253 lines
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253 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. & Marek, Paul E." docDate="2021" docId="B8D91701E813590AB3A29E03FEA6CB39" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria rhysodesmoides" docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1030</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181004718" LSID="urn:lsid:plazi:treatment:B8D91701E813590AB3A29E03FEA6CB39" httpUri="http://treatment.plazi.org/id/B8D91701E813590AB3A29E03FEA6CB39" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="8RVVX" LSID="B8D91701-E813-590A-B3A2-9E03FEA6CB39" authority="(Hennen & Shelley, 2015)" baseAuthorityName="Hennen & Shelley" baseAuthorityYear="2015" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria rhysodesmoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rhysodesmoides">Nannaria rhysodesmoides (Hennen & Shelley, 2015)</taxonomicName>
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<figureCitation captionStart="Figure 108" captionStartId="F108" captionText="Figure 108. Nannaria rhysodesmoides holotype ♂ (FSCA) left gonopod A anterior view, red arrow indicates small, pointed lateral flange B medial view; red triangle indicates slight sinuation of the acropodite apex C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure108" httpUri="https://binary.pensoft.net/fig/531211" pageId="0" pageNumber="1">Fig. 108 Vernacular name: "The Rhysodesmus-Like Twisted-Claw Millipede"</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Diplopoda" family="Xystodesmidae" genus="Mimuloria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mimuloria rhysodesmoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rhysodesmoides">Mimuloria rhysodesmoides</taxonomicName>
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Hennen & Shelley, 2015: 1-16.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Hennen & Shelley" baseAuthorityYear="2015" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria rhysodesmoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rhysodesmoides">Nannaria rhysodesmoides</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
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: S72.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355687" collectingDate="1958-04-12" collectorName="H. E. Evans" country="United States" county="Putnam County" latitude="36.1628" location="Cookeville" longLatPrecision="6" longitude="-85.5016" specimenCount="♂" stateProvince="Virginia" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingCounty>Putnam County</collectingCounty>
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,
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<location LSID="urn:lsid:plazi:treatment:B8D91701E813590AB3A29E03FEA6CB39:1FD67BD22C020F63A6D4C89221DE0BC4" country="United States" county="Putnam County" latitude="36.1628" longLatPrecision="6" longitude="-85.5016" name="Cookeville" stateProvince="Virginia">Cookeville</location>
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; [
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<geoCoordinate degrees="36.1628" direction="north" orientation="latitude" precision="5" value="36.1628">36.1628°N</geoCoordinate>
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, -
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<geoCoordinate degrees="85.5016" direction="west" orientation="longitude" precision="5" value="-85.5016">85.5016°W</geoCoordinate>
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];
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<collectingDate value="1958-04-12">12 Apr. 1958</collectingDate>
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;
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<collectorName>H.E. Evans</collectorName>
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leg.; FSCA.
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355636" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="H. E. Evans" country="United States" county="Putnam County" latitude="36.1628" location="Virginia" longLatPrecision="6" longitude="-85.5016" specimenCount="6" specimenCount-male="6" stateProvince="Virginia" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">6 ♂♂</specimenCount>
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; same collection data as holotype; NCSM 27913. For detailed collection data see Suppl. material 7
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult males of
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<taxonomicName baseAuthorityName="Hennen & Shelley" baseAuthorityYear="2015" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria rhysodesmoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rhysodesmoides">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria rhysodesmoides</emphasis>
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</taxonomicName>
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are distinct from other
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
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</taxonomicName>
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and the nearby
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov. based on the following combination of characters:
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
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.
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</emphasis>
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Gonopodal acropodite simple, gently curving dorsomedially, not with pronounced medial swelling as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov. Acropodite tip with small, triangular lateral flange (Fig.
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<figureCitation captionStart="Figure 108" captionStartId="F108" captionText="Figure 108. Nannaria rhysodesmoides holotype ♂ (FSCA) left gonopod A anterior view, red arrow indicates small, pointed lateral flange B medial view; red triangle indicates slight sinuation of the acropodite apex C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure108" httpUri="https://binary.pensoft.net/fig/531211" pageId="0" pageNumber="1">108A</figureCitation>
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, red arrow), not large, hooked lateral flange as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov. Apex slightly sinuate (Fig.
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<figureCitation captionStart="Figure 108" captionStartId="F108" captionText="Figure 108. Nannaria rhysodesmoides holotype ♂ (FSCA) left gonopod A anterior view, red arrow indicates small, pointed lateral flange B medial view; red triangle indicates slight sinuation of the acropodite apex C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure108" httpUri="https://binary.pensoft.net/fig/531211" pageId="0" pageNumber="1">108B</figureCitation>
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, red triangle). Acropodite tip terminating in small, sharp, dorsally directed point, not blunt, squared off as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov. Height of telopodite basal zone <1/2 length of prefemoral process, not ca. 1/2 length of prefemoral process as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov. Prefemoral process straight, wide, approaching laminate, not thin and acuminate as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov. Prefemoral spine completely reduced not forming small ridge as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
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.
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</emphasis>
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The preserved holotype suggests that
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. rhysodesmoides" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="rhysodesmoides">
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<emphasis italics="true" pageId="0" pageNumber="1">N. rhysodesmoides</emphasis>
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</taxonomicName>
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may have metatergal stripes connecting the paranotal spots, though living specimens have not be observed by the authors.
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</paragraph>
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<caption doi="10.3897/zookeys.1030.62544.figure108" httpUri="https://binary.pensoft.net/fig/531211" pageId="0" pageNumber="1" start="Figure 108" startId="F108">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 108.</emphasis>
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<taxonomicName baseAuthorityName="Hennen & Shelley" baseAuthorityYear="2015" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria rhysodesmoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rhysodesmoides">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria rhysodesmoides</emphasis>
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</taxonomicName>
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holotype ♂ (FSCA) left gonopod
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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anterior view, red arrow indicates small, pointed lateral flange
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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medial view; red triangle indicates slight sinuation of the acropodite apex
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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posterior view. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="1">♂ paratype (NCSM27913; ♂ holotype too fragmented for measurement): BL = 22.3, CW = 2.9, IW = 1.8, ISW = 0.8, B11W = 4.4, B11H = 2.4.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Known only from the type locality (Tennessee: Putnam County, Suppl. material 7; Fig.
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<figureCitation captionStart="Figure 129" captionStartId="F129" captionText="Figure 129. Distribution map of Nannaria minor species group collection localities west of the Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure129" httpUri="https://binary.pensoft.net/fig/531232" pageId="0" pageNumber="1">129</figureCitation>
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). Distribution area: N/A; status: MRE.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Hennen, DA" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 21" publicationUrl="http://digitalcommons.unl.edu/insectamundi/924" refId="B28" refString="Hennen, DA, Shelley, RM, 2015. A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964. Insecta Mundi 0418: 1 - 21, http://digitalcommons.unl.edu/insectamundi/924" title="A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964." url="http://digitalcommons.unl.edu/insectamundi/924" volume="0418" year="2015">Hennen and Shelley (2015)</bibRefCitation>
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do not mention ecological notes in their description of the species, most likely due to a lack of ecological information from the original collector.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Hennen, DA" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 21" publicationUrl="http://digitalcommons.unl.edu/insectamundi/924" refId="B28" refString="Hennen, DA, Shelley, RM, 2015. A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964. Insecta Mundi 0418: 1 - 21, http://digitalcommons.unl.edu/insectamundi/924" title="A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964." url="http://digitalcommons.unl.edu/insectamundi/924" volume="0418" year="2015">Hennen and Shelley (2015)</bibRefCitation>
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chose the specific name based on the presence of a lateral flange on the gonopod (Fig.
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<figureCitation captionStart="Figure 108" captionStartId="F108" captionText="Figure 108. Nannaria rhysodesmoides holotype ♂ (FSCA) left gonopod A anterior view, red arrow indicates small, pointed lateral flange B medial view; red triangle indicates slight sinuation of the acropodite apex C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure108" httpUri="https://binary.pensoft.net/fig/531211" pageId="0" pageNumber="1">108A</figureCitation>
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, red arrow), a character also seen in the genus
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<taxonomicName class="Diplopoda" family="Xystodesmidae" genus="Rhysodesmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Rhysodesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhysodesmus</emphasis>
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</taxonomicName>
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Cook, 1895.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">United States, Virginia, Putnam County, Cookeville.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="notes">
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<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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In the original publication,
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<bibRefCitation author="Hennen, DA" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 21" publicationUrl="http://digitalcommons.unl.edu/insectamundi/924" refId="B28" refString="Hennen, DA, Shelley, RM, 2015. A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964. Insecta Mundi 0418: 1 - 21, http://digitalcommons.unl.edu/insectamundi/924" title="A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964." url="http://digitalcommons.unl.edu/insectamundi/924" volume="0418" year="2015">Hennen and Shelley (2015</bibRefCitation>
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: 15) designate a male holotype and five male paratypes (FSCA) collected by H. E. Evans on 12 April 1958, and note that one male paratype from this collection was retained at the NCSM.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |