604 lines
87 KiB
XML
604 lines
87 KiB
XML
<document id="AB9A87B3249F4CCF078DC180ED9A57EF" ID-CLB-Dataset="30402" ID-DOI="10.11646/zootaxa.4402.2.3" ID-GBIF-Dataset="7139708e-922d-41d4-ad42-a5d5f7076f1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="10070885" ID-ZooBank="CEA5700E-9F94-42A8-9067-BD8F4D301B9D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522215469332" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2018" docId="03AB2952FFEC5E5B0FBDFB89FC930C1C" docLanguage="en" docName="zootaxa.4402.2.3.pdf.imf" docOrigin="Zootaxa 4402 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Unixenus intragramineus Huynh 2018, sp. n." docType="treatment" docVersion="12" lastPageNumber="292" masterDocId="FF92512AFFEE5E520F2AFFE1FFB20949" masterDocTitle="Two new species of penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam" masterLastPageNumber="302" masterPageNumber="283" pageNumber="285" updateTime="1699558794197" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A15CCE4F07D26C8D6594E8DD451B3C21">
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<mods:title id="7B10E3132A790A3982C8436FC421AD71">Two new species of penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam</mods:title>
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<mods:name id="3E816D88D63CB3C3A1EF7EF280FDC74B" type="personal">
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<mods:roleTerm id="BB1161C50D5C06BA3494774BE7BB6AED">Author</mods:roleTerm>
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<mods:namePart id="7161FB619F980E7FE50910DA6B650F32">Huynh, Cuong</mods:namePart>
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<mods:title id="B09B9F1B5449A956EA46EE5471A30294">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="47650A95A396A0D4CCD293F73FBE49F9">
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<mods:date id="496379A305967FEB07F8A8ED5D24C90D">2018</mods:date>
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<mods:number id="F517F82D8EAEEF8ADF6453121A24A20D">2018-03-28</mods:number>
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<mods:number id="541FA80493C0ECA813A73A8E0B677FDC">4402</mods:number>
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<mods:identifier id="A2AFD6EFD8A2AF3D13E26C965EE93CE9" type="DOI">10.11646/zootaxa.4402.2.3</mods:identifier>
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<mods:identifier id="E1D206149E2F548CEC990C4FF6108C7D" type="GBIF-Dataset">7139708e-922d-41d4-ad42-a5d5f7076f1a</mods:identifier>
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<treatment id="03AB2952FFEC5E5B0FBDFB89FC930C1C" ID-DOI="http://doi.org/10.5281/zenodo.5995384" ID-GBIF-Taxon="142657620" ID-Zenodo-Dep="5995384" LSID="urn:lsid:plazi:treatment:03AB2952FFEC5E5B0FBDFB89FC930C1C" httpUri="http://treatment.plazi.org/id/03AB2952FFEC5E5B0FBDFB89FC930C1C" lastPageId="9" lastPageNumber="292" pageId="2" pageNumber="285">
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<subSubSection id="C318CBCFFFEC5E500FBDFB89FDAA0DCA" box="[151,536,1128,1155]" pageId="2" pageNumber="285" type="nomenclature">
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<paragraph id="8BBD9844FFEC5E500FBDFB89FDAA0DCA" blockId="2.[151,536,1128,1189]" box="[151,536,1128,1155]" pageId="2" pageNumber="285">
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<heading id="D0F52F28FFEC5E500FBDFB89FDAA0DCA" bold="true" box="[151,536,1128,1155]" fontSize="11" level="1" pageId="2" pageNumber="285" reason="1">
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<emphasis id="B9764456FFEC5E500FBDFB89FDAA0DCA" bold="true" box="[151,536,1128,1155]" pageId="2" pageNumber="285">
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<taxonomicName id="4C02E3C7FFEC5E500FBDFB89FE7D0DCB" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[151,463,1128,1154]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
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<emphasis id="B9764456FFEC5E500FBDFB89FE7D0DCB" bold="true" box="[151,463,1128,1154]" italics="true" pageId="2" pageNumber="285">Unixenus intragramineus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A245F92DFFEC5E500EFCFB88FDAA0DCA" box="[470,536,1129,1155]" pageId="2" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C318CBCFFFEC5E500FBDFB6DFF590DED" box="[151,235,1164,1189]" pageId="2" pageNumber="285" type="description">
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<paragraph id="8BBD9844FFEC5E500FBDFB6DFF590DED" blockId="2.[151,536,1128,1189]" box="[151,235,1164,1189]" pageId="2" pageNumber="285">
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(
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<figureCitation id="133984C1FFEC5E500FB5FB6DFF510DED" box="[159,227,1164,1189]" captionStart="FIGURE 2" captionStartId="4.[151,250,575,597]" captionTargetBox="[165,1435,201,552]" captionTargetId="figure@4.[151,1436,193,553]" captionTargetPageId="4" captionText="FIGURE 2. Unixenus intragramineus sp. n., A. a digital image and B. a SEM image." figureDoi="http://doi.org/10.5281/zenodo.10092668" httpUri="https://zenodo.org/record/10092668/files/figure.png" pageId="2" pageNumber="285">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C318CBCFFFEC5E500FBDFB35FE420C8E" pageId="2" pageNumber="285" type="materials_examined">
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<paragraph id="8BBD9844FFEC5E500FBDFB35FB400C12" blockId="2.[151,1437,1236,1983]" pageId="2" pageNumber="285">
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<materialsCitation id="3B6A9219FFEC5E500FBDFB35FB400C12" ID-GBIF-Occurrence="1836914085" collectedFrom="within the stem of a creeping grass (Poaceae) (Fig. 3) growing within the intertidal zone on the beach" collectingDate="1 st July, 2015" collectionCode="QMS" country="Vietnam" county="Male" elevation="5" latitude="10.173006" location="Tranh River shore (Cua Lap hamlet, Duong To commune, Phu Quoc Island" longLatPrecision="1" longitude="103.968925" municipality="Tranh River" pageId="2" pageNumber="285" specimenCode="QMS 108520" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<emphasis id="B9764456FFEC5E500FBDFB35FEB90DA4" bold="true" box="[151,267,1236,1261]" pageId="2" pageNumber="285">
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<typeStatus id="54B926E6FFEC5E500FBDFB35FEB40DA4" box="[151,262,1236,1261]" pageId="2" pageNumber="285" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCount id="9D0453CDFFEC5E500E31FB35FEE50DA4" box="[283,343,1236,1261]" pageId="2" pageNumber="285" type="male">Male</specimenCount>
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,
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<location id="8EDDCE9FFFEC5E500E41FB35FAAB0DA4" LSID="urn:lsid:plazi:treatment:03AB2952FFEC5E5B0FBDFB89FC930C1C:8EDDCE9FFFEC5E500E41FB35FAAB0DA4" box="[363,1305,1236,1261]" country="Vietnam" county="Male" latitude="10.173006" longLatPrecision="1" longitude="103.968925" municipality="Tranh River" name="Tranh River shore (Cua Lap hamlet, Duong To commune, Phu Quoc Island" pageId="2" pageNumber="285" stateProvince="Queensland">Tranh River shore (Cua Lap hamlet, Duong To commune, Phu Quoc Island</location>
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,
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<collectingCountry id="F315D8D4FFEC5E500A07FB35FA230DA4" box="[1325,1425,1236,1261]" name="Vietnam" pageId="2" pageNumber="285">Vietnam</collectingCountry>
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),
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<geoCoordinate id="EE36FE83FFEC5E500FBDFB19FEF20C59" box="[151,320,1272,1296]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="2" pageNumber="285" precision="1" seconds="22.82" value="10.173006">10°10'22.82"N</geoCoordinate>
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,
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<geoCoordinate id="EE36FE83FFEC5E500E61FB16FDB20C59" box="[331,512,1271,1296]" degrees="103" direction="east" minutes="58" orientation="longitude" pageId="2" pageNumber="285" precision="1" seconds="08.14" value="103.968925">103°58'08.14"E</geoCoordinate>
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;
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<elevation id="002F7F77FFEC5E500D21FB16FD140C46" box="[523,678,1271,1296]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="285" unit="m" value="5.0">
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elevation
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<quantity id="4CFA35A1FFEC5E500D52FB16FD140C46" box="[632,678,1271,1296]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="285" unit="m" value="5.0">5 m</quantity>
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</elevation>
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; the specimen was collected by the first author from
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<collectedFrom id="381EDBB9FFEC5E500BDBFB16FBFF0C7E" pageId="2" pageNumber="285">
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within the stem of a creeping grass (
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<taxonomicName id="4C02E3C7FFEC5E500E5FFAFEFE660C7E" box="[373,468,1311,1335]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
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) (
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<figureCitation id="133984C1FFEC5E500EC1FAFFFD9E0C7E" box="[491,556,1310,1335]" captionStart="FIGURE 3" captionStartId="4.[151,250,1879,1901]" captionTargetBox="[264,1323,646,1857]" captionTargetId="figure@4.[264,1323,646,1857]" captionTargetPageId="4" captionText="FIGURE 3. The typical beach habitat of Unixenus intragramineus sp. n. lacking leaf litter. This species was found living within the stem of a creeping grass (Poaceae) growing on an unstable, sandy substrate within the intertidal zone of Tranh River beach, where it is subject to intermittent tidal inundation and is often covered by sand particles. A. The foreshore where U. intragramineus sp. n. was found, the white arrow indicates the grass habitat; B. The grass species showing an aggregation of its runners forming a small clump; C. The grass stems were buried in sand in a beach habitat where no leaf litter was present." figureDoi="http://doi.org/10.5281/zenodo.10092670" httpUri="https://zenodo.org/record/10092670/files/figure.png" pageId="2" pageNumber="285">Fig. 3</figureCitation>
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) growing within the intertidal zone on the beach
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</collectedFrom>
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; on
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<collectingDate id="EFF8476CFFEC5E500B50FAFEFABC0C7E" box="[1146,1294,1307,1335]" pageId="2" pageNumber="285">
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<date id="FFBCBE84FFEC5E500B50FAFEFB7A0C7E" box="[1146,1224,1307,1335]" pageId="2" pageNumber="285">
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1
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<superScript id="7C77350CFFEC5E500BA2FAFAFB200C60" attach="left" box="[1160,1170,1307,1321]" fontSize="6" pageId="2" pageNumber="285">st</superScript>
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July
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</date>
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, 2015
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</collectingDate>
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. Queensland Museum accession number:
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<specimenCode id="DBA4303FFFEC5E500EFDFAA2FDDC0C12" box="[471,622,1346,1371]" pageId="2" pageNumber="285">QMS 108520</specimenCode>
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(deposited in Queensland Museum, Brisbane, Australia).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBD9844FFEC5E500FEDFA87FE420C8E" blockId="2.[151,1437,1236,1983]" pageId="2" pageNumber="285">
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<materialsCitation id="3B6A9219FFEC5E500FEDFA87FE5E0C8E" ID-GBIF-Occurrence="2608822823" collectionCode="QMS" country="Australia" location="Museum" municipality="Brisbane" pageId="2" pageNumber="285" specimenCode="QMS 108521-108530" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Queensland" typeStatus="paratype">
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<emphasis id="B9764456FFEC5E500FEDFA87FEF40C36" bold="true" box="[199,326,1382,1407]" pageId="2" pageNumber="285">
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<typeStatus id="54B926E6FFEC5E500FEDFA87FEF30C36" box="[199,321,1382,1407]" pageId="2" pageNumber="285" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCount id="9D0453CDFFEC5E500E7AFA86FE1B0C36" box="[336,425,1382,1407]" pageId="2" pageNumber="285" type="male">4 males</specimenCount>
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and
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<specimenCount id="9D0453CDFFEC5E500ECCFA87FDE70C36" box="[486,597,1382,1407]" pageId="2" pageNumber="285" type="female">6 females</specimenCount>
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were collected in the same place and on the same date as the holotype. Queensland Museum accession numbers of 10 paratypes:
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<specimenCode id="DBA4303FFFEC5E500C76FA6AFBED0CEA" box="[860,1119,1418,1443]" pageId="2" pageNumber="285">QMS 108521–108530</specimenCode>
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(deposited in Queensland Museum, Brisbane, Australia)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318CBCFFFEC5E500FEDFA33FD200F46" pageId="2" pageNumber="285" type="etymology">
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<paragraph id="8BBD9844FFEC5E500FEDFA33FD200F46" blockId="2.[151,1437,1236,1983]" pageId="2" pageNumber="285">
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<emphasis id="B9764456FFEC5E500FEDFA33FEFC0CA2" bold="true" box="[199,334,1490,1515]" pageId="2" pageNumber="285">Etymology.</emphasis>
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The species is named intragramineus, noun in apposition, as it was found living in the stem of a creeping grass growing in the intertidal zone.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318CBCFFFEC5E500FEDF9FBFCFB0F8A" pageId="2" pageNumber="285" type="diagnosis">
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<paragraph id="8BBD9844FFEC5E500FEDF9FBFCFB0F8A" blockId="2.[151,1437,1236,1983]" pageId="2" pageNumber="285">
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<emphasis id="B9764456FFEC5E500FEDF9FBFEF30F7A" bold="true" box="[199,321,1562,1587]" pageId="2" pageNumber="285">Diagnosis.</emphasis>
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<taxonomicName id="4C02E3C7FFEC5E500E63F9FDFDAB0F7A" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[329,537,1564,1587]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
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<emphasis id="B9764456FFEC5E500E63F9FDFDAB0F7A" box="[329,537,1564,1587]" italics="true" pageId="2" pageNumber="285">U. intragramineus</emphasis>
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</taxonomicName>
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<emphasis id="B9764456FFEC5E500D0BF9FBFDD00F7A" bold="true" box="[545,610,1562,1587]" pageId="2" pageNumber="285">
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<taxonomicNameLabel id="A245F92DFFEC5E500D0BF9FBFDD00F7A" box="[545,610,1562,1587]" pageId="2" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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has the general characteristics of
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<taxonomicName id="4C02E3C7FFEC5E500CCEF9FDFBFC0F7A" authorityName="Jones" authorityYear="1944" box="[996,1102,1564,1587]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
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<emphasis id="B9764456FFEC5E500CCEF9FDFBFC0F7A" box="[996,1102,1564,1587]" italics="true" pageId="2" pageNumber="285">Unixenus</emphasis>
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</taxonomicName>
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and it shares most characters with
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<taxonomicName id="4C02E3C7FFEC5E500FF8F9A1FED40F1E" authorityName="Verhoeff" authorityYear="1924" box="[210,358,1598,1623]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="mjoebergi">
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<emphasis id="B9764456FFEC5E500FF8F9A1FED40F1E" box="[210,358,1598,1623]" italics="true" pageId="2" pageNumber="285">U. mjoebergi</emphasis>
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</taxonomicName>
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. Chaetotaxy: one seta present on prefemur and two setae on femur, a single seta on tibia and tarsus 2. The claw of
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<taxonomicName id="4C02E3C7FFEC5E500EB1F985FDDC0F32" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[411,622,1636,1659]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
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<emphasis id="B9764456FFEC5E500EB1F985FDDC0F32" box="[411,622,1636,1659]" italics="true" pageId="2" pageNumber="285">U. intragramineus</emphasis>
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</taxonomicName>
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||
<emphasis id="B9764456FFEC5E500D52F983FD6B0F32" bold="true" box="[632,729,1634,1659]" pageId="2" pageNumber="285">
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||
<taxonomicNameLabel id="A245F92DFFEC5E500D52F983FD0E0F32" box="[632,700,1634,1659]" pageId="2" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
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is
|
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</emphasis>
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different from that of
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<taxonomicName id="4C02E3C7FFEC5E500CCCF985FBCE0F32" authorityName="Verhoeff" authorityYear="1924" box="[998,1148,1634,1659]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="mjoebergi">
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<emphasis id="B9764456FFEC5E500CCCF985FBCE0F32" box="[998,1148,1634,1659]" italics="true" pageId="2" pageNumber="285">U. mjoebergi</emphasis>
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</taxonomicName>
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in being slender, with a posterior lateral process longer by half the length of the claw. A small anterior lateral process is present, anterior setiform process longer than the claw and with enlarged base.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318CBCFFFEC5E5B0FEDF92FFAD10A8E" lastPageId="9" lastPageNumber="292" pageId="2" pageNumber="285" type="description">
|
||
<paragraph id="8BBD9844FFEC5E500FEDF92FFBB50E66" blockId="2.[151,1437,1236,1983]" pageId="2" pageNumber="285">
|
||
<emphasis id="B9764456FFEC5E500FEDF92FFEEB0FAE" bold="true" box="[199,345,1742,1767]" pageId="2" pageNumber="285">Description.</emphasis>
|
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<emphasis id="B9764456FFEC5E500E75F931FDB30FAE" box="[351,513,1744,1767]" italics="true" pageId="2" pageNumber="285">Measurements</emphasis>
|
||
:
|
||
<typeStatus id="54B926E6FFEC5E500D3BF92FFDCB0FAE" box="[529,633,1742,1767]" pageId="2" pageNumber="285" type="holotype">Holotype</typeStatus>
|
||
male body length
|
||
<quantity id="4CFA35A1FFEC5E500C60F92EFC100FAE" box="[842,930,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="285" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, males (
|
||
<typeStatus id="54B926E6FFEC5E500CD4F92EFBDF0FAE" box="[1022,1133,1743,1767]" pageId="2" pageNumber="285" type="paratype">paratypes</typeStatus>
|
||
) ranging from 1.7–2.0 mm (
|
||
<emphasis id="B9764456FFEC5E500F8AF915FF1C0E42" box="[160,174,1780,1803]" italics="true" pageId="2" pageNumber="285">n</emphasis>
|
||
= 10) and females (
|
||
<emphasis id="B9764456FFEC5E500EBBF915FE2D0E42" box="[401,415,1780,1803]" italics="true" pageId="2" pageNumber="285">n</emphasis>
|
||
= 10) from
|
||
<quantity id="4CFA35A1FFEC5E500D02F913FD010E43" box="[552,691,1778,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="2" pageNumber="285" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 mm</quantity>
|
||
. Caudal bundle of male slightly narrower in width and longer (
|
||
<quantity id="4CFA35A1FFEC5E500A52F913FF760E66" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="285" unit="mm" value="0.6">0.6 mm</quantity>
|
||
) compared to female caudal bundle, the latter wider and shorter (
|
||
<quantity id="4CFA35A1FFEC5E500CB5F8F7FC480E66" box="[927,1018,1814,1839]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="285" unit="mm" value="0.5">0.5 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFEC5E500FEDF8DBFCE60EF6" blockId="2.[151,1437,1236,1983]" pageId="2" pageNumber="285">
|
||
<emphasis id="B9764456FFEC5E500FEDF8DBFEE20E1A" box="[199,336,1850,1875]" italics="true" pageId="2" pageNumber="285">Colouration</emphasis>
|
||
: Head dark brown in eye area connected to 2 brown transverse bands in vertex area. Body milky white in colour, contrasting with light brown pleural trichomes and darker colour in caudal bundle; dark brown marks on latero-posterior rosette trichomes forming a dark band along each side of body laterally; last tergite darkest in colour. Ventral side of body white in colour (
|
||
<figureCitation id="133984C1FFEC5E500C28F847FCF50EF6" box="[770,839,1958,1983]" captionStart="FIGURE 2" captionStartId="4.[151,250,575,597]" captionTargetBox="[165,1435,201,552]" captionTargetId="figure@4.[151,1436,193,553]" captionTargetPageId="4" captionText="FIGURE 2. Unixenus intragramineus sp. n., A. a digital image and B. a SEM image." figureDoi="http://doi.org/10.5281/zenodo.10092668" httpUri="https://zenodo.org/record/10092668/files/figure.png" pageId="2" pageNumber="285">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7DC8CCFFED5E510FBDF850FBC30EAF" ID-DOI="http://doi.org/10.5281/zenodo.10092666" ID-Zenodo-Dep="10092666" httpUri="https://zenodo.org/record/10092666/files/figure.png" pageId="3" pageNumber="286" startId="3.[151,250,1969,1991]" targetBox="[172,1396,196,1923]" targetPageId="3">
|
||
<paragraph id="8BBD9844FFED5E510FBDF850FBC30EAF" blockId="3.[151,1435,1969,2022]" pageId="3" pageNumber="286">
|
||
<emphasis id="B9764456FFED5E510FBDF850FEA60E8E" bold="true" box="[151,276,1969,1991]" pageId="3" pageNumber="286">FIGURE 1.</emphasis>
|
||
Phu Quoc Island with the inserted map of Vietnam indicating the location of Phu Quoc Island. Locations of Da Ban stream (·), Tranh stream (.) and Tranh river beach (▲) on Phu Quoc Island (Not to scale).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7DC8CCFFEA5E560FBDFDDEFC4E0B1C" ID-DOI="http://doi.org/10.5281/zenodo.10092668" ID-Zenodo-Dep="10092668" box="[151,1020,575,597]" httpUri="https://zenodo.org/record/10092668/files/figure.png" pageId="4" pageNumber="287" startId="4.[151,250,575,597]" targetBox="[165,1435,201,552]" targetPageId="4">
|
||
<paragraph id="8BBD9844FFEA5E560FBDFDDEFC4E0B1C" blockId="4.[151,1020,575,597]" box="[151,1020,575,597]" pageId="4" pageNumber="287">
|
||
<emphasis id="B9764456FFEA5E560FBDFDDEFEBF0B1C" bold="true" box="[151,269,575,597]" pageId="4" pageNumber="287">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C02E3C7FFEA5E560E33FDA1FDA90B1C" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[281,539,576,597]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFEA5E560E33FDA1FDA90B1C" box="[281,539,576,597]" italics="true" pageId="4" pageNumber="287">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFEA5E560D0BFDDEFDE80B1C" bold="true" box="[545,602,575,597]" pageId="4" pageNumber="287">
|
||
<taxonomicNameLabel id="A245F92DFFEA5E560D0BFDDEFDE80B1C" box="[545,602,575,597]" pageId="4" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFEA5E560D4DFDDEFDCB0B1C" bold="true" box="[615,633,575,597]" pageId="4" pageNumber="287">A</emphasis>
|
||
. a digital image and
|
||
<emphasis id="B9764456FFEA5E560C67FDDEFCEC0B1C" bold="true" box="[845,862,575,597]" pageId="4" pageNumber="287">B</emphasis>
|
||
. a SEM image.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7DC8CCFFEA5E560FBDF8B6FADF0EA1" ID-DOI="http://doi.org/10.5281/zenodo.10092670" ID-Zenodo-Dep="10092670" httpUri="https://zenodo.org/record/10092670/files/figure.png" pageId="4" pageNumber="287" startId="4.[151,250,1879,1901]" targetBox="[264,1323,646,1857]" targetPageId="4">
|
||
<paragraph id="8BBD9844FFEA5E560FBDF8B6FADF0EA1" blockId="4.[151,1436,1879,2024]" pageId="4" pageNumber="287">
|
||
<emphasis id="B9764456FFEA5E560FBDF8B6FEA70E24" bold="true" box="[151,277,1879,1901]" pageId="4" pageNumber="287">FIGURE 3.</emphasis>
|
||
The typical beach habitat of
|
||
<taxonomicName id="4C02E3C7FFEA5E560D64F8B9FCE10E24" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[590,851,1880,1901]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFEA5E560D64F8B9FCE10E24" box="[590,851,1880,1901]" italics="true" pageId="4" pageNumber="287">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFEA5E560C77F8B6FC2B0E24" bold="true" box="[861,921,1879,1901]" pageId="4" pageNumber="287">
|
||
<taxonomicNameLabel id="A245F92DFFEA5E560C77F8B6FC2B0E24" box="[861,921,1879,1901]" pageId="4" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
lacking leaf litter. This species was found living within the stem of a creeping grass (
|
||
<taxonomicName id="4C02E3C7FFEA5E560D2EF894FDE80EC2" box="[516,602,1909,1931]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="4" pageNumber="287" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||
) growing on an unstable, sandy substrate within the intertidal zone of Tranh River beach, where it is subject to intermittent tidal inundation and is often covered by sand particles.
|
||
<emphasis id="B9764456FFEA5E560BAEF875FB240EE3" bold="true" box="[1156,1174,1940,1962]" pageId="4" pageNumber="287">A</emphasis>
|
||
. The foreshore where
|
||
<taxonomicName id="4C02E3C7FFEA5E560AA9F874FE810E80" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFEA5E560AA9F874FE810E80" italics="true" pageId="4" pageNumber="287">U. intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFEA5E560E13F852FEC30E80" bold="true" box="[313,369,1971,1993]" pageId="4" pageNumber="287">
|
||
<taxonomicNameLabel id="A245F92DFFEA5E560E13F852FEC30E80" box="[313,369,1971,1993]" pageId="4" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was found, the white arrow indicates the grass habitat;
|
||
<emphasis id="B9764456FFEA5E560CB5F852FC020E80" bold="true" box="[927,944,1971,1993]" pageId="4" pageNumber="287">B</emphasis>
|
||
. The grass species showing an aggregation of its runners forming a small clump;
|
||
<emphasis id="B9764456FFEA5E560EF6F833FE5C0EA1" bold="true" box="[476,494,2002,2024]" pageId="4" pageNumber="287">C</emphasis>
|
||
. The grass stems were buried in sand in a beach habitat where no leaf litter was present.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7DC8CCFFEB5E570FBDF9A4FC160FBD" ID-DOI="http://doi.org/10.5281/zenodo.10092672" ID-Zenodo-Dep="10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="5" pageNumber="288" startId="5.[151,250,1605,1627]" targetBox="[151,1425,186,1574]" targetPageId="5">
|
||
<paragraph id="8BBD9844FFEB5E570FBDF9A4FC160FBD" blockId="5.[151,1436,1605,1780]" pageId="5" pageNumber="288">
|
||
<emphasis id="B9764456FFEB5E570FBDF9A4FEBC0F12" bold="true" box="[151,270,1605,1627]" pageId="5" pageNumber="288">FIGURE 4</emphasis>
|
||
. Holotype
|
||
<taxonomicName id="4C02E3C7FFEB5E570EA8F9A7FD340F12" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[386,646,1606,1627]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFEB5E570EA8F9A7FD340F12" box="[386,646,1606,1627]" italics="true" pageId="5" pageNumber="288">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFEB5E570DA7F9A4FD7A0F12" bold="true" box="[653,712,1605,1627]" pageId="5" pageNumber="288">
|
||
<taxonomicNameLabel id="A245F92DFFEB5E570DA7F9A4FD7A0F12" box="[653,712,1605,1627]" pageId="5" pageNumber="288" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFEB5E570DFCF9A4FD5A0F12" bold="true" box="[726,744,1605,1627]" pageId="5" pageNumber="288">A</emphasis>
|
||
. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria
|
||
<emphasis id="B9764456FFEB5E570C68F984FCFC0F33" bold="true" box="[834,846,1637,1658]" italics="true" pageId="5" pageNumber="288">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFEB5E570C77F985FCDB0F33" bold="true" box="[861,873,1636,1658]" italics="true" pageId="5" pageNumber="288">b</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFEB5E570CB7F984FC1A0F33" bold="true" box="[925,936,1637,1658]" italics="true" pageId="5" pageNumber="288">c</emphasis>
|
||
).
|
||
<emphasis id="B9764456FFEB5E570C94F985FC7D0F33" bold="true" box="[958,975,1636,1658]" pageId="5" pageNumber="288">B</emphasis>
|
||
. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets;
|
||
<emphasis id="B9764456FFEB5E570D57F962FD3D0FD0" bold="true" box="[637,655,1667,1689]" pageId="5" pageNumber="288">C</emphasis>
|
||
. Tergite 2 (T2);
|
||
<emphasis id="B9764456FFEB5E570C15F962FCE30FD0" bold="true" box="[831,849,1667,1689]" pageId="5" pageNumber="288">D</emphasis>
|
||
. Tergite 10 – the last tergite;
|
||
<emphasis id="B9764456FFEB5E570BA2F962FB2B0FD0" bold="true" box="[1160,1177,1667,1689]" pageId="5" pageNumber="288">E</emphasis>
|
||
. trichobothria
|
||
<emphasis id="B9764456FFEB5E570A19F962FA8D0FD1" bold="true" box="[1331,1343,1667,1688]" italics="true" pageId="5" pageNumber="288">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFEB5E570A65F963FAE90FD1" bold="true" box="[1359,1371,1666,1688]" italics="true" pageId="5" pageNumber="288">b</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFEB5E570ABBF962FA2E0FD1" bold="true" box="[1425,1436,1667,1688]" italics="true" pageId="5" pageNumber="288">c</emphasis>
|
||
(right hand side) (trichobothrium
|
||
<emphasis id="B9764456FFEB5E570D29F943FDBD0FFE" bold="true" box="[515,527,1698,1719]" italics="true" pageId="5" pageNumber="288">a</emphasis>
|
||
, posterior position, trichobothrium
|
||
<emphasis id="B9764456FFEB5E570CBBF940FC2F0FFE" bold="true" box="[913,925,1697,1719]" italics="true" pageId="5" pageNumber="288">b</emphasis>
|
||
, lateral position and trichobothrium
|
||
<emphasis id="B9764456FFEB5E570A05F943FA880FFE" bold="true" box="[1327,1338,1698,1719]" italics="true" pageId="5" pageNumber="288">c</emphasis>
|
||
, anterior position);
|
||
<emphasis id="B9764456FFEB5E570FD4F921FEB90F9F" bold="true" box="[254,267,1728,1750]" pageId="5" pageNumber="288">F</emphasis>
|
||
. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla;
|
||
<emphasis id="B9764456FFEB5E570BF9F921FB560F9F" bold="true" box="[1235,1252,1728,1750]" pageId="5" pageNumber="288">G</emphasis>
|
||
. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD9844FFEB5E540FEDF8FDFF7A0854" blockId="5.[151,1436,1820,2025]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="289" pageId="5" pageNumber="288">
|
||
<emphasis id="B9764456FFEB5E570FEDF8FDFEB60E7C" box="[199,260,1820,1845]" italics="true" pageId="5" pageNumber="288">Head</emphasis>
|
||
: 8 ommatidia on each side: 4 dorsal, 4 lateral (1 anterior, 2 medial and 1 posterior). Vertex with 2 posterior trichome groups and a large gap between them, each group with 2 rows. Anterior, oblique row with trichome sockets of similar size. Posterior row with slightly smaller trichome sockets. A narrow space between anterior and posterior rows (
|
||
<figureCitation id="133984C1FFEB5E570EC8F869FDF30EE9" box="[482,577,1928,1953]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="5" pageNumber="288">Figs 4A</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFEB5E570D57F869FD110EE8" box="[637,675,1928,1953]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1852]" captionTargetBox="[151,1436,331,1809]" captionTargetId="figure@6.[151,1436,329,1809]" captionTargetPageId="6" captionText="FIGURE 5. Scanning Electron Microscope images of Unixenus intragramineus sp. n.: A. Head, the posterior vertex trichomes group (pv), collum (Co) and tergite 2 (T2); B. Trichobothria a, b and c; C. Labrum showed 4 + 4 lamellae (l); D. Labrum with the pointed spherical papillae (Psp) on its surface; E. The male caudal bundle structure showing the dorsal ornamental trichome sockets (ots), barbate trichome sockets (bts), trichome socket free tissue (tsft) and caudal trichome sockets (cts) with some caudal trichomes (ct) attached. F. The caudal bundle structure of females showing a difference in the arrangement with caudal trichome sockets (cts) located dorsally, barbate trichome sockets (bts) with the largest size in the centre, trichome socket free tissue (tsft) and nest trichome sockets (nts) that were smaller located centro-ventrally." figureDoi="http://doi.org/10.5281/zenodo.10092674" httpUri="https://zenodo.org/record/10092674/files/figure.png" pageId="5" pageNumber="288">5A</figureCitation>
|
||
).
|
||
<typeStatus id="54B926E6FFEB5E570D90F869FC900EE8" box="[698,802,1928,1953]" pageId="5" pageNumber="288" type="holotype">Holotype</typeStatus>
|
||
with 13+13 trichome sockets in anterior row and 7+7 trichome sockets in posterior one.
|
||
<typeStatus id="54B926E6FFEB5E570D31F84CFD3A0E8C" box="[539,648,1965,1989]" pageId="5" pageNumber="288" type="paratype">Paratypes</typeStatus>
|
||
with variations common in this species regardless of sex, from 10–16 (anterior rows) and 5–8 (posterior rows). Trichobothria: trichobothrium
|
||
<emphasis id="B9764456FFEB5E570C96F831FC780EAE" bold="true" box="[956,970,2000,2023]" italics="true" pageId="5" pageNumber="288">a</emphasis>
|
||
(posterior position) and trichobothrium
|
||
<emphasis id="B9764456FFEB5E570AA4F82EFA2E0EA1" bold="true" box="[1422,1436,1999,2024]" italics="true" pageId="5" pageNumber="288">b</emphasis>
|
||
(lateral position) typically thin sensory hairs with narrow cylindrical funicles; trichobothrium
|
||
<emphasis id="B9764456FFE85E540B96FF79FB7A09E6" bold="true" box="[1212,1224,152,175]" italics="true" pageId="6" pageNumber="289">c</emphasis>
|
||
(anterior position) similar in structure to trichobothria
|
||
<emphasis id="B9764456FFE85E540D04FF5DFD8E099A" bold="true" box="[558,572,188,211]" italics="true" pageId="6" pageNumber="289">a</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE85E540D5CFF5AFD36099D" bold="true" box="[630,644,187,212]" italics="true" pageId="6" pageNumber="289">b</emphasis>
|
||
, but its funicle slightly enlarged distally. All these trichobothria (
|
||
<emphasis id="B9764456FFE85E540A5AFF5DFACC099A" bold="true" box="[1392,1406,188,211]" italics="true" pageId="6" pageNumber="289">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE85E540AA4FF5AFA2E099D" bold="true" box="[1422,1436,187,212]" italics="true" pageId="6" pageNumber="289">b</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE85E540FEDFF01FF6109BE" bold="true" box="[199,211,224,247]" italics="true" pageId="6" pageNumber="289">c</emphasis>
|
||
) equal in socket size and forming an isosceles triangle with equal distance between
|
||
<emphasis id="B9764456FFE85E540BB9FF3EFB1D09B1" bold="true" box="[1171,1199,223,248]" italics="true" pageId="6" pageNumber="289">ab</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE85E540BC2FF3EFAB109B1" bold="true" box="[1256,1283,223,248]" italics="true" pageId="6" pageNumber="289">bc</emphasis>
|
||
(
|
||
<figureCitation id="133984C1FFE85E540A38FF3EFAD909B1" box="[1298,1387,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="6" pageNumber="289">Figs 4E</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE85E540FBDFEE5FF080854" box="[151,186,260,285]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1852]" captionTargetBox="[151,1436,331,1809]" captionTargetId="figure@6.[151,1436,329,1809]" captionTargetPageId="6" captionText="FIGURE 5. Scanning Electron Microscope images of Unixenus intragramineus sp. n.: A. Head, the posterior vertex trichomes group (pv), collum (Co) and tergite 2 (T2); B. Trichobothria a, b and c; C. Labrum showed 4 + 4 lamellae (l); D. Labrum with the pointed spherical papillae (Psp) on its surface; E. The male caudal bundle structure showing the dorsal ornamental trichome sockets (ots), barbate trichome sockets (bts), trichome socket free tissue (tsft) and caudal trichome sockets (cts) with some caudal trichomes (ct) attached. F. The caudal bundle structure of females showing a difference in the arrangement with caudal trichome sockets (cts) located dorsally, barbate trichome sockets (bts) with the largest size in the centre, trichome socket free tissue (tsft) and nest trichome sockets (nts) that were smaller located centro-ventrally." figureDoi="http://doi.org/10.5281/zenodo.10092674" httpUri="https://zenodo.org/record/10092674/files/figure.png" pageId="6" pageNumber="289">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7DC8CCFFE85E540FBDF8C7FC480EBD" ID-DOI="http://doi.org/10.5281/zenodo.10092674" ID-Zenodo-Dep="10092674" httpUri="https://zenodo.org/record/10092674/files/figure.png" pageId="6" pageNumber="289" startId="6.[151,250,1830,1852]" targetBox="[151,1436,331,1809]" targetPageId="6">
|
||
<paragraph id="8BBD9844FFE85E540FBDF8C7FC480EBD" blockId="6.[151,1436,1830,2036]" pageId="6" pageNumber="289">
|
||
<emphasis id="B9764456FFE85E540FBDF8C7FEBE0E75" bold="true" box="[151,268,1830,1852]" pageId="6" pageNumber="289">FIGURE 5</emphasis>
|
||
. Scanning Electron Microscope images of
|
||
<taxonomicName id="4C02E3C7FFE85E540D95F8C6FC720E75" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[703,960,1831,1852]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE85E540D95F8C6FC720E75" box="[703,960,1831,1852]" italics="true" pageId="6" pageNumber="289">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE85E540CEDF8C7FBB20E75" bold="true" box="[967,1024,1830,1852]" pageId="6" pageNumber="289">
|
||
<taxonomicNameLabel id="A245F92DFFE85E540CEDF8C7FBB20E75" box="[967,1024,1830,1852]" pageId="6" pageNumber="289" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B9764456FFE85E540B27F8C7FBAD0E75" bold="true" box="[1037,1055,1830,1852]" pageId="6" pageNumber="289">A</emphasis>
|
||
. Head, the posterior vertex trichomes group (pv), collum (Co) and tergite 2 (T2);
|
||
<emphasis id="B9764456FFE85E540D78F8A4FDD10E12" bold="true" box="[594,611,1861,1883]" pageId="6" pageNumber="289">B</emphasis>
|
||
. Trichobothria
|
||
<emphasis id="B9764456FFE85E540C2AF8A4FCBF0E13" bold="true" box="[768,781,1861,1882]" italics="true" pageId="6" pageNumber="289">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE85E540C30F8A4FC950E12" bold="true" box="[794,807,1861,1883]" italics="true" pageId="6" pageNumber="289">b</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE85E540C72F8A4FCD10E13" bold="true" box="[856,867,1861,1882]" italics="true" pageId="6" pageNumber="289">c</emphasis>
|
||
;
|
||
<emphasis id="B9764456FFE85E540C5BF8A4FC310E12" bold="true" box="[881,899,1861,1883]" pageId="6" pageNumber="289">C</emphasis>
|
||
. Labrum showed 4 + 4 lamellae (l);
|
||
<emphasis id="B9764456FFE85E540BD3F8A4FAB90E12" bold="true" box="[1273,1291,1861,1883]" pageId="6" pageNumber="289">D</emphasis>
|
||
. Labrum with the pointed spherical papillae (Psp) on its surface;
|
||
<emphasis id="B9764456FFE85E540DB9F882FD160E30" bold="true" box="[659,676,1891,1913]" pageId="6" pageNumber="289">E</emphasis>
|
||
. The male caudal bundle structure showing the dorsal ornamental trichome sockets (ots), barbate trichome sockets (bts), trichome socket free tissue (tsft) and caudal trichome sockets (cts) with some caudal trichomes (ct) attached.
|
||
<emphasis id="B9764456FFE85E540EFCF840FE510EFE" bold="true" box="[470,483,1953,1975]" pageId="6" pageNumber="289">F</emphasis>
|
||
. The caudal bundle structure of females showing a difference in the arrangement with caudal trichome sockets (cts) located dorsally, barbate trichome sockets (bts) with the largest size in the centre, trichome socket free tissue (tsft) and nest trichome sockets (nts) that were smaller located centro-ventrally.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7DC8CCFFE95E550FBDFBE3FC440D1F" ID-DOI="http://doi.org/10.5281/zenodo.10092676" ID-Zenodo-Dep="10092676" httpUri="https://zenodo.org/record/10092676/files/figure.png" pageId="7" pageNumber="290" startId="7.[151,250,1026,1048]" targetBox="[164,1414,201,1007]" targetPageId="7">
|
||
<paragraph id="8BBD9844FFE95E550FBDFBE3FC440D1F" blockId="7.[151,1436,1026,1110]" pageId="7" pageNumber="290">
|
||
<emphasis id="B9764456FFE95E550FBDFBE3FEBF0D51" bold="true" box="[151,269,1026,1048]" pageId="7" pageNumber="290">FIGURE 6</emphasis>
|
||
.
|
||
<emphasis id="B9764456FFE95E550E30FBE3FE9E0D51" bold="true" box="[282,300,1026,1048]" pageId="7" pageNumber="290">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C02E3C7FFE95E550E10FBE2FD8E0D51" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[314,572,1027,1048]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE95E550E10FBE2FD8E0D51" box="[314,572,1027,1048]" italics="true" pageId="7" pageNumber="290">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE95E550D69FBE3FDCF0D51" bold="true" box="[579,637,1026,1048]" pageId="7" pageNumber="290">
|
||
<taxonomicNameLabel id="A245F92DFFE95E550D69FBE3FDCF0D51" box="[579,637,1026,1048]" pageId="7" pageNumber="290" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, scanning electron microscope image of the antennal articles VI, VII and VIII with 4 sensory cones (sc);
|
||
<emphasis id="B9764456FFE95E550E8EFBC0FE070D7E" bold="true" box="[420,437,1057,1079]" pageId="7" pageNumber="290">B</emphasis>
|
||
. Arrangement of sensilla on the antennal article VII from the holotype;
|
||
<emphasis id="B9764456FFE95E550BA6FBC0FB2C0D7E" bold="true" box="[1164,1182,1057,1079]" pageId="7" pageNumber="290">C</emphasis>
|
||
. Sensilla on the antennal article VI. (c: conical sensillum, T: thick bacilliform sensillum, s: setiform sensillum)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD9844FFE95E550FEDFB60FDE50C39" blockId="7.[151,1437,1152,2005]" pageId="7" pageNumber="290">
|
||
<emphasis id="B9764456FFE95E550FEDFB60FE830DD1" box="[199,305,1153,1176]" italics="true" pageId="7" pageNumber="290">Antennae</emphasis>
|
||
: 8 articles and 4 sensory cones (
|
||
<figureCitation id="133984C1FFE95E550D8AFB61FD480DD0" box="[672,762,1152,1177]" captionStart="FIGURE 6" captionStartId="7.[151,250,1026,1048]" captionTargetBox="[164,1414,201,1007]" captionTargetId="figure@7.[164,1427,196,1017]" captionTargetPageId="7" captionText="FIGURE 6. A. Unixenus intragramineus sp. n., scanning electron microscope image of the antennal articles VI, VII and VIII with 4 sensory cones (sc); B. Arrangement of sensilla on the antennal article VII from the holotype; C. Sensilla on the antennal article VI. (c: conical sensillum, T: thick bacilliform sensillum, s: setiform sensillum)" figureDoi="http://doi.org/10.5281/zenodo.10092676" httpUri="https://zenodo.org/record/10092676/files/figure.png" pageId="7" pageNumber="290">Fig. 6A</figureCitation>
|
||
), characteristics typical of
|
||
<taxonomicName id="4C02E3C7FFE95E550B03FB61FB050DD0" authorityName="Lucas" authorityYear="1840" box="[1065,1207,1152,1177]" class="Diplopoda" family="Polyxenidae" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="290" phylum="Arthropoda" rank="family">Polyxenidae</taxonomicName>
|
||
. Antennal article VI with 3 thick bacilliform sensilla (T) of different lengths: a short sensillum anteriorly (T1), the longest intermediate sensillum (T2), and a sensillum of medium length (T3). A setiform sensillum (s) present between T1 and T2, and a conical sensillum posteriorly (c) (
|
||
<figureCitation id="133984C1FFE95E550D3FFB0DFDDC0C4C" box="[533,622,1260,1285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1026,1048]" captionTargetBox="[164,1414,201,1007]" captionTargetId="figure@7.[164,1427,196,1017]" captionTargetPageId="7" captionText="FIGURE 6. A. Unixenus intragramineus sp. n., scanning electron microscope image of the antennal articles VI, VII and VIII with 4 sensory cones (sc); B. Arrangement of sensilla on the antennal article VII from the holotype; C. Sensilla on the antennal article VI. (c: conical sensillum, T: thick bacilliform sensillum, s: setiform sensillum)" figureDoi="http://doi.org/10.5281/zenodo.10092676" httpUri="https://zenodo.org/record/10092676/files/figure.png" pageId="7" pageNumber="290">Fig. 6C</figureCitation>
|
||
). Antennal article VII with 2 thick bacilliform sensilla (T), anterior one being slightly longer than the one located posteriorly, with one setiform sensillum (s) between them and a conical sensillum (c) located in posterior position (
|
||
<figureCitation id="133984C1FFE95E550D8AFAD5FD4C0C04" box="[672,766,1332,1357]" captionStart="FIGURE 6" captionStartId="7.[151,250,1026,1048]" captionTargetBox="[164,1414,201,1007]" captionTargetId="figure@7.[164,1427,196,1017]" captionTargetPageId="7" captionText="FIGURE 6. A. Unixenus intragramineus sp. n., scanning electron microscope image of the antennal articles VI, VII and VIII with 4 sensory cones (sc); B. Arrangement of sensilla on the antennal article VII from the holotype; C. Sensilla on the antennal article VI. (c: conical sensillum, T: thick bacilliform sensillum, s: setiform sensillum)" figureDoi="http://doi.org/10.5281/zenodo.10092676" httpUri="https://zenodo.org/record/10092676/files/figure.png" pageId="7" pageNumber="290">Fig. 6B</figureCitation>
|
||
). This pattern of sensilla on the antennal article VII is commonly seen in all
|
||
<taxonomicName id="4C02E3C7FFE95E550EA4FAB8FE4A0C39" authorityName="Jones" authorityYear="1944" box="[398,504,1369,1392]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="290" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE95E550EA4FAB8FE4A0C39" box="[398,504,1369,1392]" italics="true" pageId="7" pageNumber="290">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE95E550FEDFA9DFACD0C95" blockId="7.[151,1437,1152,2005]" pageId="7" pageNumber="290">
|
||
<emphasis id="B9764456FFE95E550FEDFA9DFED50CDC" box="[199,359,1404,1429]" italics="true" pageId="7" pageNumber="290">Clypeolabrum</emphasis>
|
||
:
|
||
<typeStatus id="54B926E6FFE95E550E5CFA9DFE6C0CDC" box="[374,478,1404,1429]" pageId="7" pageNumber="290" type="holotype">Holotype</typeStatus>
|
||
with 10 setae at posterior margin, these setae being half the width of labrum. Setae in
|
||
<typeStatus id="54B926E6FFE95E550FBDFA40FEB00CF0" box="[151,258,1441,1465]" pageId="7" pageNumber="290" type="paratype">paratypes</typeStatus>
|
||
ranging from
|
||
<quantity id="4CFA35A1FFE95E550E8DFA41FDB90CF0" box="[423,523,1440,1465]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="7" pageNumber="290" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
|
||
both sexes. Labral surface with spherical papillae with pointed ends, reduced in size posteriorly. Anterior margin of labrum with 4+4 lamellae on each side of median cleft (
|
||
<figureCitation id="133984C1FFE95E550BB0FA25FB470C95" box="[1178,1269,1476,1501]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="7" pageNumber="290">Figs 4G</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE95E550A06FA25FAC00C95" box="[1324,1394,1476,1501]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1852]" captionTargetBox="[151,1436,331,1809]" captionTargetId="figure@6.[151,1436,329,1809]" captionTargetPageId="6" captionText="FIGURE 5. Scanning Electron Microscope images of Unixenus intragramineus sp. n.: A. Head, the posterior vertex trichomes group (pv), collum (Co) and tergite 2 (T2); B. Trichobothria a, b and c; C. Labrum showed 4 + 4 lamellae (l); D. Labrum with the pointed spherical papillae (Psp) on its surface; E. The male caudal bundle structure showing the dorsal ornamental trichome sockets (ots), barbate trichome sockets (bts), trichome socket free tissue (tsft) and caudal trichome sockets (cts) with some caudal trichomes (ct) attached. F. The caudal bundle structure of females showing a difference in the arrangement with caudal trichome sockets (cts) located dorsally, barbate trichome sockets (bts) with the largest size in the centre, trichome socket free tissue (tsft) and nest trichome sockets (nts) that were smaller located centro-ventrally." figureDoi="http://doi.org/10.5281/zenodo.10092674" httpUri="https://zenodo.org/record/10092674/files/figure.png" pageId="7" pageNumber="290">5C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE95E550FEDFA09FB5A0F6D" blockId="7.[151,1437,1152,2005]" pageId="7" pageNumber="290">
|
||
<emphasis id="B9764456FFE95E550FEDFA09FD840F48" box="[199,566,1512,1537]" italics="true" pageId="7" pageNumber="290">Lateral palp of gnathochilarium</emphasis>
|
||
: Lateral palp 2.5 times as long as medial palp. 13 conical sensilla on lateral palp, medial palp with 22 slender conical sensilla observed in all
|
||
<typeStatus id="54B926E6FFE95E550C58F9ECFC6C0F6C" box="[882,990,1549,1573]" pageId="7" pageNumber="290" type="paratype">paratypes</typeStatus>
|
||
and
|
||
<typeStatus id="54B926E6FFE95E550B3EF9EDFBC40F6C" box="[1044,1142,1548,1573]" pageId="7" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="133984C1FFE95E550BAFF9EDFB690F6D" box="[1157,1243,1548,1573]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="7" pageNumber="290">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE95E5A0FEDF9D1FCEC099C" blockId="7.[151,1437,1152,2005]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="291" pageId="7" pageNumber="290">
|
||
<emphasis id="B9764456FFE95E550FEDF9D1FEBA0F00" box="[199,264,1584,1609]" italics="true" pageId="7" pageNumber="290">Trunk</emphasis>
|
||
: Body with 10 segments, 9 pleural projections, excluding both telson and caudal bundle; 13 pairs of legs. Collum (tergite 1, smallest) with trichome sockets arranged in 2 oval shapes laterally, opposite each other connected by a posterior row of trichome sockets forming a line with a gap in middle. Collum, a tergite only with lateral protuberances, bearing a small number of trichome sockets. In
|
||
<typeStatus id="54B926E6FFE95E550CE4F97DFB810FFC" box="[974,1075,1692,1717]" pageId="7" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
, collum with 46 (L) or 48 (R) trichome sockets, lateral protuberances with 5 trichome sockets on each side (
|
||
<figureCitation id="133984C1FFE95E550B1EF921FB230F91" box="[1076,1169,1728,1753]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="7" pageNumber="290">Fig. 4B</figureCitation>
|
||
). Trichome sockets on collum varying in
|
||
<typeStatus id="54B926E6FFE95E550E4CF904FE630FB4" box="[358,465,1765,1789]" pageId="7" pageNumber="290" type="paratype">paratypes</typeStatus>
|
||
and ranging from 42–60, trichome sockets ranging from 4–7 on lateral protuberances. All other tergites, from 2 to 10, each with a pair of pleural projections located anterolaterally. Tergal trichome socket arrangements from tergites 2 to 9 typically with 4 undefined rows of tergal trichomes arranged in two lateroposterior clusters, either side of midline. Anterior row often uneven, intermediate rows rarely in defined rows and posterior row of trichomes continuous and evenly distributed, forming a line with a medial gap. Tergite 10 exceptional in trichome sockets being smaller and denser. Almost no space between lateral rosette trichome sockets and those in posterior row. Trichome sockets of tergite
|
||
<quantity id="4CFA35A1FFE95E550C2BF85DFC9E0E9C" box="[769,812,1980,2005]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="290" unit="in" value="2.0">2 in</quantity>
|
||
<typeStatus id="54B926E6FFE95E550C19F85DFC270E9C" box="[819,917,1980,2005]" pageId="7" pageNumber="290" type="holotype">holotype</typeStatus>
|
||
numbered 52 (L) and 54 (R) (
|
||
<figureCitation id="133984C1FFE95E550BC3F85DFAF20E9D" box="[1257,1344,1980,2005]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="7" pageNumber="290">Fig. 4C</figureCitation>
|
||
), tergite 10 with 50 (L) and 49 (R) (
|
||
<figureCitation id="133984C1FFE65E5A0EF9FF76FD8209F9" box="[467,560,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1605,1627]" captionTargetBox="[151,1425,186,1574]" captionTargetId="figure@5.[151,1436,181,1584]" captionTargetPageId="5" captionText="FIGURE 4. Holotype Unixenus intragramineus sp. n., A. Head capsule with 8 ommatidia on each side, the posterior vertex trichome groups (pv) and 3 sockets of trichobothria (trichobothria a, b and c). B. Collum (Co) and lateral protuberances (Lp) showing the arrangement of trichome sockets; C. Tergite 2 (T2); D. Tergite 10 – the last tergite; E. trichobothria a, b and c (right hand side) (trichobothrium a, posterior position, trichobothrium b, lateral position and trichobothrium c, anterior position); F. Gnathochilarium: the lateral palp (LP) with 13 sensilla and medial palp (MP) with 22 sensilla; G. Labrum showing the pointed spherical papillae on its surface and the structures 4 + 4 lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092672" httpUri="https://zenodo.org/record/10092672/files/figure.png" pageId="8" pageNumber="291">Fig. 4D</figureCitation>
|
||
). In contrast, trichome sockets on tergite
|
||
<quantity id="4CFA35A1FFE65E5A0B22FF79FB8709F9" box="[1032,1077,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="291" unit="in" value="2.0">2 in</quantity>
|
||
<typeStatus id="54B926E6FFE65E5A0B14FF79FB1B09F9" box="[1086,1193,152,176]" pageId="8" pageNumber="291" type="paratype">paratypes</typeStatus>
|
||
ranging from 46–67, whereas tergite 10 containing between 53–66 trichome sockets.
|
||
</paragraph>
|
||
<caption id="DF7DC8CCFFE65E5A0FBDF99DFA830E03" ID-DOI="http://doi.org/10.5281/zenodo.10092678" ID-Zenodo-Dep="10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="8" pageNumber="291" startId="8.[151,250,1660,1682]" targetBox="[169,1424,284,1628]" targetPageId="8">
|
||
<paragraph id="8BBD9844FFE65E5A0FBDF99DFA830E03" blockId="8.[151,1436,1660,1866]" pageId="8" pageNumber="291">
|
||
<emphasis id="B9764456FFE65E5A0FBDF99DFEBF0FDB" bold="true" box="[151,269,1660,1682]" pageId="8" pageNumber="291">FIGURE 7</emphasis>
|
||
.
|
||
<emphasis id="B9764456FFE65E5A0E31F99DFE9F0FDB" bold="true" box="[283,301,1660,1682]" pageId="8" pageNumber="291">A</emphasis>
|
||
. Holotype
|
||
<taxonomicName id="4C02E3C7FFE65E5A0E8AF99FFD110FDA" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[416,675,1662,1683]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE65E5A0E8AF99FFD110FDA" box="[416,675,1662,1683]" italics="true" pageId="8" pageNumber="291">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A245F92DFFE65E5A0D80F99DFD560FDB" box="[682,740,1660,1682]" pageId="8" pageNumber="291" rank="species">
|
||
<emphasis id="B9764456FFE65E5A0D80F99DFD6C0FDB" bold="true" box="[682,734,1660,1682]" pageId="8" pageNumber="291">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (
|
||
<emphasis id="B9764456FFE65E5A0B9FF97AFB740FF8" bold="true" box="[1205,1222,1691,1713]" pageId="8" pageNumber="291">B</emphasis>
|
||
) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (
|
||
<emphasis id="B9764456FFE65E5A0BB2F939FB180FA7" bold="true" box="[1176,1194,1752,1774]" pageId="8" pageNumber="291">C</emphasis>
|
||
); seta on tibia (
|
||
<emphasis id="B9764456FFE65E5A0A79F939FAD70FA7" bold="true" box="[1363,1381,1752,1774]" pageId="8" pageNumber="291">D</emphasis>
|
||
); the setiform seta on tarsus 2 (
|
||
<emphasis id="B9764456FFE65E5A0E8AF916FE030E44" bold="true" box="[416,433,1783,1805]" pageId="8" pageNumber="291">E</emphasis>
|
||
).
|
||
<emphasis id="B9764456FFE65E5A0EEDF916FE660E44" bold="true" box="[455,468,1783,1805]" pageId="8" pageNumber="291">F</emphasis>
|
||
. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side).
|
||
<emphasis id="B9764456FFE65E5A0A7FF8F7FAD40E65" bold="true" box="[1365,1382,1814,1836]" pageId="8" pageNumber="291">G</emphasis>
|
||
. The dorsal ornamental trichome sockets of trichomes
|
||
<emphasis id="B9764456FFE65E5A0DA0F8D4FD240E03" bold="true" box="[650,662,1845,1866]" italics="true" pageId="8" pageNumber="291">a</emphasis>
|
||
(
|
||
<emphasis id="B9764456FFE65E5A0D8FF8D4FD000E03" bold="true" box="[677,690,1845,1866]" italics="true" pageId="8" pageNumber="291">a</emphasis>
|
||
), trichome
|
||
<emphasis id="B9764456FFE65E5A0C0DF8D5FC810E03" bold="true" box="[807,819,1844,1866]" italics="true" pageId="8" pageNumber="291">b</emphasis>
|
||
(
|
||
<emphasis id="B9764456FFE65E5A0C69F8D5FCE20E03" bold="true" box="[835,848,1844,1866]" italics="true" pageId="8" pageNumber="291">b</emphasis>
|
||
), trichomes
|
||
<emphasis id="B9764456FFE65E5A0CE5F8D4FC680E03" bold="true" box="[975,986,1845,1866]" italics="true" pageId="8" pageNumber="291">c</emphasis>
|
||
(
|
||
<emphasis id="B9764456FFE65E5A0CC3F8D4FC460E03" bold="true" box="[1001,1012,1845,1866]" italics="true" pageId="8" pageNumber="291">c</emphasis>
|
||
) and the circular indentation
|
||
<emphasis id="B9764456FFE65E5A0A34F8D5FA990E03" bold="true" box="[1310,1323,1844,1866]" italics="true" pageId="8" pageNumber="291">d</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD9844FFE65E5B0FEDF899FBE70809" blockId="8.[151,1436,1911,2008]" lastBlockId="9.[151,1437,151,1365]" lastPageId="9" lastPageNumber="292" pageId="8" pageNumber="291">
|
||
<emphasis id="B9764456FFE65E5A0FEDF899FF4E0EC6" box="[199,252,1912,1935]" italics="true" pageId="8" pageNumber="291">Legs</emphasis>
|
||
: Leg segments are named following
|
||
<bibRefCitation id="EF93E5B5FFE65E5A0DB1F899FCF50ED9" author="Manton, S. M." box="[667,839,1911,1936]" pageId="8" pageNumber="291" pagination="153 - 187" refId="ref11810" refString="Manton, S. M. (1956) The Evolution of Arthropodan Locomotory Mechanisms-Part 5. The Structure, Habits and Evolution of the Pselaphognatha (Diplopoda). Journal of the Linnean Society London, 43, 153 - 187. https: // doi. org / 10.1111 / j. 1096 - 3642.1957. tb 02516. x" type="journal article" year="1956">Manton (1956)</bibRefCitation>
|
||
. Legs 1 and 2 without trochanter, leg 1 lacking tarsus 1 as well. Chaetotaxy: coxa 1: 2 setae, coxa 2: 2 setae, coxae 3–13: 3–4 setae; prefemur, tibia and tarsus 2 with 1 seta; femur with 2 setae (
|
||
<figureCitation id="133984C1FFE65E5A0E97F85EFDAB0E91" box="[445,537,1983,2008]" captionStart="FIGURE 7" captionStartId="8.[151,250,1660,1682]" captionTargetBox="[169,1424,284,1628]" captionTargetId="figure@8.[151,1436,265,1639]" captionTargetPageId="8" captionText="FIGURE 7. A. Holotype Unixenus intragramineus sp. n., the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (B) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (C); seta on tibia (D); the setiform seta on tarsus 2 (E). F. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side). G. The dorsal ornamental trichome sockets of trichomes a (a), trichome b (b), trichomes c (c) and the circular indentation d." figureDoi="http://doi.org/10.5281/zenodo.10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="8" pageNumber="291">Fig. 7A</figureCitation>
|
||
). Setae on coxa and prefemur, and at distal edge of femur: bi-articulated with longitudinal ridges on basal funicle, each ridge extending distally into a long and thin spine, with the spines surrounding a base of flagellum (
|
||
<figureCitation id="133984C1FFE75E5B0D27FF5DFDD7099D" box="[525,613,188,213]" captionStart="FIGURE 7" captionStartId="8.[151,250,1660,1682]" captionTargetBox="[169,1424,284,1628]" captionTargetId="figure@8.[151,1436,265,1639]" captionTargetPageId="8" captionText="FIGURE 7. A. Holotype Unixenus intragramineus sp. n., the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (B) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (C); seta on tibia (D); the setiform seta on tarsus 2 (E). F. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side). G. The dorsal ornamental trichome sockets of trichomes a (a), trichome b (b), trichomes c (c) and the circular indentation d." figureDoi="http://doi.org/10.5281/zenodo.10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="9" pageNumber="292">Fig. 7B</figureCitation>
|
||
); seta of mid femur similar, but smaller (
|
||
<figureCitation id="133984C1FFE75E5B0B1EFF5DFB3E099D" box="[1076,1164,188,213]" captionStart="FIGURE 7" captionStartId="8.[151,250,1660,1682]" captionTargetBox="[169,1424,284,1628]" captionTargetId="figure@8.[151,1436,265,1639]" captionTargetPageId="8" captionText="FIGURE 7. A. Holotype Unixenus intragramineus sp. n., the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (B) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (C); seta on tibia (D); the setiform seta on tarsus 2 (E). F. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side). G. The dorsal ornamental trichome sockets of trichomes a (a), trichome b (b), trichomes c (c) and the circular indentation d." figureDoi="http://doi.org/10.5281/zenodo.10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="9" pageNumber="292">Fig. 7C</figureCitation>
|
||
), tibia and tarsus 2 each with a setiform seta (
|
||
<figureCitation id="133984C1FFE75E5B0E55FF3EFD8309B1" box="[383,561,223,248]" captionStart="FIGURE 7" captionStartId="8.[151,250,1660,1682]" captionTargetBox="[169,1424,284,1628]" captionTargetId="figure@8.[151,1436,265,1639]" captionTargetPageId="8" captionText="FIGURE 7. A. Holotype Unixenus intragramineus sp. n., the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (B) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (C); seta on tibia (D); the setiform seta on tarsus 2 (E). F. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side). G. The dorsal ornamental trichome sockets of trichomes a (a), trichome b (b), trichomes c (c) and the circular indentation d." figureDoi="http://doi.org/10.5281/zenodo.10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="9" pageNumber="292">Figs 7D and 7E</figureCitation>
|
||
). Posterior edge of last sternite with 2–4 setae similar to those present on coxa, number of these setae varying; two present in
|
||
<typeStatus id="54B926E6FFE75E5B0D88FEE5FCB60854" box="[674,772,260,285]" pageId="9" pageNumber="292" type="holotype">holotype</typeStatus>
|
||
and
|
||
<quantity id="4CFA35A1FFE75E5B0C16FEE5FC360854" box="[828,900,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="9" pageNumber="292" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
|
||
<typeStatus id="54B926E6FFE75E5B0CA6FEE4FC490854" box="[908,1019,261,285]" pageId="9" pageNumber="292" type="paratype">paratypes</typeStatus>
|
||
. Sex organs in male: A pair of penes present on coxae 2 and 2 pairs of coxal glands located on coxal plates of legs 8 and 9.
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE75E5B0FEDFEADFE5E08E5" blockId="9.[151,1437,151,1365]" pageId="9" pageNumber="292">
|
||
<emphasis id="B9764456FFE75E5B0FEDFEADFECD082C" box="[199,383,332,357]" italics="true" pageId="9" pageNumber="292">Telotarsus-Claw</emphasis>
|
||
: A slender structure bearing a posterior process, the latter longer by half the length of claw. Both a small anterior lateral process and a lamella process present, anterior setiform process enlarged at base and longer than the claw (
|
||
<figureCitation id="133984C1FFE75E5B0EA0FE75FE6D08E5" box="[394,479,404,429]" captionStart="FIGURE 7" captionStartId="8.[151,250,1660,1682]" captionTargetBox="[169,1424,284,1628]" captionTargetId="figure@8.[151,1436,265,1639]" captionTargetPageId="8" captionText="FIGURE 7. A. Holotype Unixenus intragramineus sp. n., the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (B) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (C); seta on tibia (D); the setiform seta on tarsus 2 (E). F. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side). G. The dorsal ornamental trichome sockets of trichomes a (a), trichome b (b), trichomes c (c) and the circular indentation d." figureDoi="http://doi.org/10.5281/zenodo.10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="9" pageNumber="292">Fig. 7F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE75E5B0FEDFE56FE1F0B75" blockId="9.[151,1437,151,1365]" pageId="9" pageNumber="292">
|
||
<emphasis id="B9764456FFE75E5B0FEDFE56FEBF0899" box="[199,269,439,464]" italics="true" pageId="9" pageNumber="292">Telson</emphasis>
|
||
: Dorsal ornamental trichome sockets arranged symmetrically with 4 sockets of trichomes
|
||
<emphasis id="B9764456FFE75E5B0A2BFE59FABD0886" bold="true" box="[1281,1295,440,463]" italics="true" pageId="9" pageNumber="292">a</emphasis>
|
||
in
|
||
<typeStatus id="54B926E6FFE75E5B0A18FE56FA250899" box="[1330,1431,439,464]" pageId="9" pageNumber="292" type="holotype">holotype</typeStatus>
|
||
;
|
||
<typeStatus id="54B926E6FFE75E5B0FBDFE3CFEB008BC" box="[151,258,477,501]" pageId="9" pageNumber="292" type="paratype">paratypes</typeStatus>
|
||
with 5–7 sockets of trichomes
|
||
<emphasis id="B9764456FFE75E5B0D4EFE3DFDC008BA" bold="true" box="[612,626,476,499]" italics="true" pageId="9" pageNumber="292">a</emphasis>
|
||
. A single trichome
|
||
<emphasis id="B9764456FFE75E5B0C78FE3AFCD208BD" bold="true" box="[850,864,475,500]" italics="true" pageId="9" pageNumber="292">b</emphasis>
|
||
and 3 large protruding base sockets of trichome
|
||
<emphasis id="B9764456FFE75E5B0AA2FE3DFA2608BA" bold="true" box="[1416,1428,476,499]" italics="true" pageId="9" pageNumber="292">c</emphasis>
|
||
:
|
||
<emphasis id="B9764456FFE75E5B0FBDFDE1FF110B5E" bold="true" box="[151,163,512,535]" italics="true" pageId="9" pageNumber="292">c</emphasis>
|
||
1,
|
||
<emphasis id="B9764456FFE75E5B0F95FDE1FF790B5E" bold="true" box="[191,203,512,535]" italics="true" pageId="9" pageNumber="292">c</emphasis>
|
||
2 and
|
||
<emphasis id="B9764456FFE75E5B0E3BFDE1FEAF0B5E" bold="true" box="[273,285,512,535]" italics="true" pageId="9" pageNumber="292">c</emphasis>
|
||
3, forming a triangular shape, on each side of telson. Circular indentation
|
||
<emphasis id="B9764456FFE75E5B0B76FE1EFBD80B51" bold="true" box="[1116,1130,511,536]" italics="true" pageId="9" pageNumber="292">d</emphasis>
|
||
apparent near exterior side of trichomes
|
||
<emphasis id="B9764456FFE75E5B0E01FDC5FE850B72" bold="true" box="[299,311,548,571]" italics="true" pageId="9" pageNumber="292">c</emphasis>
|
||
(
|
||
<figureCitation id="133984C1FFE75E5B0E6CFDC5FE120B75" box="[326,416,548,573]" captionStart="FIGURE 7" captionStartId="8.[151,250,1660,1682]" captionTargetBox="[169,1424,284,1628]" captionTargetId="figure@8.[151,1436,265,1639]" captionTargetPageId="8" captionText="FIGURE 7. A. Holotype Unixenus intragramineus sp. n., the 2nd leg showing seven leg articles (Co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2, C: claw) and sex organ – penis (P); (B) A biarticulated seta with a ridged funicle present on the coxa, prefemur, the distal edge of femur and the posterior edge of last sternite; a similar smaller version of a biarticulated seta with a ridged funicle is present in the middle of the femur (C); seta on tibia (D); the setiform seta on tarsus 2 (E). F. Telotarsus–Claw showing all processes: the anterior lateral process (lp), the posterior process (p), the claw (c), the setiform process (s) and the lamella process (l), SEM image of the telotarsus (right hand side). G. The dorsal ornamental trichome sockets of trichomes a (a), trichome b (b), trichomes c (c) and the circular indentation d." figureDoi="http://doi.org/10.5281/zenodo.10092678" httpUri="https://zenodo.org/record/10092678/files/figure.png" pageId="9" pageNumber="292">Fig. 7G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE75E5B0FEDFDA6FAD10A8E" blockId="9.[151,1437,151,1365]" pageId="9" pageNumber="292">
|
||
<emphasis id="B9764456FFE75E5B0FEDFDA6FEC80B29" box="[199,378,583,608]" italics="true" pageId="9" pageNumber="292">Caudal bundles</emphasis>
|
||
: In males, caudal bundle comprised of a group of caudal trichome sockets forming a single structure of uniform size; this structure split ventrally with trichome socket-free tissue and extending towards the centre with a small gap dorsally. Three rows of the largest sockets of barbate trichomes forming slightly uneven lateral rows, the latter extending towards the centre of caudal structure (
|
||
<figureCitation id="133984C1FFE75E5B0CDDFD55FBE10B84" box="[1015,1107,692,717]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1852]" captionTargetBox="[151,1436,331,1809]" captionTargetId="figure@6.[151,1436,329,1809]" captionTargetPageId="6" captionText="FIGURE 5. Scanning Electron Microscope images of Unixenus intragramineus sp. n.: A. Head, the posterior vertex trichomes group (pv), collum (Co) and tergite 2 (T2); B. Trichobothria a, b and c; C. Labrum showed 4 + 4 lamellae (l); D. Labrum with the pointed spherical papillae (Psp) on its surface; E. The male caudal bundle structure showing the dorsal ornamental trichome sockets (ots), barbate trichome sockets (bts), trichome socket free tissue (tsft) and caudal trichome sockets (cts) with some caudal trichomes (ct) attached. F. The caudal bundle structure of females showing a difference in the arrangement with caudal trichome sockets (cts) located dorsally, barbate trichome sockets (bts) with the largest size in the centre, trichome socket free tissue (tsft) and nest trichome sockets (nts) that were smaller located centro-ventrally." figureDoi="http://doi.org/10.5281/zenodo.10092674" httpUri="https://zenodo.org/record/10092674/files/figure.png" pageId="9" pageNumber="292">Fig. 5E</figureCitation>
|
||
). In females, caudal bundle different from those in males, with two obvious distinguishing features: caudal bundle structure, the main dorsal structure similar to that of male, and 2 laterosternal bundle structures with smaller sockets of nest trichomes. These smaller sockets located internally and surrounded by 2 rows of caudal trichome sockets externally. Trichome socket-free tissue present ventrally, extending with a small gap and connecting to central tissue where some barbate trichome sockets present. No separation between caudal and nest trichome sockets, all fused into a unified structure of a single caudal bundle structure (
|
||
<figureCitation id="133984C1FFE75E5B0D01FC6DFD330AEC" box="[555,641,908,933]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1852]" captionTargetBox="[151,1436,331,1809]" captionTargetId="figure@6.[151,1436,329,1809]" captionTargetPageId="6" captionText="FIGURE 5. Scanning Electron Microscope images of Unixenus intragramineus sp. n.: A. Head, the posterior vertex trichomes group (pv), collum (Co) and tergite 2 (T2); B. Trichobothria a, b and c; C. Labrum showed 4 + 4 lamellae (l); D. Labrum with the pointed spherical papillae (Psp) on its surface; E. The male caudal bundle structure showing the dorsal ornamental trichome sockets (ots), barbate trichome sockets (bts), trichome socket free tissue (tsft) and caudal trichome sockets (cts) with some caudal trichomes (ct) attached. F. The caudal bundle structure of females showing a difference in the arrangement with caudal trichome sockets (cts) located dorsally, barbate trichome sockets (bts) with the largest size in the centre, trichome socket free tissue (tsft) and nest trichome sockets (nts) that were smaller located centro-ventrally." figureDoi="http://doi.org/10.5281/zenodo.10092674" httpUri="https://zenodo.org/record/10092674/files/figure.png" pageId="9" pageNumber="292">Fig. 5F</figureCitation>
|
||
). Arrangement of caudal bundle of
|
||
<taxonomicName id="4C02E3C7FFE75E5B0B39FC6CFB500AED" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[1043,1250,909,932]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE75E5B0B39FC6CFB500AED" box="[1043,1250,909,932]" italics="true" pageId="9" pageNumber="292">U. intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE75E5B0BC0FC6DFA980AEC" bold="true" box="[1258,1322,908,933]" pageId="9" pageNumber="292">
|
||
<taxonomicNameLabel id="A245F92DFFE75E5B0BC0FC6DFA980AEC" box="[1258,1322,908,933]" pageId="9" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
similar in structure to that observed in
|
||
<taxonomicName id="4C02E3C7FFE75E5B0EFCFC50FBCA0A81" authority="(Conde and Nguyen Duy-Jacquemin, 1992)" baseAuthorityName="Conde and Nguyen Duy-Jacquemin" baseAuthorityYear="1992" box="[470,1144,943,968]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="broelemanni">
|
||
<emphasis id="B9764456FFE75E5B0EFCFC50FD370A81" box="[470,645,943,968]" italics="true" pageId="9" pageNumber="292">U. broelemanni</emphasis>
|
||
(
|
||
<bibRefCitation id="EF93E5B5FFE75E5B0DBEFC4EFBC20A81" author="Conde, B. & Nguyen Duy-Jacquemin, M." box="[660,1136,943,968]" pageId="9" pageNumber="292" pagination="57 - 62" refId="ref11323" refString="Conde, B. & Nguyen Duy-Jacquemin, M. (1992) Decouverte d'un caractѐre sexuel secondaire nouveau chez le male d'un Polyxenide (Myriapodes, Penicillates). Berichte des naturwissenschaftlichen-medizinischen Verein Innsbruck, Supplementum 10, 57 - 62." type="journal article" year="1992">Condé and Nguyen Duy-Jacquemin, 1992</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, classified as
|
||
<typeStatus id="54B926E6FFE75E5B0A38FC51FAF00A81" box="[1298,1346,944,968]" pageId="9" pageNumber="292">type</typeStatus>
|
||
II.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318CBCFFFE75E5B0FEDFC35FC930C1C" pageId="9" pageNumber="292" type="discussion">
|
||
<paragraph id="8BBD9844FFE75E5B0FEDFC35FC930C1C" blockId="9.[151,1437,151,1365]" pageId="9" pageNumber="292">
|
||
<emphasis id="B9764456FFE75E5B0FEDFC35FE890AA4" bold="true" box="[199,315,980,1005]" pageId="9" pageNumber="292">Remarks.</emphasis>
|
||
<taxonomicName id="4C02E3C7FFE75E5B0E6CFC34FE020AA5" authorityName="Jones" authorityYear="1944" box="[326,432,981,1004]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE75E5B0E6CFC34FE020AA5" box="[326,432,981,1004]" italics="true" pageId="9" pageNumber="292">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
species show very similar taxonomically important morphological characteristics: the number of sensilla on antennal article VI, the structure of the gnathochilarium and the chaetotaxy. Caudal bundle
|
||
<typeStatus id="54B926E6FFE75E5B0FBDFBFCFF750D7C" box="[151,199,1053,1077]" pageId="9" pageNumber="292">type</typeStatus>
|
||
II is common to the genus
|
||
<taxonomicName id="4C02E3C7FFE75E5B0ED5FBFCFDDB0D7D" authorityName="Jones" authorityYear="1944" box="[511,617,1053,1076]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE75E5B0ED5FBFCFDDB0D7D" box="[511,617,1053,1076]" italics="true" pageId="9" pageNumber="292">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
, as well as in other genera in the subfamily
|
||
<taxonomicName id="4C02E3C7FFE75E5B0B4AFBFDFE4D0D11" authority="(Conde and Nguyen Duy-Jacquemin 2008)" baseAuthorityName="Conde and Nguyen Duy-Jacquemin" baseAuthorityYear="2008" pageId="9" pageNumber="292" rank="subFamily" subFamily="Monographinae">
|
||
Monographinae (
|
||
<bibRefCitation id="EF93E5B5FFE75E5B0A0BFBFDFE4A0D11" author="Conde, B. & Nguyen Duy-Jacquemin, M." pageId="9" pageNumber="292" pagination="291 - 302" refId="ref11366" refString="Conde, B. & Nguyen Duy-Jacquemin, M. (2008) Classification actuelle des Diplopodes Penicillates (Myriapodes) avec nouvelles definition des taxa. Bulletin de la Societe Zoologique de France, 133, 4, 291 - 302." type="journal article" year="2008">Condé and Nguyen Duy-Jacquemin 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. If only these characteristics are used for identification, it can be difficult to confidently identify
|
||
<taxonomicName id="4C02E3C7FFE75E5B0E55FB84FE5B0D35" authorityName="Jones" authorityYear="1944" box="[383,489,1125,1148]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE75E5B0E55FB84FE5B0D35" box="[383,489,1125,1148]" italics="true" pageId="9" pageNumber="292">Unixenus</emphasis>
|
||
</taxonomicName>
|
||
specimens to the species level. The telotarsus has proved to be a useful additional structure that can help distinguish between species in this genus.
|
||
<taxonomicName id="4C02E3C7FFE75E5B0C42FB68FB340DE9" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[872,1158,1161,1184]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE75E5B0C42FB68FB340DE9" box="[872,1158,1161,1184]" italics="true" pageId="9" pageNumber="292">Unixenus intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE75E5B0BA6FB66FB780DE9" bold="true" box="[1164,1226,1159,1184]" pageId="9" pageNumber="292">
|
||
<taxonomicNameLabel id="A245F92DFFE75E5B0BA6FB66FB780DE9" box="[1164,1226,1159,1184]" pageId="9" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has characteristics that are very similar to those of
|
||
<taxonomicName id="4C02E3C7FFE75E5B0ED0FB4CFD3E0D8C" authorityName="Verhoeff" authorityYear="1924" box="[506,652,1196,1221]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="mjoebergi">
|
||
<emphasis id="B9764456FFE75E5B0ED0FB4CFD3E0D8C" box="[506,652,1196,1221]" italics="true" pageId="9" pageNumber="292">U. mjoebergi</emphasis>
|
||
</taxonomicName>
|
||
; both species have the same chaetotaxy which makes separating these species difficult. However, the structure of the telotarsus in these two species differs.
|
||
<taxonomicName id="4C02E3C7FFE75E5B0B59FB30FAB50DA1" authorityName="Verhoeff" authorityYear="1924" box="[1139,1287,1231,1256]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="mjoebergi">
|
||
<emphasis id="B9764456FFE75E5B0B59FB30FAB50DA1" box="[1139,1287,1231,1256]" italics="true" pageId="9" pageNumber="292">U. mjoebergi</emphasis>
|
||
</taxonomicName>
|
||
has a robust claw, while the claw of
|
||
<taxonomicName id="4C02E3C7FFE75E5B0E92FB14FD3E0C45" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[440,652,1269,1292]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE75E5B0E92FB14FD3E0C45" box="[440,652,1269,1292]" italics="true" pageId="9" pageNumber="292">U. intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE75E5B0DBDFB15FD6E0C44" bold="true" box="[663,732,1268,1293]" pageId="9" pageNumber="292">
|
||
<taxonomicNameLabel id="A245F92DFFE75E5B0DBDFB15FD6E0C44" box="[663,732,1268,1293]" pageId="9" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a slender structure.
|
||
<taxonomicName id="4C02E3C7FFE75E5B0CC5FB14FB710C45" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[1007,1219,1269,1292]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="intragramineus" status="sp. nov.">
|
||
<emphasis id="B9764456FFE75E5B0CC5FB14FB710C45" box="[1007,1219,1269,1292]" italics="true" pageId="9" pageNumber="292">U. intragramineus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE75E5B0BE4FB15FAA00C44" bold="true" box="[1230,1298,1268,1293]" pageId="9" pageNumber="292">
|
||
<taxonomicNameLabel id="A245F92DFFE75E5B0BE4FB15FAA00C44" box="[1230,1298,1268,1293]" pageId="9" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the only penicillate millipede found living within a grass stem, which is a highly specialized habitat that provides a food source and a refuge from the intermittent tidal inundation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |