424 lines
54 KiB
XML
424 lines
54 KiB
XML
<document id="B512159D2F6CCEFF809245A020613722" ID-DOI="https://doi.org/10.5852/ejt.2023.870.2123" ID-ISSN="2118-9773" ID-Zenodo-Dep="7986633" ID-ZooBank="508560B7-BD89-472A-B3FD-F48851B024C3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1685458305949" checkinUser="felipe" docAuthor="Nagy, Hajnalka, Dózsa-Farkas, Klára & Felföldi, Tamás" docDate="2023" docId="B857879AFFC1FFADFDA35810FD47F802" docLanguage="en" docName="ejt-870_107-145.pdf" docOrigin="European Journal of Taxonomy 870" docSource="http://dx.doi.org/10.5852/ejt.2023.870.2123" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Enchytraeus undefined-3" docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="446EFFE2FFDFFFB2FF825D44FFFEFFDC" masterDocTitle="New insights into the Enchytraeus albidus complex (Annelida, Enchytraeidae), with the description of three new species from seashores in Italy and Croatia" masterLastPageNumber="145" masterPageNumber="107" pageNumber="31" updateTime="1685474603013" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="267438578EA6EDABEB1A3BE6B959E84F">New insights into the Enchytraeus albidus complex (Annelida, Enchytraeidae), with the description of three new species from seashores in Italy and Croatia</mods:title>
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<mods:roleTerm id="C45CDFAF0C6B65381CFD6E6CCFCAEBD5">Author</mods:roleTerm>
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<mods:namePart id="98248E318AEBD0DEF7FFC0D2CC9A8722">Nagy, Hajnalka</mods:namePart>
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<mods:affiliation id="F8EC323562BE1A95FDAC93E0A507586B">Department of Microbiology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Hungarian Natural History Museum, Baross utca 13, H- 1088 Budapest, Hungary. Department of Systematic Zoology and Ecology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Institute of Aquatic Ecology, Centre for Ecological Research, Karolina út 29, H- 1113 Budapest, Hungary.</mods:affiliation>
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<mods:nameIdentifier id="A7E2E4D9D21A592CAD755CEC18CE631A" type="email">nhajni6@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="B97D6C4774E0DF1F0C2084A5265687CC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E2F454BDA6EF45CBFBD2A14C525833F9">Dózsa-Farkas, Klára</mods:namePart>
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<mods:nameIdentifier id="2E28EA2F103876949EF4A7E643A21538" type="email">kdozsafarkas@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="FE29689918E88F763C1BBD5B76A75E58" type="personal">
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</mods:role>
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<mods:namePart id="C762D77FF88F5E4C26E8B319ED461948">Felföldi, Tamás</mods:namePart>
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<mods:affiliation id="0F7435EAF5248A474D57CDF74A313B1B">Department of Microbiology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Hungarian Natural History Museum, Baross utca 13, H- 1088 Budapest, Hungary. Department of Systematic Zoology and Ecology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1 / C, H- 1117 Budapest, Hungary. Institute of Aquatic Ecology, Centre for Ecological Research, Karolina út 29, H- 1113 Budapest, Hungary.</mods:affiliation>
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<mods:nameIdentifier id="17C5CDB152E0611B7263F8F3AE366574" type="email">tamas.felfoldi@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="EF25F422992659F556AABCECF2A1D6B1">text</mods:typeOfResource>
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<mods:title id="E80320E93597C528D7165A1E25ACBAEB">European Journal of Taxonomy</mods:title>
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<mods:part id="DD106BAE2B17805E332005FFE82E3390">
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<mods:date id="FC8594DF67DC890E7A44FCCEFEEBF61E">2023</mods:date>
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<mods:number id="2E1DE6236EAF962708255E118E861D01">2023-05-26</mods:number>
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<mods:detail id="74F2E8F0E2C8DB464844794A1A3AD20B" type="volume">
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<mods:number id="E4FB31672BBD1666B6DBC18D3C8049D0">870</mods:number>
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<mods:start id="074135F822B0F77B1BD9F21197C646C4">107</mods:start>
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<mods:end id="1A226F4323D6BE05181C888CD56EEF14">145</mods:end>
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<mods:url id="29AA6117C6221E2C9CD6F9731D3F4296">http://dx.doi.org/10.5852/ejt.2023.870.2123</mods:url>
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</mods:location>
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<mods:classification id="227420B31954137F5A0906287CB75009">journal article</mods:classification>
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<mods:identifier id="37DB44E0F1FC15288EB929C0BC5CCA2B" type="DOI">https://doi.org/10.5852/ejt.2023.870.2123</mods:identifier>
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<mods:identifier id="2B392271E22BE2BE5243FA0F72994333" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="DB891AB4EC889905C7D2AF7A0FCFA447" type="Zenodo-Dep">7986633</mods:identifier>
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</mods:mods>
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<treatment id="B857879AFFC1FFADFDA35810FD47F802" LSID="urn:lsid:plazi:treatment:B857879AFFC1FFADFDA35810FD47F802" httpUri="http://treatment.plazi.org/id/B857879AFFC1FFADFDA35810FD47F802" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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<subSubSection id="78E46507FFC1FFACFDA35810FBEDFAB3" box="[545,1043,1364,1391]" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="3041368CFFC1FFACFDA35810FBEDFAB3" blockId="30.[189,1043,1364,1426]" box="[545,1043,1364,1391]" pageId="30" pageNumber="31">
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<heading id="6B0981E0FFC1FFACFDA35810FBEDFAB3" box="[545,1043,1364,1391]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
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<taxonomicName id="F7FE4D0FFFC1FFACFDA35810FD0DFAB3" box="[545,755,1364,1391]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="undefined-3">
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<emphasis id="028AEA9EFFC1FFACFDA35810FD42FAB2" bold="true" box="[545,700,1364,1390]" italics="true" pageId="30" pageNumber="31">Enchytraeus</emphasis>
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sp.3
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</taxonomicName>
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from Kerguelen Islands
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="78E46507FFC1FFACFF3F583CFA99FA1D" pageId="30" pageNumber="31" type="materials_examined">
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<paragraph id="3041368CFFC1FFACFF3F583CFE53FA4E" blockId="30.[189,1043,1364,1426]" box="[189,429,1400,1426]" pageId="30" pageNumber="31">
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<heading id="6B0981E0FFC1FFACFF3F583CFE53FA4E" bold="true" box="[189,429,1400,1426]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
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<emphasis id="028AEA9EFFC1FFACFF3F583CFE53FA4E" bold="true" box="[189,429,1400,1426]" pageId="30" pageNumber="31">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F58E3FA99FA1D" blockId="30.[189,1383,1446,1473]" box="[189,1383,1446,1473]" pageId="30" pageNumber="31">
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<materialsCitation id="80963CD1FFC1FFACFF3F58E3FA9AFA1D" box="[189,1380,1446,1473]" collectionCode="ELTE" location="Ile Guillou Island" pageId="30" pageNumber="31" specimenCode="slide No. 3035 to 3037" specimenCount="5" stateProvince="KERGUELEN ISLANDS">
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<collectingRegion id="F23AF86EFFC1FFACFF3F58E3FE0EFA1D" box="[189,496,1447,1473]" pageId="30" pageNumber="31">KERGUELEN ISLANDS</collectingRegion>
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•
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<specimenCount id="26F8FD05FFC1FFACFD8B58E2FD9CFA1D" box="[521,610,1446,1473]" pageId="30" pageNumber="31" type="generic">5 specs</specimenCount>
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;
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<location id="35216057FFC1FFACFDEC58E2FCBEFA1C" LSID="urn:lsid:plazi:treatment:B857879AFFC1FFADFDA35810FD47F802:35216057FFC1FFACFDEC58E2FCBEFA1C" box="[622,832,1446,1472]" name="Ile Guillou Island" pageId="30" pageNumber="31" stateProvince="KERGUELEN ISLANDS">Ile Guillou Island</location>
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;
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<accessionNumber id="2FADAB6FFFC1FFACFCCF58E3FC0DFA1C" box="[845,1011,1446,1473]" pageId="30" pageNumber="31">ID 260 to 261</accessionNumber>
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,
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<specimenCode id="60589EF7FFC1FFACFC7D58E2FAF0FA1D" box="[1023,1294,1446,1473]" pageId="30" pageNumber="31">slide No. 3035 to 3037</specimenCode>
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;
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<collectionCode id="56EFAE49FFC1FFACFA9958E3FA9AFA1D" box="[1307,1380,1447,1473]" pageId="30" pageNumber="31">ELTE</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="78E46507FFC1FFADFF3F58ACFD31FD8D" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
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<paragraph id="3041368CFFC1FFACFF3F58ACFEB1F9DE" blockId="30.[189,335,1512,1538]" box="[189,335,1512,1538]" pageId="30" pageNumber="31">
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<heading id="6B0981E0FFC1FFACFF3F58ACFEB1F9DE" bold="true" box="[189,335,1512,1538]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
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<emphasis id="028AEA9EFFC1FFACFF3F58ACFEB1F9DE" bold="true" box="[189,335,1512,1538]" pageId="30" pageNumber="31">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F5B52FD44F988" blockId="30.[189,1398,1558,1620]" pageId="30" pageNumber="31">
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Based on the few specimens in imperfect condition,
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<emphasis id="028AEA9EFFC1FFACFCBA5B52FC43F9EC" box="[824,957,1558,1584]" italics="true" pageId="30" pageNumber="31">Echytraeus</emphasis>
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sp.3 from Kerguelen Islands is very similar morphologically to
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<taxonomicName id="F7FE4D0FFFC1FFACFE7F5B7EFD8BF98F" authorityName="Henle" authorityYear="1837" box="[509,629,1593,1619]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="albidus">
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<emphasis id="028AEA9EFFC1FFACFE7F5B7EFD8BF98F" box="[509,629,1593,1619]" italics="true" pageId="30" pageNumber="31">E. albidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="19B957E5FFC1FFACFDFF5B7EFD44F988" box="[637,698,1594,1620]" pageId="30" pageNumber="31" sensu="stricto">s. str.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F5B38FA88F949" blockId="30.[189,1398,1659,1686]" box="[189,1398,1659,1686]" pageId="30" pageNumber="31">
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<smallCapsWord id="36A7A050FFC1FFACFF3F5B38FE90F948" baselines="1679,1679" box="[189,366,1660,1686]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="30" pageNumber="31">MEASUREMENTS</smallCapsWord>
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. Body length
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<quantity id="F7069B69FFC1FFACFD975B3FFD68F94A" box="[533,662,1659,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="30" pageNumber="31" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
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, segment number 44–54, 610–660 μm wide at clitellum, fixed.
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F5BF9FE5BF8C1" blockId="30.[189,1399,1724,1821]" pageId="30" pageNumber="31">
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<smallCapsWord id="36A7A050FFC1FFACFF3F5BF9FEDCF909" baselines="1744,1744" box="[189,290,1725,1751]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Chaetae" pageId="30" pageNumber="31">CHAETAE</smallCapsWord>
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. Chaetal formula: 3–2,3:3,4–2,3. 87–95 μm × 8 μm in ventral bundles, 77–80 μm × 7.5 μm in lateral bundles preclitellarly and 77–80 μm × 7 μm in ventral bundles and 62–70 μm × 5 μm in lateral bundles posteriorly.
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F5A01FDCDF883" blockId="30.[189,563,1860,1887]" box="[189,563,1860,1887]" pageId="30" pageNumber="31">
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<smallCapsWord id="36A7A050FFC1FFACFF3F5A01FEE8F881" baselines="1880,1880" box="[189,278,1861,1887]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dorsal" pageId="30" pageNumber="31">DORSAL</smallCapsWord>
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<smallCapsWord id="36A7A050FFC1FFACFE9C5A0EFE94F881" baselines="1880" box="[286,362,1866,1885]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="vessel" pageId="30" pageNumber="31">VESSEL</smallCapsWord>
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. from XIII–XIV.
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F5AC2FDCAF87C" blockId="30.[189,564,1926,1952]" box="[189,564,1926,1952]" pageId="30" pageNumber="31">
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<smallCapsWord id="36A7A050FFC1FFACFF3F5AC2FE95F842" baselines="1945,1945" box="[189,363,1926,1952]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coelomocytes" pageId="30" pageNumber="31">COELOMOCYTES</smallCapsWord>
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. 31–42 μm long.
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</paragraph>
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<paragraph id="3041368CFFC1FFACFF3F5A83FD10F83E" blockId="30.[189,750,1991,2018]" box="[189,750,1991,2018]" pageId="30" pageNumber="31">
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<smallCapsWord id="36A7A050FFC1FFACFF3F5A83FEFBF83C" baselines="2010,2011" box="[189,261,1991,2017]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sperm" pageId="30" pageNumber="31">SPERM</smallCapsWord>
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<smallCapsWord id="36A7A050FFC1FFACFE8F5A89FEBFF83C" baselines="2011" box="[269,321,1997,2016]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="sacs" pageId="30" pageNumber="31">SACS</smallCapsWord>
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||
. Large sperm sacs, located in X–XI.
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</paragraph>
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<paragraph id="3041368CFFC0FFADFF3F5C4DFC37FEFF" blockId="31.[189,969,265,292]" box="[189,969,265,292]" pageId="31" pageNumber="32">
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<smallCapsWord id="36A7A050FFC0FFADFF3F5C4DFEFBFEFE" baselines="284,285" box="[189,261,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sperm" pageId="31" pageNumber="32">SPERM</smallCapsWord>
|
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<smallCapsWord id="36A7A050FFC0FFADFE8F5C4BFEA1FEFE" baselines="285" box="[269,351,271,290]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="funnel" pageId="31" pageNumber="32">FUNNEL</smallCapsWord>
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. 375–550 μm long, about 4 × as long as wide, fixed.
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</paragraph>
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<paragraph id="3041368CFFC0FFADFF3F5C0FFEF9FE54" blockId="31.[189,1399,330,392]" pageId="31" pageNumber="32">
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||
<smallCapsWord id="36A7A050FFC0FFADFF3F5C0FFEA6FEBF" baselines="350,350" box="[189,344,331,357]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spermatozoa" pageId="31" pageNumber="32">SPERMATOZOA</smallCapsWord>
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. About 60 μm long, heads 18 μm. Sperm duct about same wide in all length (23–30 μm wide).
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</paragraph>
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<paragraph id="3041368CFFC0FFADFF3F5CF4FDE2FE30" blockId="31.[189,1399,431,493]" pageId="31" pageNumber="32">
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<smallCapsWord id="36A7A050FFC0FFADFF3F5CF4FF01FE14" baselines="451,451" box="[189,255,432,458]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="31" pageNumber="32">MALE</smallCapsWord>
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||
<smallCapsWord id="36A7A050FFC0FFADFE8A5CF1FE6EFE14" baselines="451" box="[264,400,437,456]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="copulatory" pageId="31" pageNumber="32">COPULATORY</smallCapsWord>
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<smallCapsWord id="36A7A050FFC0FFADFE1A5CF1FE13FE14" baselines="451" box="[408,493,437,456]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="31" pageNumber="32">ORGANS</smallCapsWord>
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. Male glands multiple: one larger rounded primary bulb (80–100 μm, fixed) and numerous smaller glands.
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</paragraph>
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<paragraph id="3041368CFFC0FFADFF3F5F50FD31FD8D" blockId="31.[189,1398,532,593]" pageId="31" pageNumber="32">
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<smallCapsWord id="36A7A050FFC0FFADFF3F5F50FE9AFDF0" baselines="551,551" box="[189,356,532,558]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spermathecae" pageId="31" pageNumber="32">SPERMATHECAE</smallCapsWord>
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. Ectal duct of spermatheca with glands around duct, not only at the orifice. Ampulla with one diverticulum (about 210 μm long, fixed).
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</paragraph>
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</subSubSection>
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<subSubSection id="78E46507FFC0FFADFF3F5F3CFD47F802" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="3041368CFFC0FFADFF3F5F3CFE0CFD4E" blockId="31.[189,498,632,658]" box="[189,498,632,658]" pageId="31" pageNumber="32">
|
||
<heading id="6B0981E0FFC0FFADFF3F5F3CFE0CFD4E" bold="true" box="[189,498,632,658]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5F3CFE0CFD4E" bold="true" box="[189,498,632,658]" pageId="31" pageNumber="32">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3041368CFFC0FFADFF3F5FE3FBC0FD1D" blockId="31.[189,1086,679,705]" box="[189,1086,679,705]" pageId="31" pageNumber="32">
|
||
Guillou Island, Kerguelen Islands, under a small clump of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFCEF5FECFBC9FD1D" box="[877,1079,679,705]" class="Liliopsida" family="Poaceae" genus="Poa" kingdom="Plantae" order="Poales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="kerguelensis">
|
||
<emphasis id="028AEA9EFFC0FFADFCEF5FECFBC9FD1D" box="[877,1079,679,705]" italics="true" pageId="31" pageNumber="32">Poa kerguelensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3041368CFFC0FFADFF3F5FACFDD5FCDE" blockId="31.[189,555,744,770]" box="[189,555,744,770]" pageId="31" pageNumber="32">
|
||
<heading id="6B0981E0FFC0FFADFF3F5FACFDD5FCDE" bold="true" box="[189,555,744,770]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5FACFDD5FCDE" bold="true" box="[189,555,744,770]" pageId="31" pageNumber="32">Results of molecular analysis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3041368CFFC0FFADFF3F5E53FD47F802" blockId="31.[188,1400,791,2014]" pageId="31" pageNumber="32">
|
||
Results of molecular analysis corroborated that
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC945E5CFC36FCED" authority="Nagy & Dózsa-Farkas & Felföldi, 2023" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[790,968,791,817]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
|
||
<emphasis id="028AEA9EFFC0FFADFC945E5CFC36FCED" box="[790,968,791,817]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFC545E53FBC4FCED" box="[982,1082,791,817]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFBCF5E5CFB31FCED" authority="Nagy & Dózsa-Farkas & Felföldi, 2023" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1101,1231,791,817]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi" status="sp. nov.">
|
||
<emphasis id="028AEA9EFFC0FFADFBCF5E5CFB31FCED" box="[1101,1231,791,817]" italics="true" pageId="31" pageNumber="32">E. andrasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFB5F5E53FAC0FCED" box="[1245,1342,791,817]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3F5E7FFE79FC88" authority="Nagy & Dózsa-Farkas & Felföldi, 2023" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[189,391,826,852]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasiformis" status="sp. nov.">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5E7FFE79FC88" box="[189,391,826,852]" italics="true" pageId="31" pageNumber="32">E. andrasiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFE105E7FFE11FC89" box="[402,495,827,853]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are new, distinct species, since the sequences obtained from the examined individuals clearly separated in the phylogenetic trees (
|
||
<figureCitation id="A8C52A09FFC0FFADFCE55E19FC0BFCAB" box="[871,1013,861,887]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="32.[189,232,1892,1918]" captionStartId-1="33.[190,233,1892,1918]" captionStartId-2="34.[190,233,1892,1918]" captionTargetBox-0="[189,1398,297,1823]" captionTargetBox-1="[279,1309,605,1855]" captionTargetBox-2="[203,1383,1356,1842]" captionTargetId-0="figure-17@32.[189,1398,297,1823]" captionTargetId-1="figure-128@33.[279,1309,605,1855]" captionTargetId-2="figure-498@34.[189,1398,1335,1854]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionText-0="Fig. 10. Maximum likelihood (ML) tree of the COI gene for specimens of the Enchytraeus albidus species complex, based on 568 nucleotide positions using the General Time Reversible substitution model. Bootstrap values greater than 50 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Scale bar: 0.05 substitutions per nucleotide." captionText-1="Fig. 11. Maximum likelihood (ML) tree of the H3 gene for specimens of the Enchytraeus albidus species complex, based on 225 nucleotide positions using the Kimura 2-parameter substitution model. Bootstrap values greater than 50 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Scale bar: 0.01 substitutions per nucleotide." captionText-2="Fig. 12. Enchytraeus Henle, 1837 species tree, based on COI, H3 and 16S rRNA gene (Bayes analysis, *BEAST). Posterior probabilities greater than 0.5 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Abbreviation: n = number of included specimens for each species. Scale bar = 0.07 substitutions per nucleotide." figureDoi-0="http://doi.org/10.5281/zenodo.7986660" figureDoi-1="http://doi.org/10.5281/zenodo.7986662" figureDoi-2="http://doi.org/10.5281/zenodo.7986666" httpUri-0="https://zenodo.org/record/7986660/files/figure.png" httpUri-1="https://zenodo.org/record/7986662/files/figure.png" httpUri-2="https://zenodo.org/record/7986666/files/figure.png" pageId="31" pageNumber="32">Figs 10–12</figureCitation>
|
||
). Our study also provided new reference sequences for
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFE625EC5FDA4FC46" authorityName="Henle" authorityYear="1837" box="[480,602,896,922]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFE625EC5FDA4FC46" box="[480,602,896,922]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFDE05EC5FD5DFC47" box="[610,675,897,923]" pageId="31" pageNumber="32" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFD315EC5FCAAFC46" authorityName="Nielsen & Christensen" authorityYear="1961" box="[691,852,896,922]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="irregularis">
|
||
<emphasis id="028AEA9EFFC0FFADFD315EC5FCAAFC46" box="[691,852,896,922]" italics="true" pageId="31" pageNumber="32">E. irregularis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFCE65EC5FBF8FC46" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[868,1030,896,922]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFCE65EC5FBF8FC46" box="[868,1030,896,922]" italics="true" pageId="31" pageNumber="32">E. krumbachi</emphasis>
|
||
</taxonomicName>
|
||
, and it provided sequences for further unnamed taxa within the
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFDB65EE0FD55FC62" authorityName="Henle" authorityYear="1837" box="[564,683,932,958]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFDB65EE0FD55FC62" box="[564,683,932,958]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<emphasis id="028AEA9EFFC0FFADFC845EE0FCE6FC61" box="[774,792,932,957]" italics="true" pageId="31" pageNumber="32">E</emphasis>
|
||
. sp.2 and
|
||
<emphasis id="028AEA9EFFC0FFADFC095EE0FC5AFC61" box="[907,932,932,957]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.3).
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC735EE0FB18FC62" authority="Nagy & Dózsa-Farkas & Felföldi, 2023" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1009,1254,932,958]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi" status="sp. nov.">
|
||
<emphasis id="028AEA9EFFC0FFADFC735EE0FB18FC62" box="[1009,1254,932,958]" italics="true" pageId="31" pageNumber="32">Enchytraeus andrasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFB6F5EE0FAB9FC62" box="[1261,1351,932,958]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3E5E8CFE7DFC3D" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[188,387,967,993]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasiformis">
|
||
<emphasis id="028AEA9EFFC0FFADFF3E5E8CFE7DFC3D" box="[188,387,967,993]" italics="true" pageId="31" pageNumber="32">E. andrasiformis</emphasis>
|
||
</taxonomicName>
|
||
formed distinct lineages, they were placed outside the clade of the
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFB105E8CFAF4FC3D" authorityName="Henle" authorityYear="1837" box="[1170,1290,967,993]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFB105E8CFAF4FC3D" box="[1170,1290,967,993]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex in the phylogenetic trees. The p-distances between the COI sequences of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFB825EAFFB84FBD8" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1024,1146,1002,1028]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi">
|
||
<emphasis id="028AEA9EFFC0FFADFB825EAFFB84FBD8" box="[1024,1146,1002,1028]" italics="true" pageId="31" pageNumber="32">E. andrasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFB2D5EAFFA8AFBD8" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1199,1396,1002,1028]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasiformis">
|
||
<emphasis id="028AEA9EFFC0FFADFB2D5EAFFA8AFBD8" box="[1199,1396,1002,1028]" italics="true" pageId="31" pageNumber="32">E. andrasiformis</emphasis>
|
||
</taxonomicName>
|
||
are 16.8–17.6%, the H3 distances between them are 3%. The COI distances between
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFB1D594AFAE4FBFB" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1183,1306,1037,1063]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi">
|
||
<emphasis id="028AEA9EFFC0FFADFB1D594AFAE4FBFB" box="[1183,1306,1037,1063]" italics="true" pageId="31" pageNumber="32">E. andrasi</emphasis>
|
||
</taxonomicName>
|
||
and the members of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFED05975FE35FB96" authorityName="Henle" authorityYear="1837" box="[338,459,1072,1098]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFED05975FE35FB96" box="[338,459,1072,1098]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex (including
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFD405975FC90FB96" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[706,878,1072,1098]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFD405975FC90FB96" box="[706,878,1072,1098]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
) are 15.9–20.6%, the H3 distances between them are 5.4–8.5%. The intraspecific COI distances between specimens of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFBA25910FB65FBB2" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1056,1179,1108,1134]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi">
|
||
<emphasis id="028AEA9EFFC0FFADFBA25910FB65FBB2" box="[1056,1179,1108,1134]" italics="true" pageId="31" pageNumber="32">E. andrasi</emphasis>
|
||
</taxonomicName>
|
||
are 0–1.8%. There are no intraspecific differences between the H3 sequences of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC0B593CFBFBFB4D" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[905,1029,1143,1169]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi">
|
||
<emphasis id="028AEA9EFFC0FFADFC0B593CFBFBFB4D" box="[905,1029,1143,1169]" italics="true" pageId="31" pageNumber="32">E. andrasi</emphasis>
|
||
</taxonomicName>
|
||
individuals. The COI distances between
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFEAB59DFFE0DFB68" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[297,499,1178,1204]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasiformis">
|
||
<emphasis id="028AEA9EFFC0FFADFEAB59DFFE0DFB68" box="[297,499,1178,1204]" italics="true" pageId="31" pageNumber="32">E. andrasiformis</emphasis>
|
||
</taxonomicName>
|
||
and the species of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFD6359DFFCA5FB68" authorityName="Henle" authorityYear="1837" box="[737,859,1178,1204]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFD6359DFFCA5FB68" box="[737,859,1178,1204]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex (including
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFBD559DFFAFBFB68" authority="Nagy & Dózsa-Farkas & Felföldi, 2023" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1111,1285,1178,1204]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
|
||
<emphasis id="028AEA9EFFC0FFADFBD559DFFAFBFB68" box="[1111,1285,1178,1204]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFA9259DFFA90FB68" box="[1296,1390,1178,1205]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) are 15.6–20.3%, the H3 distances between them are 5.2–8.1%.
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC2059F9FB36FB0B" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[930,1224,1213,1239]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFC2059F9FB36FB0B" box="[930,1224,1213,1239]" italics="true" pageId="31" pageNumber="32">Enchytraeus adrianensis</emphasis>
|
||
</taxonomicName>
|
||
and additional species missing from the study of
|
||
<bibRefCitation id="546F4B7DFFC0FFADFDEA59A5FCA1FB27" author="Erseus C. & Klinth M. J. & Rota E. & Wit P. D. & Gustafsson D. R. & Martinsson S." box="[616,863,1248,1275]" pageId="31" pageNumber="32" pagination="105 - 133" refId="ref21493" refString="Erseus C., Klinth M. J., Rota E., Wit P. D., Gustafsson D. R. & Martinsson S. 2019. The popular model annelid Enchytraeus albidus is only one species in a complex of seashore white worms (Clitellata, Enchytraeidae). Organisms Diversity & Evolution 19: 105 - 133. https: // doi. org / 10.1007 / s 13127 - 019 - 00402 - 6" type="journal article" year="2019">
|
||
Erséus
|
||
<emphasis id="028AEA9EFFC0FFADFD4059A5FCFCFB26" box="[706,770,1248,1274]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
, namely
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC5059A5FB89FB26" authorityName="Nielsen & Christensen" authorityYear="1961" box="[978,1143,1248,1274]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="irregularis">
|
||
<emphasis id="028AEA9EFFC0FFADFC5059A5FB89FB26" box="[978,1143,1248,1274]" italics="true" pageId="31" pageNumber="32">E. irregularis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="028AEA9EFFC0FFADFB0859A5FB62FB26" box="[1162,1180,1249,1274]" italics="true" pageId="31" pageNumber="32">E</emphasis>
|
||
. sp.2 and
|
||
<emphasis id="028AEA9EFFC0FFADFAA359A5FAC5FB26" box="[1313,1339,1249,1274]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.3 were placed inside the clade of the
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFDF05840FD13FAC2" authorityName="Henle" authorityYear="1837" box="[626,749,1284,1310]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFDF05840FD13FAC2" box="[626,749,1284,1310]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex. The COI distances between
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFB4A5840FA88FAC2" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[1224,1398,1284,1310]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFB4A5840FA88FAC2" box="[1224,1398,1284,1310]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
and members of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFE08586CFDFBFA9D" authorityName="Henle" authorityYear="1837" box="[394,517,1319,1345]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFE08586CFDFBFA9D" box="[394,517,1319,1345]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex are 12.8–17.8%, the H3 distances between them are 0.4–5.5%. There are no intraspecific differences between the COI and between the H3 sequences of specimens of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3F582AFE99FA5B" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[189,359,1389,1415]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F582AFE99FA5B" box="[189,359,1389,1415]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFEF45829FDBFFA5B" box="[374,577,1389,1415]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="undefined-2">
|
||
<emphasis id="028AEA9EFFC0FFADFEF45829FDF4FA5B" box="[374,522,1389,1415]" italics="true" pageId="31" pageNumber="32">Enchytraeus</emphasis>
|
||
sp.2
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="028AEA9EFFC0FFADFDFB582AFD6CFA5B" box="[633,658,1390,1415]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.3 formed one group on the gene trees. It is worth noting that the COI distance between
|
||
<emphasis id="028AEA9EFFC0FFADFE7358D5FDF5FA76" box="[497,523,1425,1450]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.2 and
|
||
<emphasis id="028AEA9EFFC0FFADFDF858D5FD6AFA76" box="[634,660,1425,1450]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.3 is very small (0.2%), while the H3 distance between them is bigger (2.5%), and comparable to interspecific differences in the rest of the H3 gene tree (
|
||
<figureCitation id="A8C52A09FFC0FFADFA9258F0FA96FA12" box="[1296,1384,1460,1486]" captionStart="Fig" captionStartId="33.[190,233,1892,1918]" captionTargetBox="[279,1309,605,1855]" captionTargetId="figure-128@33.[279,1309,605,1855]" captionTargetPageId="33" captionText="Fig. 11. Maximum likelihood (ML) tree of the H3 gene for specimens of the Enchytraeus albidus species complex, based on 225 nucleotide positions using the Kimura 2-parameter substitution model. Bootstrap values greater than 50 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Scale bar: 0.01 substitutions per nucleotide." figureDoi="http://doi.org/10.5281/zenodo.7986662" httpUri="https://zenodo.org/record/7986662/files/figure.png" pageId="31" pageNumber="32">Fig. 11</figureCitation>
|
||
).
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3F5893FE1AFA2D" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[189,484,1495,1521]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5893FE1AFA2D" box="[189,484,1495,1521]" italics="true" pageId="31" pageNumber="32">Enchytraeus adrianensis</emphasis>
|
||
</taxonomicName>
|
||
appeared as the sister group of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFCE4589CFBF6FA2D" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[870,1032,1495,1521]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFCE4589CFBF6FA2D" box="[870,1032,1495,1521]" italics="true" pageId="31" pageNumber="32">E. krumbachi</emphasis>
|
||
</taxonomicName>
|
||
in the COI and H3 gene trees, and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF6C58BFFE71F9C8" authorityName="Nielsen & Christensen" authorityYear="1961" box="[238,399,1530,1556]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="irregularis">
|
||
<emphasis id="028AEA9EFFC0FFADFF6C58BFFE71F9C8" box="[238,399,1530,1556]" italics="true" pageId="31" pageNumber="32">E. irregularis</emphasis>
|
||
</taxonomicName>
|
||
appeared as the sister group of
|
||
<emphasis id="028AEA9EFFC0FFADFC8058BFFCE2F9C8" box="[770,796,1531,1556]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.2 / sp.3 clade in both of the gene trees (
|
||
<figureCitation id="A8C52A09FFC0FFADFA8D58BEFF22F9EB" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="32.[189,232,1892,1918]" captionStartId-1="33.[190,233,1892,1918]" captionTargetBox-0="[189,1398,297,1823]" captionTargetBox-1="[279,1309,605,1855]" captionTargetId-0="figure-17@32.[189,1398,297,1823]" captionTargetId-1="figure-128@33.[279,1309,605,1855]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="Fig. 10. Maximum likelihood (ML) tree of the COI gene for specimens of the Enchytraeus albidus species complex, based on 568 nucleotide positions using the General Time Reversible substitution model. Bootstrap values greater than 50 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Scale bar: 0.05 substitutions per nucleotide." captionText-1="Fig. 11. Maximum likelihood (ML) tree of the H3 gene for specimens of the Enchytraeus albidus species complex, based on 225 nucleotide positions using the Kimura 2-parameter substitution model. Bootstrap values greater than 50 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Scale bar: 0.01 substitutions per nucleotide." figureDoi-0="http://doi.org/10.5281/zenodo.7986660" figureDoi-1="http://doi.org/10.5281/zenodo.7986662" httpUri-0="https://zenodo.org/record/7986660/files/figure.png" httpUri-1="https://zenodo.org/record/7986662/files/figure.png" pageId="31" pageNumber="32">Figs 10– 11</figureCitation>
|
||
). The COI distances between
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFDCB5B5AFD09F9EB" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[585,759,1565,1591]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFDCB5B5AFD09F9EB" box="[585,759,1565,1591]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFCB45B5AFC26F9EB" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[822,984,1565,1591]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFCB45B5AFC26F9EB" box="[822,984,1565,1591]" italics="true" pageId="31" pageNumber="32">E. krumbachi</emphasis>
|
||
</taxonomicName>
|
||
are 12.9–13.4%, the H3 distances between them are low, 0.4–1.6%. The COI distances between
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC175B05FBC8F986" authorityName="Nielsen & Christensen" authorityYear="1961" box="[917,1078,1600,1626]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="irregularis">
|
||
<emphasis id="028AEA9EFFC0FFADFC175B05FBC8F986" box="[917,1078,1600,1626]" italics="true" pageId="31" pageNumber="32">E. irregularis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="028AEA9EFFC0FFADFBEC5B05FB76F986" box="[1134,1160,1601,1626]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.2 / sp.3 clade are 12.2–12.9%, the H3 distances between them are 2.1–3.8%. According to the molecular analysis, our
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3F5BCCFE9EF97D" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[189,352,1671,1697]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5BCCFE9EF97D" box="[189,352,1671,1697]" italics="true" pageId="31" pageNumber="32">E. krumbachi</emphasis>
|
||
</taxonomicName>
|
||
individuals and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFDAE5BCCFC9CF97D" authority="CE 1684" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[556,866,1671,1697]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" isUncertain="true" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFDAE5BCCFDB8F97D" box="[556,582,1672,1697]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
cf.
|
||
<emphasis id="028AEA9EFFC0FFADFDFB5BC3FD06F97D" box="[633,760,1671,1697]" italics="true" pageId="31" pageNumber="32">krumbachi</emphasis>
|
||
CE1684
|
||
</taxonomicName>
|
||
and CE1689 specimens belong to the same species. The intraspecific COI distances between our
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFCC25BEFFC1DF918" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[832,995,1706,1732]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFCC25BEFFC1DF918" box="[832,995,1706,1732]" italics="true" pageId="31" pageNumber="32">E. krumbachi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFBA25BEFFB15F918" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[1056,1259,1706,1732]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" isUncertain="true" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">
|
||
<emphasis id="028AEA9EFFC0FFADFBA25BEFFBC4F918" box="[1056,1082,1707,1732]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
cf.
|
||
<emphasis id="028AEA9EFFC0FFADFBEE5BEEFB15F918" box="[1132,1259,1706,1732]" italics="true" pageId="31" pageNumber="32">krumbachi</emphasis>
|
||
</taxonomicName>
|
||
individuals are 0.7–3%, the intraspecific H3 distances between them are 0–0.8%. On the species tree (
|
||
<figureCitation id="A8C52A09FFC0FFADFB655B89FAC2F93B" box="[1255,1340,1741,1767]" captionStart="Fig" captionStartId="34.[190,233,1892,1918]" captionTargetBox="[203,1383,1356,1842]" captionTargetId="figure-498@34.[189,1398,1335,1854]" captionTargetPageId="34" captionText="Fig. 12. Enchytraeus Henle, 1837 species tree, based on COI, H3 and 16S rRNA gene (Bayes analysis, *BEAST). Posterior probabilities greater than 0.5 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Abbreviation: n = number of included specimens for each species. Scale bar = 0.07 substitutions per nucleotide." figureDoi="http://doi.org/10.5281/zenodo.7986666" httpUri="https://zenodo.org/record/7986666/files/figure.png" pageId="31" pageNumber="32">Fig. 12</figureCitation>
|
||
), the
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3F5BB5FEC9F8D6" authorityName="Henle" authorityYear="1837" box="[189,311,1776,1802]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5BB5FEC9F8D6" box="[189,311,1776,1802]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex formed a distinct clade with maximum posterior probability support and contains
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFF3F5A50FE99F8F2" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[189,359,1812,1838]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis">
|
||
<emphasis id="028AEA9EFFC0FFADFF3F5A50FE99F8F2" box="[189,359,1812,1838]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFEF45A50FDE8F8F2" authorityName="Nielsen & Christensen" authorityYear="1961" box="[374,534,1812,1838]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="irregularis">
|
||
<emphasis id="028AEA9EFFC0FFADFEF45A50FDE8F8F2" box="[374,534,1812,1838]" italics="true" pageId="31" pageNumber="32">E. irregularis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="028AEA9EFFC0FFADFDCF5A50FD98F8F1" box="[589,614,1812,1837]" italics="true" pageId="31" pageNumber="32">E.</emphasis>
|
||
sp.2 / sp.3, as in the gene trees.
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFC5E5A50FB78F8F2" authority="Nagy & Dózsa-Farkas & Felföldi, 2023" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[988,1158,1812,1838]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="adrianensis" status="sp. nov.">
|
||
<emphasis id="028AEA9EFFC0FFADFC5E5A50FB78F8F2" box="[988,1158,1812,1838]" italics="true" pageId="31" pageNumber="32">E. adrianensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19B957E5FFC0FFADFB0F5A50FB19F8F2" box="[1165,1255,1812,1838]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
appeared as the sister group of
|
||
<emphasis id="028AEA9EFFC0FFADFE125A7CFDAAF88D" box="[400,596,1847,1873]" italics="true" pageId="31" pageNumber="32">
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFE125A7CFDCFF88D" authorityName="CE" authorityYear="1684" baseAuthorityName="Cejka" baseAuthorityYear="1913" box="[400,561,1847,1873]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="krumbachi">E. krumbachi</taxonomicName>
|
||
, E.
|
||
</emphasis>
|
||
sp.2 / sp.3 appeared as the sister group of
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFBB75A7CFB2AF88D" authorityName="Nielsen & Christensen" authorityYear="1961" box="[1077,1236,1847,1873]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="irregularis">
|
||
<emphasis id="028AEA9EFFC0FFADFBB75A7CFB2AF88D" box="[1077,1236,1847,1873]" italics="true" pageId="31" pageNumber="32">E. irregularis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFB605A73FEE6F8A8" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi">
|
||
<emphasis id="028AEA9EFFC0FFADFB605A73FEE6F8A8" italics="true" pageId="31" pageNumber="32">Enchytraeus andrasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFED45A1FFDE0F8A8" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[342,542,1882,1908]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasiformis">
|
||
<emphasis id="028AEA9EFFC0FFADFED45A1FFDE0F8A8" box="[342,542,1882,1908]" italics="true" pageId="31" pageNumber="32">E. andrasiformis</emphasis>
|
||
</taxonomicName>
|
||
were placed outside the
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFCC85A1FFC3DF8A8" authorityName="Henle" authorityYear="1837" box="[842,963,1882,1908]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="albidus">
|
||
<emphasis id="028AEA9EFFC0FFADFCC85A1FFC3DF8A8" box="[842,963,1882,1908]" italics="true" pageId="31" pageNumber="32">E. albidus</emphasis>
|
||
</taxonomicName>
|
||
complex and they appeared there as sister groups of each other.
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFE7D5A3AFD85F84B" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[511,635,1917,1943]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasi">
|
||
<emphasis id="028AEA9EFFC0FFADFE7D5A3AFD85F84B" box="[511,635,1917,1943]" italics="true" pageId="31" pageNumber="32">E. andrasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFD365A3AFC85F84B" authorityName="Nagy & Dózsa-Farkas & Felföldi" authorityYear="2023" box="[692,891,1917,1943]" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="andrasiformis">
|
||
<emphasis id="028AEA9EFFC0FFADFD365A3AFC85F84B" box="[692,891,1917,1943]" italics="true" pageId="31" pageNumber="32">E. andrasiformis</emphasis>
|
||
</taxonomicName>
|
||
clustered with
|
||
<taxonomicName id="F7FE4D0FFFC0FFADFBAD5A3AFF06F866" authority="Abrahamsen, 1969" authorityName="Abrahamsen" authorityYear="1969" class="Clitellata" family="Enchytraeidae" genus="Enchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="norvegicus">
|
||
<emphasis id="028AEA9EFFC0FFADFBAD5A3AFB2CF84B" box="[1071,1234,1918,1943]" italics="true" pageId="31" pageNumber="32">E. norvegicus</emphasis>
|
||
Abrahamsen, 1969
|
||
</taxonomicName>
|
||
on the species tree (
|
||
<figureCitation id="A8C52A09FFC0FFADFE655AE4FDC2F867" box="[487,572,1952,1979]" captionStart="Fig" captionStartId="34.[190,233,1892,1918]" captionTargetBox="[203,1383,1356,1842]" captionTargetId="figure-498@34.[189,1398,1335,1854]" captionTargetPageId="34" captionText="Fig. 12. Enchytraeus Henle, 1837 species tree, based on COI, H3 and 16S rRNA gene (Bayes analysis, *BEAST). Posterior probabilities greater than 0.5 are shown at the nodes. Accession codes of sequences with collection information are given in Table 1. Abbreviation: n = number of included specimens for each species. Scale bar = 0.07 substitutions per nucleotide." figureDoi="http://doi.org/10.5281/zenodo.7986666" httpUri="https://zenodo.org/record/7986666/files/figure.png" pageId="31" pageNumber="32">Fig. 12</figureCitation>
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) but with only low posterior probability support, and the latter species differs from the new species in many traits.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |