treatments-xml/data/8B/63/87/8B6387AE166CFF99EB8DFA886C839C6C.xml

<document id="FF4A737905D2A8FF71B495AA30CAEB8C" ID="10.11646/zootaxa.4273.1.10SLASH11205" ID-CLB-Dataset="32921" ID-DOI="10.11646/zootaxa.4273.1.10" ID-GBIF-Dataset="bf6bbd16-4517-4814-8ec6-9b42014ed797" ID-ISSN="1175-5326" ID-Zenodo-Dep="801855" ID-ZooBank="F42A8DE1-C70E-47E1-AF93-A99A66E97476" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496357082059" checkinUser="plazi" docAuthor="Manuel, Michaël &amp; Ramahandrison, Andriamirado T." docDate="2017" docId="8B6387AE166CFF99EB8DFA886C839C6C" docLanguage="en" docName="zootaxa.4273.1.10SLASH11205.pdf" docOrigin="Zootaxa 4273 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Canthyporus reebae Manuel &amp; Ramahandrison, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="138" masterDocId="775AFFD6166DFF9EEB1AFF9C6D1D9F54" masterDocTitle="Canthyporus reebae sp. nov. from the Itremo and Andringitra mountains of central eastern Madagascar (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="140" masterPageNumber="131" pageNumber="132" updateTime="1698440136036" updateUser="ExternalLinkService">
<mods:mods id="F0A5E7BC879E8F88186932A245A3B5DA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="17616980431EA600EC82FF018823E2F1">Canthyporus reebae sp. nov. from the Itremo and Andringitra mountains of central eastern Madagascar (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="3642508AC92FB3478DC062D350273F9A">Manuel, Michaël</mods:namePart>
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<mods:name id="249A33595259ABDFCC41DD4050A1319F" type="personal">
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<mods:namePart id="41B9BDE1C730D5096096A4721A382C54">Ramahandrison, Andriamirado T.</mods:namePart>
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<mods:typeOfResource id="8EEF92E39A67C2818BD866AFFCCE9476">text</mods:typeOfResource>
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<mods:title id="E3C57A61B5F541CD162756B769588C7A">Zootaxa</mods:title>
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<mods:part id="47C12834151B8465D52EAC67001B6603">
<mods:date id="521957B0A973A79A501453A158A1E189">2017</mods:date>
<mods:detail id="0FDDE91FDCBD204C9958F0FD3D19CD07" type="volume">
<mods:number id="10910C1ED45D7B9986F450BF21DE6302">4273</mods:number>
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<mods:identifier id="1CB500F1F658A75D4E900CD2806FCA5C" type="DOI">10.11646/zootaxa.4273.1.10</mods:identifier>
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<treatment id="8B6387AE166CFF99EB8DFA886C839C6C" ID-DOI="http://doi.org/10.5281/zenodo.6038528" ID-GBIF-Taxon="131811808" ID-Zenodo-Dep="6038528" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C" httpUri="http://treatment.plazi.org/id/8B6387AE166CFF99EB8DFA886C839C6C" lastPageId="7" lastPageNumber="138" pageId="1" pageNumber="132">
<subSubSection id="4BD06533166CFF9FEB8DFA886C3E9A04" pageId="1" pageNumber="132" type="nomenclature">
<paragraph id="037536B8166CFF9FEB8DFA886CF29A7B" blockId="1.[151,495,1300,1360]" box="[151,495,1300,1327]" pageId="1" pageNumber="132">
<heading id="583D81D4166CFF9FEB8DFA886CF29A7B" bold="true" box="[151,495,1300,1327]" fontSize="11" level="1" pageId="1" pageNumber="132" reason="1">
<emphasis id="31BEEAAA166CFF9FEB8DFA886CF29A7B" bold="true" box="[151,495,1300,1327]" pageId="1" pageNumber="132">
<taxonomicName id="C4CA4D3B166CFF9FEB8DFA886C979A7A" ID-CoL="8LZ4P" box="[151,394,1300,1326]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166CFF9FEB8DFA886C979A7A" bold="true" box="[151,394,1300,1326]" italics="true" pageId="1" pageNumber="132">Canthyporus reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166CFF9FEA88FA896CF29A7B" box="[402,495,1301,1327]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="037536B8166CFF9FEB8DFAAB6C3E9A04" blockId="1.[151,495,1300,1360]" box="[151,291,1335,1360]" pageId="1" pageNumber="132">
<figureCitation id="9BF12A3D166CFF9FEB8DFAAB6C029A04" box="[151,287,1335,1360]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 6" captionStart-2="FIGURES 7 – 9" captionStartId-0="2.[151,264,1302,1325]" captionStartId-1="3.[151,264,1106,1129]" captionStartId-2="3.[151,264,1969,1992]" captionTargetBox-0="[220,1387,324,1282]" captionTargetBox-1="[192,1402,473,1089]" captionTargetBox-2="[346,1238,1201,1948]" captionTargetId-0="figure@2.[197,1387,303,1282]" captionTargetId-1="figure@3.[192,1402,473,1089]" captionTargetId-2="figure@3.[346,1238,1201,1948]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." captionText-1="FIGURES 3 – 6. Male genitalia of Canthyporus reebae sp. nov. (holotype). (3) Median lobe of aedeagus in lateral view; (4) median lobe of aedeagus in ventral view; (5) left paramere; (6) right paramere (parameres in external view). Scale bar: 0. 2 mm." captionText-2="FIGURES 7 – 9. Female reproductive structures of Canthyporus reebae sp. nov. (paratype from the locus typicus). (7) Left gonocoxosternum in ventral view; (8) gonocoxae in ventral view; (9) spermathecal tract. Scale bar: 0. 2 mm." httpUri-0="https://zenodo.org/record/801857/files/figure.png" httpUri-1="https://zenodo.org/record/801859/files/figure.png" httpUri-2="https://zenodo.org/record/801861/files/figure.png" pageId="1" pageNumber="132">Figures 1–9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4BD06533166CFF9CEB8DFAE36FBE9FAC" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132" type="materials_examined">
<paragraph id="037536B8166CFF9FEB8DFAE36C489A8B" blockId="1.[151,1437,1407,2009]" pageId="1" pageNumber="132">
<materialsCitation id="B3A23CE5166CFF9FEB8DFAE36C4C9A8B" ID-GBIF-Occurrence="1563183426" country="Madagascar" elevation="1700" latitude="-20.766666" location="Ambalamanaka" longLatPrecision="1266" longitude="47.166668" municipality="Itremo mountain range" pageId="1" pageNumber="132" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="31BEEAAA166CFF9FEB8DFAE36C329ACC" bold="true" box="[151,303,1407,1432]" pageId="1" pageNumber="132">
<typeStatus id="DC71881A166CFF9FEB8DFAE36DCF9ACC" box="[151,210,1407,1432]" pageId="1" pageNumber="132">Type</typeStatus>
locality
</emphasis>
. 
<collectingCountry id="7BDD7628166CFF9FEA24FAE36CDA9ACC" box="[318,455,1407,1432]" name="Madagascar" pageId="1" pageNumber="132">Madagascar</collectingCountry>
, former province of 
<collectingRegion id="C10EF85A166CFF9FE9ABFAE36E5E9ACC" box="[689,835,1407,1432]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="132">Fianarantsoa</collectingRegion>
, south-eastern part of the 
<collectingMunicipality id="E311ACC2166CFF9FEF70FA1C686D9ACC" box="[1130,1392,1407,1432]" pageId="1" pageNumber="132">Itremo mountain range</collectingMunicipality>
, ca. 
<quantity id="C4329B5D166CFF9FEB8DFA386DF99AE9" box="[151,228,1444,1469]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="132" unit="km" value="3.5">3.5 km</quantity>
South of 
<location id="06156063166CFF9FEA4AFA386F1E9AE9" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C:06156063166CFF9FEA4AFA386F1E9AE9" box="[336,515,1444,1469]" country="Madagascar" latitude="-20.766666" longLatPrecision="1266" longitude="47.166668" municipality="Itremo mountain range" name="Ambalamanaka" pageId="1" pageNumber="132" stateProvince="Fianarantsoa">Ambalamanaka</location>
, large densely vegetated pond close to N7 road, GPS coordinates 
<geoCoordinate id="66FE507F166CFF9FEFC4FA38682A9AE9" box="[1246,1335,1444,1469]" direction="south" orientation="latitude" pageId="1" pageNumber="132" precision="925" value="-20.766666">S20°46'</geoCoordinate>
<geoCoordinate id="66FE507F166CFF9FEE26FA3868849AE8" box="[1340,1433,1444,1468]" direction="east" orientation="longitude" pageId="1" pageNumber="132" precision="925" value="47.166668">E47°10'</geoCoordinate>
, altitude 
<elevation id="88E7D18B166CFF9FEBE8FA546C4C9A8B" box="[242,337,1480,1504]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="132" unit="m" value="1700.0">
<quantity id="C4329B5D166CFF9FEBE8FA546C4C9A8B" box="[242,337,1480,1504]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="132" unit="m" value="1700.0">1,700 m</quantity>
</elevation>
</materialsCitation>
.
</paragraph>
<paragraph id="037536B8166CFF9FEBDDFA706C7B99C0" blockId="1.[151,1437,1407,2009]" pageId="1" pageNumber="132">
<materialsCitation id="B3A23CE5166CFF9FEBDDFA706FD79951" ID-GBIF-Occurrence="1563183430" box="[199,714,1516,1541]" country="Madagascar" location="Madagascar" pageId="1" pageNumber="132" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="31BEEAAA166CFF9FEBDDFA706CED9951" bold="true" box="[199,496,1516,1541]" pageId="1" pageNumber="132">
<typeStatus id="DC71881A166CFF9FEBDDFA706C1F9951" box="[199,258,1516,1541]" pageId="1" pageNumber="132">Type</typeStatus>
material. 
<typeStatus id="DC71881A166CFF9FEA99FA706CED9951" box="[387,496,1516,1541]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="15CCFD31166CFF9FE91EFA706F019950" box="[516,540,1516,1540]" pageId="1" pageNumber="132" type="male">♂</specimenCount>
): &quot; 
<collectingCountry id="7BDD7628166CFF9FE927FA706FD79951" box="[573,714,1516,1541]" name="Madagascar" pageId="1" pageNumber="132">Madagascar</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="B3A23CE5166CFF9FE9C2FA716C7B99C0" ID-GBIF-Occurrence="1563183428" collectingDate="2016-05-19" collectionCode="MNHN" collectorName="Manuel &amp; Ramahandrison &amp; Alt. &amp; Near &amp; Close" country="Madagascar" elevation="1700" latitude="-20.766666" location="Itremo mountain range" longLatPrecision="1266" longitude="47.166668" pageId="1" pageNumber="132" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
ex-Prov. 
<collectingRegion id="C10EF85A166CFF9FE85AFA706ECE9951" box="[832,979,1516,1541]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="132">Fianarantsoa</collectingRegion>
. ca. 
<locationDeviation id="12C72F9A166CFF9FEF16FA7068279951" box="[1036,1338,1516,1541]" pageId="1" pageNumber="132">
<quantity id="C4329B5D166CFF9FEF16FA7069439951" box="[1036,1118,1516,1541]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="132" unit="km" value="3.5">3.5 km</quantity>
S Ambalamanaka.
</locationDeviation>
<geoCoordinate id="66FE507F166CFF9FEE5EFA7068809951" box="[1348,1437,1516,1541]" direction="south" orientation="latitude" pageId="1" pageNumber="132" precision="925" value="-20.766666">S20°46'</geoCoordinate>
<geoCoordinate id="66FE507F166CFF9FEB8DF98C6DE9997C" box="[151,244,1552,1576]" direction="east" orientation="longitude" pageId="1" pageNumber="132" precision="925" value="47.166668">E47°10'</geoCoordinate>
. 
<collectingDate id="6730E990166CFF9FEBE4F9936C69997C" box="[254,372,1551,1576]" pageId="1" pageNumber="132" value="2016-05-19">
<date id="77741078166CFF9FEBE4F9936C69997C" box="[254,372,1551,1576]" pageId="1" pageNumber="132" value="2016-05-19">19.V.2016</date>
</collectingDate>
. 
<collectorName id="AE3F536E166CFF9FEA65F9936CC9997C" box="[383,468,1551,1576]" pageId="1" pageNumber="132">Manuel</collectorName>
&amp; 
<collectorName id="AE3F536E166CFF9FEAE3F9936FB4997C" box="[505,681,1551,1576]" pageId="1" pageNumber="132">Ramahandrison</collectorName>
leg.&quot;, &quot; 
<collectorName id="AE3F536E166CFF9FE9E6F9936E34997C" box="[764,809,1551,1576]" pageId="1" pageNumber="132">Alt.</collectorName>
<elevation id="88E7D18B166CFF9FE82AF98C6E929973" box="[816,911,1552,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="132" unit="m" value="1700.0">
<quantity id="C4329B5D166CFF9FE82AF98C6E929973" box="[816,911,1552,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="132" unit="m" value="1700.0">1,700 m</quantity>
</elevation>
. 
<collectorName id="AE3F536E166CFF9FE880F98C6ECD9973" box="[922,976,1552,1575]" pageId="1" pageNumber="132">Near</collectorName>
end of small stream, on margins of large pond with 
<taxonomicName id="C4CA4D3B166CFF9FEA0DF9A86CDF9919" box="[279,450,1588,1613]" class="Magnoliopsida" family="Nymphaeaceae" kingdom="Plantae" order="Nymphaeales" pageId="1" pageNumber="132" phylum="Tracheophyta" rank="family">Nymphaeaceae</taxonomicName>
and peat bog with 
<taxonomicName id="C4CA4D3B166CFF9FE9B6F9A86E3F9918" box="[684,802,1588,1612]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="1" pageNumber="132" phylum="Bryophyta" rank="genus">
<emphasis id="31BEEAAA166CFF9FE9B6F9A86E3F9918" box="[684,802,1588,1612]" italics="true" pageId="1" pageNumber="132">Sphagnum</emphasis>
</taxonomicName>
and 
<taxonomicName id="C4CA4D3B166CFF9FE878F9A96EA39918" box="[866,958,1589,1612]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="31BEEAAA166CFF9FE878F9A96EA39918" box="[866,958,1589,1612]" italics="true" pageId="1" pageNumber="132">Drosera</emphasis>
</taxonomicName>
. 
<collectorName id="AE3F536E166CFF9FE8CAF9A8690D9919" box="[976,1040,1588,1613]" pageId="1" pageNumber="132">Close</collectorName>
to N7 road, south-eastern part of 
<location id="06156063166CFF9FEB8DF9C46C869924" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C:06156063166CFF9FEB8DF9C46C869924" box="[151,411,1623,1648]" country="Madagascar" latitude="-20.766666" longLatPrecision="1266" longitude="47.166668" name="Itremo mountain range" pageId="1" pageNumber="132" stateProvince="Fianarantsoa">Itremo mountain range</location>
&quot; [printed], “ 
<typeStatus id="DC71881A166CFF9FE93CF9CB6F8E9924" box="[550,659,1623,1648]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
, 
<taxonomicName id="C4CA4D3B166CFF9FE985F9CB6E639924" box="[671,894,1623,1648]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">Canthyporus reebae</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166CFF9FE89CF9C46EC2993B" box="[902,991,1624,1647]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
, 
<collectorName id="AE3F536E166CFF9FE8F1F9CB695D9924" box="[1003,1088,1623,1648]" pageId="1" pageNumber="132">Manuel</collectorName>
&amp; 
<collectorName id="AE3F536E166CFF9FEF7FF9CB68089924" box="[1125,1301,1623,1648]" pageId="1" pageNumber="132">Ramahandrison</collectorName>
2017” [red, printed] (
<collectionCode id="65DBAE7D166CFF9FEBE4F9E06C4799C0" box="[254,346,1660,1684]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="132">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="037536B8166CFF9FEBDDF9036881988C" blockId="1.[151,1437,1407,2009]" pageId="1" pageNumber="132">
<emphasis id="31BEEAAA166CFF9FEBDDF9036C2299EC" bold="true" box="[199,319,1695,1720]" pageId="1" pageNumber="132">Paratypes</emphasis>
(in total 14♂, 20♀) (CMM, MNHN, BMNH, NMPC): 
<materialsCitation id="B3A23CE5166CFF9FE8DDF93C682C99EC" ID-GBIF-Occurrence="1563183429" box="[967,1329,1695,1720]" collectingDate="2016-05-19" collectionCode="MNHN" collectorName="Manuel &amp; Ramahandrison &amp; Alt. &amp; Near &amp; Close" country="Madagascar" elevation="1700" latitude="-20.766666" location="Itremo mountain range" longLatPrecision="1266" longitude="47.166668" pageId="1" pageNumber="132" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="Fianarantsoa">
<specimenCount id="15CCFD31166CFF9FE8DDF93C6EF199EC" box="[967,1004,1696,1720]" pageId="1" pageNumber="132" type="male">4♂</specimenCount>
, 
<specimenCount id="15CCFD31166CFF9FE8E1F93C690299EC" box="[1019,1055,1696,1720]" pageId="1" pageNumber="132" type="female">8♀</specimenCount>
, same data as holotype
</materialsCitation>
. 
<materialsCitation id="B3A23CE5166CFF9FEE24F93C6C449871" ID-GBIF-Occurrence="1563183433" collectingDate="2016-05-21" collectorName="Manuel &amp; Ramahandrison &amp; Alt. &amp; Seepage &amp; Andringitra National Park" country="Madagascar" elevation="1600" latitude="-22.133333" location="Antanifotsy." longLatPrecision="1261" longitude="46.883335" pageId="1" pageNumber="132" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Fianarantsoa">
<specimenCount id="15CCFD31166CFF9FEE24F93C687999EC" box="[1342,1380,1696,1720]" pageId="1" pageNumber="132" type="male">2♂</specimenCount>
, 
<specimenCount id="15CCFD31166CFF9FEE68F93C688899EC" box="[1394,1429,1696,1720]" pageId="1" pageNumber="132" type="female">1♀</specimenCount>
, &quot; 
<collectingCountry id="7BDD7628166CFF9FEB85F9586C319989" box="[159,300,1732,1757]" name="Madagascar" pageId="1" pageNumber="132">Madagascar</collectingCountry>
. ex-Prov. 
<collectingRegion id="C10EF85A166CFF9FEAB1F9586F239989" box="[427,574,1732,1757]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="132">Fianarantsoa</collectingRegion>
. ca. 
<quantity id="C4329B5D166CFF9FE964F9586FC99989" box="[638,724,1732,1757]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="132" unit="km" value="2.5">2.5 km</quantity>
SSW 
<location id="06156063166CFF9FE831F9586EA89989" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C:06156063166CFF9FE831F9586EA89989" box="[811,949,1732,1757]" country="Madagascar" latitude="-22.133333" longLatPrecision="1261" longitude="46.883335" name="Antanifotsy." pageId="1" pageNumber="132" stateProvince="Fianarantsoa">Antanifotsy.</location>
<geoCoordinate id="66FE507F166CFF9FE8D9F95869009988" box="[963,1053,1732,1756]" direction="south" orientation="latitude" pageId="1" pageNumber="132" precision="925" value="-22.133333">S22°08'</geoCoordinate>
<geoCoordinate id="66FE507F166CFF9FEF30F95869959989" box="[1066,1160,1732,1757]" direction="east" orientation="longitude" pageId="1" pageNumber="132" precision="925" value="46.883335">E46°53'</geoCoordinate>
. 
<collectingDate id="6730E990166CFF9FEF83F958680D9989" box="[1177,1296,1732,1757]" pageId="1" pageNumber="132" value="2016-05-21">
<date id="77741078166CFF9FEF83F958680D9989" box="[1177,1296,1732,1757]" pageId="1" pageNumber="132" value="2016-05-21">21.V.2016</date>
</collectingDate>
. 
<collectorName id="AE3F536E166CFF9FEE38F95868659989" box="[1314,1400,1732,1757]" pageId="1" pageNumber="132">Manuel</collectorName>
&amp; 
<collectorName id="AE3F536E166CFF9FEB8DF97B6C5A9854" box="[151,327,1767,1792]" pageId="1" pageNumber="132">Ramahandrison</collectorName>
leg.&quot;, &quot; 
<collectorName id="AE3F536E166CFF9FEA80F97B6CD59854" box="[410,456,1767,1792]" pageId="1" pageNumber="132">Alt.</collectorName>
<elevation id="88E7D18B166CFF9FEAD5F97B6F3399AB" box="[463,558,1767,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="132" unit="m" value="1600.0">
<quantity id="C4329B5D166CFF9FEAD5F97B6F3399AB" box="[463,558,1767,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="132" unit="m" value="1600.0">1,600 m</quantity>
</elevation>
. 
<collectorName id="AE3F536E166CFF9FE920F9746F8A9854" box="[570,663,1768,1792]" pageId="1" pageNumber="132">Seepage</collectorName>
in meadow surrounded by forest. 
<collectorName id="AE3F536E166CFF9FEF03F97B685C9854" box="[1049,1345,1767,1792]" pageId="1" pageNumber="132">Andringitra National Park</collectorName>
, eastern slope of massif&quot;
</materialsCitation>
. 
<materialsCitation id="B3A23CE5166CFF9FEA72F89069719839" ID-GBIF-Occurrence="1563183427" collectingDate="2016-05-22" collectorName="Manuel &amp; Ramahandrison &amp; Alt. &amp; Small &amp; Andringitra National Park" country="Madagascar" elevation="1580" latitude="-22.116667" location="Antanifotsy." longLatPrecision="1261" longitude="46.883335" pageId="1" pageNumber="132" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<specimenCount id="15CCFD31166CFF9FEA72F8906C919870" box="[360,396,1804,1828]" pageId="1" pageNumber="132" type="male">1♂</specimenCount>
, 
<specimenCount id="15CCFD31166CFF9FEA84F8906CDC9870" box="[414,449,1804,1828]" pageId="1" pageNumber="132" type="female">1♀</specimenCount>
, &quot; 
<collectingCountry id="7BDD7628166CFF9FEAC6F8906F749871" box="[476,617,1804,1829]" name="Madagascar" pageId="1" pageNumber="132">Madagascar</collectingCountry>
. ex-Prov. 
<collectingRegion id="C10EF85A166CFF9FE9FBF8906E699871" box="[737,884,1804,1829]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="132">Fianarantsoa</collectingRegion>
. ca. 
<quantity id="C4329B5D166CFF9FE8B5F890691F9871" box="[943,1026,1804,1829]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="132" unit="km" value="2.3">2.3 km</quantity>
SW 
<location id="06156063166CFF9FEF59F89069D09871" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C:06156063166CFF9FEF59F89069D09871" box="[1091,1229,1804,1829]" country="Madagascar" latitude="-22.116667" longLatPrecision="1261" longitude="46.883335" name="Antanifotsy." pageId="1" pageNumber="132" stateProvince="Fianarantsoa">Antanifotsy.</location>
<geoCoordinate id="66FE507F166CFF9FEFC2F890682C9870" box="[1240,1329,1804,1828]" direction="south" orientation="latitude" pageId="1" pageNumber="132" precision="925" value="-22.116667">S22°07'</geoCoordinate>
<geoCoordinate id="66FE507F166CFF9FEE26F89068879871" box="[1340,1434,1804,1829]" direction="east" orientation="longitude" pageId="1" pageNumber="132" precision="925" value="46.883335">E46°53'</geoCoordinate>
. 
<collectingDate id="6730E990166CFF9FEB8DF8B36C11981C" box="[151,268,1839,1864]" pageId="1" pageNumber="132" value="2016-05-22">
<date id="77741078166CFF9FEB8DF8B36C11981C" box="[151,268,1839,1864]" pageId="1" pageNumber="132" value="2016-05-22">22.V.2016</date>
</collectingDate>
. 
<collectorName id="AE3F536E166CFF9FEA06F8B36C6C981C" box="[284,369,1839,1864]" pageId="1" pageNumber="132">Manuel</collectorName>
&amp; 
<collectorName id="AE3F536E166CFF9FEA87F8B36F50981C" box="[413,589,1839,1864]" pageId="1" pageNumber="132">Ramahandrison</collectorName>
leg.&quot;, &quot; 
<collectorName id="AE3F536E166CFF9FE9B3F8B36FCB981C" box="[681,726,1839,1864]" pageId="1" pageNumber="132">Alt.</collectorName>
<elevation id="88E7D18B166CFF9FE9FBF8B36E599813" box="[737,836,1839,1864]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="132" unit="m" value="1580.0">
<quantity id="C4329B5D166CFF9FE9FBF8B36E599813" box="[737,836,1839,1864]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="132" unit="m" value="1580.0">1,580 m</quantity>
</elevation>
. 
<collectorName id="AE3F536E166CFF9FE849F8B36E89981C" box="[851,916,1839,1864]" pageId="1" pageNumber="132">Small</collectorName>
ditch with clear circulating water, near trail between bridge and campground 1. 
<collectorName id="AE3F536E166CFF9FE933F8C86E4C9839" box="[553,849,1876,1901]" pageId="1" pageNumber="132">Andringitra National Park</collectorName>
, eastern slope of massif&quot;
</materialsCitation>
. 
<materialsCitation id="B3A23CE5166CFF9FEF6DF8C86E7C988D" ID-GBIF-Occurrence="1563183432" collectingDate="2016-05-23" collectorName="Manuel &amp; Ramahandrison &amp; Alt. &amp; Small &amp; Andringitra National Park" country="Madagascar" elevation="2140" latitude="-22.133333" location="Antanifotsy." longLatPrecision="1261" longitude="46.866665" pageId="1" pageNumber="132" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Fianarantsoa">
<specimenCount id="15CCFD31166CFF9FEF6DF8C869819838" box="[1143,1180,1876,1900]" pageId="1" pageNumber="132" type="male">2♂</specimenCount>
, 
<specimenCount id="15CCFD31166CFF9FEFB2F8C869D19838" box="[1192,1228,1876,1900]" pageId="1" pageNumber="132" type="female">3♀</specimenCount>
, &quot; 
<collectingCountry id="7BDD7628166CFF9FEFFBF8C868739839" box="[1249,1390,1876,1901]" name="Madagascar" pageId="1" pageNumber="132">Madagascar</collectingCountry>
. ex- Prov. 
<collectingRegion id="C10EF85A166CFF9FEBCDF8EB6C7798C4" box="[215,362,1911,1936]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="132">Fianarantsoa</collectingRegion>
. ca. 
<quantity id="C4329B5D166CFF9FEA81F8E46CF598C4" box="[411,488,1911,1936]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="132" unit="km" value="4.2">4.2 km</quantity>
SW 
<location id="06156063166CFF9FE905F8EB6FB798C4" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C:06156063166CFF9FE905F8EB6FB798C4" box="[543,682,1911,1936]" country="Madagascar" latitude="-22.133333" longLatPrecision="1261" longitude="46.866665" name="Antanifotsy." pageId="1" pageNumber="132" stateProvince="Fianarantsoa">Antanifotsy.</location>
<geoCoordinate id="66FE507F166CFF9FE9B5F8E46E1598C4" box="[687,776,1912,1936]" direction="south" orientation="latitude" pageId="1" pageNumber="132" precision="925" value="-22.133333">S22°08'</geoCoordinate>
<geoCoordinate id="66FE507F166CFF9FE814F8EB6E7198C4" box="[782,876,1911,1936]" direction="east" orientation="longitude" pageId="1" pageNumber="132" precision="925" value="46.866665">E46°52'</geoCoordinate>
. 
<collectingDate id="6730E990166CFF9FE86FF8EB6EF698C4" box="[885,1003,1911,1936]" pageId="1" pageNumber="132" value="2016-05-23">
<date id="77741078166CFF9FE86FF8EB6EF698C4" box="[885,1003,1911,1936]" pageId="1" pageNumber="132" value="2016-05-23">23.V.2016</date>
</collectingDate>
. 
<collectorName id="AE3F536E166CFF9FE8EFF8EB695698C4" box="[1013,1099,1911,1936]" pageId="1" pageNumber="132">Manuel</collectorName>
&amp; 
<collectorName id="AE3F536E166CFF9FEF77F8EB680098C4" box="[1133,1309,1911,1936]" pageId="1" pageNumber="132">Ramahandrison</collectorName>
leg.&quot;, &quot; 
<collectorName id="AE3F536E166CFF9FEE75F8EB688198C4" box="[1391,1436,1911,1936]" pageId="1" pageNumber="132">Alt.</collectorName>
<elevation id="88E7D18B166CFF9FEB8DF8006DEA98E0" box="[151,247,1948,1972]" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" pageId="1" pageNumber="132" unit="m" value="2140.0">
<quantity id="C4329B5D166CFF9FEB8DF8006DEA98E0" box="[151,247,1948,1972]" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" pageId="1" pageNumber="132" unit="m" value="2140.0">2,140 m</quantity>
</elevation>
. 
<collectorName id="AE3F536E166CFF9FEA1EF8006C5898E1" box="[260,325,1948,1973]" pageId="1" pageNumber="132">Small</collectorName>
stream with many 
<taxonomicName id="C4CA4D3B166CFF9FE939F8006FBB98E0" box="[547,678,1948,1972]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="1" pageNumber="132" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
and peaty substrate, very slowly flowing clear water, in subalpine grassland. 
<collectorName id="AE3F536E166CFF9FEA0BF85C6F20988D" box="[273,573,1984,2009]" pageId="1" pageNumber="132">Andringitra National Park</collectorName>
, eastern slope of massif&quot;
</materialsCitation>
. 
<materialsCitation id="B3A23CE5166CFF9FE877F85C6881988C" ID-GBIF-Occurrence="1563183434" box="[877,1436,1984,2009]" country="Madagascar" location="Fianarantsoa" pageId="1" pageNumber="132" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Fianarantsoa">
<specimenCount id="15CCFD31166CFF9FE877F85C6E8F988D" box="[877,914,1984,2009]" pageId="1" pageNumber="132" type="male">5♂</specimenCount>
, 
<specimenCount id="15CCFD31166CFF9FE8BBF85D6ED8988C" box="[929,965,1985,2008]" pageId="1" pageNumber="132" type="female">7♀</specimenCount>
, &quot; 
<collectingCountry id="7BDD7628166CFF9FE8C6F85C6974988D" box="[988,1129,1984,2009]" name="Madagascar" pageId="1" pageNumber="132">Madagascar</collectingCountry>
. ex-Prov. 
<collectingRegion id="C10EF85A166CFF9FEFC7F85C686D988D" box="[1245,1392,1984,2009]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="132">Fianarantsoa</collectingRegion>
. ca.
</materialsCitation>
</paragraph>
<paragraph id="037536B8166FFF9CEB8DFF046FBE9FAC" blockId="2.[151,1436,151,248]" pageId="2" pageNumber="133">
<materialsCitation id="B3A23CE5166FFF9CEB8DFF046FBE9FAC" ID-GBIF-Occurrence="1563183431" collectingDate="2016-05-23" collectorName="Manuel &amp; Ramahandrison &amp; Alt. &amp; Grassy &amp; Andringitra National Park &amp; All" elevation="1990" latitude="-22.133333" location="Antanifotsy." longLatPrecision="1261" longitude="46.883335" pageId="2" pageNumber="133" specimenCount="1" typeStatus="paratype">
<quantity id="C4329B5D166FFF9CEB8DFF046DF49FE4" box="[151,233,151,176]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="133" unit="km" value="4.3">4.3 km</quantity>
SW 
<location id="06156063166FFF9CEA31FF0B6CA89FE4" LSID="urn:lsid:plazi:treatment:8B6387AE166CFF99EB8DFA886C839C6C:06156063166FFF9CEA31FF0B6CA89FE4" box="[299,437,151,176]" latitude="-22.133333" longLatPrecision="1261" longitude="46.883335" name="Antanifotsy." pageId="2" pageNumber="133">Antanifotsy.</location>
<geoCoordinate id="66FE507F166FFF9CEAA5FF046F049FE4" box="[447,537,152,176]" direction="south" orientation="latitude" pageId="2" pageNumber="133" precision="925" value="-22.133333">S22°08'</geoCoordinate>
<geoCoordinate id="66FE507F166FFF9CE939FF0B6F9C9FE4" box="[547,641,151,176]" direction="east" orientation="longitude" pageId="2" pageNumber="133" precision="925" value="46.883335">E46°53'</geoCoordinate>
. 
<collectingDate id="6730E990166FFF9CE995FF0B6E189FE4" box="[655,773,151,176]" pageId="2" pageNumber="133" value="2016-05-23">
<date id="77741078166FFF9CE995FF0B6E189FE4" box="[655,773,151,176]" pageId="2" pageNumber="133" value="2016-05-23">23.V.2016</date>
</collectingDate>
. 
<collectorName id="AE3F536E166FFF9CE80EFF0B6E749FE4" box="[788,873,151,176]" pageId="2" pageNumber="133">Manuel</collectorName>
&amp; 
<collectorName id="AE3F536E166FFF9CE88DFF0B695A9FE4" box="[919,1095,151,176]" pageId="2" pageNumber="133">Ramahandrison</collectorName>
leg.&quot;, &quot; 
<collectorName id="AE3F536E166FFF9CEFB9FF0B69CD9FE4" box="[1187,1232,151,176]" pageId="2" pageNumber="133">Alt.</collectorName>
<elevation id="88E7D18B166FFF9CEFC1FF0468229FFB" box="[1243,1343,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="2" pageNumber="133" unit="m" value="1990.0">
<quantity id="C4329B5D166FFF9CEFC1FF0468229FFB" box="[1243,1343,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="2" pageNumber="133" unit="m" value="1990.0">1,990 m</quantity>
</elevation>
. 
<collectorName id="AE3F536E166FFF9CEE54FF0468819FE4" box="[1358,1436,152,176]" pageId="2" pageNumber="133">Grassy</collectorName>
brooklet, very shallow, in subalpine grassland. 
<collectorName id="AE3F536E166FFF9CE9D4FF20691F9F81" box="[718,1026,188,213]" pageId="2" pageNumber="133">Andringitra National Park</collectorName>
, eastern slope of the massif&quot;. 
<collectorName id="AE3F536E166FFF9CEE62FF2068819F81" box="[1400,1436,188,213]" pageId="2" pageNumber="133">All</collectorName>
<typeStatus id="DC71881A166FFF9CEB8DFF7C6C1F9FAC" box="[151,258,224,248]" pageId="2" pageNumber="133" type="paratype">paratypes</typeStatus>
with the respective printed red label.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="57B56630166FFF9CEB8DFA8A68509A79" box="[151,1357,1302,1325]" httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="2" pageNumber="133" targetBox="[220,1387,324,1282]" targetPageId="2">
<paragraph id="037536B8166FFF9CEB8DFA8A68509A79" blockId="2.[151,1357,1302,1325]" box="[151,1357,1302,1325]" pageId="2" pageNumber="133">
<emphasis id="31BEEAAA166FFF9CEB8DFA8A6C279A78" bold="true" box="[151,314,1302,1325]" pageId="2" pageNumber="133">FIGURES 1–2.</emphasis>
Habitus of 
<taxonomicName id="C4CA4D3B166FFF9CEAABFA8B6F639A78" box="[433,638,1303,1324]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166FFF9CEAABFA8B6F639A78" box="[433,638,1303,1324]" italics="true" pageId="2" pageNumber="133">Canthyporus reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166FFF9CE99EFA8B6FC89A78" box="[644,725,1303,1324]" pageId="2" pageNumber="133" rank="species">
<emphasis id="31BEEAAA166FFF9CE99EFA8B6FC89A78" bold="true" box="[644,725,1303,1324]" pageId="2" pageNumber="133">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm).
</paragraph>
</caption>
<subSubSection id="4BD06533166FFF9BEBDDFAC8688498C1" lastPageId="5" lastPageNumber="136" pageId="2" pageNumber="133" type="description">
<paragraph id="037536B8166FFF9CEBDDFAC868229A39" blockId="2.[151,1437,1364,2036]" box="[199,1343,1364,1389]" pageId="2" pageNumber="133">
<emphasis id="31BEEAAA166FFF9CEBDDFAC86C4C9A39" bold="true" box="[199,337,1364,1389]" pageId="2" pageNumber="133">Description</emphasis>
. The description refers only to the male 
<typeStatus id="DC71881A166FFF9CE802FAC86E609A39" box="[792,893,1364,1389]" pageId="2" pageNumber="133" type="holotype">holotype</typeStatus>
, until paragraph &quot;aedeagus&quot; inclusively.
</paragraph>
<paragraph id="037536B8166FFF9CEBDDFAE46FC89A8C" blockId="2.[151,1437,1364,2036]" pageId="2" pageNumber="133">
Habitus (
<figureCitation id="9BF12A3D166FFF9CEA35FAE46C699AC5" box="[303,372,1400,1425]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1302,1325]" captionTargetBox="[220,1387,324,1282]" captionTargetId="figure@2.[197,1387,303,1282]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="2" pageNumber="133">Fig. 1</figureCitation>
): Oval-oblong, with sides widely rounded, posteriorly rather strongly attenuated to narrowly rounded apex; maximum width slightly before midlength; lateral outline continuous between pronotum and elytron; dorsal surface strongly and evenly convex.
</paragraph>
<paragraph id="037536B8166FFF9CEBDDFA786C5C9848" blockId="2.[151,1437,1364,2036]" pageId="2" pageNumber="133">
Colour (
<figureCitation id="9BF12A3D166FFF9CEA39FA786CAB9AA9" box="[291,438,1508,1533]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1302,1325]" captionTargetBox="[220,1387,324,1282]" captionTargetId="figure@2.[197,1387,303,1282]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="2" pageNumber="133">Figs. 1 and 2</figureCitation>
): Dorsal surface of head uniformly dark brown except diffusely delimited rufo-testaceous area along clypeal margin. Pronotum rufo-testaceous; disk with diffuse brown central longitudinal marking, extending from shortly behind anterior margin to posterior margin, vaguely and shortly extended transversally along posterior margin. Elytron dark brown; with rather broad diffusely delimited rufo-testaceous area along lateral margin (visible only in lateral view); within pale lateral area with narrow submarginal dark brown stripe in posterior half; near apex diffusely paler except along suture. Venter of head, pronotal and elytral epipleura, propleuron and posterior third of abdominal ventrite VI rufo-testaceous; prosternum, abdominal ventrite I and metacoxal lobes ferrugineous; rest of ventral surface black, vaguely paler along posterior margin of abdominal ventrites III–V.
</paragraph>
<paragraph id="037536B8166FFF9CEBDDF8B46C0F98A0" blockId="2.[151,1437,1364,2036]" pageId="2" pageNumber="133">Head: Anterior margin of clypeus broadly and evenly rounded, without clypeal bead or rim. Clypeal depressions broad and shallow, with small and inconspicuous punctures. Surface shining, with moderately impressed isodiametric reticulation; punctation sparse and extremely fine; without setae. Along inner eye margins with shallow longitudinal depressions and inconspicuous punctation. Antennae testaceous with narrow dark rims around distal extremity of antennomeres V–X; antennomeres V-X slightly longer than broad. Palpi entirely testaceous.</paragraph>
<paragraph id="037536B8166EFF9DEBDDFF046DC29EF9" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="134">
Pronotum: Rather broad; with maximum width at posterior angles. Anterior margin weakly arcuate. Posterior margin bisinuate around centre; laterally weakly curved anteriorad. Lateral margins moderately convergent and evenly arcuate from posterior to anterior angle (
<figureCitation id="9BF12A3D166EFF9DE9ADFF436FE79FAC" box="[695,762,223,248]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1302,1325]" captionTargetBox="[220,1387,324,1282]" captionTargetId="figure@2.[197,1387,303,1282]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="3" pageNumber="134">Fig. 1</figureCitation>
), with narrow but well-defined bead. Anterior angles acute and narrowly rounded; posterior angles rectangular and sharp. Surface shining, with weakly impressed isodiametric reticulation; extremely finely and sparsely punctate; without setae. Close to lateral margins with larger, weakly impressed punctures in shallow depression; along anterior margin with transverse row of small coarse punctures, interrupted in middle; larger but shallower punctures present along posterior margin in outer two thirds.
</paragraph>
<caption id="57B56630166EFF9DEB8DFBCE68819BD3" httpUri="https://zenodo.org/record/801859/files/figure.png" pageId="3" pageNumber="134" targetBox="[192,1402,473,1089]" targetPageId="3">
<paragraph id="037536B8166EFF9DEB8DFBCE68819BD3" blockId="3.[151,1436,1106,1159]" pageId="3" pageNumber="134">
<emphasis id="31BEEAAA166EFF9DEB8DFBCE6C219B3C" bold="true" box="[151,316,1106,1129]" pageId="3" pageNumber="134">FIGURES 3–6.</emphasis>
Male genitalia of 
<taxonomicName id="C4CA4D3B166EFF9DEAE7FBCF6FD09B3C" box="[509,717,1107,1128]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166EFF9DEAE7FBCF6FD09B3C" box="[509,717,1107,1128]" italics="true" pageId="3" pageNumber="134">Canthyporus reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166EFF9DE9CFFBCF6E359B3C" box="[725,808,1107,1128]" pageId="3" pageNumber="134" rank="species">
<emphasis id="31BEEAAA166EFF9DE9CFFBCF6E359B3C" bold="true" box="[725,808,1107,1128]" pageId="3" pageNumber="134">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype). (3) Median lobe of aedeagus in lateral view; (4) median lobe of aedeagus in ventral view; (5) left paramere; (6) right paramere (parameres in external view). Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption id="57B56630166EFF9DEB8DF82D69C098B3" httpUri="https://zenodo.org/record/801861/files/figure.png" pageId="3" pageNumber="134" targetBox="[346,1238,1201,1948]" targetPageId="3">
<paragraph id="037536B8166EFF9DEB8DF82D69C098B3" blockId="3.[151,1436,1969,2023]" pageId="3" pageNumber="134">
<emphasis id="31BEEAAA166EFF9DEB8DF82D6C209893" bold="true" box="[151,317,1969,1992]" pageId="3" pageNumber="134">FIGURES 7–9.</emphasis>
Female reproductive structures of 
<taxonomicName id="C4CA4D3B166EFF9DE9B6F82E6E619893" box="[684,892,1970,1991]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166EFF9DE9B6F82E6E619893" box="[684,892,1970,1991]" italics="true" pageId="3" pageNumber="134">Canthyporus reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166EFF9DE89FF82E6EC59893" box="[901,984,1970,1991]" pageId="3" pageNumber="134" rank="species">
<emphasis id="31BEEAAA166EFF9DE89FF82E6EC59893" bold="true" box="[901,984,1970,1991]" pageId="3" pageNumber="134">sp. nov.</emphasis>
</taxonomicNameLabel>
(paratype from the 
<emphasis id="31BEEAAA166EFF9DEFB7F82E682E989C" box="[1197,1331,1970,1992]" italics="true" pageId="3" pageNumber="134">locus typicus</emphasis>
). (7) Left gonocoxosternum in ventral view; (8) gonocoxae in ventral view; (9) spermathecal tract. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption id="57B566301669FF9AEB8DF9186F7B98FA" httpUri="https://zenodo.org/record/801863/files/figure.png" pageId="4" pageNumber="135" targetBox="[151,1436,193,1647]" targetPageId="4">
<paragraph id="037536B81669FF9AEB8DF9186F7B98FA" blockId="4.[151,1436,1668,1966]" pageId="4" pageNumber="135">
<emphasis id="31BEEAAA1669FF9AEB8DF9186C4E99CE" bold="true" box="[151,339,1668,1691]" pageId="4" pageNumber="135">FIGURES 10–15.</emphasis>
Habitat of 
<taxonomicName id="C4CA4D3B1669FF9AEADFF9196F8F99CE" box="[453,658,1669,1690]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA1669FF9AEADFF9196F8F99CE" box="[453,658,1669,1690]" italics="true" pageId="4" pageNumber="135">Canthyporus reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D11669FF9AE982F9196FF599CE" box="[664,744,1669,1690]" pageId="4" pageNumber="135" rank="species">
<emphasis id="31BEEAAA1669FF9AE982F9196FF599CE" bold="true" box="[664,744,1669,1690]" pageId="4" pageNumber="135">sp. nov.</emphasis>
</taxonomicNameLabel>
(10) Pond with 
<taxonomicName id="C4CA4D3B1669FF9AE897F918693A99CE" box="[909,1063,1668,1690]" class="Magnoliopsida" family="Nymphaeaceae" kingdom="Plantae" order="Nymphaeales" pageId="4" pageNumber="135" phylum="Tracheophyta" rank="family">Nymphaeaceae</taxonomicName>
plants, with peat bog on the margins (
<taxonomicName id="C4CA4D3B1669FF9AEB85F93F6C1499EC" box="[159,265,1699,1720]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="4" pageNumber="135" phylum="Bryophyta" rank="genus">
<emphasis id="31BEEAAA1669FF9AEB85F93F6C1499EC" box="[159,265,1699,1720]" italics="true" pageId="4" pageNumber="135">Sphagnum</emphasis>
</taxonomicName>
spp. and 
<taxonomicName id="C4CA4D3B1669FF9AEA6EF9386F9599EC" box="[372,648,1699,1721]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="135" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis id="31BEEAAA1669FF9AEA6EF9386F9599EC" box="[372,648,1699,1721]" italics="true" pageId="4" pageNumber="135">Drosera madagascariensis</emphasis>
</taxonomicName>
), close to the N7 road between Ambositra and Fianarantsoa (ca. 3.5 km S Ambalamanakana; altitude 1,700 m). This place is the type locality of the new species. The specimens of 
<taxonomicName id="C4CA4D3B1669FF9AEFDFF95E683A9983" box="[1221,1319,1730,1751]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="31BEEAAA1669FF9AEFDFF95E683A9983" box="[1221,1319,1730,1751]" italics="true" pageId="4" pageNumber="135">C. reebae</emphasis>
</taxonomicName>
were taken towards the bottom right corner of the photo, in the grass, close to the end of a small stream with slowly-flowing water (not visible on the photo). (11) Grassy meadow surrounded by forest, in the Andringitra massif (ca 2.5 km SSW Antanifotsy; altitude 1,600 m). The specimens were taken towards the centre-right of the photo, in water slowly seeping through the grass, as shown in the close-up view in (12). The substrate was very muddy. (13) Brooklet with very slowly flowing clear water, with dense grass (Andringitra massif, ca. 2.3 km SW Antanifotsy; altitude 1,580 m). (14) Small slowly-flowing brook in wet grassland above tree line in the Andringitra massif (ca 4.3 km SW Antanifotsy; altitude 1,990 m). (15) Close-up view corresponding to the centre-right area of (14).
</paragraph>
</caption>
<paragraph id="037536B81668FF9BEBDDFF0B6CCE9E31" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">Elytra: Surface shining, with weakly impressed reticulation; on disk meshes slightly elongate in longitudinal direction; with extremely fine and sparse punctation; with few larger weakly impressed punctures near suture and between discal and lateral puncture rows; with very sparse short setae. Discal and lateral puncture rows discernible but rather inconspicuous. Lateral margin of elytron in lateral view almost straight behind shoulder, with narrow bead; anterior part of lateral bead visible in dorsal view. Just before apex with small, shallow depression, but without apico-lateral carina.</paragraph>
<paragraph id="037536B81668FF9BEBDDFEF36C0B9B44" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">
Ventral surface: Entirely very shining except on genae and median region of prosternum. Gula with very obsolete reticulation; genae with strongly impressed isodiametric reticulation. Antero-medial region of prosternum with surface microgranulose and with very short setae. Prosternal column (declivitous part of prosternum between procoxae) moderately ascending posteriad, with surface microgranulose, between procoxae with minute protuberance at level of anterior third of procoxae. Prosternal process weakly inclined; rather broad, oval; in proximal half laterally bordered by narrow bead; surface almost flat, near margins microgranulose, in centre smooth, impunctate; apex broadly rounded, located in impression in metaventrite. Metaventrite and metepisternum with obsolete reticulation, impunctate. Metaventrite without longitudinal lines adjacent to midline. Elytral epipleura with obsolete and irregular reticulation, impunctate; at shoulders as broad as mesotibia distally, narrowing rather abruptly between anterior third and midlength of elytra, then continuing as narrow ridge to apex. Metacoxal plates with weakly impressed longitudinally elongate reticulation, impunctate. Metacoxal lines finely impressed, subparallel (very weakly divergent anteriad) (
<figureCitation id="9BF12A3D1668FF9BE80CFD606E469C40" box="[790,859,764,789]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1302,1325]" captionTargetBox="[220,1387,324,1282]" captionTargetId="figure@2.[197,1387,303,1282]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="5" pageNumber="136">Fig. 2</figureCitation>
); almost reaching posterior margin of metaventrite. Between metacoxal lines, surface with weakly impressed isodiametric reticulation, with few micropunctures; midline strongly impressed. Metacoxal lobes rather broad, rounded and strongly diverging laterally. Abdominal ventrites I-III fused; junction between ventrites I and II visible; junction between ventrites II and III visible except laterally and medially. Surface of ventrites with obsolete reticulation, almost impunctate; on ventrites III-VI with sparse short setae. Reticulation on ventrites I and II longitudinally elongate, on ventrites III-V transversally elongate, on ventrite VI isodiametric except along anterior margin. Ventrite VI with very shallow transverse depression.
</paragraph>
<paragraph id="037536B81668FF9BEBDDFB816F189BBC" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">
Legs (
<figureCitation id="9BF12A3D1668FF9BEA16FB806C4C9B60" box="[268,337,1052,1077]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1302,1325]" captionTargetBox="[220,1387,324,1282]" captionTargetId="figure@2.[197,1387,303,1282]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="5" pageNumber="136">Fig. 2</figureCitation>
): Entirely rufo-testaceous. Metafemur without apical row of short spines; without medio-posterior expansion. Protibia distally moderately broad, with anterior margin very slightly sinuate. Mesotibia triangular, distally very broad, with anterior and posterior margins straight. Pro- and mesotarsi pseudo-tetramerous. Pro- and mesotarsomeres I–III distinctly enlarged, ventrally with few large, flat and circular suction setae. Mesotarsomere V distinctly longer than protarsomere V. Pro- and mesotarsal claws small, very narrow, weakly curved; anterior and posterior claws of all tarsi equal.
</paragraph>
<paragraph id="037536B81668FF9BEBDDFB686E4D9AC8" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">
Aedeagus: Median lobe in lateral view (
<figureCitation id="9BF12A3D1668FF9BE99EFB686FDB9A58" box="[644,710,1268,1293]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1106,1129]" captionTargetBox="[192,1402,473,1089]" captionTargetId="figure@3.[192,1402,473,1089]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Male genitalia of Canthyporus reebae sp. nov. (holotype). (3) Median lobe of aedeagus in lateral view; (4) median lobe of aedeagus in ventral view; (5) left paramere; (6) right paramere (parameres in external view). Scale bar: 0. 2 mm." httpUri="https://zenodo.org/record/801859/files/figure.png" pageId="5" pageNumber="136">Fig. 3</figureCitation>
) distinctly constricted above basal piece; with rather short dorsal process located in apical half. Dorsal process straight, perpendicular to main axis of median lobe. Apical part weakly curved in dorsal direction, rather evenly narrowed to apex; apex with small dorsally-directed hook. Median lobe in ventral view as in 
<figureCitation id="9BF12A3D1668FF9BEAA2FAC36CE09A2C" box="[440,509,1375,1400]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1106,1129]" captionTargetBox="[192,1402,473,1089]" captionTargetId="figure@3.[192,1402,473,1089]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Male genitalia of Canthyporus reebae sp. nov. (holotype). (3) Median lobe of aedeagus in lateral view; (4) median lobe of aedeagus in ventral view; (5) left paramere; (6) right paramere (parameres in external view). Scale bar: 0. 2 mm." httpUri="https://zenodo.org/record/801859/files/figure.png" pageId="5" pageNumber="136">Fig. 4</figureCitation>
. Parameres as in 
<figureCitation id="9BF12A3D1668FF9BE9A5FAC36E379A2C" box="[703,810,1375,1400]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1106,1129]" captionTargetBox="[192,1402,473,1089]" captionTargetId="figure@3.[192,1402,473,1089]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Male genitalia of Canthyporus reebae sp. nov. (holotype). (3) Median lobe of aedeagus in lateral view; (4) median lobe of aedeagus in ventral view; (5) left paramere; (6) right paramere (parameres in external view). Scale bar: 0. 2 mm." httpUri="https://zenodo.org/record/801859/files/figure.png" pageId="5" pageNumber="136">Figs. 5–6</figureCitation>
; apex of parameres without setae; apex of left paramere distinctly more broadly truncated than apex of right paramere.
</paragraph>
<paragraph id="037536B81668FF9BEBDDFA3B6FB89978" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">
Females: As males, except: some females with dorsal reticulation distinctly more impressed than in males; pro- and mesotarsomeres I–III narrower than in males, without suction setae. External genitalia as in 
<figureCitation id="9BF12A3D1668FF9BEE3DFA5068859AB0" box="[1319,1432,1484,1509]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[346,1238,1201,1948]" captionTargetId="figure@3.[346,1238,1201,1948]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Female reproductive structures of Canthyporus reebae sp. nov. (paratype from the locus typicus). (7) Left gonocoxosternum in ventral view; (8) gonocoxae in ventral view; (9) spermathecal tract. Scale bar: 0. 2 mm." httpUri="https://zenodo.org/record/801861/files/figure.png" pageId="5" pageNumber="136">Figs. 7–8</figureCitation>
. Spermathecal tract as in 
<figureCitation id="9BF12A3D1668FF9BEAA9FA736CE5995C" box="[435,504,1519,1544]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,1969,1992]" captionTargetBox="[346,1238,1201,1948]" captionTargetId="figure@3.[346,1238,1201,1948]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Female reproductive structures of Canthyporus reebae sp. nov. (paratype from the locus typicus). (7) Left gonocoxosternum in ventral view; (8) gonocoxae in ventral view; (9) spermathecal tract. Scale bar: 0. 2 mm." httpUri="https://zenodo.org/record/801861/files/figure.png" pageId="5" pageNumber="136">Fig. 9</figureCitation>
; spermatheca long and extended, with complex loop; sclerotisation of tubuliform ductus divided by soft, almost transparent part.
</paragraph>
<paragraph id="037536B81668FF9BEBDDF9A5693899CC" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">
Measurements: 
<typeStatus id="DC71881A1668FF9BEA61F9AB6CF59904" box="[379,488,1591,1616]" pageId="5" pageNumber="136" type="holotype">Holotype</typeStatus>
: total length (TL) = 
<quantity id="C4329B5D1668FF9BE9CEF9A46E339904" box="[724,814,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="136" unit="mm" value="2.2">2.2 mm</quantity>
, total length without head = 
<quantity id="C4329B5D1668FF9BEF60F9A469C99904" box="[1146,1236,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="136" unit="mm" value="1.9">1.9 mm</quantity>
, maximum width (MW) = 
<quantity id="C4329B5D1668FF9BEBE1F9C06C7C9920" box="[251,353,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="136" unit="mm" value="1.15">1.15 mm</quantity>
, ratio TL/MW = 1.93. 
<typeStatus id="DC71881A1668FF9BE97BF9C16FCE9921" box="[609,723,1629,1653]" pageId="5" pageNumber="136" type="paratype">Paratypes</typeStatus>
: TL = 
<quantity id="C4329B5D1668FF9BE807F9C06EB99920" box="[797,932,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="5" pageNumber="136" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1–2.3 mm</quantity>
(2.2 ± 0.05), TL without head = 1.8–2.0 mm (1.9 ± 0.05), MW = 
<quantity id="C4329B5D1668FF9BEA60F91C6F1C99CC" box="[378,513,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="5" pageNumber="136" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
(1.15 ± 0.05), TL/MW = 1.85–2.0 (1.95 ± 0.05).
</paragraph>
<paragraph id="037536B81668FF9BEBDDF938688498C1" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="136">
Variation: 
<typeStatus id="DC71881A1668FF9BEA5AF9396CB099E9" box="[320,429,1701,1725]" pageId="5" pageNumber="136" type="paratype">Paratypes</typeStatus>
vary slightly in habitus, some specimens being somewhat narrower or wider than the 
<typeStatus id="DC71881A1668FF9BEB8DF95B6DE499B4" box="[151,249,1735,1760]" pageId="5" pageNumber="136" type="holotype">holotype</typeStatus>
and some being posteriorly a bit more strongly attenuated. All specimens have the pronotum much lighter than the head and elytra and with a conspicuous dark central marking, the latter of slightly variable extension and colour intensity. The extension of the pale area near elytral apex is also slightly variable. The discal puncture row on elytra is always rather inconspicuous but its punctures vary somewhat in size and depth among specimens. In all females from the 
<typeStatus id="DC71881A1668FF9BEA7BF8C56C8C9825" box="[353,401,1881,1905]" pageId="5" pageNumber="136">type</typeStatus>
locality and in some females collected in the Andringitra massif, the dorsal reticulation is distinctly more impressed than in males; however, in some females from Andringitra it is more or less as in males.
</paragraph>
</subSubSection>
<subSubSection id="4BD065331668FF98EBDDF83C6CDE9F81" lastPageId="6" lastPageNumber="137" pageId="5" pageNumber="136" type="diagnosis">
<paragraph id="037536B81668FF98EBDDF83C6CDE9F81" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,429]" lastPageId="6" lastPageNumber="137" pageId="5" pageNumber="136">
<emphasis id="31BEEAAA1668FF9BEBDDF83C6CDB98ED" bold="true" box="[199,454,1952,1977]" pageId="5" pageNumber="136">Differential diagnosis</emphasis>
. Within the genus 
<emphasis id="31BEEAAA1668FF9BE987F83C6EB598EC" box="[669,936,1952,1976]" italics="true" pageId="5" pageNumber="136">
<taxonomicName id="C4CA4D3B1668FF9BE987F83C6E2C98EC" box="[669,817,1952,1976]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">Canthyporus</taxonomicName>
, 
<taxonomicName id="C4CA4D3B1668FF9BE826F83D6EB598EC" box="[828,936,1952,1976]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">C. reebae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="2A8D57D11668FF9BE8AAF83C691698ED" box="[944,1035,1952,1977]" pageId="5" pageNumber="136" rank="species">
<emphasis id="31BEEAAA1668FF9BE8AAF83C691698ED" bold="true" box="[944,1035,1952,1977]" pageId="5" pageNumber="136">sp. nov.</emphasis>
</taxonomicNameLabel>
is a member of the 
<taxonomicName id="C4CA4D3B1668FF9BEFF7F83D688F98EC" box="[1261,1426,1952,1976]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="hottentottus">
<emphasis id="31BEEAAA1668FF9BEFF7F83D688F98EC" box="[1261,1426,1952,1976]" italics="true" pageId="5" pageNumber="136">C. hottentottus</emphasis>
</taxonomicName>
- group. Species of this group are characterised (in males) by the median lobe with a dorsal process and the parameres without apical incision and apical setae, and (in females) by an extended spermathecal tract with a loop (
<bibRefCitation id="675B4B49166BFF98EB84FF206CA59F81" author="Bistrom" box="[158,440,188,213]" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4BD06533166BFF98EBDDFF7C69C49EA0" pageId="6" pageNumber="137" type="reference_group">
<paragraph id="037536B8166BFF98EBDDFF7C6E3D9EF8" blockId="6.[151,1437,151,429]" pageId="6" pageNumber="137">
<taxonomicName id="C4CA4D3B166BFF98EBDDFF7C6CA19FAC" ID-CoL="QPJT" box="[199,444,224,248]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EBDDFF7C6CA19FAC" box="[199,444,224,248]" italics="true" pageId="6" pageNumber="137">Canthyporus pauliani</emphasis>
</taxonomicName>
, the only species of the genus 
<taxonomicName id="C4CA4D3B166BFF98E80DFF7C6EB59FAC" ID-CoL="3GWP" box="[791,936,224,248]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="31BEEAAA166BFF98E80DFF7C6EB59FAC" box="[791,936,224,248]" italics="true" pageId="6" pageNumber="137">Canthyporus</emphasis>
</taxonomicName>
known from Madagascar prior to this study, also belongs to the 
<taxonomicName id="C4CA4D3B166BFF98EA61FE986F3F9E48" ID-CoL="QPJG" box="[379,546,260,284]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="hottentottus">
<emphasis id="31BEEAAA166BFF98EA61FE986F3F9E48" box="[379,546,260,284]" italics="true" pageId="6" pageNumber="137">C. hottentottus</emphasis>
</taxonomicName>
-group. Detailed descriptions of this species, with illustrations of habitus and male genitalia, can be found in 
<bibRefCitation id="675B4B49166BFF98EAECFEBB6FBF9E14" author="Guignot" box="[502,674,295,320]" pageId="6" pageNumber="137" refString="Guignot, F. (1951) Description de deux dytiscides malgaches nouveaux (Coleopteres). Le Naturaliste Malgache, 3, 125 - 128." type="journal article" year="1951">Guignot (1951)</bibRefCitation>
and in 
<bibRefCitation id="675B4B49166BFF98E9EFFEBB69079E14" author="Bistrom" box="[757,1050,295,320]" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson (2006)</bibRefCitation>
(the median lobe in lateral view is also reproduced in 
<bibRefCitation id="675B4B49166BFF98EA74FED06F109E31" author="Guignot" box="[366,525,332,357]" pageId="6" pageNumber="137" refString="Guignot, F. (1959) Revision des hydrocanthares d'Afrique (Coleoptera Dytiscoidea). 2. Annales du Musee Royal du Congo Belge Serie 8 ° (Sciences Zoologiques), 78, 323 - 648." type="journal article" year="1959">Guignot 1959</bibRefCitation>
). In addition, the senior author (MM) has recently studied the type material of 
<taxonomicName id="C4CA4D3B166BFF98EE98FED06DE99EDC" ID-CoL="QPJT" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EE98FED06DE99EDC" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
(one male and one female syntypes; MNHN; specimen data in 
<bibRefCitation id="675B4B49166BFF98E8C2FEF369EA9EDC" author="Bistrom" box="[984,1271,367,392]" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson 2006</bibRefCitation>
). 
<taxonomicName id="C4CA4D3B166BFF98EE11FEEC6DFF9EF8" ID-CoL="8LZ4P" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166BFF98EE11FEEC6DFF9EF8" italics="true" pageId="6" pageNumber="137">Canthyporus reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166BFF98EBF3FE086C5E9EF9" box="[233,323,404,429]" pageId="6" pageNumber="137" rank="species">
<emphasis id="31BEEAAA166BFF98EBF3FE086C5E9EF9" bold="true" box="[233,323,404,429]" pageId="6" pageNumber="137">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="C4CA4D3B166BFF98EACCFE086F499EF8" ID-CoL="QPJT" box="[470,596,404,428]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EACCFE086F499EF8" box="[470,596,404,428]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
in many respects:
</paragraph>
<paragraph id="037536B8166BFF98EB8DFE4169C49EA0" blockId="6.[151,1436,476,1184]" box="[151,1241,476,501]" pageId="6" pageNumber="137">
- size larger (according to 
<bibRefCitation id="675B4B49166BFF98EAC6FE406FF29EA1" author="Bistrom" box="[476,751,476,501]" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson 2006</bibRefCitation>
body length of 
<taxonomicName id="C4CA4D3B166BFF98E8BAFE4069039EA0" ID-CoL="QPJT" box="[928,1054,476,500]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98E8BAFE4069039EA0" box="[928,1054,476,500]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
<quantity id="C4329B5D166BFF98EF3FFE4069D09EA0" box="[1061,1229,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" metricValueMax="1.94" metricValueMin="1.92" pageId="6" pageNumber="137" unit="mm" value="1.93" valueMax="1.94" valueMin="1.92">1.92–1.94 mm</quantity>
);
</paragraph>
</subSubSection>
<subSubSection id="4BD06533166BFF99EB8DFD9C6F519DD0" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137" type="description">
<paragraph id="037536B8166BFF98EB8DFD9C6F4E9D34" blockId="6.[151,1436,476,1184]" pageId="6" pageNumber="137">
- habitus wider, with the sides more strongly rounded and the apex more narrowly rounded, with the maximum width more anterior and the dorsal surface more strongly and evenly convex (
<taxonomicName id="C4CA4D3B166BFF98EF43FDB869C79D68" box="[1113,1242,548,572]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EF43FDB869C79D68" box="[1113,1242,548,572]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
with body sides subparallel and dorsum rather flat);
</paragraph>
<paragraph id="037536B8166BFF98EB8DFDF1691B9DFC" blockId="6.[151,1436,476,1184]" pageId="6" pageNumber="137">
- colour much lighter on pronotum (in 
<taxonomicName id="C4CA4D3B166BFF98E971FDF06FF49DD0" box="[619,745,620,644]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98E971FDF06FF49DD0" box="[619,745,620,644]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
, pronotum entirely dark brown except rufo-testaceous lateral margins), and pale areas on elytral sides and apical region more extended;
</paragraph>
<paragraph id="037536B8166BFF98EB8DFD296EC89D98" blockId="6.[151,1436,476,1184]" box="[151,981,692,717]" pageId="6" pageNumber="137">
- clypeal margin more rounded (subtruncated in middle in 
<taxonomicName id="C4CA4D3B166BFF98E85FFD286EDE9D98" box="[837,963,692,716]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98E85FFD286EDE9D98" box="[837,963,692,716]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="037536B8166BFF98EB8DFD446ED49DA4" blockId="6.[151,1436,476,1184]" box="[151,969,727,752]" pageId="6" pageNumber="137">
- antennomeres VI-XI much lighter (entirely darkened in 
<taxonomicName id="C4CA4D3B166BFF98E820FD446EA59DA4" box="[826,952,728,752]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98E820FD446EA59DA4" box="[826,952,728,752]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="037536B8166BFF98EB8DFD616F0A9C6C" blockId="6.[151,1436,476,1184]" pageId="6" pageNumber="137">
- pronotum sides less strongly arcuate; lateral outline continuous between pronotum and elytron (distinctly discontinuous in 
<taxonomicName id="C4CA4D3B166BFF98EA92FCBC6F1B9C6C" box="[392,518,800,824]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EA92FCBC6F1B9C6C" box="[392,518,800,824]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="037536B8166BFF98EB8DFCD96FE19CD4" blockId="6.[151,1436,476,1184]" pageId="6" pageNumber="137">- on head, pronotum and elytra, surface more shiny, reticulation much less impressed and punctation finer; discal puncture row on elytra much more inconspicuous;</paragraph>
<paragraph id="037536B8166BFF98EB8DFC11686D9CF0" blockId="6.[151,1436,476,1184]" box="[151,1392,908,933]" pageId="6" pageNumber="137">
- colour of ventral surface darker (areas that are black in 
<taxonomicName id="C4CA4D3B166BFF98E829FC106E829CF0" box="[819,927,908,932]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166BFF98E829FC106E829CF0" box="[819,927,908,932]" italics="true" pageId="6" pageNumber="137">C. reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166BFF98E8BCFC106EE29CF1" box="[934,1023,908,933]" pageId="6" pageNumber="137" rank="species">
<emphasis id="31BEEAAA166BFF98E8BCFC106EE29CF1" bold="true" box="[934,1023,908,933]" pageId="6" pageNumber="137">sp. nov.</emphasis>
</taxonomicNameLabel>
are ferrugineous in 
<taxonomicName id="C4CA4D3B166BFF98EFFBFC1068429CF0" box="[1249,1375,908,932]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EFFBFC1068429CF0" box="[1249,1375,908,932]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="037536B8166BFF98EB8DFC2C6ED39C9C" blockId="6.[151,1436,476,1184]" box="[151,974,943,968]" pageId="6" pageNumber="137">- ventral surface more shiny, with reticulation more weakly impressed;</paragraph>
<paragraph id="037536B8166BFF98EB8DFC496CEB9CB9" blockId="6.[151,1436,476,1184]" box="[151,502,980,1005]" pageId="6" pageNumber="137">- prosternal process broader;</paragraph>
<paragraph id="037536B8166BFF98EB8DFC646FFC9B44" blockId="6.[151,1436,476,1184]" box="[151,737,1015,1040]" pageId="6" pageNumber="137">- metacoxal lines slightly more weakly divergent;</paragraph>
<paragraph id="037536B8166BFF98EB8DFB816FB39B61" blockId="6.[151,1436,476,1184]" box="[151,686,1052,1077]" pageId="6" pageNumber="137">- pro- and mesotarsi less strongly broadened;</paragraph>
<paragraph id="037536B8166BFF98EB8DFBDC6EF39B0C" blockId="6.[151,1436,476,1184]" box="[151,1006,1087,1112]" pageId="6" pageNumber="137">
- protarsal claws narrower and straighter (strongly curved in 
<taxonomicName id="C4CA4D3B166BFF98E845FBDC6EC19B0C" box="[863,988,1088,1112]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98E845FBDC6EC19B0C" box="[863,988,1088,1112]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="037536B8166BFF98EB8DFBF96FDD9BF4" blockId="6.[151,1436,476,1184]" pageId="6" pageNumber="137">
- different shape of the median lobe of aedeagus (see 
<figureCitation id="9BF12A3D166BFF98E808FBF86E499B28" box="[786,852,1124,1149]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1302,1325]" captionTargetBox="[220,1387,324,1282]" captionTargetId="figure@2.[197,1387,303,1282]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Canthyporus reebae sp. nov. (holotype). (1) Dorsal side; (2) ventral side (scale bar: 1 mm)." httpUri="https://zenodo.org/record/801857/files/figure.png" pageId="6" pageNumber="137">
Fig. 
<quantity id="C4329B5D166BFF98E85CFBF86E499B28" box="[838,852,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="137" unit="in" value="2.0">2</quantity>
</figureCitation>
in 
<bibRefCitation id="675B4B49166BFF98E863FBF8690B9B29" author="Guignot" box="[889,1046,1124,1149]" pageId="6" pageNumber="137" refString="Guignot, F. (1951) Description de deux dytiscides malgaches nouveaux (Coleopteres). Le Naturaliste Malgache, 3, 125 - 128." type="journal article" year="1951">Guignot 1951</bibRefCitation>
, and Figs. 131 and 
<quantity id="C4329B5D166BFF98EFE9FBF868279B29" box="[1267,1338,1124,1149]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" pageId="6" pageNumber="137" unit="in" value="132.0">132 in</quantity>
<bibRefCitation id="675B4B49166BFF98EE58FBF86C639BF4" author="Bistrom" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson 2006</bibRefCitation>
), particularly in lateral view.
</paragraph>
<paragraph id="037536B8166BFF98EBDDFB536C469819" blockId="6.[151,1437,1231,2013]" pageId="6" pageNumber="137">
Despite these numerous morphological differences, 
<taxonomicName id="C4CA4D3B166BFF98E83AFB4C6E8D9BBC" box="[800,912,1232,1256]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166BFF98E83AFB4C6E8D9BBC" box="[800,912,1232,1256]" italics="true" pageId="6" pageNumber="137">C. reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166BFF98E881FB536EE59BBC" box="[923,1016,1231,1256]" pageId="6" pageNumber="137" rank="species">
<emphasis id="31BEEAAA166BFF98E881FB536EE59BBC" bold="true" box="[923,1016,1231,1256]" pageId="6" pageNumber="137">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="C4CA4D3B166BFF98EF2CFB4C69AA9BBC" box="[1078,1207,1232,1256]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="31BEEAAA166BFF98EF2CFB4C69AA9BBC" box="[1078,1207,1232,1256]" italics="true" pageId="6" pageNumber="137">C. pauliani</emphasis>
</taxonomicName>
have in common a dorsal process of the median lobe which in lateral view is straight, rather thin, and perpendicular with respect to the main part of the median lobe. Within the 
<taxonomicName id="C4CA4D3B166BFF98E965FA846E359A64" box="[639,808,1304,1328]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="hottentottus">
<emphasis id="31BEEAAA166BFF98E965FA846E359A64" box="[639,808,1304,1328]" italics="true" pageId="6" pageNumber="137">C. hottentottus</emphasis>
</taxonomicName>
-group, according to the figures in 
<bibRefCitation id="675B4B49166BFF98EFD8FA8B6DF99A01" author="Bistrom" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson (2006)</bibRefCitation>
, only three additional species share this particular configuration of the median lobe: 
<taxonomicName id="C4CA4D3B166BFF98EFB7FAA06DD29A2C" authority="Guignot, 1936" authorityName="Guignot" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="alpestris">
<emphasis id="31BEEAAA166BFF98EFB7FAA0682D9A00" box="[1197,1328,1340,1364]" italics="true" pageId="6" pageNumber="137">C. alpestris</emphasis>
Guignot, 1936
</taxonomicName>
(from Kilimanjaro, Tanzania), 
<taxonomicName id="C4CA4D3B166BFF98E92EFAFC6E7C9A2C" authority="Guignot, 1951" authorityName="Guignot" authorityYear="1951" box="[564,865,1375,1400]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="petulans">
<emphasis id="31BEEAAA166BFF98E92EFAFC6FA89A2C" box="[564,693,1376,1400]" italics="true" pageId="6" pageNumber="137">C. petulans</emphasis>
<bibRefCitation id="675B4B49166BFF98E9A4FAC36E7C9A2C" author="Guignot" box="[702,865,1375,1400]" pageId="6" pageNumber="137" refString="Guignot, F. (1951) Description de deux dytiscides malgaches nouveaux (Coleopteres). Le Naturaliste Malgache, 3, 125 - 128." type="journal article" year="1951">Guignot, 1951</bibRefCitation>
</taxonomicName>
(from South Africa and Lesotho), and 
<taxonomicName id="C4CA4D3B166BFF98EE3BFAFC6C5A9AC8" authority="Wewalka, 1981" authorityName="Wewalka" authorityYear="1981" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="31BEEAAA166BFF98EE3BFAFC68819A2C" box="[1313,1436,1376,1400]" italics="true" pageId="6" pageNumber="137">C. loeffleri</emphasis>
Wewalka, 1981
</taxonomicName>
(from Ethiopia). According to published descriptions, 
<taxonomicName id="C4CA4D3B166BFF98E8D9FA1869229AC8" box="[963,1087,1412,1436]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="31BEEAAA166BFF98E8D9FA1869229AC8" box="[963,1087,1412,1436]" italics="true" pageId="6" pageNumber="137">C. loeffleri</emphasis>
</taxonomicName>
has many similarities with 
<taxonomicName id="C4CA4D3B166BFF98EE98FA186DFF9A94" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166BFF98EE98FA186DFF9A94" italics="true" pageId="6" pageNumber="137">C. reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166BFF98EBF0FA3B6C599A94" box="[234,324,1447,1472]" pageId="6" pageNumber="137" rank="species">
<emphasis id="31BEEAAA166BFF98EBF0FA3B6C599A94" bold="true" box="[234,324,1447,1472]" pageId="6" pageNumber="137">sp. nov.</emphasis>
</taxonomicNameLabel>
(notably in size, habitus, colour, reticulation and punctation), but the two species differ at least by the colour of the pronotum (black in 
<taxonomicName id="C4CA4D3B166BFF98E935FA506FB59AB0" box="[559,680,1484,1508]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="31BEEAAA166BFF98E935FA506FB59AB0" box="[559,680,1484,1508]" italics="true" pageId="6" pageNumber="137">C. loeffleri</emphasis>
</taxonomicName>
) and antennomeres V-XI (brownish in 
<taxonomicName id="C4CA4D3B166BFF98EF40FA5069CE9AB0" box="[1114,1235,1484,1508]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="31BEEAAA166BFF98EF40FA5069CE9AB0" box="[1114,1235,1484,1508]" italics="true" pageId="6" pageNumber="137">C. loeffleri</emphasis>
</taxonomicName>
) and by the shape of the median lobe of aedeagus. Type material of 
<taxonomicName id="C4CA4D3B166BFF98E9D8FA6C6E5E995C" box="[706,835,1520,1544]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="alpestris">
<emphasis id="31BEEAAA166BFF98E9D8FA6C6E5E995C" box="[706,835,1520,1544]" italics="true" pageId="6" pageNumber="137">C. alpestris</emphasis>
</taxonomicName>
(the male holotype and the single male paratype) and 
<taxonomicName id="C4CA4D3B166BFF98EB8DF9886CDD9979" authority="Guignot, 1951" authorityName="Guignot" authorityYear="1951" box="[151,448,1556,1581]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="petulans">
<emphasis id="31BEEAAA166BFF98EB8DF9886C0A9978" box="[151,279,1556,1580]" italics="true" pageId="6" pageNumber="137">C. petulans</emphasis>
<bibRefCitation id="675B4B49166BFF98EA04F9886CDD9979" author="Guignot" box="[286,448,1556,1581]" pageId="6" pageNumber="137" refString="Guignot, F. (1951) Description de deux dytiscides malgaches nouveaux (Coleopteres). Le Naturaliste Malgache, 3, 125 - 128." type="journal article" year="1951">Guignot, 1951</bibRefCitation>
</taxonomicName>
(two paratypes, one male and one female), and several non-type specimens of the latter species, have been studied by the senior author (MNHN; specimen data in 
<bibRefCitation id="675B4B49166BFF98E8FAF9AB69E49904" author="Bistrom" box="[992,1273,1591,1616]" pageId="6" pageNumber="137" refString="Bistrom, O. &amp; Nilsson, A. N. (2006) Taxonomic revision of the Ethiopian genus Canthyporus (Coleoptera Dytiscidae). Memorie della Societa Entomologica Italiana, 85, 209 - 306." type="journal article" year="2006">Biström &amp; Nilsson 2006</bibRefCitation>
). 
<taxonomicName id="C4CA4D3B166BFF98EE11F9A46DE59920" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="alpestris">
<emphasis id="31BEEAAA166BFF98EE11F9A46DE59920" italics="true" pageId="6" pageNumber="137">Canthyporus alpestris</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="C4CA4D3B166BFF98EA94F9C06CE69920" box="[398,507,1628,1652]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166BFF98EA94F9C06CE69920" box="[398,507,1628,1652]" italics="true" pageId="6" pageNumber="137">C. reebae</emphasis>
</taxonomicName>
<emphasis id="31BEEAAA166BFF98E91EF9C06F439921" bold="true" box="[516,606,1628,1653]" pageId="6" pageNumber="137">
<taxonomicNameLabel id="2A8D57D1166BFF98E91EF9C06F439921" box="[516,606,1628,1653]" pageId="6" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mainly by: antennomeres V-X darker and slightly more elongated; much more strongly impressed dorsal reticulation; slightly larger punctures on pronotum and elytra; more strongly impressed discal and lateral puncture rows; lateral margins of pronotum less strongly converging; ventral surface paler; and shape of the median lobe. 
<taxonomicName id="C4CA4D3B166BFF98E928F9546E3599B4" box="[562,808,1736,1760]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="petulans">
<emphasis id="31BEEAAA166BFF98E928F9546E3599B4" box="[562,808,1736,1760]" italics="true" pageId="6" pageNumber="137">Canthyporus petulans</emphasis>
</taxonomicName>
is similar in habitus to the species described here, but is otherwise very distinct (much larger, with central marking of pronotum more contrasted, with elytra paler, with dorsal reticulation more impressed and dorsal and ventral punctures much larger and coarser, and different shape of the median lobe).
</paragraph>
<paragraph id="037536B8166BFF99EBDDF8CB6EEE9D69" blockId="6.[151,1437,1231,2013]" lastBlockId="7.[151,1437,151,824]" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<emphasis id="31BEEAAA166BFF98EBDDF8CB6C3E9824" bold="true" box="[199,291,1879,1904]" pageId="6" pageNumber="137">Habitat</emphasis>
. All specimens of the 
<typeStatus id="DC71881A166BFF98E904F8C46F539824" box="[542,590,1880,1904]" pageId="6" pageNumber="137">type</typeStatus>
series were found in 
<date id="77741078166BFF98E827F8C46EAF9824" box="[829,946,1879,1904]" pageId="6" pageNumber="137" value="2016-05">May 2016</date>
, in two distinct areas: the south-eastern part of the Itremo mountain range (a single collecting site which is the 
<typeStatus id="DC71881A166BFF98E89BF8E16EAC98C1" box="[897,945,1917,1941]" pageId="6" pageNumber="137">type</typeStatus>
locality of the new species) and the eastern slope of the Andringitra massif (four collecting sites). The altitude range of the five collecting sites was 
<quantity id="C4329B5D166BFF98EE31F83C689A98EC" box="[1323,1415,1952,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="6" pageNumber="137" unit="m" value="1580.0">1,580 m</quantity>
– 
<quantity id="C4329B5D166BFF98EB8DF8586DEA9888" box="[151,247,1988,2012]" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" pageId="6" pageNumber="137" unit="m" value="2140.0">2,140 m</quantity>
. The 
<typeStatus id="DC71881A166BFF98EA2CF8596C7B9889" box="[310,358,1989,2013]" pageId="6" pageNumber="137">type</typeStatus>
locality is a large pond with dense vegetation of 
<taxonomicName id="C4CA4D3B166BFF98E88FF858695D9889" box="[917,1088,1988,2013]" class="Magnoliopsida" family="Nymphaeaceae" kingdom="Plantae" order="Nymphaeales" pageId="6" pageNumber="137" phylum="Tracheophyta" rank="family">Nymphaeaceae</taxonomicName>
and helophytes, bordered by a peat bog with 
<taxonomicName id="C4CA4D3B166AFF99EA22FF046CB39FE4" box="[312,430,152,176]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="7" pageNumber="138" phylum="Bryophyta" rank="genus">
<emphasis id="31BEEAAA166AFF99EA22FF046CB39FE4" box="[312,430,152,176]" italics="true" pageId="7" pageNumber="138">Sphagnum</emphasis>
</taxonomicName>
(indicative of high acidity) and 
<taxonomicName id="C4CA4D3B166AFF99E806FF0569549FE4" box="[796,1097,152,176]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="138" phylum="Tracheophyta" rank="species" species="madagascariensis">
<emphasis id="31BEEAAA166AFF99E806FF0569549FE4" box="[796,1097,152,176]" italics="true" pageId="7" pageNumber="138">Drosera madagascariensis</emphasis>
</taxonomicName>
(
<figureCitation id="9BF12A3D166AFF99EF40FF0B69B39FE4" box="[1114,1198,151,176]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@4.[151,1436,193,1647]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Habitat of Canthyporus reebae sp. nov. (10) Pond with Nymphaeaceae plants, with peat bog on the margins (Sphagnum spp. and Drosera madagascariensis), close to the N 7 road between Ambositra and Fianarantsoa (ca. 3. 5 km S Ambalamanakana; altitude 1,700 m). This place is the type locality of the new species. The specimens of C. reebae were taken towards the bottom right corner of the photo, in the grass, close to the end of a small stream with slowly-flowing water (not visible on the photo). (11) Grassy meadow surrounded by forest, in the Andringitra massif (ca 2. 5 km SSW Antanifotsy; altitude 1,600 m). The specimens were taken towards the centre-right of the photo, in water slowly seeping through the grass, as shown in the close-up view in (12). The substrate was very muddy. (13) Brooklet with very slowly flowing clear water, with dense grass (Andringitra massif, ca. 2. 3 km SW Antanifotsy; altitude 1,580 m). (14) Small slowly-flowing brook in wet grassland above tree line in the Andringitra massif (ca 4. 3 km SW Antanifotsy; altitude 1,990 m). (15) Close-up view corresponding to the centre-right area of (14)." httpUri="https://zenodo.org/record/801863/files/figure.png" pageId="7" pageNumber="138">Fig. 10</figureCitation>
). Here, specimens of 
<taxonomicName id="C4CA4D3B166AFF99EB8DFF206C199F80" box="[151,260,188,212]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166AFF99EB8DFF206C199F80" box="[151,260,188,212]" italics="true" pageId="7" pageNumber="138">C. reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166AFF99EA16FF206C759F81" box="[268,360,188,213]" pageId="7" pageNumber="138" rank="species">
<emphasis id="31BEEAAA166AFF99EA16FF206C759F81" bold="true" box="[268,360,188,213]" pageId="7" pageNumber="138">sp. nov.</emphasis>
</taxonomicNameLabel>
were taken at small depth, not everywhere along margins but close to a small stream that was ending into the pond (not visible on 
<figureCitation id="9BF12A3D166AFF99E934FF436F9D9FAC" box="[558,640,223,248]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@4.[151,1436,193,1647]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Habitat of Canthyporus reebae sp. nov. (10) Pond with Nymphaeaceae plants, with peat bog on the margins (Sphagnum spp. and Drosera madagascariensis), close to the N 7 road between Ambositra and Fianarantsoa (ca. 3. 5 km S Ambalamanakana; altitude 1,700 m). This place is the type locality of the new species. The specimens of C. reebae were taken towards the bottom right corner of the photo, in the grass, close to the end of a small stream with slowly-flowing water (not visible on the photo). (11) Grassy meadow surrounded by forest, in the Andringitra massif (ca 2. 5 km SSW Antanifotsy; altitude 1,600 m). The specimens were taken towards the centre-right of the photo, in water slowly seeping through the grass, as shown in the close-up view in (12). The substrate was very muddy. (13) Brooklet with very slowly flowing clear water, with dense grass (Andringitra massif, ca. 2. 3 km SW Antanifotsy; altitude 1,580 m). (14) Small slowly-flowing brook in wet grassland above tree line in the Andringitra massif (ca 4. 3 km SW Antanifotsy; altitude 1,990 m). (15) Close-up view corresponding to the centre-right area of (14)." httpUri="https://zenodo.org/record/801863/files/figure.png" pageId="7" pageNumber="138">Fig. 10</figureCitation>
). In Andringitra, specimens were collected below tree line at two sites: seepage in meadow surrounded by forest (
<figureCitation id="9BF12A3D166AFF99E962FE986E049E48" box="[632,793,260,285]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@4.[151,1436,193,1647]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Habitat of Canthyporus reebae sp. nov. (10) Pond with Nymphaeaceae plants, with peat bog on the margins (Sphagnum spp. and Drosera madagascariensis), close to the N 7 road between Ambositra and Fianarantsoa (ca. 3. 5 km S Ambalamanakana; altitude 1,700 m). This place is the type locality of the new species. The specimens of C. reebae were taken towards the bottom right corner of the photo, in the grass, close to the end of a small stream with slowly-flowing water (not visible on the photo). (11) Grassy meadow surrounded by forest, in the Andringitra massif (ca 2. 5 km SSW Antanifotsy; altitude 1,600 m). The specimens were taken towards the centre-right of the photo, in water slowly seeping through the grass, as shown in the close-up view in (12). The substrate was very muddy. (13) Brooklet with very slowly flowing clear water, with dense grass (Andringitra massif, ca. 2. 3 km SW Antanifotsy; altitude 1,580 m). (14) Small slowly-flowing brook in wet grassland above tree line in the Andringitra massif (ca 4. 3 km SW Antanifotsy; altitude 1,990 m). (15) Close-up view corresponding to the centre-right area of (14)." httpUri="https://zenodo.org/record/801863/files/figure.png" pageId="7" pageNumber="138">Figs. 11 &amp; 12</figureCitation>
); small, grassy slowly-flowing ditch, exposed to sun but close to forest edge (
<figureCitation id="9BF12A3D166AFF99EA9FFEBB6CC79E14" box="[389,474,295,320]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@4.[151,1436,193,1647]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Habitat of Canthyporus reebae sp. nov. (10) Pond with Nymphaeaceae plants, with peat bog on the margins (Sphagnum spp. and Drosera madagascariensis), close to the N 7 road between Ambositra and Fianarantsoa (ca. 3. 5 km S Ambalamanakana; altitude 1,700 m). This place is the type locality of the new species. The specimens of C. reebae were taken towards the bottom right corner of the photo, in the grass, close to the end of a small stream with slowly-flowing water (not visible on the photo). (11) Grassy meadow surrounded by forest, in the Andringitra massif (ca 2. 5 km SSW Antanifotsy; altitude 1,600 m). The specimens were taken towards the centre-right of the photo, in water slowly seeping through the grass, as shown in the close-up view in (12). The substrate was very muddy. (13) Brooklet with very slowly flowing clear water, with dense grass (Andringitra massif, ca. 2. 3 km SW Antanifotsy; altitude 1,580 m). (14) Small slowly-flowing brook in wet grassland above tree line in the Andringitra massif (ca 4. 3 km SW Antanifotsy; altitude 1,990 m). (15) Close-up view corresponding to the centre-right area of (14)." httpUri="https://zenodo.org/record/801863/files/figure.png" pageId="7" pageNumber="138">Fig. 13</figureCitation>
). The species was also found above tree line, in two very slowly-flowing streams in subalpine wet grasslands. The first one was about 
<quantity id="C4329B5D166AFF99E9EAFED06E279E30" box="[752,826,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="138" unit="cm" value="50.0">50 cm</quantity>
deep, with peaty substratum and many 
<taxonomicName id="C4CA4D3B166AFF99EE00FED068809E30" box="[1306,1437,332,356]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="138" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
helophytes (locality not illustrated). In the second one, 
<taxonomicName id="C4CA4D3B166AFF99E815FEEC6E619EDC" box="[783,892,368,392]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="31BEEAAA166AFF99E815FEEC6E619EDC" box="[783,892,368,392]" italics="true" pageId="7" pageNumber="138">C. reebae</emphasis>
</taxonomicName>
specimens were more abundant; it was a much smaller and shallower stream with dense short grass (
<figureCitation id="9BF12A3D166AFF99E9EFFE086E8B9EF9" box="[757,918,404,429]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@4.[151,1436,193,1647]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Habitat of Canthyporus reebae sp. nov. (10) Pond with Nymphaeaceae plants, with peat bog on the margins (Sphagnum spp. and Drosera madagascariensis), close to the N 7 road between Ambositra and Fianarantsoa (ca. 3. 5 km S Ambalamanakana; altitude 1,700 m). This place is the type locality of the new species. The specimens of C. reebae were taken towards the bottom right corner of the photo, in the grass, close to the end of a small stream with slowly-flowing water (not visible on the photo). (11) Grassy meadow surrounded by forest, in the Andringitra massif (ca 2. 5 km SSW Antanifotsy; altitude 1,600 m). The specimens were taken towards the centre-right of the photo, in water slowly seeping through the grass, as shown in the close-up view in (12). The substrate was very muddy. (13) Brooklet with very slowly flowing clear water, with dense grass (Andringitra massif, ca. 2. 3 km SW Antanifotsy; altitude 1,580 m). (14) Small slowly-flowing brook in wet grassland above tree line in the Andringitra massif (ca 4. 3 km SW Antanifotsy; altitude 1,990 m). (15) Close-up view corresponding to the centre-right area of (14)." httpUri="https://zenodo.org/record/801863/files/figure.png" pageId="7" pageNumber="138">Figs. 14 &amp; 15</figureCitation>
). Common features of all five collecting sites include: high altitude, open environment, clear water, very slow water circulation, substratum rich in organic matter (mineral substratum invisible), small depth, and dense marginal vegetation of 
<taxonomicName id="C4CA4D3B166AFF99EF45FE4169A69EA0" box="[1119,1211,477,500]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="138" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
and/or 
<taxonomicName id="C4CA4D3B166AFF99EE09FE4068849EA0" box="[1299,1433,476,500]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="138" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
. Within the study area, the species was never encountered in purely stagnant water bodies or in streams and rivers with dominant mineral substrate and/or noticeably unidirectional water flow.
</paragraph>
<paragraph id="037536B8166AFF99EBDDFDDB6F519DD0" blockId="7.[151,1437,151,824]" pageId="7" pageNumber="138">
<emphasis id="31BEEAAA166AFF99EBDDFDDB6C1B9D34" bold="true" box="[199,262,583,608]" pageId="7" pageNumber="138">Note.</emphasis>
The 
<typeStatus id="DC71881A166AFF99EA57FDD46C609D34" box="[333,381,584,608]" pageId="7" pageNumber="138">type</typeStatus>
locality of 
<taxonomicName id="C4CA4D3B166AFF99E908FDD46F999D34" box="[530,644,584,608]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="reebae" status="sp. nov.">
<emphasis id="31BEEAAA166AFF99E908FDD46F999D34" box="[530,644,584,608]" italics="true" pageId="7" pageNumber="138">C. reebae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2A8D57D1166AFF99E988FDDB6FEF9D34" box="[658,754,583,608]" pageId="7" pageNumber="138" rank="species">
<emphasis id="31BEEAAA166AFF99E988FDDB6FEF9D34" bold="true" box="[658,754,583,608]" pageId="7" pageNumber="138">sp. nov.</emphasis>
</taxonomicNameLabel>
is located about 
<quantity id="C4329B5D166AFF99E8D1FDD469049D0B" box="[971,1049,584,608]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="138" unit="m" value="400.0">400 m</quantity>
south from the 
<typeStatus id="DC71881A166AFF99EFFFFDD468089D34" box="[1253,1301,584,608]" pageId="7" pageNumber="138">type</typeStatus>
locality of 
<taxonomicName id="C4CA4D3B166AFF99EB8DFDF06F559DD0" authority="Stastny, 2012" authorityName="Stastny" authorityYear="2012" box="[151,584,620,645]" class="Insecta" family="Dytiscidae" genus="Herophydrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="travniceki">
<emphasis id="31BEEAAA166AFF99EB8DFDF06CBC9DD0" box="[151,417,620,644]" italics="true" pageId="7" pageNumber="138">Herophydrus travniceki</emphasis>
Šťastný, 2012
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4BD06533166AFF99EBDDFD1369A49D98" pageId="7" pageNumber="138" type="distribution">
<paragraph id="037536B8166AFF99EBDDFD1369A49D98" blockId="7.[151,1437,151,824]" pageId="7" pageNumber="138">
<emphasis id="31BEEAAA166AFF99EBDDFD136C449DFC" bold="true" box="[199,345,655,680]" pageId="7" pageNumber="138">Distribution</emphasis>
(
<figureCitation id="9BF12A3D166AFF99EA73FD136CA09DFC" box="[361,445,655,680]" captionStart="FIGURE 16" captionStartId="7.[151,250,1915,1937]" captionTargetBox="[371,1179,894,1869]" captionTargetId="figure@7.[357,1199,877,1885]" captionTargetPageId="7" captionText="FIGURE 16. Map of Madagascar showing known localities of Canthyporus pauliani (black empty square) and C. reebae sp. nov. (black empty stars)." httpUri="https://zenodo.org/record/801865/files/figure.png" pageId="7" pageNumber="138">Fig. 16</figureCitation>
). So far only known from the south-eastern part of the Itremo mountain range and from the Andringitra massif, in central eastern 
<collectingCountry id="7BDD7628166AFF99E97DFD286FED9D99" box="[615,752,692,717]" name="Madagascar" pageId="7" pageNumber="138">Madagascar</collectingCountry>
, at altitudes between 1,580 and 
<quantity id="C4329B5D166AFF99EF4CFD2869A89D98" box="[1110,1205,692,716]" metricMagnitude="3" metricUnit="m" metricValue="2.1399999999999997" pageId="7" pageNumber="138" unit="m" value="2140.0">2,140 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4BD06533166AFF99EBDDFD4B6C839C6C" pageId="7" pageNumber="138" type="etymology">
<paragraph id="037536B8166AFF99EBDDFD4B6C839C6C" blockId="7.[151,1437,151,824]" pageId="7" pageNumber="138">
<emphasis id="31BEEAAA166AFF99EBDDFD4B6CBA9DA4" bold="true" box="[199,423,727,752]" pageId="7" pageNumber="138">Derivatio nominis</emphasis>
. This species is dedicated to Catherine Reeb, who organised and supervised the RicciaTeam expedition in 
<date id="77741078166AFF99EAA6FD616F6F9C41" box="[444,626,764,789]" pageId="7" pageNumber="138" value="2016-05" valueMax="2016-06-31" valueMin="2016-05-00">May–June 2016</date>
, during which the specimens were collected. The species name is a noun in the genitive singular.
</paragraph>
</subSubSection>
</treatment>
</document>