. Tumpeesuwan" authorityYear="2017" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" lsidName="Aenigmatoconcha" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="9CB31EB52C41D52CBA706EF4D4844077" italics="true" pageId="0" pageNumber="1">Aenigmatoconcha</emphasis>
</taxonomicName>
has a small flagellum and no dart apparatus, and
<taxonomicName id="2D2E16537642755F155C1E3ECA11E179" authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepotaxis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="4C3838913475422F3C04733B728EE8CD" italics="true" pageId="0" pageNumber="1">Chalepotaxis</emphasis>
</taxonomicName>
has neither flagellum nor dart apparatus (
<bibRefCitation id="DE4330FE348F7B04905992ECA289B81E" DOI="https://doi.org/10.1080/13235818.2016.1234996" author="Pall-Gergely, B" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="113 - 119" refId="B20" refString="Pall-Gergely, B, Feher, Z, Otani, JU, Asami, T, 2016. An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylommatophora: Helicarionidae). Molluscan Research 37 (2): 113 - 119, DOI: https://doi.org/10.1080/13235818.2016.1234996" title="An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylommatophora: Helicarionidae)." url="https://doi.org/10.1080/13235818.2016.1234996" volume="37" year="2016">
<normalizedToken id="A5A01038B3D894389D5C09294CE9A4FA" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2016
</bibRefCitation>
;
<bibRefCitation id="F8B8059D82096029FB6037C82D68B68B" DOI="https://doi.org/10.1111/jzs.12420" author="Sutcharit, C" journalOrPublisher="Journal of Zoological Systematics and Evolutionary Research" pageId="0" pageNumber="1" pagination="957 - 981" refId="B34" refString="Sutcharit, C, Jeratthitikul, E, Pholyotha, A, Lin, A, Panha, S, 2020. Molecular phylogeny reveals high diversity and endemism in the limestone karst-restricted land snail genus Sophina Benson, 1859 from Myanmar (Eupulmonata: Helicarionidae), with description of four new species. Journal of Zoological Systematics and Evolutionary Research 58 (4): 957 - 981, DOI: https://doi.org/10.1111/jzs.12420" title="Molecular phylogeny reveals high diversity and endemism in the limestone karst-restricted land snail genus Sophina Benson, 1859 from Myanmar (Eupulmonata: Helicarionidae), with description of four new species." url="https://doi.org/10.1111/jzs.12420" volume="58" year="2020">Sutcharit et al. 2020</bibRefCitation>
;
<bibRefCitation id="39ABA5E3B0BF9A7E48887973B65899DE" DOI="https://doi.org/10.5852/ejt.2021.767.1487" author="Pholyotha, A" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="55 - 82" refId="B27" refString="Pholyotha, A, Sutcharit, C, Tongkerd, P, Panha, S, 2021b. Systematic revision of the limestone karst-restricted land snail genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with description of a new species. European Journal of Taxonomy 767: 55 - 82, DOI: https://doi.org/10.5852/ejt.2021.767.1487" title="Systematic revision of the limestone karst-restricted land snail genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with description of a new species." url="https://doi.org/10.5852/ejt.2021.767.1487" volume="767" year="2021 b">Pholyotha et al. 2021b</bibRefCitation>
).
</paragraph>
<paragraph id="AAE85482F776E57BAA95207CECC57525" pageId="0" pageNumber="1">
In addition,
<taxonomicName id="F09A0DC141C1E016E3C7A23282E7FDCB" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="6C018072BA57F3FC0A8B43811CEFE1DD" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. is clearly discriminated from other helicarionid and ariophantid genera with or without shell lobes by the presence of the monocuspid radular teeth (see Table
<tableCitation id="E848E0608BC4786822A501AAD87F251E" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of the morpho-anatomical characteristics of Burmochlamys gen. nov. and the possibly related genera of the Helicarionidae and Ariophantidae in mainland Southeast Asia. References: 1 = this study, 2 = Stoliczka (1871), 3 = Blanford and Godwin-Austen (1908), 4 = van Benthem Jutting (1950), 5 = Solem (1966), 6 = Schileyko (2002), 7 = Schileyko (2003), 8 = Pall-Gergely et al. (2016), 9 = Tanmuangpak et al. (2017), 10 = Pholyotha et al. (2018), 11 = Pholyotha et al. (2020 b), 12 = Pholyotha et al. (2020 c), 13 = Pholyotha et al. (2021 a), 14 = Pholyotha et al. (2021 b), and 15 = Sutcharit et al. (2020)." httpUri="http://table.plazi.org/id/1522F13CA73C2C435B1AFC2599055978" pageId="0" pageNumber="1" tableUuid="1522F13CA73C2C435B1AFC2599055978">1</tableCitation>
). In comparison, the radular morphology of most genera in these families possess bicuspid, tricuspid, or multicuspid teeth (
<bibRefCitation id="B87381D4055587F7FC181727A02B1E1A" author="Stoliczka, F" journalOrPublisher="The Veliger" pageId="0" pageNumber="1" refId="B32" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim provinces), with description of new species. Journal of the Asiatic Society of Bengal 40: 143-177, 217-259." title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim provinces), with description of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177, 217 - 259." year="1871">Stoliczka 1871</bibRefCitation>
;
<bibRefCitation id="F56CC0EC7CF212C8F9D776A6C7748422" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B3" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
;
<bibRefCitation id="D34E0FA5CCDB8825EC0AFD1D8AA1ED50" author="van Benthem Jutting, WSS" journalOrPublisher="Treubia" pageId="0" pageNumber="1" pagination="381 - 505" publicationUrl="http://e-journal.biologi.lipi.go.id/index.php/treubia/article/view/2634/2270" refId="B42" refString="van Benthem Jutting, WSS, 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian Archipelago. II Critical revision of the Javanese pulmonate landshells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505, http://e-journal.biologi.lipi.go.id/index.php/treubia/article/view/2634/2270" title="Systematic studies on the non-marine Mollusca of the Indo-Australian Archipelago. II Critical revision of the Javanese pulmonate landshells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae." url="http://e-journal.biologi.lipi.go.id/index.php/treubia/article/view/2634/2270" volume="20" year="1950">van Benthem Jutting 1950</bibRefCitation>
;
<bibRefCitation id="ACBF68D3C9DA17D7C5701422DF84E545" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B30" refString="Solem, A, 1966. Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem 1966</bibRefCitation>
;
<bibRefCitation id="F35A2F0C5D5F6FBF23C863069F0817F8" author="Schileyko, AA" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" refId="B28" refString="Schileyko, AA, 2002. Treatise on recent terrestrial pulmonated mollusks. Part 9. Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica (Supplement 2): 1167-1307." title="Treatise on recent terrestrial pulmonated mollusks. Part 9. Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica (Supplement 2): 1167 - 1307." year="2002">Schileyko 2002</bibRefCitation>
,
<bibRefCitation id="99A0A2133C81D35524CA6E24DAED3B17" author="Schileyko, AA" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" refId="B29" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonate molluscs. Part 10. Ariophantidae, Ostracolethidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica (Supplement 2): 1309-1466." title="Treatise on recent terrestrial pulmonate molluscs. Part 10. Ariophantidae, Ostracolethidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica (Supplement 2): 1309 - 1466." year="2003">2003</bibRefCitation>
;
<bibRefCitation id="15154F7A6D7BD4419097F8A1D85405C1" DOI="https://doi.org/10.1080/13235818.2016.1234996" author="Pall-Gergely, B" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="113 - 119" refId="B20" refString="Pall-Gergely, B, Feher, Z, Otani, JU, Asami, T, 2016. An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylommatophora: Helicarionidae). Molluscan Research 37 (2): 113 - 119, DOI: https://doi.org/10.1080/13235818.2016.1234996" title="An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylommatophora: Helicarionidae)." url="https://doi.org/10.1080/13235818.2016.1234996" volume="37" year="2016">
<normalizedToken id="092CEFC1BEF1B3103ABA903156DFF737" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2016
</bibRefCitation>
;
<bibRefCitation id="6DA61C3DAD6BEE64DCC28A15CC1BF5B7" DOI="https://doi.org/10.1111/jzs.12420" author="Sutcharit, C" journalOrPublisher="Journal of Zoological Systematics and Evolutionary Research" pageId="0" pageNumber="1" pagination="957 - 981" refId="B34" refString="Sutcharit, C, Jeratthitikul, E, Pholyotha, A, Lin, A, Panha, S, 2020. Molecular phylogeny reveals high diversity and endemism in the limestone karst-restricted land snail genus Sophina Benson, 1859 from Myanmar (Eupulmonata: Helicarionidae), with description of four new species. Journal of Zoological Systematics and Evolutionary Research 58 (4): 957 - 981, DOI: https://doi.org/10.1111/jzs.12420" title="Molecular phylogeny reveals high diversity and endemism in the limestone karst-restricted land snail genus Sophina Benson, 1859 from Myanmar (Eupulmonata: Helicarionidae), with description of four new species." url="https://doi.org/10.1111/jzs.12420" volume="58" year="2020">Sutcharit et al. 2020</bibRefCitation>
;
<bibRefCitation id="AD92A68270706D0F0D516ECE622FB82F" DOI="https://doi.org/10.5852/ejt.2021.767.1487" author="Pholyotha, A" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="55 - 82" refId="B27" refString="Pholyotha, A, Sutcharit, C, Tongkerd, P, Panha, S, 2021b. Systematic revision of the limestone karst-restricted land snail genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with description of a new species. European Journal of Taxonomy 767: 55 - 82, DOI: https://doi.org/10.5852/ejt.2021.767.1487" title="Systematic revision of the limestone karst-restricted land snail genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with description of a new species." url="https://doi.org/10.5852/ejt.2021.767.1487" volume="767" year="2021 b">Pholyotha et al. 2021b</bibRefCitation>
).
</paragraph>
<caption id="1522F13CA73C2C435B1AFC2599055978" ID-Table-UUID="1522F13CA73C2C435B1AFC2599055978" httpUri="http://table.plazi.org/id/1522F13CA73C2C435B1AFC2599055978" pageId="0" pageNumber="1" start="Table 1" startId="T1">
<paragraph id="6B4AC7CBA6764A470359E6BB4D53CBF2" pageId="0" pageNumber="1">
<emphasis id="1CBEF4CBA873408E95B8A8BD3B6C4D18" bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
Comparison of the morpho-anatomical characteristics of
<taxonomicName id="A7E4B1E14C878DBED99C38C0C481D084" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="1E928F79E8F87779ECF884AF719D7693" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. and the possibly related genera of the
<taxonomicName id="506C2894BA4B6B642B1FE019D3AF97D8" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
and
<taxonomicName id="F911F3F92BB39EA217ED310B22D638F1" authorityName="Godwin-Austen" authorityYear="1883" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
in mainland Southeast Asia. References: 1 = this study, 2 =
<bibRefCitation id="B4FFAEAE77B9802EE8D747A77B5BA484" author="Stoliczka, F" journalOrPublisher="The Veliger" pageId="0" pageNumber="1" refId="B32" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim provinces), with description of new species. Journal of the Asiatic Society of Bengal 40: 143-177, 217-259." title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim provinces), with description of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177, 217 - 259." year="1871">Stoliczka (1871)</bibRefCitation>
, 3 =
<bibRefCitation id="C3EF4C609F6657939F4E86DE31863D12" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B3" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen (1908)</bibRefCitation>
, 4 =
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<normalizedToken id="15348419F41FC0BE013C4E6972EB520D" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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.
</paragraph>
</caption>
<paragraph id="540CE0D54D79F870AB56601664FD59AF" pageId="0" pageNumber="1">
<table id="A90EE2C99E32FAA361580C8A0C6D92D1" pageId="0" pageNumber="1">
<tr id="D0E3B6604039BD6E52A6121753ECC884" pageId="0" pageNumber="1">
<th id="D116B39CC7443240C41727DE6431902F" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</th>
<th id="820F890C84D2B8F0ECC2F870E8A52E2C" colspan="1" pageId="0" pageNumber="1" rowspan="1">Shell size (diameter)</th>
<th id="B6406B1C1688A9A1B15E608ABA028691" colspan="1" pageId="0" pageNumber="1" rowspan="1">Periphery</th>
<th id="80DDE52BF741D0357CB87F4A2D67C4E6" colspan="1" pageId="0" pageNumber="1" rowspan="1">Shell lobe</th>
<th id="F4361C4B3AA6A82B57E4E44104FE4A70" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="FE13B34DC62FEFF7722B60949D005FC9" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">Radula</taxonomicName>
teeth
</th>
<th id="00785AA74849EEA8E1349166720EB930" colspan="1" pageId="0" pageNumber="1" rowspan="1">Epiphallic caecum</th>
<th id="F55DC62A9B73FB0C019E13F660FBBAF4" colspan="1" pageId="0" pageNumber="1" rowspan="1">Flagellum</th>
<th id="7BFD285D2122C16C6818E30B2F60B83B" colspan="1" pageId="0" pageNumber="1" rowspan="1">Dart apparatus</th>
<th id="3222C1763B6AC3D7BFFF205A4E6AC9B9" colspan="1" pageId="0" pageNumber="1" rowspan="1">Ref.</th>
</tr>
<tr id="C78E5AA19F522B1039E8311631CC5B8D" pageId="0" pageNumber="1">
<td id="A2B5B9242954B3D44378B00827D9ECE5" colspan="9" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="62816CA02CE649E84F048993B495FE30" bold="true" pageId="0" pageNumber="1">
Family
<taxonomicName id="75379D4B4BE8725E211F1A0E37A5BBFB" authorityName="Godwin-Austen" authorityYear="1883" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
Godwin-Austen, 1883
</emphasis>
</td>
</tr>
<tr id="EE600163285A05B0EEDB730C0F103ED8" pageId="0" pageNumber="1">
<td id="E42402E2D1EABE56737CFE09444E787D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="78714D2E39CDBC049AFDB7729850BFFF" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="853F2BE94A8BED372545455A029A0A9A" authorityName="Godwin-Austen" authorityYear="1899" class="Gastropoda" family="Ariophantidae" genus="Khasiella" kingdom="Animalia" lsidName="Khasiella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="7394E981EE0BECB7E9398D287E86135A" italics="true" pageId="0" pageNumber="1">Khasiella</emphasis>
</taxonomicName>
Godwin-Austen, 1899 (
<taxonomicName id="72B81748A1555D1B9268C61DE600D3DD" authorityName="Hanley &amp; Theobald" authorityYear="1876" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix vidua" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="vidua">
<emphasis id="546776D8C80299D2DA45B2E91136692B" italics="true" pageId="0" pageNumber="1">Helix vidua</emphasis>
</taxonomicName>
Hanley &amp; Theobald, 1876)
</emphasis>
*
</td>
<td id="E76EA31EC9A7E35AB3EED45224852E2A" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="39C70305277A90915D38A2C1096E00AF" colspan="1" pageId="0" pageNumber="1" rowspan="1">keeled or subangulate</td>
<td id="B5489EA136DAA4A57DDF2A7A3A1A8891" colspan="1" pageId="0" pageNumber="1" rowspan="1">short, slightly extended</td>
<td id="8687D405F3F536DE6DBE0DB7E955E707" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="B2DABF117620465F549C29C17908F137" colspan="1" pageId="0" pageNumber="1" rowspan="1">free bent</td>
<td id="4F26E8E9D0FECC717B390DDA570EC6F3" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="18AE8C7AEF048B3BD96CC6E3AAB1328F" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="1E9B24ABA6F5B0057A3A23EF5B7AF198" colspan="1" pageId="0" pageNumber="1" rowspan="1">3,6</td>
</tr>
<tr id="814432132E285B7DC8FC253D9E547568" pageId="0" pageNumber="1">
<td id="2A774744B1AD45A05D0B3C53221FC1FD" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="C84374185098FE30637D613E293ED084" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="55CB08F331E557D95F2E1F75D1FC40F7" authorityName="Gray" authorityYear="1847" family="Helicarionidae" genus="Macrochlamys" kingdom="Animalia" lsidName="Macrochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="42071989EA0DBDEA58E5C61959B94268" italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
</taxonomicName>
Gray, 1847 (
<taxonomicName id="F66915D92E19E90D8393C4E147D60C0F" authorityName="Deshayes" authorityYear="1831" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix vitrinoides" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="vitrinoides">
<emphasis id="26EFD2BC96067399C63E2D25EF96A027" italics="true" pageId="0" pageNumber="1">Helix vitrinoides</emphasis>
</taxonomicName>
Deshayes, 1831)
</emphasis>
</td>
<td id="97E20B19662486EF8E3A1E5567377A25" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to large</td>
<td id="E5410D3C5EE34C25CB1C8C6BD4137394" colspan="1" pageId="0" pageNumber="1" rowspan="1">round or anglulate</td>
<td id="D2C5E64C558D5E6AA7F6C78966D1DF8D" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate to long, slender</td>
<td id="918CC2A9F157257754C3C80EFCC49192" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="5457B4F214BDA33A5A4357E4A0F9B4E6" colspan="1" pageId="0" pageNumber="1" rowspan="1">coiled</td>
<td id="67EE6C8EE3FA998BE17A80CBF3214276" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="035EEB6FE774B67A8868C6A2A5DC7958" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="D75F9DFED81F70E4BD52688131A2B418" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, 7, 10, 11</td>
</tr>
<tr id="4A55C1B094250A3AA44FC0818DC2345B" pageId="0" pageNumber="1">
<td id="BCB777C0727F1DA09CF01F83BD1265B3" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="27E23AFAAC2C3FCF98242F674062A7A1" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="846EF60197CDB892A35F3BA6783626C6" authorityName="Morch" authorityYear="1872" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="E78DB0A7DA36766590C5FB6A0CC4E60C" italics="true" pageId="0" pageNumber="1">Microcystina</emphasis>
</taxonomicName>
<normalizedToken id="1FB80E76A81796190026056DD6676455" originalValue="Mörch">Moerch</normalizedToken>
, 1872 (
<taxonomicName id="BFC6902196BB9E649957011FB8C1DF54" authorityName="Morch" authorityYear="1872" family="Helicarionidae" genus="Nanina" kingdom="Animalia" lsidName="Nanina rinki" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="rinki">
<emphasis id="5DE38264DEF71E1CCC3353C18ADAE9AC" italics="true" pageId="0" pageNumber="1">Nanina rinki</emphasis>
</taxonomicName>
<normalizedToken id="ED4081628547746D9855DBEFD68B8BD5" originalValue="Mörch">Moerch</normalizedToken>
, 1872)
</emphasis>
*
</td>
<td id="9408D02EA3EE1C977057C59F3BE4C0A1" colspan="1" pageId="0" pageNumber="1" rowspan="1">small</td>
<td id="C282DB82614BAC3E3EC04CF54C54C1D0" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="AB9934F1E582E83847FBC1E4D294B8DD" colspan="1" pageId="0" pageNumber="1" rowspan="1">short to moderate, slender</td>
<td id="54FB61470803D32BEA60A91ADB7EF805" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, laterals tricuspid, marginals bicuspid</td>
<td id="784751FA5510AA9D77729CB3387C8CD3" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="6B87E2CE5F5AE62EC0D488392AC495E7" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="6069D59D411424AF5BA08D0B6DC241BD" colspan="1" pageId="0" pageNumber="1" rowspan="1">present1</td>
<td id="7A08FE044701E32E13F380587263AB39" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, 4, 7</td>
</tr>
<tr id="AF2E27AAC0577564A4E32125612BD806" pageId="0" pageNumber="1">
<td id="6D660357B67F7A356B29BA25855CD40D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="77591029E010F224733EE04E17884493" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="D41FCB74D6C71EA00EE95B56E03A8CBF" authorityName="Godwin-Austen" authorityYear="1908" class="Gastropoda" family="Ariophantidae" genus="Sakiella" kingdom="Animalia" lsidName="Sakiella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="8734077E7A8F30ECF5B16474FA50D4C6" italics="true" pageId="0" pageNumber="1">Sakiella</emphasis>
</taxonomicName>
Godwin-Austen, 1908 (
<taxonomicName id="694A1711178A2FBA394E4277B7787A88" authorityName="A. A. Gould" authorityYear="1846" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix honesta" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="honesta">
<emphasis id="59218893F4F8891B1E3D7899DFB01A0E" italics="true" pageId="0" pageNumber="1">Helix honesta</emphasis>
</taxonomicName>
Gould, 1846)
</emphasis>
*
</td>
<td id="369288B347D556F50155F0095EF102EB" colspan="1" pageId="0" pageNumber="1" rowspan="1">medium</td>
<td id="E105FA25791FA9DEBEB2972EB810AB27" colspan="1" pageId="0" pageNumber="1" rowspan="1">subangulate</td>
<td id="A86DAD83A9DD6CBD0D6F74D0578B874C" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate, slender</td>
<td id="9F8CC865F55DA6DB77FD94D6B77B049A" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="2F59CCDEAFC5ED1B67787A1BF7A09FBE" colspan="1" pageId="0" pageNumber="1" rowspan="1">n.a.</td>
<td id="0016B8083AA1AAF7A71DB8A0B93B6519" colspan="1" pageId="0" pageNumber="1" rowspan="1">n.a.</td>
<td id="611572D54D681D7CE046B966E2E06D79" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="475D354B3900AEF42940EFBDF2620694" colspan="1" pageId="0" pageNumber="1" rowspan="1">2, 3, 7</td>
</tr>
<tr id="01C2A579BD94CD2AEBAD44EF575D15BE" pageId="0" pageNumber="1">
<td id="4F258CCBE25BE76C9947648B6D01CAB2" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="99E2F78EDF0BFCC0E02AE0C90E20E778" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="C378ED94174A9218BBF7FF7DB9CC608B" authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="B9B173C28F2305E4520BDBC56A9FB9A3" italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
</taxonomicName>
Godwin-Austen, 1907 (
<taxonomicName id="55965C44A2FEE78C38BCD350FBE556E8" authorityName="Philippi" authorityYear="1847" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix resplendens" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="resplendens">
<emphasis id="3F48F21EB4131DAD907F6F44AC4267DA" italics="true" pageId="0" pageNumber="1">Helix resplendens</emphasis>
</taxonomicName>
Philippi, 1847)
</emphasis>
*
</td>
<td id="B31F1CEAA44230E522D6D0B144277EAC" colspan="1" pageId="0" pageNumber="1" rowspan="1">medium to large</td>
<td id="FB72270B402CE5FED4DAF2887605B656" colspan="1" pageId="0" pageNumber="1" rowspan="1">round or angulate</td>
<td id="B3E28887AD2137409805AC41127DB8E7" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate to long, slender</td>
<td id="4561641521731815D24260335E6F2365" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="4BE0B162CFE8062F68512B35E5CE8BC1" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="728C706D08FD3661CAC2A75926654044" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="BF9E984885A154B096A23EB02C4A43EC" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="2071EFCA0EF6F8C3A4A64C4DBAC7516E" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, 11, 6, 12</td>
</tr>
<tr id="9CF0781ED6EC2B1F1C0016FA80895DF8" pageId="0" pageNumber="1">
<td id="B7887237BDCABAE0E03DE667026D6BEE" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="253B518E70EAAEC27BCB10BF5C457D3E" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="07D9EDF800E626C60649BAE93A263F8A" authorityName="Albers" authorityYear="1860" class="Gastropoda" family="Ariophantidae" genus="Sesara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sesara" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="37781FC3C753CFC87B16E3EAB3471C03" italics="true" pageId="0" pageNumber="1">Sesara</emphasis>
</taxonomicName>
Albers, 1860 (
<taxonomicName id="F30D98730DDA19D92BE91B0CAC259970" authorityName="Gould" authorityYear="1843" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix infrendens" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="infrendens">
<emphasis id="BDA1D6CC0FC17FB22AA5420AEF82826D" italics="true" pageId="0" pageNumber="1">Helix infrendens</emphasis>
</taxonomicName>
Gould, 1843)
</emphasis>
*
</td>
<td id="CAA415BFC291B3123798A1632644E332" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="B8CCDB08188E7C532BDABEC28418DC25" colspan="1" pageId="0" pageNumber="1" rowspan="1">round, angulate, or carinate</td>
<td id="1EBED14AA9E24EC7A629FFF2C4060716" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="4576C63D5FBFF5EE5B0A38F022330EB1" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="EEC89103DC0299F95A36360CBDA92E36" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight2</td>
<td id="5956E83EC2F814A99A2F2BA0E2498EBE" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="ADEF31A4951717984B9A411F0F354ED7" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="7FA0A81918C7579DAFCD0C3368E6B8C1" colspan="1" pageId="0" pageNumber="1" rowspan="1">2, 3, 5, 6, 9</td>
</tr>
<tr id="2DFEFEB28487FC28781BEDEF3A64AADD" pageId="0" pageNumber="1">
<td id="A757583A91FDC4CB05D6C93A65485D0B" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="3EE8041262CAB25D3D21770B24867D11" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="776B3BFCCEF20636C8CFF0DA7468C013" authorityName="Pholyotha &amp; Panha" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Taphrenalla" kingdom="Animalia" lsidName="Taphrenalla" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="0941F289A8358450FEECB21152271344" italics="true" pageId="0" pageNumber="1">Taphrenalla</emphasis>
</taxonomicName>
Pholyotha &amp; Panha, 2020 (
<taxonomicName id="87E2DC7C144A946EE026B81276D0283F" authorityName="Dall" authorityYear="1897" family="Helicarionidae" genus="Nanina" kingdom="Animalia" lsidName="Nanina diadema" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="diadema">
<emphasis id="B7DF563AED047B7A437F103B7941E40D" italics="true" pageId="0" pageNumber="1">Nanina diadema</emphasis>
</taxonomicName>
Dall, 1897)
</emphasis>
*
</td>
<td id="31BF819C6AE50FC5D617FF51D95C8C29" colspan="1" pageId="0" pageNumber="1" rowspan="1">medium to large</td>
<td id="77FC97CF50913EC37919314FDCE2F265" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="76FE662AFB085C7B00A90ECF2D961128" colspan="1" pageId="0" pageNumber="1" rowspan="1">long, slender</td>
<td id="DA738A7B23FEAB8CC0E52439F1764059" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="8869E3B12FBBD3362D4B9ABFC5AC3B20" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight to little bent</td>
<td id="C37632A00D28CA70B9E6136D29F043A4" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="F9380746FD516F8C916F8DEF7437CB92" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="DEE626372EF9B2B3DC3066373042803B" colspan="1" pageId="0" pageNumber="1" rowspan="1">13</td>
</tr>
<tr id="33F9AB045B11148E2DE1E188D318E857" pageId="0" pageNumber="1">
<td id="C12A2DB0971C3DCF92D29894F694B8AC" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="F0551AA062E29F819F0349B8E575A432" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="EB033948F9C04E00AD65A4F68ACC1E23" authorityName="W. T. Blanford" authorityYear="1904" class="Gastropoda" family="Ariophantidae" genus="Taphrospira" kingdom="Animalia" lsidName="Taphrospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="A7E93D898EEE5F47D42143811CACB370" italics="true" pageId="0" pageNumber="1">Taphrospira</emphasis>
</taxonomicName>
Blanford, 1904 (
<taxonomicName id="6F8181F7C90FCAE579719EAD337C02DF" authorityName="Benson" authorityYear="1856" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix convallata" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="convallata">
<emphasis id="B0D025DDB9F23C987FCAFAD10BFF4C63" italics="true" pageId="0" pageNumber="1">Helix convallata</emphasis>
</taxonomicName>
Benson, 1856)
</emphasis>
</td>
<td id="F1E28452FB86B1E1C05D101C60F32EC3" colspan="1" pageId="0" pageNumber="1" rowspan="1">medium</td>
<td id="0ECEAF6C02A7968DAC7351A4CD7FE3EB" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="DACE90EE448EBA3A3C893FB7AB35EE52" colspan="1" pageId="0" pageNumber="1" rowspan="1">long and fairly broad, covering partially</td>
<td id="1FEB40987BFDFF5FB554D92801B55511" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, laterals tricuspid, marginals bicuspid</td>
<td id="1645ADA3F6FD7E65F67184F69E9190FA" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="DD034C0A8CEA1F0DEA34D1235676E97B" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="6B73B679048F96A5F08BD291DB56DDB8" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="B4A2DEEBE34C03C22152079F5FC07332" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, 6</td>
</tr>
<tr id="FDF4D157EEA50B58ACB85C7253EB6286" pageId="0" pageNumber="1">
<td id="37725D09C1C72E6C129C152CE50AE20B" colspan="9" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="7ADD3281B8D70ABF6402EF988BF83D4E" bold="true" pageId="0" pageNumber="1">
Family
<taxonomicName id="B4BC24E51C1A5C1C43FEC2E4E4557FD4" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
Bourguignat, 1877
</emphasis>
</td>
</tr>
<tr id="32879090B1B8AA56E794468AB3BCC2AB" pageId="0" pageNumber="1">
<td id="F4E7851773D92AF34D124465D187907F" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="F26330653FA6BB61ECBF01BEEA8BAA8C" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="F3CD5C8E2299BF81C524B141FAD9DF14" authorityName="C. Tumpeesuwan &amp; S. Tumpeesuwan" authorityYear="2017" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" lsidName="Aenigmatoconcha" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="B2FA56241CD5AE07C3D7CC69C764F67C" italics="true" pageId="0" pageNumber="1">Aenigmatoconcha</emphasis>
</taxonomicName>
Tumpeesuwan &amp; Tumpeesuwan, 2017 (
<taxonomicName id="9063CC3745C4C660ABF2EC918B07A428" authorityName="C. Tumpeesuwan &amp; S. Tumpeesuwan" authorityYear="2017" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" lsidName="Aenigmatoconcha clivicola" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="clivicola">
<emphasis id="25437A62025B88289E893950A4A20276" italics="true" pageId="0" pageNumber="1">Aenigmatoconcha clivicola</emphasis>
</taxonomicName>
Tumpeesuwan &amp; Tumpeesuwan, 2017)
</emphasis>
*
</td>
<td id="3207153E1C1BF3908D86E713776CD1C2" colspan="1" pageId="0" pageNumber="1" rowspan="1">medium</td>
<td id="B3C0F0D3DFD65BB2A983F825BFD1F03E" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="6FBBE8B2595B3B33EC9EA4E88B2E5E88" colspan="1" pageId="0" pageNumber="1" rowspan="1">long, broad, enlarged and covering most of shell</td>
<td id="B134AC28DA41B9C71D828F79B4B5E7BB" colspan="1" pageId="0" pageNumber="1" rowspan="1">central monocuspid, lateromarginal monocuspid</td>
<td id="5ADF202A4195CD2F15B49FCCF6C5863B" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="74B448054B902D654D68E01EA761E03F" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="3323F74A992FBB21886FC2633179708E" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="E9A5A3C0467C7C26F46779267CDA0B64" colspan="1" pageId="0" pageNumber="1" rowspan="1">14</td>
</tr>
<tr id="D900EF762E9693FD2A8A238B0303A6D6" pageId="0" pageNumber="1">
<td id="CFB1AC5C0CD00B00B1C3098023522D3C" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="213F2BF4827BC8F3CFE90A43653DA768" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="5F2769F1415B44393F99F6A387695636" authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalepotaxis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="8F86A6F3A923E37BB33DACC6BB7D3BB4" italics="true" pageId="0" pageNumber="1">Chalepotaxis</emphasis>
</taxonomicName>
Ancey, 1887 (
<taxonomicName id="1650314E674396D5B6B51CEA5BEDFF94" authorityName="Gredler" authorityYear="1881" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix similaris subsp. var. var. infantilis" order="Eupulmonata" pageId="0" pageNumber="1" rank="variety" species="similaris" subSpecies="var." variety="infantilis">Helix similaris var. infantilis</taxonomicName>
Gredler, 1881)
</emphasis>
*
</td>
<td id="5EAF5798497F525A2136484A2CB7F20A" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="1C287B461840C2E77BECAA4DBA01E089" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="203529135CBBFC1355C9AB11F9178AD3" colspan="1" pageId="0" pageNumber="1" rowspan="1">long, broad, enlarged and covering most of shell</td>
<td id="B22B2BE6419F90956FAE610B31215853" colspan="1" pageId="0" pageNumber="1" rowspan="1">central monocuspid, lateromarginal monocuspid</td>
<td id="B7B5B9D3A58F83C55E62869E8046D745" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="6CB2EA5C28EC8F83D6840F9CF5575814" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="275425CA2FD7D11334EEA88051EB155E" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="D153B442A3F9B61E7656D5B127A9EA6F" colspan="1" pageId="0" pageNumber="1" rowspan="1">8</td>
</tr>
<tr id="55290C6E61E33E3C8B54363AE7A24925" pageId="0" pageNumber="1">
<td id="5CA4E2CF261808B6B83DA2B1A70177A7" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="6443441C7E1B31677A5E7E4ED0741D5C" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="8DC574D41B6EDF620F1551D4FB6090E4" authorityName="Cockerell" authorityYear="1891" class="Gastropoda" family="Helicarionidae" genus="Cryptaustenia" kingdom="Animalia" lsidName="Cryptaustenia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="552F3641BA01B2E53E89956AB607CE03" italics="true" pageId="0" pageNumber="1">Cryptaustenia</emphasis>
</taxonomicName>
Cockerell, 1891 (
<taxonomicName id="20E8A27D3A64B9E634E630DBAD76BF55" authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Vitrinidae" genus="Vitrina" kingdom="Animalia" lsidName="Vitrina planospira" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="planospira">
<emphasis id="CB321748CD7E7CD6177633A1E9E005D7" italics="true" pageId="0" pageNumber="1">Vitrina planospira</emphasis>
</taxonomicName>
Benson, 1859 (=
<taxonomicName id="98DB2EB866375CB7FBA091273C631BBA" authorityName="Reeve" authorityYear="1862" class="Gastropoda" family="Vitrinidae" genus="Vitrina" kingdom="Animalia" lsidName="Vitrina succina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="succina">
<emphasis id="A76B8842F550EC70500C4042033B5AF3" italics="true" pageId="0" pageNumber="1">Vitrina succina</emphasis>
</taxonomicName>
Reeve, 1862))
</emphasis>
*
</td>
<td id="6B159753E8CFFEB9844B8DDA132096B4" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="E89DF0B3D3C492F5CEA32667545EB5A5" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="695A367CF6CB456D823C5DC1D5A95404" colspan="1" pageId="0" pageNumber="1" rowspan="1">long, broad, enlarged and covering most of shell</td>
<td id="B82C6A1631B5487705D385E150A2D313" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral bi- or tricuspid, marginal bicuspid</td>
<td id="87F8F2AD32B040CE939C9E2C046B12CA" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="71F8EE0C0A60BEA8A0B6DB21F10D75AD" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="34C2D4A16C73DE29102F958F2C695B5C" colspan="1" pageId="0" pageNumber="1" rowspan="1">present3</td>
<td id="7E445AADA5D417C59DFBD2FE98D5BB2B" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, 5, 7</td>
</tr>
<tr id="D13DE20AA1BCF4935882564E59EAA75A" pageId="0" pageNumber="1">
<td id="E4F421136F392DB5719D4CC9C01D489D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="3429DC969901B92D2A303918082C5119" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="E6330744A65813DFBD7DEEE2931076C8" authorityName="W. T. Blanford" authorityYear="1863" class="Gastropoda" family="Helicarionidae" genus="Durgella" kingdom="Animalia" lsidName="Durgella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="4E8CB3E149A269EC1DB8B2DC3783FD11" italics="true" pageId="0" pageNumber="1">Durgella</emphasis>
</taxonomicName>
Blanford, 1863 (
<taxonomicName id="88166F1BFF4F59AE21789F21077A1997" authorityName="Benson" authorityYear="1859" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix levicula" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="levicula">
<emphasis id="2AC72F0F9313D988CD63376AB3CC263D" italics="true" pageId="0" pageNumber="1">Helix levicula</emphasis>
</taxonomicName>
Benson, 1859)
</emphasis>
*
</td>
<td id="B0FA8F27E323A553BD1E38DB2FAFF414" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="1F939E78C155BC098B5B876DE9CC41B9" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="ACA393F2626DB2090DFEEA10407FC4CC" colspan="1" pageId="0" pageNumber="1" rowspan="1">long, broad, enlarged and covering most of shell</td>
<td id="A17F4BE2F0153531EA81DF75F596A441" colspan="1" pageId="0" pageNumber="1" rowspan="1">central unicuspid, lateromarginal teeth bicuspid with numerous cusps on the outer side</td>
<td id="E58C6779CB97C166C64C5BBBA8FFA888" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="76D5FB281AC6072A10D4AB55313F8225" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="D83E6D231AF8EEBD386ED7BDBB299B82" colspan="1" pageId="0" pageNumber="1" rowspan="1">present4</td>
<td id="F8117CB86F8A32EBF6BD36356D8FB9B7" colspan="1" pageId="0" pageNumber="1" rowspan="1">3, 5, 6</td>
</tr>
<tr id="E7EE243B01D609F85BCBD61671DA9F60" pageId="0" pageNumber="1">
<td id="1D52A8C130A3F652414DF95EAEC57BC8" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="D84DB3E1FD5DA3F4D98F9F77369E2FEC" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="9B80A25A5858284E0BD1FC69FA83A893" authorityName="Godwin-Austen" authorityYear="1908" class="Gastropoda" family="Helicarionidae" genus="Holkeion" kingdom="Animalia" lsidName="Holkeion" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="BC16C5359DB0AA9B20116F7C66CB5A29" italics="true" pageId="0" pageNumber="1">Holkeion</emphasis>
</taxonomicName>
Godwin-Austen, 1908 (
<taxonomicName id="631BD9C7381566ED3CF7143AB9FDEA50" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix anceps" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="anceps">
<emphasis id="C53134AB50B249AA50173E0A6AF52F82" italics="true" pageId="0" pageNumber="1">Helix anceps</emphasis>
</taxonomicName>
Gould, 1843)
</emphasis>
*
</td>
<td id="C0F612BDB41DC3C91506AE2B3E1A09B7" colspan="1" pageId="0" pageNumber="1" rowspan="1">medium to large</td>
<td id="D4F751892FDB5447C7D4779D257BC15E" colspan="1" pageId="0" pageNumber="1" rowspan="1">sharply angulate</td>
<td id="09E0B781D4C8599E2C4522818135977B" colspan="1" pageId="0" pageNumber="1" rowspan="1">short to moderate, slender</td>
<td id="7A030981F3401703FB7FC7DB9ADE480E" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateral tricuspid, marginal bicuspid</td>
<td id="A11F5BAA5360F04EFF88DBB46F78F6C4" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="CBCD53171E91EA3AFD18178CFD367F01" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="3A6149D4599F995F40462606C7439095" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="D291FC3871660D8BD969CE5FE0CB1760" colspan="1" pageId="0" pageNumber="1" rowspan="1">3,7</td>
</tr>
<tr id="A1606FF4CC6D927721D7BBD904B12B71" pageId="0" pageNumber="1">
<td id="5D0AB66E8C0491EAB262C226BBA62E73" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="8CF9054DA7B17EC78FB3AD75568DCA67" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="9F63C0A745FA1AEEBF1F0A68F74CB9E0" authorityName="H. Adams" authorityYear="1865" class="Gastropoda" family="Helicarionidae" genus="Sitala" kingdom="Animalia" lsidName="Sitala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="FC00614D55FD73A8F2EB41C38F6D3CF2" italics="true" pageId="0" pageNumber="1">Sitala</emphasis>
</taxonomicName>
Adams, 1865 (
<taxonomicName id="5A457930DD476D4EF67A0CA4AA01CE71" authorityName="Benson" authorityYear="1848" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix infula" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="infula">
<emphasis id="24885BE80CDBF054BD35D96FF3029F19" italics="true" pageId="0" pageNumber="1">Helix infula</emphasis>
</taxonomicName>
Benson, 1848)
</emphasis>
*
</td>
<td id="786483F0B577048C14155A5E6AE1F3C0" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="3E851236FAAE8AF0D6EFED67A508ED8A" colspan="1" pageId="0" pageNumber="1" rowspan="1">subangulate or carinate</td>
<td id="5AB2F2C95FA5C077DE3EB54A77767F05" colspan="1" pageId="0" pageNumber="1" rowspan="1">short, slightly extended</td>
<td id="981B9270A68D490B06256111E9C3F0CD" colspan="1" pageId="0" pageNumber="1" rowspan="1">central tricuspid, lateromarginal teeth pointed with 2-5 cusps on the outer side</td>
<td id="16A07A1DED70E2AAECA43A3A96EF39A0" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="EF99F3EBEA1A55659A28FD2D7ABCE879" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="94F4596D721B4167DE0D173564D989D8" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent5</td>
<td id="3E7D00F83D658AFFED0CBF251784E0E3" colspan="1" pageId="0" pageNumber="1" rowspan="1">2,3</td>
</tr>
<tr id="AF4918AB52534045E9990B385BA3DC7A" pageId="0" pageNumber="1">
<td id="1B8FFF6DD24016D420AB3BD3E0836790" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="FD64D519EDEA0BB5EF89F03191AA133B" bold="true" pageId="0" pageNumber="1">
<emphasis id="352B0C8FA13E42767978DF282973205B" italics="true" pageId="0" pageNumber="1">Sophina</emphasis>
Benson, 1859 (
<taxonomicName id="322E84CBF7F6609E6647A0E61D5C27C3" authorityName="Benson" authorityYear="1859" family="Helicarionidae" genus="Helix" kingdom="Animalia" lsidName="Helix schistostelis" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="schistostelis">
<emphasis id="E8AFC377765FFBFB8E72FB8E2B5F9DD8" italics="true" pageId="0" pageNumber="1">Helix schistostelis</emphasis>
</taxonomicName>
Benson, 1859)
</emphasis>
*
</td>
<td id="7C9F29137499551CAEDC4E25899F5D50" colspan="1" pageId="0" pageNumber="1" rowspan="1">small to medium</td>
<td id="28744D590282CE47BBB931A20158C22F" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="A9AB81BF96FD109B7526CEEB3A86D3A0" colspan="1" pageId="0" pageNumber="1" rowspan="1">long, broad, enlarged and covering most of shell</td>
<td id="6350E197C0C1C17963FFB0D7A9287B62" colspan="1" pageId="0" pageNumber="1" rowspan="1">central monocuspid, lateromarginal monocuspid</td>
<td id="1EAB98E8F5DE4053370FB8580F28CDD7" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="251CDB0ED0F1856279489BE77E0500A8" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="3E76324F218026F3A810D60DD76B0F1B" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="0B9C720C8216B9659B930C46D66A18F9" colspan="1" pageId="0" pageNumber="1" rowspan="1">2,3,15</td>
</tr>
<tr id="AB696E4B26F4594FCD3224AC6F409804" pageId="0" pageNumber="1">
<td id="961E5CDB6064F278AA6DD6385FE63F40" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="21DB6D47DBFAC17ECCFC7C7A1FD6B6AA" bold="true" pageId="0" pageNumber="1">
<taxonomicName id="20B710971F9DF171CBBE5A8BFDDA695C" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="30E4B64A0867A165A2E55D1AF9B56D1A" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. (
<taxonomicName id="A4FFC5D746003F1DFFB46CAD0EDAE985" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys fasciola" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="3143D59FC8A18A02B54095858DB42915" italics="true" pageId="0" pageNumber="1">Burmochlamys fasciola</emphasis>
</taxonomicName>
sp. nov.)
</emphasis>
*
</td>
<td id="B0A2C9DDDB2324545CF792584ECF9564" colspan="1" pageId="0" pageNumber="1" rowspan="1">small</td>
<td id="0A2707F4FD71EA53C163BAC9749DC6F8" colspan="1" pageId="0" pageNumber="1" rowspan="1">round</td>
<td id="B367A91ADF0494083ED3C25CC600C70A" colspan="1" pageId="0" pageNumber="1" rowspan="1">short to moderate, slender</td>
<td id="5B326A6CFC230EB7362CF38008569276" colspan="1" pageId="0" pageNumber="1" rowspan="1">central monocuspid, lateromarginal monocuspid</td>
<td id="6938C8B22A357582F3DF5635BD7F4AA4" colspan="1" pageId="0" pageNumber="1" rowspan="1">straight</td>
<td id="47FCD176D35ECF56041FDF2509A1C42F" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="2B37D11639D1E7A299553F2290A546AE" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="62DD160601E04876973EB2DB29970506" colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
</tr>
</table>
</paragraph>
<paragraph id="51512A8F5226C7FAFF42AFAF1A57B4B4" pageId="0" pageNumber="1">
<tableNote id="A981F7A7B151AE6AACFA2FDCAA5B31F3" pageId="0" pageNumber="1">
* Genitalia of the type species were examined. 1
<taxonomicName id="D818A97EA0C80163B232916BE35BB99D" authorityName="Godwin-Austen" authorityYear="1899" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina bintennensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bintennensis">
<emphasis id="AFF93CB8697334FE8BF6AD8A3F634DAD" italics="true" pageId="0" pageNumber="1">Microcystina bintennensis</emphasis>
</taxonomicName>
Godwin-Austen, 1899 does not have dart apparatus (see
<bibRefCitation id="ECCAF00899A22CDECD814859C98AEE8C" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B3" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
). 2
<taxonomicName id="1813668BE055796DAE8E50A4413D8E94" authorityName="Solem" authorityYear="1966" class="Gastropoda" family="Ariophantidae" genus="Sesara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sesara parva" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="parva">
<emphasis id="8FF7050F273A270951E770F7922B5219" italics="true" pageId="0" pageNumber="1">Sesara parva</emphasis>
</taxonomicName>
Solem, 1966 has a spirally coiled epiphallic caecum (
<bibRefCitation id="B955306329EAC44FE1DC67C11BA2195E" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B30" refString="Solem, A, 1966. Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem 1966</bibRefCitation>
). 3
<taxonomicName id="579349989B89B5C36F7AE5353FB43DEE" authorityName="Solem" authorityYear="1966" class="Gastropoda" family="Helicarionidae" genus="Cryptaustenia" kingdom="Animalia" lsidName="Cryptaustenia gadinodromica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gadinodromica">
<emphasis id="286E5EC24059C3AB97A216D820F0BF54" italics="true" pageId="0" pageNumber="1">Cryptaustenia gadinodromica</emphasis>
</taxonomicName>
Solem, 1966 does not have dart apparatus (
<bibRefCitation id="DA34BC6188ED390F724AE58D547CDB57" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B30" refString="Solem, A, 1966. Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem 1966</bibRefCitation>
). 4
<taxonomicName id="C4AEFC5D543FAE207B1E840458B19947" authorityName="Gondwin-Austen" authorityYear="1881" class="Gastropoda" family="Helicarionidae" genus="Durgella" kingdom="Animalia" lsidName="Durgella assamica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="assamica">
<emphasis id="0C54EBEA55CA1264866CFF14F0847C0E" italics="true" pageId="0" pageNumber="1">Durgella assamica</emphasis>
</taxonomicName>
Godwin-Austen, 1881 and
<taxonomicName id="803F83D5D508CBEE5753500D5EC2E473" lsidName="D. rogersi" pageId="0" pageNumber="1" rank="species" species="rogersi">
<emphasis id="7F25ADB85A0A2F419126BC8098D400BB" italics="true" pageId="0" pageNumber="1">D. rogersi</emphasis>
</taxonomicName>
Godwin-Austen, 1907 do not have dart apparatus (
<bibRefCitation id="BD9A0AD00277500B10FCA9F994E85DD6" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B3" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
). 5
<taxonomicName id="AE65F8E5488482C667025F44662181D1" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Helicarionidae" genus="Sitala" kingdom="Animalia" lsidName="Sitala attegia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="attegia">
<emphasis id="F0F8B4AAB5763857A14DDFA3FE9A1FD9" italics="true" pageId="0" pageNumber="1">Sitala attegia</emphasis>
</taxonomicName>
(Benson, 1859) has a well-developed dart apparatus (
<bibRefCitation id="91C6773876BEC113EE450BF3AA9F6650" author="Stoliczka, F" journalOrPublisher="The Veliger" pageId="0" pageNumber="1" refId="B32" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim provinces), with description of new species. Journal of the Asiatic Society of Bengal 40: 143-177, 217-259." title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim provinces), with description of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177, 217 - 259." year="1871">Stoliczka 1871</bibRefCitation>
;
<bibRefCitation id="9E9E6332EED58D8F00710AA2833AA5AE" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B3" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen 1908</bibRefCitation>
).
</tableNote>
</paragraph>
<paragraph id="1CD74842F75464D8DE26BD26C455DE02" pageId="0" pageNumber="1">
As observed in the field, we searched after rain and found the snails climbing on the limestone walls or hiding under the slope of rocks (Fig.
<figureCitation id="B11152EEBAA1BEE4CA924C3FB891F888" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A adult of B. moulmeinica sp. nov. while climbing slowly on the karst wall after raining at Lun Nya Pagoda B mating pairs of B. versicolor sp. nov. on the karst wall at Bayin Nyi Cave C B. versicolor sp. nov. eaten by the carnivorous snail Carinartemis sp. on the karst wall at Bayin Nyi Cave D B. cassidula eaten by the carnivorous snail Haploptychius sp. on the karst wall at Kaw Gon Cave." figureDoi="10.3897/zookeys.1110.82461.figure4" httpUri="https://binary.pensoft.net/fig/709570" pageId="0" pageNumber="1">4A</figureCitation>
). Regarding the simultaneous hermaphroditism, many copulating pairs were also discovered (Fig.
<figureCitation id="DFF4271DC7E76F85625D2DC50E3DC517" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A adult of B. moulmeinica sp. nov. while climbing slowly on the karst wall after raining at Lun Nya Pagoda B mating pairs of B. versicolor sp. nov. on the karst wall at Bayin Nyi Cave C B. versicolor sp. nov. eaten by the carnivorous snail Carinartemis sp. on the karst wall at Bayin Nyi Cave D B. cassidula eaten by the carnivorous snail Haploptychius sp. on the karst wall at Kaw Gon Cave." figureDoi="10.3897/zookeys.1110.82461.figure4" httpUri="https://binary.pensoft.net/fig/709570" pageId="0" pageNumber="1">4B</figureCitation>
). Information on its natural predators and parasites remains scarce, but the carnivorous streptaxid snails were found sympatrically in some localities (Fig.
<figureCitation id="A1A8B9451A6D7B469D72BD2E71D83B3C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A adult of B. moulmeinica sp. nov. while climbing slowly on the karst wall after raining at Lun Nya Pagoda B mating pairs of B. versicolor sp. nov. on the karst wall at Bayin Nyi Cave C B. versicolor sp. nov. eaten by the carnivorous snail Carinartemis sp. on the karst wall at Bayin Nyi Cave D B. cassidula eaten by the carnivorous snail Haploptychius sp. on the karst wall at Kaw Gon Cave." figureDoi="10.3897/zookeys.1110.82461.figure4" httpUri="https://binary.pensoft.net/fig/709570" pageId="0" pageNumber="1">4C, D</figureCitation>
).
</paragraph>
<caption id="B98A27F6223E9A44A1AE4FA1CAAE92A6" doi="10.3897/zookeys.1110.82461.figure4" httpUri="https://binary.pensoft.net/fig/709570" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="A531BF2BC509A8FB434ABCE697743233" pageId="0" pageNumber="1">
<emphasis id="0870A44F29DF9060B003E639A11DD420" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<emphasis id="8D2AC8BD154485FD2B8BF3C2377D0F98" bold="true" pageId="0" pageNumber="1">A</emphasis>
adult of
<taxonomicName id="FB9EC104A66C5A0CDBE18C2E34808D55" lsidName="B. moulmeinica" pageId="0" pageNumber="1" rank="species" species="moulmeinica">
<emphasis id="77C7BD93CB65F1F2A333F6D076040F84" italics="true" pageId="0" pageNumber="1">B. moulmeinica</emphasis>
</taxonomicName>
sp. nov. while climbing slowly on the karst wall after raining at Lun Nya Pagoda
<emphasis id="488146F60C009E385A476CD2832F10E8" bold="true" pageId="0" pageNumber="1">B</emphasis>
mating pairs of
<taxonomicName id="18B615C8ACD825AC54718091AA88A4E0" lsidName="B. versicolor" pageId="0" pageNumber="1" rank="species" species="versicolor">
<emphasis id="123B8A699EC79957AF0307D810F52077" italics="true" pageId="0" pageNumber="1">B. versicolor</emphasis>
</taxonomicName>
sp. nov. on the karst wall at Bayin Nyi Cave
<emphasis id="E1CFF91FF7AA962E49695AACEE271C6F" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="40D7B4A2D533BD5FC391B91BFCD172D9" lsidName="B. versicolor" pageId="0" pageNumber="1" rank="species" species="versicolor">
<emphasis id="DDCF1FA2DBD35F14AFFEA2EE48459C83" italics="true" pageId="0" pageNumber="1">B. versicolor</emphasis>
</taxonomicName>
sp. nov. eaten by the carnivorous snail
<taxonomicName id="F46D2E40E74E7DF6B34E936B35EC954A" authorityName="Siriboon &amp; Panha" authorityYear="2014" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" kingdom="Animalia" lsidName="Carinartemis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="915BD72316B0EF24DC6531529109DBD2" italics="true" pageId="0" pageNumber="1">Carinartemis</emphasis>
</taxonomicName>
sp. on the karst wall at Bayin Nyi Cave
<emphasis id="8816D1D4B41FFEBE67B3F27C05AD9646" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="ADB86539B142AD501892F0AA66ECCDDE" lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis id="F048918487E95D1AF549C873BC8A8385" italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
eaten by the carnivorous snail
<taxonomicName id="3420D1FB3D2DAF452D7342FD9AC39CC6" authorityName="Kobelt" authorityYear="1905" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" kingdom="Animalia" lsidName="Haploptychius" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="73B347F2C94482846DE64AB756E4C1C0" italics="true" pageId="0" pageNumber="1">Haploptychius</emphasis>
</taxonomicName>
sp. on the karst wall at Kaw Gon Cave.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>