treatments-xml/data/E8/5D/87/E85D87D2FF914310389DE625FC78FB77.xml

<document id="30CF0881BCB457ED1B4B44B1E13279AA" ID-DOI="10.11646/zootaxa.3835.2.5" ID-GBIF-Dataset="28a9f342-dd22-44c3-bfba-8dfb8c245e9c" ID-ISSN="1175-5326" ID-Zenodo-Dep="249634" ID-ZooBank="017DC939-E929-4A29-B99A-D5D9C6B64787" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460456427685" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; García, Mauricio" docDate="2014" docId="E85D87D2FF914310389DE625FC78FB77" docLanguage="en" docName="zt03835p262.pdf" docOrigin="Zootaxa 3835 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Radicitus granitum Short &amp; Garcia, sp. n." docType="treatment" docVersion="7" lastPageNumber="257" masterDocId="1464FFAAFF924316380AE462FFF1FFB3" masterDocTitle="A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)" masterLastPageNumber="262" masterPageNumber="251" pageNumber="254" updateTime="1698302202375" updateUser="plazi">
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<mods:title id="0E227B1A283F429EB75FF2621080FFDD">A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
</mods:titleInfo>
<mods:name id="6AC3A2519FE827C7A092CE314BD189A0" type="personal">
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<mods:namePart id="DFC9650EA6C74906D4AE2B4DBDFD824C">Short, Andrew Edward Z.</mods:namePart>
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<mods:namePart id="6B6FBA208DD843D804A5619173A0C53E">García, Mauricio</mods:namePart>
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<mods:title id="6AF095E776476C76500863C6980D0B0C">Zootaxa</mods:title>
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<mods:part id="13B4BB46E5B07F65B076FC9F6B8E2908">
<mods:date id="81C766AAAF7D82263938F203CC00B807">2014</mods:date>
<mods:detail id="D29E7D20F6921794D77CAFB08E8A359A" type="volume">
<mods:number id="27B0A2D78E7B830549665657825D94C8">3835</mods:number>
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</mods:mods>
<treatment id="E85D87D2FF914310389DE625FC78FB77" ID-DOI="http://doi.org/10.5281/zenodo.6133259" ID-GBIF-Taxon="119511038" ID-Zenodo-Dep="6133259" LSID="urn:lsid:plazi:treatment:E85D87D2FF914310389DE625FC78FB77" httpUri="http://treatment.plazi.org/id/E85D87D2FF914310389DE625FC78FB77" lastPageId="6" lastPageNumber="257" pageId="3" pageNumber="254">
<subSubSection id="28EE654FFF914315389DE625FDF7FD37" pageId="3" pageNumber="254" type="nomenclature">
<paragraph id="604B36C4FF914315389DE625FD8FFDD3" blockId="3.[151,638,583,645]" box="[151,638,583,608]" pageId="3" pageNumber="254">
<heading id="3B0381A8FF914315389DE625FD8FFDD3" bold="true" box="[151,638,583,608]" fontSize="10" level="2" pageId="3" pageNumber="254" reason="2">
<emphasis id="5280EAD6FF914315389DE625FD8FFDD3" bold="true" box="[151,638,583,608]" pageId="3" pageNumber="254">
<taxonomicName id="A7F44D47FF914315389DE625FDC6FDD3" ID-CoL="78RV7" authority="Short &amp; Garcia" authorityName="Short &amp; Garcia" box="[151,567,583,608]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="granitum" status="sp. nov.">
<emphasis id="5280EAD6FF914315389DE625FE86FDD3" bold="true" box="[151,375,583,608]" italics="true" pageId="3" pageNumber="254">Radicitus granitum</emphasis>
Short &amp; García
</taxonomicName>
<taxonomicNameLabel id="49B357ADFF9143153A34E625FD8FFDD3" box="[574,638,583,608]" pageId="3" pageNumber="254" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="604B36C4FF914315389DE60EFDF7FD37" blockId="3.[151,638,583,645]" box="[151,518,620,645]" pageId="3" pageNumber="254">
(
<figureCitation id="F8CF2A41FF9143153895E60EFF1FFD37" box="[159,238,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1972,1994]" captionTargetBox="[170,1421,794,1910]" captionTargetId="figure@3.[151,1436,769,1950]" captionTargetPageId="3" captionText="FIGURE 2. Radicitus granitum, dorsal, ventral, and lateral habitus (paratype)." httpUri="https://zenodo.org/record/249636/files/figure.png" pageId="3" pageNumber="254">Figs. 2</figureCitation>
, 
<figureCitation id="F8CF2A41FF91431538F0E60EFEF9FD37" box="[250,264,620,644]" captionStart="FIGURE 3" captionStartId="4.[151,250,1003,1025]" captionTargetBox="[151,1436,193,980]" captionTargetId="figure@4.[151,1436,193,981]" captionTargetPageId="4" captionText="FIGURE 3. Heads of Radicitus spp., ventral view. A) R. ayacucho, B) R. granitum." httpUri="https://zenodo.org/record/249637/files/figure.png" pageId="3" pageNumber="254">3</figureCitation>
B, 4B, 5, 6B, 7D, 8, 9)
</paragraph>
</subSubSection>
<subSubSection id="28EE654FFF914312389DE6D6FB86FA7E" lastPageId="4" lastPageNumber="255" pageId="3" pageNumber="254" type="materials_examined">
<paragraph id="604B36C4FF914315389DE6D6FB8BFD7F" blockId="3.[151,1146,692,717]" box="[151,1146,692,717]" pageId="3" pageNumber="254">
<emphasis id="5280EAD6FF914315389DE6D6FEC4FD7E" bold="true" box="[151,309,692,717]" pageId="3" pageNumber="254">
<typeStatus id="BF4F8866FF914315389DE6D6FF23FD7E" box="[151,210,692,717]" pageId="3" pageNumber="254">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="18E37654FF9143153936E6D6FE43FD7E" box="[316,434,692,717]" name="Venezuela" pageId="3" pageNumber="254">Venezuela</collectingCountry>
, Bolívar State, Los Pijiguaos, 
<geoCoordinate id="05C05003FF9143153B0FE6D6FC7AFD7E" box="[773,907,692,717]" direction="north" orientation="latitude" pageId="3" pageNumber="254" precision="1" value="6.5936165">6°35.617’N</geoCoordinate>
, 
<geoCoordinate id="05C05003FF9143153B9DE6D6FBDFFD7E" box="[919,1070,692,717]" direction="west" orientation="longitude" pageId="3" pageNumber="254" precision="1" value="-66.82063">66°49.238’W</geoCoordinate>
, 
<specimenCount id="76F2FD4DFF9143153C30E6D6FB8BFD7F" box="[1082,1146,692,716]" pageId="3" pageNumber="254" type="generic">
<quantity id="A70C9B21FF9143153C30E6D6FB87FD7F" box="[1082,1142,692,716]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="254" unit="m" value="80.0">80 m</quantity>
.
</specimenCount>
</paragraph>
<caption id="348B664CFF914315389DE3D6FC44F879" box="[151,949,1972,1994]" httpUri="https://zenodo.org/record/249636/files/figure.png" pageId="3" pageNumber="254" targetBox="[170,1421,794,1910]" targetPageId="3">
<paragraph id="604B36C4FF914315389DE3D6FC44F879" blockId="3.[151,949,1972,1994]" box="[151,949,1972,1994]" pageId="3" pageNumber="254">
<emphasis id="5280EAD6FF914315389DE3D6FEE3F879" bold="true" box="[151,274,1972,1994]" pageId="3" pageNumber="254">FIGURE 2.</emphasis>
<taxonomicName id="A7F44D47FF9143153913E3D6FE2DF879" box="[281,476,1972,1994]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="granitum">
<emphasis id="5280EAD6FF9143153913E3D6FE2DF879" box="[281,476,1972,1994]" italics="true" pageId="3" pageNumber="254">Radicitus granitum</emphasis>
</taxonomicName>
, dorsal, ventral, and lateral habitus (paratype).
</paragraph>
</caption>
<caption id="348B664CFF964312389DE789FC1BFBB2" box="[151,1002,1003,1026]" httpUri="https://zenodo.org/record/249637/files/figure.png" pageId="4" pageNumber="255" targetBox="[151,1436,193,980]" targetPageId="4">
<paragraph id="604B36C4FF964312389DE789FC1BFBB2" blockId="4.[151,1002,1003,1026]" box="[151,1002,1003,1026]" pageId="4" pageNumber="255">
<emphasis id="5280EAD6FF964312389DE789FEE3FBB2" bold="true" box="[151,274,1003,1025]" pageId="4" pageNumber="255">FIGURE 3.</emphasis>
Heads of 
<taxonomicName id="A7F44D47FF9643123970E789FE2BFBB3" box="[378,474,1003,1024]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="251" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5280EAD6FF9643123970E789FE2BFBB3" box="[378,474,1003,1024]" italics="true" pageId="4" pageNumber="255">Radicitus</emphasis>
</taxonomicName>
spp., ventral view. A) 
<taxonomicName id="A7F44D47FF9643123ACAE78EFCCCFBB3" box="[704,829,1003,1025]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="ayacucho">
<emphasis id="5280EAD6FF9643123ACAE78EFCCCFBB3" box="[704,829,1003,1025]" italics="true" pageId="4" pageNumber="255">R. ayacucho</emphasis>
</taxonomicName>
, B) 
<taxonomicName id="A7F44D47FF9643123B63E78EFC14FBB2" box="[873,997,1004,1025]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="granitum">R. granitum</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="604B36C4FF96431238CDE04AFB86FA7E" blockId="4.[151,1437,1064,2025]" pageId="4" pageNumber="255">
<emphasis id="5280EAD6FF96431238CDE04AFE8AFBF2" bold="true" box="[199,379,1064,1089]" pageId="4" pageNumber="255">
<typeStatus id="BF4F8866FF96431238CDE04AFEF3FBF2" box="[199,258,1064,1089]" pageId="4" pageNumber="255">Type</typeStatus>
material.
</emphasis>
<typeStatus id="BF4F8866FF9643123982E04AFE01FBF2" box="[392,496,1064,1089]" pageId="4" pageNumber="255" type="holotype">Holotype</typeStatus>
&lt;male&gt;: “ 
<collectingCountry id="18E37654FF9643123A78E04BFCEEFBF2" box="[626,799,1065,1089]" name="Venezuela" pageId="4" pageNumber="255">VENEZUELA</collectingCountry>
: Bolivar State/ 
<geoCoordinate id="05C05003FF9643123BD7E04AFB92FBF2" box="[989,1123,1064,1089]" direction="north" orientation="latitude" pageId="4" pageNumber="255" precision="1" value="6.5936165">6°35.617’N</geoCoordinate>
, 
<geoCoordinate id="05C05003FF9643123C7FE04AFAFCFBF2" box="[1141,1293,1064,1089]" direction="west" orientation="longitude" pageId="4" pageNumber="255" precision="1" value="-66.82063">66°49.238’W</geoCoordinate>
, 
<quantity id="A70C9B21FF9643123D14E04BFAACFBF2" box="[1310,1373,1065,1089]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="255" unit="m" value="80.0">80 m</quantity>
/ Los Pijiguaos; 
<date id="144A1004FF9643123905E02EFE77FBD6" box="[271,390,1100,1125]" pageId="4" pageNumber="255" value="2007-09-16">16.ix.2007</date>
/ leg. A. Short, M. Garcia, L. Joly/ AS-07-015; morichal/rock outcrop” (
<collectionCode id="06E5AE01FF9643123CC8E02FFAE1FBD6" LSID="urn:lsid:biocol.org:col:33834" box="[1218,1296,1101,1125]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="255">MIZA</collectionCode>
). 
<emphasis id="5280EAD6FF9643123D2EE02EFD97FB3A" bold="true" pageId="4" pageNumber="255">
<typeStatus id="BF4F8866FF9643123D2EE02EFA6DFBD6" box="[1316,1436,1100,1125]" pageId="4" pageNumber="255" type="paratype">Paratypes</typeStatus>
(40): 
<collectingCountry id="18E37654FF96431238D3E012FE7DFB3A" box="[217,396,1136,1161]" name="Venezuela" pageId="4" pageNumber="255">VENEZUELA</collectingCountry>
: Amazonas State:
</emphasis>
Tobogan de la Selva, 
<geoCoordinate id="05C05003FF9643123B65E012FC04FB3A" box="[879,1013,1136,1161]" direction="north" orientation="latitude" pageId="4" pageNumber="255" precision="1" value="5.386783">5°23.207’N</geoCoordinate>
, 
<geoCoordinate id="05C05003FF9643123C0EE012FB6AFB3A" box="[1028,1179,1136,1161]" direction="west" orientation="longitude" pageId="4" pageNumber="255" precision="1" value="-67.615364">67°36.922’W</geoCoordinate>
, 
<quantity id="A70C9B21FF9643123CA0E012FB06FB3B" box="[1194,1271,1136,1161]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="255" unit="m" value="125.0">125 m</quantity>
, 
<date id="144A1004FF9643123D0CE012FA92FB3A" box="[1286,1379,1136,1161]" pageId="4" pageNumber="255" value="2006-01-07">7.i.2006</date>
, leg. A.E.Z. Short &amp; M. García, AS-06-017 (1 ex.: 
<collectionCode id="06E5AE01FF9643123ABAE0F7FCF0FB1E" box="[688,769,1173,1197]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
); same locality but “old tobogancito”, 
<date id="144A1004FF9643123CCAE0F6FACAFB1E" box="[1216,1339,1172,1197]" pageId="4" pageNumber="255" value="2008-08-08">8.viii.2008</date>
, AS-08- 080b, (2 exs.: 
<collectionCode id="06E5AE01FF964312393EE0DBFE74FB62" box="[308,389,1209,1233]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
); same locality but 
<date id="144A1004FF9643123A57E0DAFD38FB62" box="[605,713,1208,1233]" pageId="4" pageNumber="255" value="2009-01-14">14.i.2009</date>
, “upstream slide”, leg. Short, García, Miller &amp; Joly, VZ09-0114- 01F (1 ex.: 
<collectionCode id="06E5AE01FF964312392EE0BFFE84FB46" box="[292,373,1245,1269]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
). 
<emphasis id="5280EAD6FF9643123981E0BEFDC2FB46" bold="true" box="[395,563,1244,1269]" pageId="4" pageNumber="255">Bolívar State:</emphasis>
Same data as 
<typeStatus id="BF4F8866FF9643123AEEE0BEFCB7FB46" box="[740,838,1244,1269]" pageId="4" pageNumber="255" type="holotype">holotype</typeStatus>
(24 exs.: 
<collectionCode id="06E5AE01FF9643123BCAE0BFFBFEFB46" box="[960,1039,1245,1269]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
, 
<collectionCode id="06E5AE01FF9643123C14E0BFFB83FB46" LSID="urn:lsid:biocol.org:col:34871" box="[1054,1138,1245,1269]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="255">USNM</collectionCode>
, 
<collectionCode id="06E5AE01FF9643123C8BE0BFFB19FB46" box="[1153,1256,1245,1269]" pageId="4" pageNumber="255">MALUZ</collectionCode>
, 
<collectionCode id="06E5AE01FF9643123CFDE0BFFAB0FB46" LSID="urn:lsid:biocol.org:col:33834" box="[1271,1345,1245,1269]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="255">MIZA</collectionCode>
, 
<collectionCode id="06E5AE01FF9643123D5AE0BFFA69FB47" box="[1360,1432,1245,1268]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="255">NMW</collectionCode>
, 
<collectionCode id="06E5AE01FF964312389DE163FF1DFAAA" box="[151,236,1281,1305]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="255">NMPC</collectionCode>
); Same data as 
<typeStatus id="BF4F8866FF96431239ABE162FDF2FAAA" box="[417,515,1280,1305]" pageId="4" pageNumber="255" type="holotype">holotype</typeStatus>
but 
<date id="144A1004FF9643123A33E162FD44FAAA" box="[569,693,1280,1305]" pageId="4" pageNumber="255" value="2008-08-06">6.viii.2008</date>
, AS-08-076 (1 ex.: 
<collectionCode id="06E5AE01FF9643123B96E163FC1CFAAA" box="[924,1005,1281,1305]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
); ca. 
<quantity id="A70C9B21FF9643123C21E162FB85FAAA" box="[1067,1140,1280,1305]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="255" unit="km" value="25.0">25 km</quantity>
E. El Burro, 
<geoCoordinate id="05C05003FF9643123D19E162FA66FAAA" box="[1299,1431,1280,1305]" direction="north" orientation="latitude" pageId="4" pageNumber="255" precision="1" value="6.2179446">6°13’4.6”N</geoCoordinate>
, 
<geoCoordinate id="05C05003FF964312389DE146FECDFA8E" box="[151,316,1316,1341]" direction="west" orientation="longitude" pageId="4" pageNumber="255" precision="1" value="-67.24066">67°14’26.4”W</geoCoordinate>
, 
<quantity id="A70C9B21FF9643123942E146FE8CFA8E" box="[328,381,1316,1341]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="255" unit="m" value="60.0">60m</quantity>
, 
<date id="144A1004FF9643123983E146FDF4FA8E" box="[393,517,1316,1341]" pageId="4" pageNumber="255" value="2008-08-07">7.viii.2008</date>
, leg. A. Short, M. García, &amp; L. Joly, rocky morichal, AS-08-077 (1 ex.: 
<collectionCode id="06E5AE01FF9643123D35E147FA61FA8E" box="[1343,1424,1317,1341]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
); same locality but 
<date id="144A1004FF9643123956E12AFE39FAD2" box="[348,456,1352,1377]" pageId="4" pageNumber="255" value="2009-01-12">12.i.2009</date>
, leg. A. Short &amp; M. García, VZ09-0113- 
<date id="144A1004FF9643123B84E12BFC4CFAD2" box="[910,957,1353,1377]" pageId="4" pageNumber="255" value="2001-10">01X</date>
(8 exs.: 
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, 
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, 
<collectionCode id="06E5AE01FF9643123CC0E12BFAC2FAD2" box="[1226,1331,1353,1377]" pageId="4" pageNumber="255">MALUZ</collectionCode>
); 
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E. Rio Cuchivero, 
<geoCoordinate id="05C05003FF9643123947E10FFE11FA36" box="[333,480,1389,1413]" direction="north" orientation="latitude" pageId="4" pageNumber="255" precision="1" value="7.496472">7°29’47.3”N</geoCoordinate>
, 
<geoCoordinate id="05C05003FF96431239E7E10EFD60FA36" box="[493,657,1388,1413]" direction="west" orientation="longitude" pageId="4" pageNumber="255" precision="1" value="-65.86244">65°51’44.8”W</geoCoordinate>
, 
<quantity id="A70C9B21FF9643123A95E10EFD22FA36" box="[671,723,1388,1413]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="255" unit="m" value="45.0">45m</quantity>
, 
<date id="144A1004FF9643123AEAE10EFCADFA36" box="[736,860,1388,1413]" pageId="4" pageNumber="255" value="2008-08-06">6.viii.2008</date>
, leg. A. Short, M. García, &amp; L. Joly, rock outcrop seeps, AS-08-075 (1 ex.: 
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); rock outcrop ca. 
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NE Pijiguaos, 
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, 
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, 
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, 
<date id="144A1004FF964312389DE1D6FEFAFA7E" box="[151,267,1460,1485]" pageId="4" pageNumber="255" value="2010-07-09">9.vii.2010</date>
, leg. A. Short &amp; M. Tellez, detritus flotation, VZ10-0709-01B (1 ex.: 
<collectionCode id="06E5AE01FF9643123C10E1D7FB9AFA7E" box="[1050,1131,1461,1485]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="255">SEMC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="28EE654FFF96431238CDE1BAFC83F933" pageId="4" pageNumber="255" type="diagnosis">
<paragraph id="604B36C4FF96431238CDE1BAFE78F98A" blockId="4.[151,1437,1064,2025]" pageId="4" pageNumber="255">
<emphasis id="5280EAD6FF96431238CDE1BAFEB0FA42" bold="true" box="[199,321,1496,1521]" pageId="4" pageNumber="255">Diagnosis.</emphasis>
Size 4.5–5.0 mm. Elytra with extremely fine serial punctures, with punctures smaller than surrounding ground punctation (
<figureCitation id="F8CF2A41FF9643123A1CE19EFD93F9A6" box="[534,610,1532,1557]" captionStart="FIGURE 4" captionStartId="5.[151,250,1934,1956]" captionTargetBox="[151,1436,589,1912]" captionTargetId="figure@5.[151,1436,589,1913]" captionTargetPageId="5" captionText="FIGURE 4. Elytra of Radicitus spp. A) R. ayacucho, B) R. granitum." httpUri="https://zenodo.org/record/249638/files/figure.png" pageId="4" pageNumber="255">Fig. 4</figureCitation>
B). Fifth ventrite entirely pubescent. Parameres not divided into two projections (
<figureCitation id="F8CF2A41FF9643123929E242FE97F98A" box="[291,358,1568,1593]" captionStart="FIGURE 7" captionStartId="8.[151,250,1631,1653]" captionTargetBox="[242,1344,303,1609]" captionTargetId="figure@8.[242,1344,303,1609]" captionTargetPageId="8" captionText="FIGURE 7. Aedeagi of Radicitus spp. A) R. ayacucho, B) R. surinamensis, C) R. cf. granitum (Suriname), D) R. granitum (Venezuela)." httpUri="https://zenodo.org/record/249641/files/figure.png" pageId="4" pageNumber="255">Fig. 7</figureCitation>
D).
</paragraph>
<paragraph id="604B36C4FF96431238CDE226FC83F933" blockId="4.[151,1437,1064,2025]" pageId="4" pageNumber="255">
The smaller size and lack of coarsely punctate elytral striae, and completely pubescent fifth ventrite will easily separate this species from both 
<taxonomicName id="A7F44D47FF96431239FDE20BFD73F932" box="[503,642,1640,1665]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="ayacucho">
<emphasis id="5280EAD6FF96431239FDE20BFD73F932" box="[503,642,1640,1665]" italics="true" pageId="4" pageNumber="255">R. ayacucho</emphasis>
</taxonomicName>
and 
<taxonomicName id="A7F44D47FF9643123AB2E20BFC9AF933" box="[696,875,1641,1664]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="surinamensis">
<emphasis id="5280EAD6FF9643123AB2E20BFC9AF933" box="[696,875,1641,1664]" italics="true" pageId="4" pageNumber="255">R. surinamensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="28EE654FFF96431338CDE2EEFA84FE1F" lastPageId="5" lastPageNumber="256" pageId="4" pageNumber="255" type="description">
<paragraph id="604B36C4FF96431238CDE2EEFE59F886" blockId="4.[151,1437,1064,2025]" pageId="4" pageNumber="255">
<emphasis id="5280EAD6FF96431238CDE2EEFEA8F916" bold="true" box="[199,345,1676,1701]" pageId="4" pageNumber="255">Description.</emphasis>
<emphasis id="5280EAD6FF9643123955E2EEFDA1F917" box="[351,592,1676,1701]" italics="true" pageId="4" pageNumber="255">Color and punctation</emphasis>
. Dorsum of head, pronotum and elytra very dark brown with all margins of pronotum and lateral margins of the elytra slightly paler (
<figureCitation id="F8CF2A41FF9643123B12E2D2FCACF97A" box="[792,861,1712,1737]" captionStart="FIGURE 2" captionStartId="3.[151,250,1972,1994]" captionTargetBox="[170,1421,794,1910]" captionTargetId="figure@3.[151,1436,769,1950]" captionTargetPageId="3" captionText="FIGURE 2. Radicitus granitum, dorsal, ventral, and lateral habitus (paratype)." httpUri="https://zenodo.org/record/249636/files/figure.png" pageId="4" pageNumber="255">Fig. 2</figureCitation>
). Clypeus without pale preocular patches. Thoracic and abdominal sterna and epipleura dark chestnut brown, with legs very slightly paler. Ground punctation on head and elytra moderately coarse; moderately coarse on elytral disc, and becoming extremely fine to almost obsolete laterally and posteriorly.
</paragraph>
<paragraph id="604B36C4FF96431238CDE322FBE1F812" blockId="4.[151,1437,1064,2025]" pageId="4" pageNumber="255">
<emphasis id="5280EAD6FF96431238CDE322FEF5F8EA" box="[199,260,1856,1881]" italics="true" pageId="4" pageNumber="255">Head</emphasis>
. Antennae (
<figureCitation id="F8CF2A41FF9643123984E322FE21F8EA" box="[398,464,1856,1881]" captionStart="FIGURE 3" captionStartId="4.[151,250,1003,1025]" captionTargetBox="[151,1436,193,980]" captionTargetId="figure@4.[151,1436,193,981]" captionTargetPageId="4" captionText="FIGURE 3. Heads of Radicitus spp., ventral view. A) R. ayacucho, B) R. granitum." httpUri="https://zenodo.org/record/249637/files/figure.png" pageId="4" pageNumber="255">Fig. 3</figureCitation>
B) with scape c. three times as long as pedicel, antennomere 3 subequal in length as 4 and 5 combined. Maxillary palps subequal in length to the width of the anterior margin of the clypeus, with segments two and three subequal in length with apical segment slightly shorter.
</paragraph>
<paragraph id="604B36C4FF96431338CDE3CEFD58FEF3" blockId="4.[151,1437,1064,2025]" lastBlockId="5.[151,1436,151,536]" lastPageId="5" lastPageNumber="256" pageId="4" pageNumber="255">
<emphasis id="5280EAD6FF96431238CDE3CEFEEEF876" box="[199,287,1964,1989]" italics="true" pageId="4" pageNumber="255">Thorax.</emphasis>
Elytra with rows of serial punctures weakly impressed along entire length (
<figureCitation id="F8CF2A41FF9643123C71E3CEFB30F876" box="[1147,1217,1964,1989]" captionStart="FIGURE 4" captionStartId="5.[151,250,1934,1956]" captionTargetBox="[151,1436,589,1912]" captionTargetId="figure@5.[151,1436,589,1913]" captionTargetPageId="5" captionText="FIGURE 4. Elytra of Radicitus spp. A) R. ayacucho, B) R. granitum." httpUri="https://zenodo.org/record/249638/files/figure.png" pageId="4" pageNumber="255">Fig. 4</figureCitation>
B); serial punctures subequal in size as surrounding ground punctuation, making them almost appear absent. Mesoventrite with distinctive, large laterally compressed projection, which appears as a large “tooth”; the projection strongly elevated, rising to the same plane as the ventral surface of the mesocoxae; apex of projection set with a several coarse spine-like setae. Metaventrite almost entirely pubescent, with posteromedial glabrous region represented by a narrow line in the posterior half of the metaventrite width. Metafemora with pubescence restricted to anterior third and extending distally to basal four-fifths.
</paragraph>
<paragraph id="604B36C4FF97431338CDE52EFA84FE1F" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="256">
<emphasis id="5280EAD6FF97431338CDE52EFEC0FED6" box="[199,305,332,357]" italics="true" pageId="5" pageNumber="256">Abdomen</emphasis>
. Pubescence on ventrites 1-4 extremely fine and dense. Fifth ventrite with pubescence slightly less dense at apex, but without a glabrous patch (
<figureCitation id="F8CF2A41FF9743133A9FE50DFD2BFE3B" box="[661,730,367,392]" captionStart="FIGURE 6" captionStartId="7.[151,250,1282,1304]" captionTargetBox="[151,1436,193,1260]" captionTargetId="figure@7.[151,1436,193,1261]" captionTargetPageId="7" captionText="FIGURE 6. Abdominal ventrites of Radicitus spp. A) R. ayacucho, B) R. granitum." httpUri="https://zenodo.org/record/249640/files/figure.png" pageId="5" pageNumber="256">Fig. 6</figureCitation>
B). Aedeagus (
<figureCitation id="F8CF2A41FF9743133B89E50DFC36FE3B" box="[899,967,367,392]" captionStart="FIGURE 7" captionStartId="8.[151,250,1631,1653]" captionTargetBox="[242,1344,303,1609]" captionTargetId="figure@8.[242,1344,303,1609]" captionTargetPageId="8" captionText="FIGURE 7. Aedeagi of Radicitus spp. A) R. ayacucho, B) R. surinamensis, C) R. cf. granitum (Suriname), D) R. granitum (Venezuela)." httpUri="https://zenodo.org/record/249641/files/figure.png" pageId="5" pageNumber="256">Fig. 7</figureCitation>
D) with parameres forming a single lobe, tapering apically. Lateral margins of median lobe gradually tapering, with apex slightly shorter than parameres.
</paragraph>
</subSubSection>
<subSubSection id="28EE654FFF97431338CDE5D5FA61FE63" box="[199,1424,439,464]" pageId="5" pageNumber="256" type="etymology">
<paragraph id="604B36C4FF97431338CDE5D5FA61FE63" blockId="5.[151,1436,151,536]" box="[199,1424,439,464]" pageId="5" pageNumber="256">
<emphasis id="5280EAD6FF97431338CDE5D5FEBFFE63" bold="true" box="[199,334,439,464]" pageId="5" pageNumber="256">Etymology.</emphasis>
Named after the Latin word for granite, reflecting the habitat of this species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="28EE654FFF97431038CDE5BEFC78FB77" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256" type="distribution">
<paragraph id="604B36C4FF97431338CDE5BEFE21FDAB" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="256">
<emphasis id="5280EAD6FF97431338CDE5BEFE91FE46" bold="true" box="[199,352,476,501]" pageId="5" pageNumber="256">Distribution.</emphasis>
This species is known from a series of localities along the northwestern edge of the 
<collectingCountry id="18E37654FF9743133D41E5BEFA6DFE46" box="[1355,1436,476,501]" name="Guyana" pageId="5" pageNumber="256">Guiana</collectingCountry>
Shield in 
<collectingCountry id="18E37654FF9743133908E59DFE84FDAB" box="[258,373,511,536]" name="Venezuela" pageId="5" pageNumber="256">Venezuela</collectingCountry>
(
<figureCitation id="F8CF2A41FF974313398FE59DFE36FDAB" box="[389,455,511,536]" captionStart="FIGURE 8" captionStartId="9.[151,250,1348,1370]" captionTargetBox="[151,1436,481,1326]" captionTargetId="figure@9.[151,1436,481,1326]" captionTargetPageId="9" captionText="FIGURE 8. Distribution of Radicitus spp." httpUri="https://zenodo.org/record/249642/files/figure.png" pageId="5" pageNumber="256">Fig. 8</figureCitation>
)
</paragraph>
<caption id="348B664CFF974313389DE3ECFCA9F817" box="[151,856,1934,1956]" httpUri="https://zenodo.org/record/249638/files/figure.png" pageId="5" pageNumber="256" targetBox="[151,1436,589,1912]" targetPageId="5">
<paragraph id="604B36C4FF974313389DE3ECFCA9F817" blockId="5.[151,856,1934,1956]" box="[151,856,1934,1956]" pageId="5" pageNumber="256">
<emphasis id="5280EAD6FF974313389DE3ECFEE3F817" bold="true" box="[151,274,1934,1956]" pageId="5" pageNumber="256">FIGURE 4.</emphasis>
Elytra of 
<taxonomicName id="A7F44D47FF9743133972E3ECFE29F810" box="[376,472,1934,1955]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="251" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5280EAD6FF9743133972E3ECFE29F810" box="[376,472,1934,1955]" italics="true" pageId="5" pageNumber="256">Radicitus</emphasis>
</taxonomicName>
spp. A) 
<taxonomicName id="A7F44D47FF9743133A25E3EDFD5DF810" box="[559,684,1934,1956]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="ayacucho">
<emphasis id="5280EAD6FF9743133A25E3EDFD5DF810" box="[559,684,1934,1956]" italics="true" pageId="5" pageNumber="256">R. ayacucho</emphasis>
</taxonomicName>
, B) 
<taxonomicName id="A7F44D47FF9743133AD3E3EDFCA3F817" box="[729,850,1935,1956]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="granitum">
<emphasis id="5280EAD6FF9743133AD3E3EDFCA3F817" box="[729,850,1935,1956]" italics="true" pageId="5" pageNumber="256">R. granitum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="348B664CFF944310389DE79FFC4DFBA0" box="[151,956,1021,1044]" httpUri="https://zenodo.org/record/249639/files/figure.png" pageId="6" pageNumber="257" targetBox="[151,1436,193,999]" targetPageId="6">
<paragraph id="604B36C4FF944310389DE79FFC4DFBA0" blockId="6.[151,956,1021,1044]" box="[151,956,1021,1044]" pageId="6" pageNumber="257">
<emphasis id="5280EAD6FF944310389DE79FFEE3FBA0" bold="true" box="[151,274,1021,1043]" pageId="6" pageNumber="257">FIGURE 5.</emphasis>
Details of 
<taxonomicName id="A7F44D47FF9443103988E79FFDB4FBA0" box="[386,581,1021,1043]" class="Insecta" family="Hydrophilidae" genus="Radicitus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="granitum">
<emphasis id="5280EAD6FF9443103988E79FFDB4FBA0" box="[386,581,1021,1043]" italics="true" pageId="6" pageNumber="257">Radicitus granitum</emphasis>
</taxonomicName>
. A) thoracic sternum. B) metafemur.
</paragraph>
</caption>
<paragraph id="604B36C4FF94431038CDE05DFC78FB77" blockId="6.[151,1436,1087,1220]" pageId="6" pageNumber="257">
Biology. This species, the most frequently encountered of the genus to date, has been found on a variety of habitats associated with streams and seeps on rock outcrops. It has been found by collecting detrital debris from seepages and floating them in pans of water (
<figureCitation id="F8CF2A41FF9443103A87E0E5FD21FB13" box="[653,720,1159,1184]" captionStart="FIGURE 9" captionStartId="10.[151,250,1789,1811]" captionTargetBox="[151,1436,805,1768]" captionTargetId="figure@10.[151,1436,805,1768]" captionTargetPageId="10" captionText="FIGURE 9. Habitat of Radicitus spp. in Venezuela. A) Venezuela, Bolivar State, 25 km E. El Burro (collecting event AS- 08 - 077). B) Specimen of R. granitum clinging to root mat that was pulled from the margin of the seepage shown in A. C) Venezuela: Bolivar State, 15 km NE Pijiguaos (collecting event VZ 10 - 0709 - 01 B). D) Venezuela, Amazonas State, Tobogan de la Selva (collecting event AS- 06 - 017)." httpUri="https://zenodo.org/record/249643/files/figure.png" pageId="6" pageNumber="257">Fig. 9</figureCitation>
C), and also in the roots vegetation along the margins of shallow streams and seeps that are flowing over open granite (e.g. 
<figureCitation id="F8CF2A41FF9443103B28E0C9FC94FB77" box="[802,869,1195,1220]" captionStart="FIGURE 9" captionStartId="10.[151,250,1789,1811]" captionTargetBox="[151,1436,805,1768]" captionTargetId="figure@10.[151,1436,805,1768]" captionTargetPageId="10" captionText="FIGURE 9. Habitat of Radicitus spp. in Venezuela. A) Venezuela, Bolivar State, 25 km E. El Burro (collecting event AS- 08 - 077). B) Specimen of R. granitum clinging to root mat that was pulled from the margin of the seepage shown in A. C) Venezuela: Bolivar State, 15 km NE Pijiguaos (collecting event VZ 10 - 0709 - 01 B). D) Venezuela, Amazonas State, Tobogan de la Selva (collecting event AS- 06 - 017)." httpUri="https://zenodo.org/record/249643/files/figure.png" pageId="6" pageNumber="257">Fig. 9</figureCitation>
A).
</paragraph>
</subSubSection>
</treatment>
</document>