205 lines
23 KiB
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205 lines
23 KiB
XML
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<mods:title id="9AC97F39EAA3811B09544DA0D55D4CA8">The first fossil spider (Araneae: Palpimanoidea) from the Lower Jurassic (Grimmen, Germany)</mods:title>
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<mods:namePart id="FBFFED3EE24F9277731884C3A7D4C500">Selden, Paul A.</mods:namePart>
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<mods:namePart id="920ECBFC0CA0BA000D492D6484835232">Dunlop, Jason A.</mods:namePart>
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<mods:title id="AF8BC60027A2D39066CB7ABEE862D73E">Zootaxa</mods:title>
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<mods:part id="8C1235BB5FD4700570309F5FFB312353">
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<mods:date id="74365E6F60D225D2D31A66B905C7F7DD">2014</mods:date>
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<mods:number id="0F80AFADE2B0BC069C1EDF17075409ED">3894</mods:number>
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<treatment id="03AB3E384B57FFECD2B18ED0FB3454D2" ID-DOI="http://doi.org/10.5281/zenodo.6136229" ID-GBIF-Taxon="119597666" ID-Zenodo-Dep="6136229" LSID="urn:lsid:plazi:treatment:03AB3E384B57FFECD2B18ED0FB3454D2" httpUri="http://treatment.plazi.org/id/03AB3E384B57FFECD2B18ED0FB3454D2" lastPageNumber="163" pageId="2" pageNumber="163">
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<subSubSection id="C318DCA54B57FFECD2B18ED0FE6B508C" box="[151,400,648,675]" pageId="2" pageNumber="163" type="nomenclature">
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<paragraph id="8BBD8F2E4B57FFECD2B18ED0FE6B508C" blockId="2.[151,400,648,675]" box="[151,400,648,675]" pageId="2" pageNumber="163">
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<heading id="D0F538424B57FFECD2B18ED0FE6B508C" bold="true" box="[151,400,648,675]" fontSize="11" level="1" pageId="2" pageNumber="163" reason="1">
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<emphasis id="B976533C4B57FFECD2B18ED0FE6B508C" bold="true" box="[151,400,648,675]" pageId="2" pageNumber="163">
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Genus
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<taxonomicName id="4C02F4AD4B57FFECD2C88ED1FEC3508D" box="[238,312,648,674]" class="Arachnida" family="Archaeidae" genus="Seppo" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B976533C4B57FFECD2C88ED1FEC3508D" bold="true" box="[238,312,648,674]" italics="true" pageId="2" pageNumber="163">Seppo</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A245EE474B57FFECD3188ED0FE6B508C" box="[318,400,649,675]" pageId="2" pageNumber="163" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C318DCA54B57FFECD2B18E96FDD550C8" box="[151,558,719,744]" pageId="2" pageNumber="163" type="materials_examined">
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<paragraph id="8BBD8F2E4B57FFECD2B18E96FDD550C8" blockId="2.[151,1437,719,1789]" box="[151,558,719,744]" pageId="2" pageNumber="163">
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||
<emphasis id="B976533C4B57FFECD2B18E96FEC950C7" bold="true" box="[151,306,719,744]" pageId="2" pageNumber="163">
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<typeStatus id="54B9318C4B57FFECD2B18E96FF2950C7" box="[151,210,719,744]" pageId="2" pageNumber="163">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C02F4AD4B57FFECD31F8E89FE1050C7" box="[313,491,720,744]" class="Arachnida" family="Archaeidae" genus="Seppo" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="koponeni" status="sp. nov.">
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<emphasis id="B976533C4B57FFECD31F8E89FE1050C7" box="[313,491,720,744]" italics="true" pageId="2" pageNumber="163">Seppo koponeni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A245EE474B57FFECD3D48E89FDD550C8" box="[498,558,720,743]" pageId="2" pageNumber="163" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C318DCA54B57FFECD2E18EADFE6A5100" pageId="2" pageNumber="163" type="diagnosis">
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||
<paragraph id="8BBD8F2E4B57FFECD2E18EADFE6A5100" blockId="2.[151,1437,719,1789]" pageId="2" pageNumber="163">
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<emphasis id="B976533C4B57FFECD2E18EADFEC15122" bold="true" box="[199,314,756,781]" pageId="2" pageNumber="163">Diagnosis</emphasis>
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. Palpimanoid without a cheliceral foramen; enlarged, forwardly directed leg I; short leg III; curvature of femur IV.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318DCA54B57FFECD2E18F65FC5F517A" box="[199,932,828,853]" pageId="2" pageNumber="163" type="etymology">
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<paragraph id="8BBD8F2E4B57FFECD2E18F65FC5F517A" blockId="2.[151,1437,719,1789]" box="[199,932,828,853]" pageId="2" pageNumber="163">
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<emphasis id="B976533C4B57FFECD2E18F65FEBD517A" bold="true" box="[199,326,828,853]" pageId="2" pageNumber="163">Etymology</emphasis>
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. After
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<taxonomicName id="4C02F4AD4B57FFECD3B08F65FE27517B" box="[406,476,828,852]" class="Arachnida" family="Archaeidae" genus="Seppo" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus" status="gen. nov.">Seppo</taxonomicName>
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Koponen, to celebrate his 70th birthday.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318DCA54B57FFECD2E18F06FB3454D2" pageId="2" pageNumber="163" type="discussion">
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<paragraph id="8BBD8F2E4B57FFECD2E18F06FD2157F2" blockId="2.[151,1437,719,1789]" pageId="2" pageNumber="163">
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<emphasis id="B976533C4B57FFECD2E18F06FECF5157" bold="true" box="[199,308,863,888]" pageId="2" pageNumber="163">Remarks</emphasis>
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. The new genus is accommodated in Palpimanoidea (
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<taxonomicName id="4C02F4AD4B57FFECD19E8F06FBAE5157" box="[952,1109,863,888]" class="Arachnida" family="Palpimanidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Palpimanidae</taxonomicName>
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,
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<taxonomicName id="4C02F4AD4B57FFECD6408F06FAFA5157" box="[1126,1281,863,888]" class="Arachnida" family="Stenochilidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Stenochilidae</taxonomicName>
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,
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<taxonomicName id="4C02F4AD4B57FFECD7378F06FA635157" box="[1297,1432,863,888]" class="Arachnida" family="Huttoniidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Huttoniidae</taxonomicName>
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,
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<taxonomicName id="4C02F4AD4B57FFECD2B18FDDFE7B51B2" box="[151,384,900,925]" class="Arachnida" family="Mecysmaucheniidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Mecysmaucheniidae</taxonomicName>
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,
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<taxonomicName id="4C02F4AD4B57FFECD3B58FDDFDEE51B2" box="[403,533,900,925]" class="Arachnida" family="Archaeidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Archaeidae</taxonomicName>
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, †
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<taxonomicName id="4C02F4AD4B57FFECD0128FDDFCF151B2" box="[564,778,900,925]" class="Arachnida" family="Lagonomegopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Lagonomegopidae</taxonomicName>
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, †
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<taxonomicName id="4C02F4AD4B57FFECD10F8FDDFC3951B2" box="[809,962,900,925]" class="Arachnida" family="Spatiatoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Spatiatoridae</taxonomicName>
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, †Micropalpimanidae) on account of a variety of characters which are commonly found in, but not exclusive to, this superfamily: enlarged chelicerae with peg teeth, and a lack of true teeth, along the cheliceral furrow, the presence of a distal trichobothrium on metatarsus I, paucity of large macrosetae on the legs, curved femur of leg IV, ventral anterior sclerotization and scattered bristles on the opisthosoma. Some Palpimanoidea have large chelicerae, and peg teeth are a characteristic of the superfamily (although these are also found in other spiders, e.g.
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<taxonomicName id="4C02F4AD4B57FFECD14B886EFC17567F" box="[877,1004,1079,1104]" class="Arachnida" family="Mimetidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Mimetidae</taxonomicName>
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). A distal metatarsal trichobothrium is characteristic of mecysmaucheniids and archaeids (
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<bibRefCitation id="EF93F2DF4B57FFECD0CF8805FC6E565B" author="Lehtinen" box="[745,917,1116,1141]" pageId="2" pageNumber="163" refString="Lehtinen, P. T. (1980) Trichobothrial patterns in high level taxonomy of spiders. Proceedings of the VIIIth International Arachnological Congress, Vienna, 1980, 493 - 498." type="proceedings paper" year="1980">Lehtinen 1980</bibRefCitation>
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), although we cannot be sure if others were present on this podomere, A claw on the female pedipalp tarsus occurs in many spiders; in palpimanoids, for example, it is found in
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<taxonomicName id="4C02F4AD4B57FFECD39288FDFC5D5693" authority="O. Pickard-Cambridge, 1881" authorityName="O. Pickard-Cambridge" authorityYear="1881" box="[436,934,1188,1213]" class="Arachnida" family="Archaeidae" genus="Eriauchenius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="B976533C4B57FFECD39288FDFDB25693" box="[436,585,1188,1212]" italics="true" pageId="2" pageNumber="163">Eriauchenius</emphasis>
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<collectingCountry id="F315CFBE4B57FFECD07388FDFD975693" box="[597,620,1188,1212]" name="Iceland" pageId="2" pageNumber="163">O</collectingCountry>
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. Pickard-Cambridge, 1881
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</taxonomicName>
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,
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<taxonomicName id="4C02F4AD4B57FFECD19188FDFA9C5693" authority="Forster & Platnick, 1984" authorityName="Forster & Platnick" authorityYear="1984" box="[951,1383,1188,1213]" class="Arachnida" family="Archaeidae" genus="Afrarchaea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="B976533C4B57FFECD19188FDFBCD5693" box="[951,1078,1188,1212]" italics="true" pageId="2" pageNumber="163">Afrarchaea</emphasis>
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Forster & Platnick, 1984
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</taxonomicName>
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and
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<taxonomicName id="4C02F4AD4B57FFECD2B18891FDA356CF" authority="Forster & Platnick, 1984" authorityName="Forster & Platnick" authorityYear="1984" box="[151,600,1223,1248]" class="Arachnida" family="Archaeidae" genus="Austrarchaea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="B976533C4B57FFECD2B18891FED456CF" box="[151,303,1224,1248]" italics="true" pageId="2" pageNumber="163">Austrarchaea</emphasis>
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Forster & Platnick, 1984
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</taxonomicName>
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(
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<bibRefCitation id="EF93F2DF4B57FFECD04C889EFCBF56CF" author="Wood" box="[618,836,1223,1248]" pageId="2" pageNumber="163" refString="Wood, H. M., Griswold, C. E. & Gillespie, R. G. (2012 a) Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the ' neck' and predatory behaviours of the superfamily Palpimanoidea. Cladistics, 28, 598 - 626. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2012.00411. x" type="journal article" year="2012" yearSuffix="a">
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Wood
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||
<emphasis id="B976533C4B57FFECD09E8890FD0F56CF" box="[696,756,1224,1248]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
|
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2012
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<emphasis id="B976533C4B57FFECD1108890FCBF56CF" box="[822,836,1225,1248]" italics="true" pageId="2" pageNumber="163">a</emphasis>
|
||
</bibRefCitation>
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||
). The lack of macrosetae on the legs is an unusual feature of some spider families. In the new genus, numerous bristles (thin macrosetae) are present. Reduced leg spination occurs throughout Palpimanoidea and only a few other families (
|
||
<bibRefCitation id="EF93F2DF4B57FFECD1F88956FB495707" author="Wood" box="[990,1202,1295,1320]" pageId="2" pageNumber="163" refString="Wood, H. M., Griswold, C. E. & Gillespie, R. G. (2012 a) Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the ' neck' and predatory behaviours of the superfamily Palpimanoidea. Cladistics, 28, 598 - 626. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2012.00411. x" type="journal article" year="2012" yearSuffix="a">
|
||
Wood
|
||
<emphasis id="B976533C4B57FFECD60F8948FB995707" box="[1065,1122,1296,1320]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
|
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2012a
|
||
</bibRefCitation>
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||
). A curved femur IV is characteristic of all living
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<taxonomicName id="4C02F4AD4B57FFECD3F0896DFDAD5762" box="[470,598,1332,1357]" class="Arachnida" family="Archaeidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Archaeidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF93F2DF4B57FFECD043896DFCC35763" author="Wood" box="[613,824,1332,1357]" pageId="2" pageNumber="163" refString="Wood, H. M., Griswold, C. E. & Gillespie, R. G. (2012 a) Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the ' neck' and predatory behaviours of the superfamily Palpimanoidea. Cladistics, 28, 598 - 626. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2012.00411. x" type="journal article" year="2012" yearSuffix="a">
|
||
Wood
|
||
<emphasis id="B976533C4B57FFECD089896CFD1C5763" box="[687,743,1332,1356]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
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||
2012a
|
||
</bibRefCitation>
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||
). Sclerotization on the opisthosoma occurs in a variety of spider families, often in the form of a dorsal scutum; however, ventral sclerotization occurs in the epigynal area in
|
||
<taxonomicName id="4C02F4AD4B57FFECD2918925FEB757BB" box="[183,332,1404,1428]" class="Arachnida" family="Archaeidae" genus="Eriauchenius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976533C4B57FFECD2918925FEB757BB" box="[183,332,1404,1428]" italics="true" pageId="2" pageNumber="163">Eriauchenius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C02F4AD4B57FFECD3AD8925FDF157BB" box="[395,522,1404,1428]" class="Arachnida" family="Archaeidae" genus="Afrarchaea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B976533C4B57FFECD3AD8925FDF157BB" box="[395,522,1404,1428]" italics="true" pageId="2" pageNumber="163">Afrarchaea</emphasis>
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||
</taxonomicName>
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||
, and a reduced palpal claw is present in
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||
<taxonomicName id="4C02F4AD4B57FFECD1D78925FB1557BB" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[1009,1262,1404,1429]" class="Arachnida" family="Stenochilidae" genus="Colopea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B976533C4B57FFECD1D78925FBAB57BB" box="[1009,1104,1404,1428]" italics="true" pageId="2" pageNumber="163">Colopea</emphasis>
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||
Simon, 1893
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</taxonomicName>
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||
and in several mecysmaucheniid genera (
|
||
<bibRefCitation id="EF93F2DF4B57FFECD3E489C6FDB25797" author="Wood" box="[450,585,1439,1464]" pageId="2" pageNumber="163" refString="Wood, H. M. (2008) A revision of the assassin spiders of the Eriauchenius gracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae). Zoological Journal of the Linnean Society, 152 (2), 255 - 296. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2007.00359. x" type="journal article" year="2008">Wood 2008</bibRefCitation>
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||
;
|
||
<bibRefCitation id="EF93F2DF4B57FFECD07089C6FCD15797" author="Wood" box="[598,810,1439,1464]" pageId="2" pageNumber="163" refString="Wood, H. M., Griswold, C. E. & Gillespie, R. G. (2012 a) Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the ' neck' and predatory behaviours of the superfamily Palpimanoidea. Cladistics, 28, 598 - 626. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2012.00411. x" type="journal article" year="2012" yearSuffix="a">
|
||
Wood
|
||
<emphasis id="B976533C4B57FFECD08689F8FD225797" box="[672,729,1440,1464]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
|
||
2012a
|
||
</bibRefCitation>
|
||
). Sparse bristles on the opisthosoma is also a feature of
|
||
<taxonomicName id="4C02F4AD4B57FFECD2B1899DFD2D57F2" authority="Jocque & Dippenaar-Schoeman 2006" authorityName="Jocque & Dippenaar-Schoeman" authorityYear="2006" box="[151,726,1476,1501]" class="Arachnida" family="Archaeidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">
|
||
Archaeidae (
|
||
<bibRefCitation id="EF93F2DF4B57FFECD303899DFD3557F2" author="Jocque" box="[293,718,1476,1501]" pageId="2" pageNumber="163" refString="Jocque, R. & Dippenaar-Schoeman, A. S. (2006) Spider families of the world. Royal Museum for Central Africa, Tervueren, Belgium, 336 pp." type="book" year="2006">Jocqué & Dippenaar-Schoeman 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBD8F2E4B57FFECD2E189BEFB575467" blockId="2.[151,1437,719,1789]" pageId="2" pageNumber="163">
|
||
Arguing against Palpimanoidea is the possibility of patella–tibia autospasy, which is best known from
|
||
<taxonomicName id="4C02F4AD4B57FFECD2B18A55FEDA540A" box="[151,289,1548,1573]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Linyphiidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C02F4AD4B57FFECD3128A55FE5A540A" box="[308,417,1548,1573]" class="Arachnida" family="Pimoidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Pimoidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C02F4AD4B57FFECD3958A55FDC8540A" box="[435,563,1548,1573]" class="Arachnida" family="Filistatidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Filistatidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C02F4AD4B57FFECD0638A55FD2C540A" box="[581,727,1548,1573]" class="Arachnida" family="Leptonetidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Leptonetidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C02F4AD4B57FFECD13B8A55FC60540A" box="[797,923,1548,1573]" class="Arachnida" family="Hersiliidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Hersiliidae</taxonomicName>
|
||
, and some genera in the
|
||
<taxonomicName id="4C02F4AD4B57FFECD6FA8A55FA9E540A" box="[1244,1381,1548,1573]" class="Arachnida" family="Clubionidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Clubionidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C02F4AD4B57FFECD2B18A76FEC75467" box="[151,316,1583,1608]" class="Arachnida" family="Philodromidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Philodromidae</taxonomicName>
|
||
also show a tendency for the legs to separate at this joint (
|
||
<bibRefCitation id="EF93F2DF4B57FFECD1EC8A76FB645467" author="Roth" box="[970,1183,1583,1608]" pageId="2" pageNumber="163" refString="Roth, V. D. & Roth, B. M. (1984) A review of appendotomy in spiders and other arachnids. Bulletin of the British Arachnological Society, 6, 137 - 146." type="journal article" year="1984">Roth & Roth 1984</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD8F2E4B57FFECD2E18A0DFB3454D2" blockId="2.[151,1437,719,1789]" pageId="2" pageNumber="163">
|
||
The genus differs from all palpimanoid families except
|
||
<taxonomicName id="4C02F4AD4B57FFECD16E8A0DFC375442" box="[840,972,1620,1645]" class="Arachnida" family="Huttoniidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Huttoniidae</taxonomicName>
|
||
and the extinct families in the lack of a cheliceral foramen. A bent fourth femur is found in
|
||
<taxonomicName id="4C02F4AD4B57FFECD0C68A2EFC9854BF" box="[736,867,1655,1680]" class="Arachnida" family="Archaeidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="family">Archaeidae</taxonomicName>
|
||
, but the femora of the fossil lack the hump seen in this family (
|
||
<bibRefCitation id="EF93F2DF4B57FFECD3018AC5FDF2549B" author="Wood" box="[295,521,1692,1717]" pageId="2" pageNumber="163" refString="Wood, H. M., Griswold, C. E. & Gillespie, R. G. (2012 a) Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the ' neck' and predatory behaviours of the superfamily Palpimanoidea. Cladistics, 28, 598 - 626. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2012.00411. x" type="journal article" year="2012" yearSuffix="a">
|
||
Wood
|
||
<emphasis id="B976533C4B57FFECD3518AC4FE4F549B" box="[375,436,1692,1716]" italics="true" pageId="2" pageNumber="163">et al.</emphasis>
|
||
2012a
|
||
</bibRefCitation>
|
||
). While the combination of characters suggest Palpimanoidea, there are some characters of the superfamily which cannot be seen in the fossil, e.g. cheliceral gland mound, and sclerotization around the spinnerets. Nor it is not clear to which existing family, if any, the new genus belongs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |