271 lines
20 KiB
XML
271 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. & Marek, Paul E." docDate="2021" docId="9D6047026D955AD5865DB4B414B09236" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria caverna Means & Hennen & Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1030</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181004730" LSID="urn:lsid:zoobank.org:act:41468A49-456A-41C0-B973-D66E8B757919" httpUri="http://treatment.plazi.org/id/9D6047026D955AD5865DB4B414B09236" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="8RVV2" LSID="http://zoobank.org/41468A49-456A-41C0-B973-D66E8B757919" authority="Means & Hennen & Marek, 2021" authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria caverna" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caverna" status="sp. nov.">Nannaria caverna</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Nannaria caverna sp. nov. holotype ♂ (VTEC, MPE 03139) left gonopod A anterior view B medial view; red arrow indicates ventrally directed prefemoral process C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure24" httpUri="https://binary.pensoft.net/fig/531127" pageId="0" pageNumber="1">Figs 24</figureCitation>
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Nannaria caverna sp. nov. holotype ♂ (VTEC, MPE 03139) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure25" httpUri="https://binary.pensoft.net/fig/531128" pageId="0" pageNumber="1">, 25 Vernacular name: "The Carter Caves Twisted-Claw Millipede"</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355767" accessionNumber="MPE03139" collectingDate="2017-09-24" collectorName="J. Means, D. Hennen" country="United States" county="Carter Co." latitude="38.3738" location="Carter Caves State Park" longLatPrecision="6" longitude="-83.1142" specimenCount="♂" stateProvince="Kentucky" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Kentucky">Kentucky</collectingRegion>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingCounty>Carter Co.</collectingCounty>
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,
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<location LSID="urn:lsid:plazi:treatment:9D6047026D955AD5865DB4B414B09236:2419BC8ECDFBDB58905F7D11594A82DE" country="United States" county="Carter Co." latitude="38.3738" longLatPrecision="6" longitude="-83.1142" name="Carter Caves State Park" stateProvince="Kentucky">Carter Caves State Park</location>
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;
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<geoCoordinate degrees="38.3738" direction="north" orientation="latitude" precision="5" value="38.3738">38.3738°N</geoCoordinate>
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, -
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<geoCoordinate degrees="83.1142" direction="west" orientation="longitude" precision="5" value="-83.1142">83.1142°W</geoCoordinate>
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;
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<collectingDate value="2017-09-24">24 Sep. 2017</collectingDate>
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; hand collected;
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<collectorName>J. Means</collectorName>
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,
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<collectorName>D. Hennen</collectorName>
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leg.; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03139">MPE03139</accessionNumber>
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.
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355551" accessionNumber="MPE03157" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Carter Co." latitude="38.3738" location="Kentucky" longLatPrecision="6" longitude="-83.1142" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Kentucky">Kentucky</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">1 ♂</specimenCount>
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; same collection data as holotype; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03157">MPE03157</accessionNumber>
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</materialsCitation>
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•
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<materialsCitation ID-GBIF-Occurrence="3071355548" accessionNumber="MPE03158" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Carter Co." latitude="38.3738" location="Carter Caves State Park" longLatPrecision="6" longitude="-83.1142" specimenCount="1" specimenCount-female="1" stateProvince="Kentucky" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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; same collection data as holotype; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03158">MPE03158</accessionNumber>
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</materialsCitation>
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•
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<materialsCitation ID-GBIF-Occurrence="3071355419" accessionNumber="MPE03160" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Carter Co." latitude="38.3738" location="Carter Caves State Park" longLatPrecision="6" longitude="-83.1142" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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; same collection data as holotype; VMNH
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03160">MPE03160</accessionNumber>
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</materialsCitation>
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•
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<materialsCitation ID-GBIF-Occurrence="3071355403" accessionNumber="MPE03159" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Carter Co." latitude="38.3738" location="Carter Caves State Park" longLatPrecision="6" longitude="-83.1142" specimenCount="1" specimenCount-female="1" stateProvince="Kentucky" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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; same collection data as holotye; VMNH
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03159">MPE03159</accessionNumber>
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</materialsCitation>
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. For detailed collection data see Suppl. material 7.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult males of
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria caverna" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caverna">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria caverna</emphasis>
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</taxonomicName>
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sp. nov. are distinct from other
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<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
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</taxonomicName>
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and nearby
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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, based on the following combination of characters:
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
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.
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</emphasis>
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Gonopodal acropodite linear, bent medially forming 130° angle with telopodite basal zone bent medially at tip, not continually curving throughout as
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. serpens" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="serpens">
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<emphasis italics="true" pageId="0" pageNumber="1">N. serpens</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Tip simple, rectangular, without laminate flanges as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Telopodite basal zone tall,> 1/2 length of acropodite, not <1/2 as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. serpens" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="serpens">
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<emphasis italics="true" pageId="0" pageNumber="1">N. serpens</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Prefemur with sinuous, ventrally directed prefemoral process, bent at a 90° angle, arising from large, pronounced prefemoral spine (Fig.
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Nannaria caverna sp. nov. holotype ♂ (VTEC, MPE 03139) left gonopod A anterior view B medial view; red arrow indicates ventrally directed prefemoral process C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure24" httpUri="https://binary.pensoft.net/fig/531127" pageId="0" pageNumber="1">24B</figureCitation>
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, red arrow), not directed cephalically as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. serpens" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="serpens">
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<emphasis italics="true" pageId="0" pageNumber="1">N. serpens</emphasis>
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</taxonomicName>
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sp. nov., or curving laterally as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
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.
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</emphasis>
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Tergites with light orange paranotal spots (Fig.
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Nannaria caverna sp. nov. holotype ♂ (VTEC, MPE 03139) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure25" httpUri="https://binary.pensoft.net/fig/531128" pageId="0" pageNumber="1">25</figureCitation>
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). Light grey background, likely due to teneral condition of specimens. Collum smooth with orange margin.
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</paragraph>
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<caption doi="10.3897/zookeys.1030.62544.figure24" httpUri="https://binary.pensoft.net/fig/531127" pageId="0" pageNumber="1" start="Figure 24" startId="F24">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 24.</emphasis>
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria caverna" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caverna">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria caverna</emphasis>
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</taxonomicName>
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sp. nov. holotype ♂ (VTEC, MPE03139) left gonopod
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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anterior view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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medial view; red arrow indicates ventrally directed prefemoral process
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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posterior view. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1030.62544.figure25" httpUri="https://binary.pensoft.net/fig/531128" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria caverna" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caverna">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria caverna</emphasis>
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</taxonomicName>
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sp. nov. holotype ♂ (VTEC, MPE03139) coloration. Scale bar: 4.0 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="1">♂ holotype (VTEC, MPE03139): BL = 24.7, CW = 3.8, IW = 2.5, ISW = 0.8, B11W = 4.2, B11H = 2.9. ♀ paratype (VMNH, MPE03159): BL = 25.7, CW = 3.2, IW = 2.2, ISW = 1.2, B11W = 4.0, B11H = 2.9.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Known only from the type locality (Kentucky: Carter Caves State Park; Suppl. material 7; Fig.
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<figureCitation captionStart="Figure 127" captionStartId="F127" captionText="Figure 127. Distribution map of Nannaria minor species group collection localities in the northern Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure127" httpUri="https://binary.pensoft.net/fig/531230" pageId="0" pageNumber="1">127</figureCitation>
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). Distribution area: N/A; status: MRE.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Individuals of
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. caverna" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="caverna">
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<emphasis italics="true" pageId="0" pageNumber="1">N. caverna</emphasis>
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</taxonomicName>
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sp. nov. were collected as they emerged from hardpacked soil on the edge of a creek running through Carter Caves State Park. The surrounding forest was made up of hardwood trees and rhododendron thickets.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is named for its type locality. The specific name is a noun in apposition from the Latin
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<emphasis italics="true" pageId="0" pageNumber="1">caverna</emphasis>
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for cave.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
|
|
United States, Kentucky, Carter Co., Carter Caves State Park,
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|
<geoCoordinate degrees="38.3738" direction="north" orientation="latitude" precision="5" value="38.3738">38.3738°N</geoCoordinate>
|
|
, -
|
|
<geoCoordinate degrees="83.1142" direction="west" orientation="longitude" precision="5" value="-83.1142">83.1142°W</geoCoordinate>
|
|
.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |