134 lines
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134 lines
20 KiB
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<document id="BE866E7F597AD96743AABFF377D09EBF" ID-CLB-Dataset="35386" ID-DOI="http://dx.doi.org/10.3897/zookeys.132.1508" ID-GBIF-Dataset="4e1ed198-07d4-4297-acfb-574a27c10709" ID-PMC="PMC3208432" ID-Pensoft-Pub="1313-2970-132-33" ID-PubMed="22140332" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-132-33" ModsDocOrigin="ZooKeys 132" ModsDocTitle="Taxonomic remarks about Semiclivina (Kult, 1947) new status, with description of Uroclivina subgen. n., and of two new species from South America (Coleoptera, Carabidae, Scaritinae, Clivinini)" checkinTime="1451249827898" checkinUser="pensoft" docAuthor="Dostal, Alexander" docDate="2011" docId="DBBA8CEB4C8CC0771367CAF57D066A20" docLanguage="en" docName="ZooKeys 132: 33-50" docOrigin="ZooKeys 132" docSource="http://dx.doi.org/10.3897/zookeys.132.1508" docTitle="Semiclivina Kult 1947, stat. n." docType="treatment" docVersion="6" lastPageNumber="35" masterDocId="FFE3FFACFFB0FF82FFAC0B32C709FFA3" masterDocTitle="Taxonomic remarks about Semiclivina (Kult, 1947) new status, with description of Uroclivina subgen. n., and of two new species from South America (Coleoptera, Carabidae, Scaritinae, Clivinini)" masterLastPageNumber="50" masterPageNumber="33" pageNumber="34" updateTime="1732753535401" updateUser="ExternalLinkService">
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<mods:title id="4054CC7A3896D48B0DB564C0D33F50E1">Taxonomic remarks about Semiclivina (Kult, 1947) new status, with description of Uroclivina subgen. n., and of two new species from South America (Coleoptera, Carabidae, Scaritinae, Clivinini)</mods:title>
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<mods:namePart id="8077BC950A5769DAA3A277AF22612A12">Dostal, Alexander</mods:namePart>
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<mods:typeOfResource id="3B6DF600CB26694E7858E34240494141">text</mods:typeOfResource>
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<mods:title id="FFACD0740358B92DA4AF0C07FA8EE5BC">ZooKeys</mods:title>
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<mods:date id="16430578C3DB2A1D293CF6931D348C39">2011</mods:date>
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<mods:detail id="391A523BA2E9E326B9AA0C7778BA4D01" type="volume">
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<mods:number id="DA0356D7E78E9F7D98B322BAD00E5980">132</mods:number>
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<mods:start id="E10A284351CE6D6902672C1E57A1B98D">33</mods:start>
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<mods:url id="440D774B408B0065705901A0607E2EF0">http://dx.doi.org/10.3897/zookeys.132.1508</mods:url>
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<mods:classification id="9AADE009E9A01DE831C3CC33BA8F23DD">journal article</mods:classification>
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<mods:identifier id="2F3A42FF0B960804007153096D844980" type="DOI">http://dx.doi.org/10.3897/zookeys.132.1508</mods:identifier>
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<mods:identifier id="EB0DA206B643DA1AFB192ABDEC7041C2" type="Pensoft-Pub">1313-2970-132-33</mods:identifier>
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<treatment id="DBBA8CEB4C8CC0771367CAF57D066A20" ID-GBIF-Taxon="152031188" LSID="urn:lsid:zoobank.org:act:581B1ABC-14ED-4D8B-B216-7119CF90DCDC" httpUri="http://treatment.plazi.org/id/DBBA8CEB4C8CC0771367CAF57D066A20" lastPageId="3" lastPageNumber="35" pageId="1" pageNumber="34" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Scaritinae" scope_tribe="Clivinini">
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<subSubSection id="A0578D848546807E513AB2030F267A20" pageId="1" pageNumber="34" type="nomenclature">
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<paragraph id="EA773C5FC2C5DDC334B4A1B862A6A4AD" pageId="1" pageNumber="34">
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Genus
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<taxonomicName id="88A3623747FDF201F76DAA0E28E5FABC" ID-CoL="8KYVK" LSID="http://species-id.net/wiki/Semiclivina" authority="Kult, 1947" authorityName="Kult" authorityYear="1947" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Semiclivina Kult, 1947</taxonomicName>
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<taxonomicNameLabel id="73F50EB820C903D5C1C8E66A09B70A62" pageId="1" pageNumber="34">stat. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="15604932EF34BD9365EE1B3E417DD202" pageId="1" pageNumber="34" type="reference_group">
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<paragraph id="5ABDD5F0FEF8F68992AFF27F471A4DCA" pageId="1" pageNumber="34">
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<taxonomicName id="4E8F91DEA4B054D3C303C9E278F8E46E" class="Insecta" family="Carabidae" genus="Clivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Clivina</taxonomicName>
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Subgenus
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<taxonomicName id="4A6344D5D86253FFC9CF8E04B90B0606" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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<bibRefCitation id="E0347AEF8A27CFA4E1289E222AF9BDB5" author="Kult, K" journalOrPublisher="Spolecnosti Entomologicke)" pageId="11" pageNumber="44" pagination="26 - 37" title="Treti studie o strevlicich tribu Clivini (Col.). The 3 rd study to the knowledge of tribus Clivinini (Col., Carab.). (18 th Contribution to the knowledge of Carabidae.) Acta Societatis Entomologicae Cechoslovakiae (Casopis Csl." volume="44" year="1947">Kult 1947</bibRefCitation>
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: 31-32; Reichardt 1977: 391;
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<bibRefCitation id="30CB95EBBF13B35E510D56690CF63F2B" pageId="1" pageNumber="34">Nichols 1988a</bibRefCitation>
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: 154;
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<bibRefCitation id="D41243A9EB4CCD0F728F0DD9C0550CEE" pageId="1" pageNumber="34">1988b</bibRefCitation>
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: 91;
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<bibRefCitation id="B856C6B63E4EFE7A147194C156CB9487" author="Ball, GE" journalOrPublisher="Osaka" pageId="10" pageNumber="43" pagination="129 - 156" title="The subgenera of Clivina Latreille in the Western Hemisphere, and a Revision of Subgenus Antroforceps Barr (new status), with Notes about Evolutionary Aspects (Coleoptera: Carabidae: Clivinini). Spec. Publ. Japan Coleopt. Soc." volume="1" year="2001">Ball 2001</bibRefCitation>
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: 136;
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<bibRefCitation id="9D82BDEED66F4B33863F80C639685D74" author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae), Second edition, typeset and printed by the author, Tutzing" pageId="11" pageNumber="44" title="Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
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: 145;
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<bibRefCitation id="114D583410927FE5CECFD383BE4327F9" author="Baehr, M" journalOrPublisher="Coleoptera" pageId="10" pageNumber="43" pagination="1 - 220" title="The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta-, orbitalis-, planiceps-, sulcaticeps-, tranquebarica-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae)." volume="12" year="2008">Baehr 2008</bibRefCitation>
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: 23;
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<bibRefCitation id="115A213769076FE1D341C8C3C1DA98C1" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 41.
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</paragraph>
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</subSubSection>
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<subSubSection id="B84ED456A2AB0612736DD93F992A4E4E" lastPageId="3" lastPageNumber="36" pageId="1" pageNumber="34" type="type species">
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<paragraph id="9AD8E28E2935A633E0C63F03F16F7D47" pageId="1" pageNumber="34">Type Species.</paragraph>
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<paragraph id="B9B2987DE3B99B69B4B3885872AC7094" pageId="1" pageNumber="34">
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<taxonomicName id="2D8C1185EC5991268B56064B39CE8A4B" class="Insecta" family="Carabidae" genus="Clivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clivina dentipes" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="dentipes">Clivina dentipes</taxonomicName>
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Dejaen, 1825, by original designation:
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<bibRefCitation id="F1F864F0E237536427D123C27F9DA27C" author="Kult, K" journalOrPublisher="Spolecnosti Entomologicke)" pageId="11" pageNumber="44" pagination="26 - 37" title="Treti studie o strevlicich tribu Clivini (Col.). The 3 rd study to the knowledge of tribus Clivinini (Col., Carab.). (18 th Contribution to the knowledge of Carabidae.) Acta Societatis Entomologicae Cechoslovakiae (Casopis Csl." volume="44" year="1947">Kult 1947</bibRefCitation>
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: 31.
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</paragraph>
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<paragraph id="7440BA37CA90533666F5F229014BE0C0" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
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The genus
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<taxonomicName id="CB333158B60DACA8856B78B6FFC46DAF" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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(Kult, 1947) is readily recognized by the sculptured band of the proepisternum, extended more or less parallel to the proepisternal margin in the basal part of the proepisternum, curved inward in the anterior part, ended at the anterior end of the proepisternal-prosternal suture (Fig. 1). This structure was differently interpreted by diverse authors (
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<bibRefCitation id="BCE6CC3C9F49AAF77E509333EAB1991D" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 38) as furrow, ridge or carina or elongate striole (
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<bibRefCitation id="F76CC639378F081E87426D710C25522C" author="Baehr, M" journalOrPublisher="Coleoptera" pageId="10" pageNumber="43" pagination="1 - 220" title="The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta-, orbitalis-, planiceps-, sulcaticeps-, tranquebarica-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae)." volume="12" year="2008">Baehr 2008</bibRefCitation>
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: 23). In fact it is a slightly elevated band like structure consisting of
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<pageBreakToken id="217D06A396C764B1EEE5E83DCA0D0E3A" pageId="2" pageNumber="35" start="start">very</pageBreakToken>
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fine, regular transverse ridges. This sculptured band (
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<bibRefCitation id="A8159B700393B510E2DD74EF52487B9C" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 38) is part of a stridulation organ in the sense of a pars stridens, whereas the corresponding plectrum is a fine, sharp longitudinal ridge, just above the distal part of the lower inner edge of the profemur (Fig. 2). In some species the outer edge of the profemur is produced to a remarkable denticle, but this structure does not interact with the stridulation organ in any way. The function of this stridulation organ has not been observed so far.
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</paragraph>
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<caption id="6E0168F248AEA0F4E2313493C6789D2C" pageId="2" pageNumber="35">
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<paragraph id="C26C7BA64C6F1240E135679D3C882BB2" pageId="2" pageNumber="35">
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Figure 1. Photograph of prothorax and base of metathorax of
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<taxonomicName id="434FA917DF8E703963D57B3628E15A1A" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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species; right lateral view, showing the stridulation ridge (arrow); Scale bar: 0,5 mm.
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</paragraph>
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</caption>
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<caption id="90E86CD1F80E4402135F411C07595982" pageId="2" pageNumber="35">
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<paragraph id="F7142E6BCD6C437994C21F1557D33A1A" pageId="2" pageNumber="35">
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Figure 2. Photograph of right leg, posterior view, of base of trochanter, femur, and base of tibia of
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<taxonomicName id="71A55DA88AF2FF5AA43542F7439305DB" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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species, showing stridulation ridge on inner side of profemur (arrow); Scale bar: 0,5 mm.
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</paragraph>
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</caption>
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<paragraph id="B0B111A58D4BB1C6BB6428448046320D" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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This organ is not unique within the Tribe
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<taxonomicName id="01C918226042EA10C6FFFC5B45BFC4A3" lsidName="" pageId="2" pageNumber="35" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
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: exactly the same structures (the stridulation ridge on proepisterna and the ridge on the distal part of the inner edge of the profemur) together with other features separates the ardistomine genus
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<taxonomicName id="432DA1D9F1B912E6B0CBAE3B6B66CA72" class="Insecta" family="Carabidae" genus="Ardistomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ardistomis" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ardistomis</taxonomicName>
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Putzeys, 1846 from
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<taxonomicName id="45F2F8EDAED30FCEC31FB6D8025CFB2F" class="Insecta" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiardistomis" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiardistomis</taxonomicName>
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Kult, 1950, but it is unique within
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<taxonomicName id="4418EE6D80536EEF2AD6E70B0158A84A" class="Insecta" family="Carabidae" genus="Clivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Clivina</taxonomicName>
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(sensu lato), forming a very isolated, distinct group, restricted to the western hemisphere and to a single species from Australia (New South Wales) described by
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<bibRefCitation id="4F9EEBE9DB30EC3A1CBAC811D72C539C" author="Baehr, M" journalOrPublisher="Coleoptera" pageId="10" pageNumber="43" pagination="1 - 220" title="The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta-, orbitalis-, planiceps-, sulcaticeps-, tranquebarica-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae)." volume="12" year="2008">Baehr 2008</bibRefCitation>
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: 23-25. Within the tribe
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<taxonomicName id="DF7AD6195A5842DD635B5B9CF90C8AE7" lsidName="" pageId="2" pageNumber="35" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
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no other genera than
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<taxonomicName id="DF962DA196AA8A0F5A63487EC17EE52C" class="Insecta" family="Carabidae" genus="Ardistomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ardistomis" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ardistomis</taxonomicName>
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Putzeys, 1846 and
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<taxonomicName id="E8B71ADC32299A2F9C53C0E435043ED0" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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(Kult, 1947) carry a stridulation organ on the proepisternum. Beside this,
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<taxonomicName id="855851C74B5A3736E7C2AF248178919F" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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is characterized by a distal spine-like projection at the outer end of the mesotibia, which is small in some taxa, but clearly identifiable by the apical seta
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<pageBreakToken id="A880FAF8CDDCFD7F0103F6234D1B0595" pageId="3" pageNumber="36" start="start">excentrically</pageBreakToken>
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inserted, in contradiction to a tubercle, which carries the seta at the top of the tubercle (see also
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<bibRefCitation id="BB810EAD55324DE19544D24C2A5E178C" author="Ball, GE" journalOrPublisher="Osaka" pageId="10" pageNumber="43" pagination="129 - 156" title="The subgenera of Clivina Latreille in the Western Hemisphere, and a Revision of Subgenus Antroforceps Barr (new status), with Notes about Evolutionary Aspects (Coleoptera: Carabidae: Clivinini). Spec. Publ. Japan Coleopt. Soc." volume="1" year="2001">Ball 2001</bibRefCitation>
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: 136, 140;
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<bibRefCitation id="E7B8CEEC96C0B61F6425A81DC553F2AB" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 41, 43); usually five setae in elytral interval 3 (including preapical puncture); elytral striae 1 to 5 free at base (between humeral channel and suture, the basal keel of some species and the flat tubercles at the end of intervals 2 to 5 do not limit this definition); profemur with a sometimes small dentiform projection at the outer edge toward apex; anal ventrite (abdominal sternum VII) with two marginal punctures at each side relatively close together. The species included in
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<taxonomicName id="81364DBD880192A834BDCD439FE159BB" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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are listed in Bousquet (2009: 39-41), where the assignment of some species remains uncertain. It is most likely, that species not included in the subgen. n.
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<taxonomicName id="B7622C6EB5E9E6B05B161952719DBD43" genus="Uroclivina" lsidName="Uroclivina" pageId="3" pageNumber="36" rank="genus">Uroclivina</taxonomicName>
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comprise a more or less heterogeneous group which form a probably paraphyletic group, defined as
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<taxonomicName id="F08E00F68282A79BD31A4B0B80655B80" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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(s. str.). This subgenus will be the objective of further investigations in the future.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |