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<document id="95962AB2AEFD15C2BCCFC70F40BF1C7C" ID-CLB-Dataset="7964" ID-DOI="10.11646/zootaxa.4903.2.1" ID-GBIF-Dataset="37ab0d60-c931-4718-b85a-00d5087fca50" ID-ISSN="1175-5326" ID-Zenodo-Dep="4422874" ID-ZooBank="339B8883-E0FA-4310-968B-81E7AAC76376" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610012447887" checkinUser="plazi" docAuthor="Yeshwanth, H. M., Chérot, F. &amp; Henry, T. J." docDate="2021" docId="C24787C5BF5AFFFDFF74F61D6D986126" docLanguage="en" docName="zootaxa.4903.2.1.pdf" docOrigin="Zootaxa 4903 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isometopus wolskii Yeshwanth &amp; Chérot &amp; Henry 2021, n. sp." docType="treatment" docVersion="11" lastPageNumber="161" masterDocId="3E7EFFBDBF53FFF7FFE3F062697E627B" masterDocTitle="The Isometopinae (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka: A Review of the Subfamily, with Descriptions of Six New Species" masterLastPageNumber="193" masterPageNumber="151" pageNumber="160" updateTime="1698920153217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="05D2D6B7D7ED89B9C35ED69829FC1AC1">The Isometopinae (Hemiptera: Heteroptera: Miridae) of India and Sri Lanka: A Review of the Subfamily, with Descriptions of Six New Species</mods:title>
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<mods:namePart id="CE2431D86B9A84D2583306D4B3899C21">Yeshwanth, H. M.</mods:namePart>
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<mods:affiliation id="B13706E0BE7845F9D12C6E9A4CC897DB">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560 065, India.</mods:affiliation>
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<mods:namePart id="EB694A096B8496B7E0BB2948D7356E14">Chérot, F.</mods:namePart>
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<mods:affiliation id="C0C846BCE6E6244F69BDE79287BA7542">Département de lEtude du Milieu Naturel et Agricole, Service Public de Wallonie, Gembloux, BE- 5030, Belgium. frederic. cherot @ spw. wallonie. be; https: // orcid. org / 0000 - 0001 - 7918 - 7012</mods:affiliation>
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<mods:namePart id="ACA6BC6632EDBCCA48077E3A3EE7EAE0">Henry, T. J.</mods:namePart>
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<mods:affiliation id="77D85728134933016980A23405DE6524">Systematic Entomology Laboratory, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum of Natural History, MRC- 168, Smithsonian Institution, Washington, DC 20013 - 7013 USA.</mods:affiliation>
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<paragraph id="4A5136D3BF5AFFFEFF74F61D68B164E2" blockId="9.[151,465,1662,1725]" box="[151,463,1662,1689]" pageId="9" pageNumber="160">
<heading id="111981BFBF5AFFFEFF74F61D68B164E2" bold="true" box="[151,463,1662,1689]" fontSize="11" level="1" pageId="9" pageNumber="160" reason="1">
<emphasis id="789AEAC1BF5AFFFEFF74F61D68B164E2" bold="true" box="[151,463,1662,1689]" pageId="9" pageNumber="160">
<taxonomicName id="8DEE4D50BF5AFFFEFF74F61D680064E2" authority="Yeshwanth &amp; Chérot &amp; Henry, 2021" authorityName="Yeshwanth &amp; Chérot &amp; Henry" authorityYear="2021" box="[151,382,1662,1689]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="wolskii" status="sp. nov.">
<emphasis id="789AEAC1BF5AFFFEFF74F61D680064E2" bold="true" box="[151,382,1662,1689]" italics="true" pageId="9" pageNumber="160">Isometopus wolskii</emphasis>
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,
<taxonomicNameLabel id="63A957BABF5AFFFEFE6FF61D68B164E2" box="[396,463,1663,1689]" pageId="9" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="02F46558BF5AFFFEFF74F6C168AF64C6" box="[151,465,1699,1725]" pageId="9" pageNumber="160" type="description">
<paragraph id="4A5136D3BF5AFFFEFF74F6C168AF64C6" blockId="9.[151,465,1662,1725]" box="[151,465,1699,1725]" pageId="9" pageNumber="160">
(
<figureCitation id="D2D52A56BF5AFFFEFF7CF6C1687D64C6" box="[159,259,1699,1725]" captionStart="FIGURES 2" captionStartId="8.[151,264,1759,1783]" captionTargetBox="[199,1412,212,1679]" captionTargetId="figure-22@8.[154,1434,179,1739]" captionTargetPageId="8" captionText="FIGURES 2. Indian Isometopinae: Isometopini. Habitus in dorsal view.A. Isometopus cuneatus (Distant, 1904). B. Isometopus mahal Distant, 1911. C. Isometopus webbi, n. sp. D. Isometopus wolskii, n. sp. E. Isometopus yoshizawai Yasunaga and Duwal, 2006. F. Paloniella feana (Distant, 1904)." figureDoi="http://doi.org/10.5281/zenodo.4422882" httpUri="https://zenodo.org/record/4422882/files/figure.png" pageId="9" pageNumber="160">Figs. 2D</figureCitation>
,
<figureCitation id="D2D52A56BF5AFFFEFEECF6C1685364C6" box="[271,301,1699,1725]" captionStart="FIGURES 6" captionStartId="23.[151,264,1863,1887]" captionTargetBox="[156,1420,192,1827]" captionTargetId="figure-19@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 6. Indian Isometopinae: Gigantometopini, Isometopini and Myiommini. Head in frontal view.A. Astroscopometopus hesaraghattaensis, n. sp. B. Nymph of Astroscopometopus hesaraghattaensis, n sp. C. Isometopus cuneatus (Distant, 1904). D. Isometopus mahal Distant, 1911. E. Isometopus webbi, n. sp. F. Isometopus wolskii, n. sp. G. Isometopus yoshizawai Yasunaga and Duwal, 2006. H. Paloniella feana (Distant, 1904). I. Paloniella parallela Yasunaga &amp; Hayashi, 2002. J. Isometopidea viraktamathi, n. sp. K. Myiomma belavadii, n. sp. L. Myiomma ramamurthyi, n sp." figureDoi="http://doi.org/10.5281/zenodo.4422892" httpUri="https://zenodo.org/record/4422892/files/figure.png" pageId="9" pageNumber="160">6F</figureCitation>
,
<figureCitation id="D2D52A56BF5AFFFEFEDBF6C1682C64C6" box="[312,338,1699,1725]" captionStart="FIGURES 13" captionStartId="30.[151,264,1859,1883]" captionTargetBox="[164,1423,193,1825]" captionTargetId="figure-22@30.[151,1436,181,1836]" captionTargetPageId="30" captionText="FIGURES 13. Indian Isometopinae: Isometopini. Male genital structures and evaporative area. Isometopus wolskii, n. sp. A. Head, pronotum and scutellum in dorsal view. B. Metathoracic scent gland. C. Pygophore. D. Right paramere. E. Left paramere. FG. Phallus." figureDoi="http://doi.org/10.5281/zenodo.4422914" httpUri="https://zenodo.org/record/4422914/files/figure.png" pageId="9" pageNumber="160">13</figureCitation>
AG, 20B)
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF5AFFFEFF74F6886D946552" pageId="9" pageNumber="160" type="diagnosis">
<paragraph id="4A5136D3BF5AFFFEFF74F6886D946552" blockId="9.[151,1437,1770,2013]" pageId="9" pageNumber="160">
<emphasis id="789AEAC1BF5AFFFEFF74F688686C657F" bold="true" box="[151,274,1770,1796]" pageId="9" pageNumber="160">Diagnosis.</emphasis>
Recognized by the pale brown to yellow head, the dorsally pale brown and ventrally dark brown antennal segment II, the ventrally brown pronotum, the apically white scutellum, and the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF5AFFFEFF24F7516D6D650A" pageId="9" pageNumber="160" type="materials_examined">
<paragraph id="4A5136D3BF5AFFFEFF24F7516D6D650A" blockId="9.[151,1437,1770,2013]" pageId="9" pageNumber="160">
<emphasis id="789AEAC1BF5AFFFEFF24F751680F6537" bold="true" box="[199,369,1842,1869]" pageId="9" pageNumber="160">Type material</emphasis>
.
<materialsCitation id="FA863C8EBF5AFFFEFE60F75168B5650A" ID-GBIF-Occurrence="3043834318" collectingDate="2015-07-18" collectorName="Yeshwanth, H. M." country="India" location="Nandi Hills" municipality="Chikkaballapura" pageId="9" pageNumber="160" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="95558871BF5AFFFEFE60F75168956536" box="[387,491,1843,1869]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
(³):
<subScript id="D66A3496BF5AFFFEFDCCF7516B176531" attach="left" box="[559,617,1843,1869]" fontSize="7" pageId="9" pageNumber="160">
<collectingCountry id="32F97643BF5AFFFEFDCCF7516B176531" box="[559,617,1843,1869]" name="India" pageId="9" pageNumber="160">InDIa</collectingCountry>
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:
<collectingRegion id="882AF831BF5AFFFEFD9FF7516B8C6536" box="[636,754,1843,1869]" country="India" name="Karnataka" pageId="9" pageNumber="160">Karnataka</collectingRegion>
,
<collectingMunicipality id="AA35ACA9BF5AFFFEFCE3F7516AC26536" box="[768,956,1843,1869]" pageId="9" pageNumber="160">Chikkaballapura</collectingMunicipality>
,
<location id="4F316008BF5AFFFEFC28F7516D2C6536" LSID="urn:lsid:plazi:treatment:C24787C5BF5AFFFDFF74F61D6D986126:4F316008BF5AFFFEFC28F7516D2C6536" box="[971,1106,1843,1869]" country="India" municipality="Chikkaballapura" name="Nandi Hills" pageId="9" pageNumber="160" stateProvince="Karnataka">Nandi Hills</location>
,
<date id="3E501013BF5AFFFEFB82F7506D9A6536" box="[1121,1252,1842,1869]" pageId="9" pageNumber="160" value="2015-07-18">
<collectingDate id="2E14E9FBBF5AFFFEFB82F7506D9A6536" box="[1121,1252,1842,1869]" pageId="9" pageNumber="160" value="2015-07-18">18.vii.2015</collectingDate>
</date>
, ex:
<taxonomicName id="8DEE4D50BF5AFFFEFAFCF7516C0C6536" box="[1311,1394,1843,1869]" class="Magnoliopsida" family="Malvaceae" genus="Grewia" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="genus">
<emphasis id="789AEAC1BF5AFFFEFAFCF7516C0C6536" box="[1311,1394,1843,1869]" italics="true" pageId="9" pageNumber="160">Grewia</emphasis>
</taxonomicName>
sp. bark,
<collectorName id="E71B5305BF5AFFFEFF35F73568E4650B" box="[214,410,1879,1905]" pageId="9" pageNumber="160">Yeshwanth, H.M.</collectorName>
leg.
</materialsCitation>
<materialsCitation id="FA863C8EBF5AFFFEFE32F7356D71650A" ID-GBIF-Occurrence="3043834313" box="[465,1039,1879,1907]" collectingDate="2015-07-18" collectionCode="UASB" collectorName="Yeshwanth, H. M." country="India" location="Nandi Hills" municipality="Chikkaballapura" pageId="9" pageNumber="160" specimenCount="1" stateProvince="Karnataka" typeStatus="paratype">
<typeStatus id="95558871BF5AFFFEFE32F7356B40650A" box="[465,574,1879,1905]" pageId="9" pageNumber="160" type="paratype">Paratypes</typeStatus>
(3³), same data as for holotype (
<collectionCode id="2CFFAE16BF5AFFFEFC55F7356D79650A" box="[950,1031,1879,1905]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="9" pageNumber="160">UASB</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="02F46558BF5AFFFDFF24F718681F60FE" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160" type="description">
<paragraph id="4A5136D3BF5AFFFDFF24F718681F60FE" blockId="9.[151,1437,1770,2013]" lastBlockId="10.[151,1437,151,861]" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160">
<emphasis id="789AEAC1BF5AFFFEFF24F718682765EF" bold="true" box="[199,345,1914,1940]" pageId="9" pageNumber="160">Description.</emphasis>
<emphasis id="789AEAC1BF5AFFFEFEBCF71968A465EE" box="[351,474,1915,1941]" italics="true" pageId="9" pageNumber="160">Coloration</emphasis>
: Head pale yellow with ocelli reddish brown. Antennal segment II dorsally with basal ¾ pale brown, apical ¼ brown, and evenly brown ventrally. Pronotum ventrally dark brown, dorsally yellowish brown. Mesoscutum and scutellum brown, scutellum with white patches laterally and at apex. Corium pale brown or smoky. Legs usually brown with apical 1/3 of each femur pale brown; all tibiae and tarsi pale brown.
<emphasis id="789AEAC1BF59FFFDFB1CF0F5682662AF" italics="true" pageId="10" pageNumber="161">Surface structure and vestiture</emphasis>
: Head shiny, punctate, dorsum shiny with golden-yellow, flattened, erect setae; distinctly punctate on hemelytra; antennae with short, stout, erect setae. Body oval, length 1.8 times longer than wide. Head in frontal view flat, 1.2 times wider than long, with clypeal margin sinuate, frons with a transverse carina above the clypeus, just below the ventral margins of eyes. Eyes relatively large, oval, occupying 3/4
<superScript id="BD9B9B9BBF59FFFDFBFBF1446D5B634F" attach="left" box="[1048,1061,294,308]" fontSize="6" pageId="10" pageNumber="161">th</superScript>
of the head height in lateral view. Ocelli close to or nearly touching inner margins of eyes. Antenna arising on ventral margin of head; segment I short, rounded; segment II tubular, longest, longer than head width; segment III and IV subequal in length. Labium long, extending to apex of hind coxae. Pronotum 2.7 times wider than long; lateral margins flattened, rounded; posterior margin slightly concave. Mesoscutum broad, with raised lateral margins. Scutellum slightly raised at middle, length subequal to width. Metathoracic scent gland evaporatory area large with a prominent opening (
<figureCitation id="D2D52A56BF59FFFDFB4DF1B96C6A638E" box="[1198,1300,475,501]" captionStart="FIGURES 13" captionStartId="30.[151,264,1859,1883]" captionTargetBox="[164,1423,193,1825]" captionTargetId="figure-22@30.[151,1436,181,1836]" captionTargetPageId="30" captionText="FIGURES 13. Indian Isometopinae: Isometopini. Male genital structures and evaporative area. Isometopus wolskii, n. sp. A. Head, pronotum and scutellum in dorsal view. B. Metathoracic scent gland. C. Pygophore. D. Right paramere. E. Left paramere. FG. Phallus." figureDoi="http://doi.org/10.5281/zenodo.4422914" httpUri="https://zenodo.org/record/4422914/files/figure.png" pageId="10" pageNumber="161">Fig. 13B</figureCitation>
). Hemelytra transparent, with lateral margins weakly rounded. Cuneus triangular with a prominent cuneal fracture. Membrane with a single cell. Hind femur greatly enlarged, cylindrical; tibia slender, elongate; tarsi two segmented. Abdomen rounded, extending to apex of membrane.
<emphasis id="789AEAC1BF59FFFDFD92F2256A68601A" box="[625,790,583,609]" italics="true" pageId="10" pageNumber="161">Male genitalia</emphasis>
: as in
<figureCitation id="D2D52A56BF59FFFDFCBCF2256AC6601A" box="[863,952,583,609]" captionStart="FIGURES 13" captionStartId="30.[151,264,1859,1883]" captionTargetBox="[164,1423,193,1825]" captionTargetId="figure-22@30.[151,1436,181,1836]" captionTargetPageId="30" captionText="FIGURES 13. Indian Isometopinae: Isometopini. Male genital structures and evaporative area. Isometopus wolskii, n. sp. A. Head, pronotum and scutellum in dorsal view. B. Metathoracic scent gland. C. Pygophore. D. Right paramere. E. Left paramere. FG. Phallus." figureDoi="http://doi.org/10.5281/zenodo.4422914" httpUri="https://zenodo.org/record/4422914/files/figure.png" pageId="10" pageNumber="161">Figs. 13</figureCitation>
CG. Detailed measurements: See
<tableCitation id="076C0368BF59FFFDFAA3F2256CEA601A" box="[1344,1428,583,609]" captionStart="TABLE 1" captionStartId="4.[151,239,872,896]" captionTargetPageId="4" captionText="TABLE 1. Measurements of Indian Isometopinae (species collected during present work)" httpUri="http://table.plazi.org/id/1E91665BBF57FFF3FF74F30A6D2A61FB" pageId="10" pageNumber="161" tableUuid="1E91665BBF57FFF3FF74F30A6D2A61FB">Table 1</tableCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F2EC6B4B60D1" box="[199,565,654,682]" pageId="10" pageNumber="161" type="distribution">
<paragraph id="4A5136D3BF59FFFDFF24F2EC6B4B60D1" blockId="10.[151,1437,151,861]" box="[199,565,654,682]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F2EC681F60D3" bold="true" box="[199,353,654,680]" pageId="10" pageNumber="161">Distribution.</emphasis>
<collectingCountry id="32F97643BF59FFFDFE84F2ED68DF60D2" box="[359,417,655,681]" name="India" pageId="10" pageNumber="161">India</collectingCountry>
(
<collectingRegion id="882AF831BF59FFFDFE53F2ED6B5460D1" box="[432,554,655,682]" country="India" name="Karnataka" pageId="10" pageNumber="161">Karnataka</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F2D06A6C60B6" box="[199,786,690,718]" pageId="10" pageNumber="161" type="biology_ecology">
<paragraph id="4A5136D3BF59FFFDFF24F2D06A6C60B6" blockId="10.[151,1437,151,861]" box="[199,786,690,718]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F2D0685F60B7" bold="true" box="[199,289,690,716]" pageId="10" pageNumber="161">Biology</emphasis>
. Collected on
<taxonomicName id="8DEE4D50BF59FFFDFE26F2D16B6660B6" box="[453,536,691,717]" class="Magnoliopsida" family="Malvaceae" genus="Grewia" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="genus">
<emphasis id="789AEAC1BF59FFFDFE26F2D16B6660B6" box="[453,536,691,717]" italics="true" pageId="10" pageNumber="161">Grewia</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="8DEE4D50BF59FFFDFDAEF2D16BB460B5" box="[589,714,691,718]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="161" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
) bark.
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F2B56A66616E" pageId="10" pageNumber="161" type="etymology">
<paragraph id="4A5136D3BF59FFFDFF24F2B56A66616E" blockId="10.[151,1437,151,861]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F2B56830608A" bold="true" box="[199,334,727,753]" pageId="10" pageNumber="161">Etymology.</emphasis>
We are pleased to name this species after our colleague Andrzej Wolski (University of
<collectingRegion id="882AF831BF59FFFDFAB2F2B56CE6608A" box="[1361,1432,727,753]" country="Poland" name="Opolskie" pageId="10" pageNumber="161">Opole</collectingRegion>
,
<collectingRegion id="882AF831BF59FFFDFF74F29969A0616E" box="[151,222,763,789]" country="Poland" name="Opolskie" pageId="10" pageNumber="161">Opole</collectingRegion>
,
<collectingCountry id="32F97643BF59FFFDFF09F2996842616D" box="[234,316,763,790]" name="Poland" pageId="10" pageNumber="161">Poland</collectingCountry>
), Polish specialist of
<taxonomicName id="8DEE4D50BF59FFFDFDCAF2996BE5616E" authorityName="Kirkaldy" authorityYear="1903" box="[553,667,763,789]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subFamily" subFamily="Cylapinae">Cylapinae</taxonomicName>
taxonomy.
</paragraph>
</subSubSection>
<subSubSection id="02F46558BF59FFFDFF24F37C6D986126" pageId="10" pageNumber="161" type="discussion">
<paragraph id="4A5136D3BF59FFFDFF24F37C6D986126" blockId="10.[151,1437,151,861]" pageId="10" pageNumber="161">
<emphasis id="789AEAC1BF59FFFDFF24F37C68326143" bold="true" box="[199,332,798,824]" pageId="10" pageNumber="161">Discussion.</emphasis>
This new species is separated from other species of Indian
<taxonomicName id="8DEE4D50BF59FFFDFC3DF37D6D206143" authorityName="Fieber" authorityYear="1860" box="[990,1118,799,824]" class="Insecta" family="Miridae" genus="Isometopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="genus">
<emphasis id="789AEAC1BF59FFFDFC3DF37D6D206143" box="[990,1118,799,824]" italics="true" pageId="10" pageNumber="161">Isometopus</emphasis>
</taxonomicName>
by the characters mentioned in the diagnosis and the key, mainly the total length, dorsal coloration, and male genital structures.
</paragraph>
</subSubSection>
</treatment>
</document>