treatments-xml/data/03/B9/3E/03B93E4AFFEF12191DB8DFE5FAECFEAF.xml

<document id="BF219EC0A076D0B6D62DD413012C5E44" ID-DOI="10.11646/zootaxa.5537.2.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239411" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732800235064" checkinUser="plazi" docAuthor="Kim, Junggon, Phan, Quoc Toan, Pham, Hong Thai &amp; Jung, Sunghoon" docDate="2024" docId="03B93E4AFFEF12191DB8DFE5FAECFEAF" docLanguage="en" docName="zootaxa.5537.2.10.pdf" docOrigin="Zootaxa 5537 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5537.2.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liocapsus vietnamensis Kim et Jung 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="298" masterDocId="FF804632FFEE121D1D2FDB7FFFF9FFCC" masterDocTitle="First record of the genus Liocapsus Poppius, 1915 (Hemiptera: Heteroptera: Miridae) from Vietnam, with description of a new species" masterLastPageNumber="300" masterPageNumber="295" pageNumber="296" updateTime="1732838100487" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="85B44282F55483766574A08178C969B3">First record of the genus Liocapsus Poppius, 1915 (Hemiptera: Heteroptera: Miridae) from Vietnam, with description of a new species</mods:title>
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<mods:namePart id="8D38AF1F2E69ACE3706E8C95960E5DCB">Kim, Junggon</mods:namePart>
<mods:affiliation id="F8647AA2AD97FFC6CFD77B859FF5E87E">The Center for Entomology &amp; Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, Da Nang 550000, Vietnam</mods:affiliation>
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<mods:namePart id="8E8AF34489E0B6B81772783D38B75799">Phan, Quoc Toan</mods:namePart>
<mods:affiliation id="9D78F018C95739668B05FD0C4D6B36D2">The Center for Entomology &amp; Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, Da Nang 550000, Vietnam</mods:affiliation>
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<mods:namePart id="C00C0DCA46F0C70C28E837195DAFD333">Pham, Hong Thai</mods:namePart>
<mods:nameIdentifier id="FA60BADE1F3A4722D6780262D760C33F" type="ORCID">0000-0002-4763-3679</mods:nameIdentifier>
<mods:affiliation id="F6015FA3C2AB8DCB103EA77682CE30F7">Graduate School of Science and Technology, VAST, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="B1717870A621EA3C5E4E351880E61EB7">Jung, Sunghoon</mods:namePart>
<mods:affiliation id="1EBE947CED386B9889721F0FF0A0D398">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and Life Sciences, Chungnam National University, Daejeon, Korea &amp; Department of Smart Agriculture Systems, College of Agriculture and Life Sciences, Chungnam National University, Daejeon, Korea * Contributed equally as the corresponding author</mods:affiliation>
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<mods:date id="DCCFE4B2B1DEDAEB9803E98B665716E4">2024</mods:date>
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<subSubSection id="C30ADCD7FFEF121C1DB8DFE5FD45FB78" box="[151,700,1177,1204]" pageId="1" pageNumber="296" type="nomenclature">
<paragraph id="8BAF8F5CFFEF121C1DB8DFE5FD45FB78" blockId="1.[151,1437,1177,2032]" box="[151,700,1177,1204]" pageId="1" pageNumber="296">
<heading id="D0E73830FFEF121C1DB8DFE5FD45FB78" bold="true" box="[151,700,1177,1204]" fontSize="11" level="1" pageId="1" pageNumber="296" reason="1">
<emphasis id="B964534EFFEF121C1DB8DFE5FD45FB78" bold="true" box="[151,700,1177,1204]" pageId="1" pageNumber="296">
<taxonomicName id="4C10F4DFFFEF121C1DB8DFE5FDAEFB78" authority="Kim et Jung" authorityName="Kim et Jung" authorityYear="2024" box="[151,599,1177,1204]" class="Insecta" family="Miridae" genus="Liocapsus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B964534EFFEF121C1DB8DFE5FE4CFB78" bold="true" box="[151,437,1178,1204]" italics="true" pageId="1" pageNumber="296">Liocapsus vietnamensis</emphasis>
Kim et Jung
</taxonomicName>
<taxonomicNameLabel id="A257EE35FFEF121C1F70DFE5FD45FB78" box="[607,700,1178,1204]" pageId="1" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BAF8F5CFFEF121C1DB8DFC0FEF5FB14" blockId="1.[151,1437,1177,2032]" box="[151,268,1215,1240]" pageId="1" pageNumber="296">
(
<figureCitation id="132B93D9FFEF121C1DB0DFC0FEFAFB14" box="[159,259,1215,1240]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1485,1509]" captionStartId-1="3.[151,250,1045,1069]" captionStartId-2="3.[151,250,1612,1636]" captionTargetBox-0="[159,1429,604,1454]" captionTargetBox-1="[160,1427,353,1013]" captionTargetBox-2="[360,1235,1138,1575]" captionTargetId-0="figure-251@2.[151,1436,596,1462]" captionTargetId-1="figure-66@3.[151,1436,344,1022]" captionTargetId-2="figure-92@3.[347,1241,1130,1588]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. Dorsal and lateral habitus and head and thorax in ventral view of Liocapsus vietnamensis sp. nov. (A–D) dorsal habitus; (A) Holotype, Male; (B) Paratype, Male; (C) Paratype, Female; (D) Paratype, Female; (E) lateral habitus, Female; (F) head and thorax in ventral view, Female. Scale bars: 1 mm." captionText-1="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." captionText-2="FIGURE 3. Female genitalia of Liocapsus vietnamensis sp. nov. (A) genital chamber and adjacent structures; (B) posterior wall. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14239413" figureDoi-1="http://doi.org/10.5281/zenodo.14239415" figureDoi-2="http://doi.org/10.5281/zenodo.14239417" httpUri-0="https://zenodo.org/record/14239413/files/figure.png" httpUri-1="https://zenodo.org/record/14239415/files/figure.png" httpUri-2="https://zenodo.org/record/14239417/files/figure.png" pageId="1" pageNumber="296">Figs. 1–3</figureCitation>
)
</paragraph>
<paragraph id="8BAF8F5CFFEF121C1DB8DF9BFB26FA7C" blockId="1.[151,1437,1177,2032]" pageId="1" pageNumber="296">
<emphasis id="B964534EFFEF121C1DB8DF9BFEFAFB37" box="[151,259,1252,1275]" italics="true" pageId="1" pageNumber="296">Diagnosis</emphasis>
. Recognized by dorsal habitus being mostly pale; head mostly pale brown with red markings on frons; second antennal segment is bicolor; the pronotum is mostly pale brown; callosites with a pair of dark markings, the variable size; scutellum mostly pale with brownish markings; hemelytra mostly pale brown; inner part of clavus somewhat darker; posterior part of corium darker; cuneus subhyaline with apical dark marking tinged with reddish brown (
<figureCitation id="132B93D9FFEF121C19DBDE30FACBFAA4" box="[1268,1330,1359,1384]" captionStart="FIGURE 1" captionStartId="2.[151,250,1485,1509]" captionTargetBox="[159,1429,604,1454]" captionTargetId="figure-251@2.[151,1436,596,1462]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and lateral habitus and head and thorax in ventral view of Liocapsus vietnamensis sp. nov. (A–D) dorsal habitus; (A) Holotype, Male; (B) Paratype, Male; (C) Paratype, Female; (D) Paratype, Female; (E) lateral habitus, Female; (F) head and thorax in ventral view, Female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239413" httpUri="https://zenodo.org/record/14239413/files/figure.png" pageId="1" pageNumber="296">Fig. 1</figureCitation>
); sensory lobe of left paramere blunt and slightly projected (
<figureCitation id="132B93D9FFEF121C1F80DE0CFCDCFA40" box="[687,805,1395,1420]" captionStart="FIGURE 2" captionStartId="3.[151,250,1045,1069]" captionTargetBox="[160,1427,353,1013]" captionTargetId="figure-66@3.[151,1436,344,1022]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="1" pageNumber="296">Fig. 2A–B</figureCitation>
); right paramere with broadly developed sensory lobe (
<figureCitation id="132B93D9FFEF121C185EDE0CFF21FA7C" captionStart="FIGURE 2" captionStartId="3.[151,250,1045,1069]" captionTargetBox="[160,1427,353,1013]" captionTargetId="figure-66@3.[151,1436,344,1022]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="1" pageNumber="296">Fig. 2C–E</figureCitation>
); endosoma with distinctly long, thick and sinuous spicule; median sclerite lacking (
<figureCitation id="132B93D9FFEF121C1974DEE8FB2AFA7C" box="[1115,1235,1431,1456]" captionStart="FIGURE 2" captionStartId="3.[151,250,1045,1069]" captionTargetBox="[160,1427,353,1013]" captionTargetId="figure-66@3.[151,1436,344,1022]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="1" pageNumber="296">Fig. 2G–H</figureCitation>
).
</paragraph>
<paragraph id="8BAF8F5CFFEF121F1DE8DEC3FB30FDF0" blockId="1.[151,1437,1177,2032]" lastBlockId="2.[151,1437,151,572]" lastPageId="2" lastPageNumber="297" pageId="1" pageNumber="296">
<emphasis id="B964534EFFEF121C1DE8DEC3FEBCFA1F" box="[199,325,1468,1491]" italics="true" pageId="1" pageNumber="296">Description</emphasis>
. 
<emphasis id="B964534EFFEF121C1C7DDEC4FE52FA1F" bold="true" box="[338,427,1467,1491]" pageId="1" pageNumber="296">MALE:</emphasis>
Body elongated, length 6.27–6.31. 
<emphasis id="B964534EFFEF121C1E07DEC4FC64FA1F" box="[808,925,1467,1491]" italics="true" pageId="1" pageNumber="296">Coloration</emphasis>
: mostly pale brown with dark parts. 
<emphasis id="B964534EFFEF121C180BDEC3FA90FA18" bold="true" box="[1316,1385,1468,1492]" pageId="1" pageNumber="296">Head:</emphasis>
pale brown with reddish transverse stripes on frons, in dark individuals reddish transverse stripes more distinct; clypeus entirely brown; antennae partly brown and dark brown; first antennal segment mostly pale brown except for dark base and apex, dark individuals first segment mostly dark brown; second antennal segment mostly brown, apical part dark brown; third and fourth segments mostly dark brown with pale basal part. 
<emphasis id="B964534EFFEF121C1E06DD33FC7EF9A8" bold="true" box="[809,903,1612,1636]" pageId="1" pageNumber="296">Thorax:</emphasis>
pronotum mostly pale brown with dark markings variable in size; pronotal collar entirely pale brown; calli variable, margin dark brown or calli entirely dark brown; scutellum mostly pale brown with dark spots variable in size, dark specimens mostly dark brown; hemelytra mostly pale brown with dark areas; clavus mostly dark brown, inner area darker; corium pale brown and dark brown; anterior part pale brown, posterior part dark brown; cuneus subhyaline with dark marking at posterior part, dark marking tinged with red; vein reddish brown; legs mostly pale brown with small dark spots (hindleg missing); femur entirely pale brown with tiny darker spots, very weakly tinged with reddish brown ventrally; tibia mostly pale brown except for darker apical part, with small spots where spines originating; tarsus entirely brown. 
<emphasis id="B964534EFFEF121C1E7FDC38FC31F893" bold="true" box="[848,968,1863,1887]" pageId="1" pageNumber="296">Abdomen:</emphasis>
pale brown with small dark spots ventrally. 
<emphasis id="B964534EFFEF121C1DB8DC13FE8FF84F" box="[151,374,1899,1924]" italics="true" pageId="1" pageNumber="296">Surface and vestiture</emphasis>
: body mostly glossy, covered with short silvery pubescence; head with transverse wrinkle; pronotum weakly and slightly punctate; scutellum impunctate; hemelytra sparsely covered with silvery pubescence, weakly punctate. 
<emphasis id="B964534EFFEF121C1DB8DCCBFF00F807" box="[151,249,1972,1995]" italics="true" pageId="1" pageNumber="296">Structure</emphasis>
: 
<emphasis id="B964534EFFEF121C1C26DCCBFEB7F800" bold="true" box="[265,334,1972,1996]" pageId="1" pageNumber="296">Head:</emphasis>
prognathous; vertex not sulcate, width subequal to single compound eye width; antennae shorter than body length; first antennal segment as thick as second antennal segment (except for the base of second segment), longer than vertex width; second antennal segment slightly clavate, equal to 3 x first segment; third antennal segment longer than first segment, and fourth segment, respectively; fourth segment shorter than first segment; proportion of first to fourth antennal segments 0.6: 1.8: 0.9: 0.4; labrum subequal to 1/2 first labial segment length; labium remarkably short, apex not reaching base of midcoxa (
<figureCitation id="132B93D9FFEC121F1F33DA7CFD75FED0" box="[540,652,259,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1485,1509]" captionTargetBox="[159,1429,604,1454]" captionTargetId="figure-251@2.[151,1436,596,1462]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal and lateral habitus and head and thorax in ventral view of Liocapsus vietnamensis sp. nov. (A–D) dorsal habitus; (A) Holotype, Male; (B) Paratype, Male; (C) Paratype, Female; (D) Paratype, Female; (E) lateral habitus, Female; (F) head and thorax in ventral view, Female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239413" httpUri="https://zenodo.org/record/14239413/files/figure.png" pageId="2" pageNumber="297">Fig. 1E–F</figureCitation>
). 
<emphasis id="B964534EFFEC121F1FB0DA7BFD04FED0" bold="true" box="[671,765,260,284]" pageId="2" pageNumber="297">Thorax:</emphasis>
pronotum trapezoid, longitudinal length shorter than 1/2 basal maximal width, posterior margin sinuate; calli distinctly swollen; pronotal collar medial length shorter than first antennal segment diameter; scutellum somewhat large, anterior width longer than 1/2 basal pronotal width, longitudinal length longer than commissure; lateral margin of hemelytra almost straight; cuneus elongate; legs long and slender. 
<emphasis id="B964534EFFEC121F180BDA10FA65FE4B" bold="true" box="[1316,1436,367,391]" pageId="2" pageNumber="297">Abdomen:</emphasis>
rounded, not reaching to apex of cuneus. 
<emphasis id="B964534EFFEC121F1F68DAECFD40FE67" bold="true" box="[583,697,403,427]" pageId="2" pageNumber="297">Genitalia:</emphasis>
phallotheca with fin-like process (
<emphasis id="B964534EFFEC121F190ADAEBFBC3FE60" box="[1061,1082,404,428]" italics="true" pageId="2" pageNumber="297">fp</emphasis>
); left paramere with hooked apex of hypophysis (
<emphasis id="B964534EFFEC121F1C11DAC8FEA0FE03" box="[318,345,439,463]" italics="true" pageId="2" pageNumber="297">hp</emphasis>
) and narrow sensory lobe with blunt outer margin (
<emphasis id="B964534EFFEC121F1EAEDAC8FC6AFE03" box="[897,915,439,463]" italics="true" pageId="2" pageNumber="297">sl</emphasis>
); (
<figureCitation id="132B93D9FFEC121F1E9DDAC8FBD1FE1C" box="[946,1064,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1045,1069]" captionTargetBox="[160,1427,353,1013]" captionTargetId="figure-66@3.[151,1436,344,1022]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="2" pageNumber="297">Fig. 2A–B</figureCitation>
); right paramere with hooked apex of hypophysis (
<emphasis id="B964534EFFEC121F1C6FDAA4FEA2FE3F" box="[320,347,475,499]" italics="true" pageId="2" pageNumber="297">hp</emphasis>
) and with remarkably developed sensory lobe (
<emphasis id="B964534EFFEC121F1E4CDAA4FC8CFE3F" box="[867,885,475,499]" italics="true" pageId="2" pageNumber="297">sl</emphasis>
), its outer margin rounded (
<figureCitation id="132B93D9FFEC121F1988DAA4FAE5FE38" box="[1191,1308,475,500]" captionStart="FIGURE 2" captionStartId="3.[151,250,1045,1069]" captionTargetBox="[160,1427,353,1013]" captionTargetId="figure-66@3.[151,1436,344,1022]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="2" pageNumber="297">Fig. 2C–E</figureCitation>
); endosoma membranous with spicule and sclerites, spicule (
<emphasis id="B964534EFFEC121F1FB2D97FFD4CFDDB" box="[669,693,512,535]" italics="true" pageId="2" pageNumber="297">sp</emphasis>
) long and curved near secondary gonopore (
<emphasis id="B964534EFFEC121F19BCD97FFB57FDDB" box="[1171,1198,512,535]" italics="true" pageId="2" pageNumber="297">gp</emphasis>
), its apical part sharp; lateral sclerite I (
<emphasis id="B964534EFFEC121F1C62D95CFEA6FDF7" box="[333,351,547,571]" italics="true" pageId="2" pageNumber="297">ls</emphasis>
I) thin and long near 
<emphasis id="B964534EFFEC121F1F6AD95BFDA4FDF7" box="[581,605,548,571]" italics="true" pageId="2" pageNumber="297">sp</emphasis>
; lateral sclerite II (
<emphasis id="B964534EFFEC121F1E06D95CFCC2FDF7" box="[809,827,547,571]" italics="true" pageId="2" pageNumber="297">ls</emphasis>
II) thin and rod-shaped (
<figureCitation id="132B93D9FFEC121F1969D95CFB44FDF0" box="[1094,1213,547,572]" captionStart="FIGURE 2" captionStartId="3.[151,250,1045,1069]" captionTargetBox="[160,1427,353,1013]" captionTargetId="figure-66@3.[151,1436,344,1022]" captionTargetPageId="3" captionText="FIGURE 2. Male genitalia of Liocapsus vietnamensis sp. nov. (A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="2" pageNumber="297">Fig. 2G–H</figureCitation>
).
</paragraph>
<caption id="DF6FDFD4FFEC121F1DB8DEB2FD09F9E1" ID-DOI="http://doi.org/10.5281/zenodo.14239413" ID-Zenodo-Dep="14239413" httpUri="https://zenodo.org/record/14239413/files/figure.png" pageId="2" pageNumber="297" startId="2.[151,250,1485,1509]" targetBox="[159,1429,604,1454]" targetPageId="2" targetType="figure">
<paragraph id="8BAF8F5CFFEC121F1DB8DEB2FD09F9E1" blockId="2.[151,1437,1485,1581]" pageId="2" pageNumber="297">
<emphasis id="B964534EFFEC121F1DB8DEB2FEEDFA29" bold="true" box="[151,276,1485,1509]" pageId="2" pageNumber="297">FIGURE 1.</emphasis>
Dorsal and lateral habitus and head and thorax in ventral view of 
<taxonomicName id="4C10F4DFFFEC121F1EEDDEB1FB4AFA29" authorityName="Kim &amp; Phan &amp; Pham &amp; Jung" authorityYear="2024" box="[962,1203,1486,1509]" class="Insecta" family="Miridae" genus="Liocapsus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B964534EFFEC121F1EEDDEB1FB4AFA29" box="[962,1203,1486,1509]" italics="true" pageId="2" pageNumber="297">Liocapsus vietnamensis</emphasis>
</taxonomicName>
<emphasis id="B964534EFFEC121F1994DEB2FAF4FA29" bold="true" box="[1211,1293,1485,1509]" pageId="2" pageNumber="297">
<taxonomicNameLabel id="A257EE35FFEC121F1994DEB2FAF4FA29" box="[1211,1293,1485,1509]" pageId="2" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A–D) dorsal habitus; (A) Holotype, Male; (B) Paratype, Male; (C) Paratype, Female; (D) Paratype, Female; (E) lateral habitus, Female; (F) head and thorax in ventral view, Female. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BAF8F5CFFEC121F1DE8DDFEFAA5F881" blockId="2.[151,1437,1665,2014]" pageId="2" pageNumber="297">
<emphasis id="B964534EFFEC121F1DE8DDFEFEBBF955" bold="true" box="[199,322,1665,1689]" pageId="2" pageNumber="297">FEMALE:</emphasis>
Body elongated, length 6.65–6.77. 
<emphasis id="B964534EFFEC121F1F94DDFEFCC9F955" box="[699,816,1665,1689]" italics="true" pageId="2" pageNumber="297">Coloration</emphasis>
: as in male. 
<emphasis id="B964534EFFEC121F1E9CDDFEFB6AF955" box="[947,1171,1665,1689]" italics="true" pageId="2" pageNumber="297">Surface and vestiture</emphasis>
: as in male. 
<emphasis id="B964534EFFEC121F1838DDFDFA80F955" box="[1303,1401,1666,1689]" italics="true" pageId="2" pageNumber="297">Structure</emphasis>
: as in male except for the proportion of the antennal segments (0.6: 1.7: 0.9: 0.4). 
<emphasis id="B964534EFFEC121F1EE3DDDAFBC6F971" bold="true" box="[972,1087,1701,1725]" pageId="2" pageNumber="297">Genitalia:</emphasis>
genital chamber with sclerotized rings (
<emphasis id="B964534EFFEC121F1DF5DDB5FF16F92D" box="[218,239,1738,1761]" italics="true" pageId="2" pageNumber="297">sr</emphasis>
), paired sclerotized structure (
<emphasis id="B964534EFFEC121F1F03DDB5FDB8F92D" box="[556,577,1738,1761]" italics="true" pageId="2" pageNumber="297">ss</emphasis>
), and inflated dorsal sac (
<emphasis id="B964534EFFEC121F1E65DDB6FC96F92D" box="[842,879,1737,1761]" italics="true" pageId="2" pageNumber="297">dos</emphasis>
) (
<figureCitation id="132B93D9FFEC121F1EAADDB6FC21F92D" box="[901,984,1737,1761]" captionStart="FIGURE 3" captionStartId="3.[151,250,1612,1636]" captionTargetBox="[360,1235,1138,1575]" captionTargetId="figure-92@3.[347,1241,1130,1588]" captionTargetPageId="3" captionText="FIGURE 3. Female genitalia of Liocapsus vietnamensis sp. nov. (A) genital chamber and adjacent structures; (B) posterior wall. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14239417" httpUri="https://zenodo.org/record/14239417/files/figure.png" pageId="2" pageNumber="297">Fig. 3A</figureCitation>
); 
<emphasis id="B964534EFFEC121F1EC6DDB5FC07F92D" box="[1001,1022,1738,1761]" italics="true" pageId="2" pageNumber="297">sr</emphasis>
distinctly thick and subtriangular, outer two margins long, inner margin short; 
<emphasis id="B964534EFFEC121F1F17DD91FDB4F8C9" box="[568,589,1774,1797]" italics="true" pageId="2" pageNumber="297">ss</emphasis>
trapezoidal, contiguous each other; 
<emphasis id="B964534EFFEC121F1EFADD92FC03F8C9" box="[981,1018,1773,1797]" italics="true" pageId="2" pageNumber="297">dos</emphasis>
swollen; posterior wall with relatively simple interramal lobe (
<emphasis id="B964534EFFEC121F1CB6DC6EFE4BF8E5" box="[409,434,1809,1833]" italics="true" pageId="2" pageNumber="297">irl</emphasis>
) and sclerite (
<emphasis id="B964534EFFEC121F1F61DC6DFD93F8E5" box="[590,618,1810,1833]" italics="true" pageId="2" pageNumber="297">irs</emphasis>
) and with distinctly widened dorsal structure (
<emphasis id="B964534EFFEC121F194EDC6EFB80F8E5" box="[1121,1145,1809,1833]" italics="true" pageId="2" pageNumber="297">ds</emphasis>
) (
<figureCitation id="132B93D9FFEC121F19BEDC6EFB1CF8E5" box="[1169,1253,1809,1833]" captionStart="FIGURE 3" captionStartId="3.[151,250,1612,1636]" captionTargetBox="[360,1235,1138,1575]" captionTargetId="figure-92@3.[347,1241,1130,1588]" captionTargetPageId="3" captionText="FIGURE 3. Female genitalia of Liocapsus vietnamensis sp. nov. (A) genital chamber and adjacent structures; (B) posterior wall. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14239417" httpUri="https://zenodo.org/record/14239417/files/figure.png" pageId="2" pageNumber="297">Fig. 3B</figureCitation>
); 
<emphasis id="B964534EFFEC121F19D4DC6EFAEDF8E5" box="[1275,1300,1809,1833]" italics="true" pageId="2" pageNumber="297">irl</emphasis>
with minute spinules, its apex curved, not surpassing 
<emphasis id="B964534EFFEC121F1F64DC49FD9EF881" box="[587,615,1846,1869]" italics="true" pageId="2" pageNumber="297">irs</emphasis>
; 
<emphasis id="B964534EFFEC121F1F5ADC49FD68F881" box="[629,657,1846,1869]" italics="true" pageId="2" pageNumber="297">irs</emphasis>
rounded, not distinctly contiguous medially; 
<emphasis id="B964534EFFEC121F1959DC4AFB77F881" box="[1142,1166,1845,1869]" italics="true" pageId="2" pageNumber="297">ds</emphasis>
distinctly bilobate.
</paragraph>
<paragraph id="8BAF8F5CFFEC121E1DE8DC25FD76FE8C" blockId="2.[151,1437,1665,2014]" lastBlockId="3.[151,1437,151,320]" lastPageId="3" lastPageNumber="298" pageId="2" pageNumber="297">
<emphasis id="B964534EFFEC121F1DE8DC25FE44F8BD" box="[199,445,1882,1905]" italics="true" pageId="2" pageNumber="297">Measurements (in mm).</emphasis>
Male (n=2)/Female (n=2) Body length, clypeus–apex of membrane:6.17–6.31/6.65–6.77; head length, excluding collar: 0.40–0.41/0.42–0.43; head width, including compound eyes: 0.96–1.09/1.12–1.17; vertex width: 0.40–0.41/0.48–0.49; 1 
<superScript id="7C652214FFEC121F1CBDDCDFFE65F862" attach="left" box="[402,412,1952,1966]" fontSize="6" pageId="2" pageNumber="297">st</superScript>
antennal segment length: 0.56–0.57/0.62–0.63; 2 
<superScript id="7C652214FFEC121F1E94DCDFFC32F862" attach="left" box="[955,971,1952,1966]" fontSize="6" pageId="2" pageNumber="297">nd</superScript>
antennal segment length: 1.83–1.85/1.70– 1.71; 3 
<superScript id="7C652214FFEC121F1DCEDCBBFF17F81E" attach="left" box="[225,238,1988,2002]" fontSize="6" pageId="2" pageNumber="297">rd</superScript>
antennal segment length: 0.93–0.94/0.88–0.90; 4 
<superScript id="7C652214FFEC121F1FD1DCBBFCF3F81E" attach="left" box="[766,778,1988,2002]" fontSize="6" pageId="2" pageNumber="297">th</superScript>
antennal segment length: 0.39–0.42/0.42–0.43; total antennal length: 3.71–3.78/3.62–3.67; anterior pronotal maximal width (straight): 0.82–0.86/0.87–0.89; mesal pronotal length: 0.89–0.97/1.14–1.15; posterior pronotal maximal width (straight): 2.06–2.26/2.51–2.64; anterior scutellar width: 1.31– 1.43/1.66–1.74; mesal scutellar length: 1.10–1.29/1.28–1.29; commissure length: 1.05–1.06/1.01–1.13; outer embolial margin length (straight): 2.65–3.16/3.44–3.55; outer cuneal margin length (straight): 0.96–1.01/1.07–1.25; maximal width across hemelytron: 1.10–1.11/1.14–1.21.
</paragraph>
<caption id="DF6FDFD4FFED121E1DB8DF6AFDD8FB9D" ID-DOI="http://doi.org/10.5281/zenodo.14239415" ID-Zenodo-Dep="14239415" httpUri="https://zenodo.org/record/14239415/files/figure.png" pageId="3" pageNumber="298" startId="3.[151,250,1045,1069]" targetBox="[160,1427,353,1013]" targetPageId="3" targetType="figure">
<paragraph id="8BAF8F5CFFED121E1DB8DF6AFDD8FB9D" blockId="3.[151,1437,1045,1106]" pageId="3" pageNumber="298">
<emphasis id="B964534EFFED121E1DB8DF6AFEECFBE1" bold="true" box="[151,277,1045,1069]" pageId="3" pageNumber="298">FIGURE 2.</emphasis>
Male genitalia of 
<taxonomicName id="4C10F4DFFFED121E1CF8DF69FD30FBE1" authorityName="Kim &amp; Phan &amp; Pham &amp; Jung" authorityYear="2024" box="[471,713,1046,1069]" class="Insecta" family="Miridae" genus="Liocapsus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B964534EFFED121E1CF8DF69FD30FBE1" box="[471,713,1046,1069]" italics="true" pageId="3" pageNumber="298">Liocapsus vietnamensis</emphasis>
</taxonomicName>
<emphasis id="B964534EFFED121E1FFEDF69FCDDFBE2" bold="true" box="[721,804,1046,1070]" pageId="3" pageNumber="298">
<taxonomicNameLabel id="A257EE35FFED121E1FFEDF69FCDDFBE2" box="[721,804,1046,1070]" pageId="3" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A–B) left paramere; (C–E) right paramere; (F) phallotheca; (G–H) Endosoma. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="DF6FDFD4FFED121E1DB8DD33FE60F94B" ID-DOI="http://doi.org/10.5281/zenodo.14239417" ID-Zenodo-Dep="14239417" httpUri="https://zenodo.org/record/14239417/files/figure.png" pageId="3" pageNumber="298" startId="3.[151,250,1612,1636]" targetBox="[360,1235,1138,1575]" targetPageId="3" targetType="figure">
<paragraph id="8BAF8F5CFFED121E1DB8DD33FE60F94B" blockId="3.[151,1436,1612,1672]" pageId="3" pageNumber="298">
<emphasis id="B964534EFFED121E1DB8DD33FEECF9A8" bold="true" box="[151,277,1612,1636]" pageId="3" pageNumber="298">FIGURE 3.</emphasis>
Female genitalia of 
<taxonomicName id="4C10F4DFFFED121E1CC1DD32FD19F9A8" authorityName="Kim &amp; Phan &amp; Pham &amp; Jung" authorityYear="2024" box="[494,736,1613,1636]" class="Insecta" family="Miridae" genus="Liocapsus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B964534EFFED121E1CC1DD32FD19F9A8" box="[494,736,1613,1636]" italics="true" pageId="3" pageNumber="298">Liocapsus vietnamensis</emphasis>
</taxonomicName>
<emphasis id="B964534EFFED121E1FC7DD33FCC2F9A8" bold="true" box="[744,827,1612,1636]" pageId="3" pageNumber="298">
<taxonomicNameLabel id="A257EE35FFED121E1FC7DD33FCC2F9A8" box="[744,827,1612,1636]" pageId="3" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) genital chamber and adjacent structures; (B) posterior wall. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BAF8F5CFFED12191DE8DDA3FAECFEAF" blockId="3.[151,1437,1756,2032]" lastBlockId="4.[151,1437,151,356]" lastPageId="4" lastPageNumber="299" pageId="3" pageNumber="298">
<emphasis id="B964534EFFED121E1DE8DDA3FE5DF938" box="[199,420,1756,1780]" italics="true" pageId="3" pageNumber="298">Specimen examined.</emphasis>
<emphasis id="B964534EFFED121E1C84DDA3FD6DF938" bold="true" box="[427,660,1756,1780]" pageId="3" pageNumber="298">
[
<materialsCitation id="3B788501FFED121E1C9DDDA3FD76F938" box="[434,655,1756,1780]" collectionCode="ZCDTU" pageId="3" pageNumber="298" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED011799FFED121E1C9DDDA3FDEDF938" box="[434,532,1756,1780]" pageId="3" pageNumber="298">ZCDTU</collectionCode>
] 
<typeStatus id="54AB31FEFFED121E1F0CDDA3FD76F938" box="[547,655,1756,1780]" pageId="3" pageNumber="298" type="holotype">Holotype</typeStatus>
</materialsCitation>
:
</emphasis>
<materialsCitation id="3B788501FFED121E1FB3DDA3FCC1F8D4" collectingDate="2017-05-10" collectionCode="ZCDTU" collectorName="J. Kim" country="Vietnam" latitude="20.318611" location="Cuc Phuong National Park" longLatPrecision="21" longitude="105.604996" pageId="3" pageNumber="298" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh" typeStatus="holotype">
<specimenCount id="9D1644D5FFED121E1FB3DDA3FD46F938" box="[668,703,1756,1780]" count="1" pageId="3" pageNumber="298" type="male">1♂</specimenCount>
, 
<location id="8ECFD987FFED121E1FE3DDA3FC16F938" LSID="urn:lsid:plazi:treatment:03B93E4AFFEF12191DB8DFE5FAECFEAF:8ECFD987FFED121E1FE3DDA3FC16F938" box="[716,1007,1756,1780]" country="Vietnam" latitude="20.318611" longLatPrecision="21" longitude="105.604996" name="Cuc Phuong National Park" pageId="3" pageNumber="298" stateProvince="Ninh Binh">Cuc Phuong National Park</location>
(
<geoCoordinate id="EE24E99BFFED121E1ED0DDA3FB79F938" box="[1023,1152,1756,1780]" degrees="20" direction="north" minutes="19" orientation="latitude" pageId="3" pageNumber="298" precision="15" seconds="07" value="20.318611">20°19´07˝N</geoCoordinate>
<geoCoordinate id="EE24E99BFFED121E19A7DDA3FAECF938" box="[1160,1301,1756,1780]" degrees="105" direction="east" minutes="36" orientation="longitude" pageId="3" pageNumber="298" precision="15" seconds="18" value="105.604996">105°36´18˝E</geoCoordinate>
), 
<collectingRegion id="49D441BEFFED121E1804DDA3FF2BF8DB" country="Vietnam" name="Ninh Binh" pageId="3" pageNumber="298">Ninh Binh Prov.</collectingRegion>
, 
<collectingCountry id="F307CFCCFFED121E1DF4DC7FFEC1F8D4" box="[219,312,1792,1816]" name="Vietnam" pageId="3" pageNumber="298">Vietnam</collectingCountry>
, 
<date id="FFAEA99CFFED121E1C6DDC7FFE54F8D4" box="[322,429,1792,1816]" pageId="3" pageNumber="298" value="2017-05-10">
<collectingDate id="EFEA5074FFED121E1C6DDC7FFE54F8D4" box="[322,429,1792,1816]" pageId="3" pageNumber="298" value="2017-05-10">10.v.2017</collectingDate>
</date>
, 
<collectingMethod id="5251F74BFFED121E1C97DC7FFDC5F8DB" box="[440,572,1792,1816]" pageId="3" pageNumber="298">by light trap</collectingMethod>
, 
<collectorName id="26E5EA8AFFED121E1F68DC7FFD74F8D4" box="[583,653,1792,1816]" pageId="3" pageNumber="298">J. Kim</collectorName>
leg.; 
<emphasis id="B964534EFFED121E1FE6DC7FFCC1F8D4" bold="true" box="[713,824,1792,1816]" pageId="3" pageNumber="298">
[
<collectionCode id="ED011799FFED121E1FFFDC7FFCCBF8D4" box="[720,818,1792,1816]" pageId="3" pageNumber="298">ZCDTU</collectionCode>
]
</emphasis>
</materialsCitation>
<materialsCitation id="3B788501FFED121E1E10DC7FFADEF8D4" box="[831,1319,1792,1816]" collectingDate="2017-05-10" collectionCode="CNU" collectorName="J. Kim" country="Vietnam" latitude="20.318611" location="Cuc Phuong National Park" longLatPrecision="21" longitude="105.604996" pageId="3" pageNumber="298" specimenCount="1" specimenCount-female="1" stateProvince="Ninh Binh" typeStatus="paratype">
<emphasis id="B964534EFFED121E1E10DC7FFC56F8D4" bold="true" box="[831,943,1792,1816]" pageId="3" pageNumber="298">
<typeStatus id="54AB31FEFFED121E1E10DC7FFC52F8D4" box="[831,939,1792,1816]" pageId="3" pageNumber="298" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D1644D5FFED121E1E9ADC7FFC2EF8D4" box="[949,983,1792,1816]" count="1" pageId="3" pageNumber="298" type="female">1♀</specimenCount>
, same data as holotype; 
<emphasis id="B964534EFFED121E19F4DC7FFADEF8D4" bold="true" box="[1243,1319,1792,1816]" pageId="3" pageNumber="298">
[
<collectionCode id="ED011799FFED121E19CDDC7FFAD9F8D4" box="[1250,1312,1792,1816]" country="China" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Capital Normal University, College of Life Sciences" pageId="3" pageNumber="298">CNU</collectionCode>
]
</emphasis>
</materialsCitation>
<materialsCitation id="3B788501FFED12191802DC7FFAE8FEAF" collectingDate="2017-05-10" collectionCode="ZCDTU" collectorName="J. Kim" country="Vietnam" county="This" lastPageId="4" lastPageNumber="299" latitude="20.318611" location="Host. Unknown. Distribution." longLatPrecision="21" longitude="105.604996" municipality="Remarks. This" pageId="3" pageNumber="298" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ninh Binh" typeStatus="paratype">
<typeStatus id="54AB31FEFFED121E1802DC7FFA60F8D4" box="[1325,1433,1792,1816]" pageId="3" pageNumber="298" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D1644D5FFED121E1DB8DC5BFF44F8F0" box="[151,189,1828,1852]" count="1" pageId="3" pageNumber="298" type="male">1♂</specimenCount>
<specimenCount id="9D1644D5FFED121E1D92DC5BFF22F8F0" box="[189,219,1828,1852]" count="1" pageId="3" pageNumber="298" type="female">1♀</specimenCount>
, same data as holotype. 
<location id="8ECFD987FFED121E1DE8DC36FEA8F848" LSID="urn:lsid:plazi:treatment:03B93E4AFFEF12191DB8DFE5FAECFEAF:8ECFD987FFED121E1DE8DC36FEA8F848" country="Vietnam" county="This" municipality="Remarks. This" name="Host. Unknown. Distribution." pageId="3" pageNumber="298">
<emphasis id="B964534EFFED121E1DE8DC36FF00F8AC" box="[199,249,1865,1888]" italics="true" pageId="3" pageNumber="298">Host</emphasis>
. Unknown. 
<emphasis id="B964534EFFED121E1DE8DC13FEA8F848" box="[199,337,1900,1924]" italics="true" pageId="3" pageNumber="298">Distribution.</emphasis>
</location>
<collectingCountry id="F307CFCCFFED121E1C79DC13FE49F848" box="[342,432,1900,1924]" name="Vietnam" pageId="3" pageNumber="298">Vietnam</collectingCountry>
(
<location id="8ECFD987FFED121E1C90DC13FDEDF848" LSID="urn:lsid:plazi:treatment:03B93E4AFFEF12191DB8DFE5FAECFEAF:8ECFD987FFED121E1C90DC13FDEDF848" box="[447,532,1900,1924]" country="Vietnam" county="This" municipality="Remarks. This" name="Central" pageId="3" pageNumber="298">Central</location>
). 
<location id="8ECFD987FFED121E1DE8DCEFFE6BF864" LSID="urn:lsid:plazi:treatment:03B93E4AFFEF12191DB8DFE5FAECFEAF:8ECFD987FFED121E1DE8DCEFFE6BF864" box="[199,402,1936,1960]" country="Vietnam" county="This" municipality="Remarks. This" name="Etymology. Named" pageId="3" pageNumber="298">
<emphasis id="B964534EFFED121E1DE8DCEFFEC6F864" box="[199,319,1936,1960]" italics="true" pageId="3" pageNumber="298">Etymology.</emphasis>
Named
</location>
after type locality, 
<collectingCountry id="F307CFCCFFED121E1F4FDCEFFD47F864" box="[608,702,1936,1960]" name="Vietnam" pageId="3" pageNumber="298">Vietnam</collectingCountry>
. 
<collectingMunicipality id="6BCB1526FFED121E1DE8DCCBFEA6F800" box="[199,351,1972,1996]" pageId="3" pageNumber="298">
<emphasis id="B964534EFFED121E1DE8DCCBFEDDF800" box="[199,292,1972,1996]" italics="true" pageId="3" pageNumber="298">Remarks</emphasis>
. This
</collectingMunicipality>
species is easily distinguished from all known congeners by: pale head with reddish transverse stripes; mostly pale hemelytra with dark area in posterior part of corium; and subhyaline cuneus with narrow reddish marking at posterior part; sensory lobe of left paramere blunt; outer margin of sensory lobe of right paramere rounded; spicule thin and long, as long as membranous lobes. 
<collectingCounty id="62CEF7D0FFEA12191F10DBC4FD97FF1F" box="[575,622,187,211]" pageId="4" pageNumber="299">This</collectingCounty>
new species is similar in general appearance to the congener from 
<collectingCountry id="F307CFCCFFEA12191803DBC4FA84FF1F" box="[1324,1405,187,211]" name="Taiwan" pageId="4" pageNumber="299">Taiwan</collectingCountry>
, 
<taxonomicName id="4C10F4DFFFEA121918A8DBC3FDF9FF3B" authority="Yasunaga &amp; Schwartz" authorityName="Yasunaga &amp; Schwartz" authorityYear="2007" class="Insecta" family="Miridae" genus="Liocapsus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="ochromelas">
<emphasis id="B964534EFFEA121918A8DBC3FEEBFF3B" italics="true" pageId="4" pageNumber="299">L. ochromelas</emphasis>
Yasunaga &amp; Schwartz
</taxonomicName>
, but it can be distinguished by generally pale pronotum (vs. distinctly bicolor pronotum); generally pale hemelytra (vs. strongly contrasted hemelytra); and subhyaline cuneus with reddish marking apically (vs. ocherous cuneus with blackish dark marking apically); blunt sensory lobe of left paramere (vs. projected sensory lobe of left paramere); and spicule as long as membranous lobes (vs. much longer spicule than membranous lobes)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>