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<mods:title id="289C1F7505456BF5E6AB03655CDBA013">Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae)</mods:title>
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<mods:namePart id="196E1AE005B276D9F50D264E0A2413D1">Roell, Talita</mods:namePart>
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<mods:affiliation id="B0890316B7D5404C3107F1914B1C25A3">Universidade Federal do Rio Grande do Sul, Instituto de Biociências, Departamento de Zoologia, Av. Bento Gonçalves, 9500, Agronomia, 91501 - 970 Porto Alegre, RS, Brasil.</mods:affiliation>
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<mods:namePart id="B06360A005ACEA2DD5C2E18992EA85BA">Lemaître, Valérie A.</mods:namePart>
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<taxonomicName id="C6E7F4C9D56BFF9EF9AE9177B493F9F9" authority="Lemaitre, Roell &amp; Brugnera" authorityName="Lemaitre, Roell &amp; Brugnera" authorityYear="2021" box="[151,688,1626,1653]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="33935358D56BFF9EF9AE9177B777F9F9" bold="true" box="[151,340,1626,1653]" italics="true" pageId="8" pageNumber="578">Podisus thomasi</emphasis>
Lemaître, Roell &amp; Brugnera
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, 
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(
<figureCitation id="99DC93CFD56BFF9EF9A69152B703F915" box="[159,288,1662,1689]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="8" pageNumber="578">Figs 13–15</figureCitation>
, 
<figureCitation id="99DC93CFD56BFF9EF8159152B756F915" box="[300,373,1663,1689]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="8" pageNumber="578">22–24</figureCitation>
, 
<figureCitation id="99DC93CFD56BFF9EF8B99152B7EBF915" box="[384,456,1663,1689]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">31–33</figureCitation>
, 
<figureCitation id="99DC93CFD56BFF9EF8ED9152B43EF915" box="[468,541,1663,1689]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="8" pageNumber="578">38–39</figureCitation>
, 
<figureCitation id="99DC93CFD56BFF9EFB119152B453F915" box="[552,624,1663,1689]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="8" pageNumber="578">46–48</figureCitation>
, 
<figureCitation id="99DC93CFD56BFF9EFB459153B4E7F915" box="[636,708,1662,1689]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">53–54</figureCitation>
, 
<figureCitation id="99DC93CFD56BFF9EFBE99153B53BF915" box="[720,792,1662,1689]" captionStart="FIGURES 55–60" captionStartId="12.[151,264,1290,1314]" captionTargetBox="[151,1436,453,1267]" captionTargetId="figure-150@12.[151,1436,453,1267]" captionTargetPageId="12" captionText="FIGURES 55–60. Live specimens of Podisus graziae sp. n. (55–57), and Podisus thomasi sp. n. (58–60). 55–57, photos by Ricardo Brugnera, Porto Alegre, Brazil. 58–60, photos by Erika Schulz, Mindo, Pichincha, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.4692483" httpUri="https://zenodo.org/record/4692483/files/figure.png" pageId="8" pageNumber="578">58–60</figureCitation>
)
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<taxonomicName id="C6E7F4C9D56BFF9EF9AE91EEB7BAF957" authority="(Dallas)" baseAuthorityName="Dallas" box="[151,409,1731,1755]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="33935358D56BFF9EF9AE91EEB71DF957" box="[151,318,1731,1755]" italics="true" pageId="8" pageNumber="578">Podisus pallipes</emphasis>
(Dallas)
</taxonomicName>
: 
<treatmentCitation id="8046A95BD56BFF9EF89A91EEB447F957" author="Thomas, D. B." box="[419,612,1731,1755]" page="101" pageId="8" pageNumber="578" year="1992">
<bibRefCitation id="6576F2BBD56BFF9EF89A91EEB447F957" author="Thomas, D. B." box="[419,612,1731,1755]" pageId="8" pageNumber="578" refId="ref10128" refString="Thomas, D. B. (1992) Taxonomic Synopsis of the Asopine Pentatomidae (Heteroptera) of the Western Hemisphere. Entomological Society of America, Lanham, Maryland, 156 pp." type="book" year="1992">Thomas 1992: 101</bibRefCitation>
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; 
<treatmentCitation id="8046A95BD56BFF9EFB5791EEB57AF957" author="Weiler, L. &amp; Bianchi, F. M. &amp; Grazia, J." box="[622,857,1731,1755]" page="188" pageId="8" pageNumber="578" year="2012">
<bibRefCitation id="6576F2BBD56BFF9EFB5791EEB57AF957" author="Weiler, L. &amp; Bianchi, F. M. &amp; Grazia, J." box="[622,857,1731,1755]" pageId="8" pageNumber="578" pagination="186 - 191" refId="ref10287" refString="Weiler, L., Bianchi, F. M. &amp; Grazia, J. (2012) Percevejos-do-mato (Hemiptera, Heteroptera, Pentatomoidea) da Floresta Nacional de Sao Francisco de Paula, RS, Brasil. Brazilian Journal of Biosciences, 10 (2), 186 - 191." type="journal article" year="2012">
Weiler 
<emphasis id="33935358D56BFF9EFB8E91E9B4C7F957" box="[695,740,1731,1755]" italics="true" pageId="8" pageNumber="578">et al</emphasis>
. 2012: 188
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="8046A95BD56BFF9EFA5A91EEB274F957" author="Dellape, G. &amp; Rider, D. &amp; Dellape, P. M." box="[867,1111,1731,1755]" page="170" pageId="8" pageNumber="578" year="2015">
<bibRefCitation id="6576F2BBD56BFF9EFA5A91EEB274F957" author="Dellape, G. &amp; Rider, D. &amp; Dellape, P. M." box="[867,1111,1731,1755]" pageId="8" pageNumber="578" pagination="169 - 176" refId="ref8853" refString="Dellape, G., Rider, D. &amp; Dellape, P. M. (2015) Notes on distributions for Argentinean Pentatomidae (Heteroptera: Pentatomoidea), with new records in the country. Revista Brasileira de Entomologia, 59, 169 - 176. https: // doi. org / 10.1016 / j. rbe. 2015.06.001" type="journal article" year="2015">
Dellapé 
<emphasis id="33935358D56BFF9EFA8191E9B5C6F957" box="[952,997,1731,1755]" italics="true" pageId="8" pageNumber="578">et al</emphasis>
. 2015: 170
</bibRefCitation>
</treatmentCitation>
(misapplication).
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</paragraph>
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<paragraph id="01588F4AD56BFF9EF9AE9024B791F8E7" blockId="8.[151,1436,1801,2008]" pageId="8" pageNumber="578">
<materialsCitation id="B18F8517D56BFF9EF9AE9024B791F8E7" ID-GBIF-Occurrence="3083796312" collectingDate="1986-07-09" collectionCode="NHMUK" collectorName="R. S. Miller &amp; J. E. Eger Collection" country="Venezuela" location="Las Cruces" pageId="8" pageNumber="578" specimenCount="2" stateProvince="Merida" typeStatus="holotype">
<emphasis id="33935358D56BFF9EF9AE9024B728F8AF" bold="true" box="[151,267,1801,1827]" pageId="8" pageNumber="578">
<typeStatus id="DE5C31E8D56BFF9EF9AE9024B725F8AF" box="[151,262,1801,1827]" pageId="8" pageNumber="578" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Labels: “ 
<collectingCountry id="79F0CFDAD56BFF9EF8809024B444F8AF" box="[441,615,1801,1827]" name="Venezuela" pageId="8" pageNumber="578">VENEZUELA</collectingCountry>
: 
<collectingRegion id="C32341A8D56BFF9EFB499024B4E2F8AF" box="[624,705,1801,1827]" country="Venezuela" name="Merida" pageId="8" pageNumber="578">Mérida</collectingRegion>
<location id="0438D991D56BFF9EFBFE9027B560F8AF" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56BFF9EFBFE9027B560F8AF" box="[711,835,1801,1828]" country="Venezuela" name="Las Cruces" pageId="8" pageNumber="578" stateProvince="Merida">Las Cruces</location>
<date id="7559A98AD56BFF9EFA709024B5EAF8AF" box="[841,969,1801,1827]" pageId="8" pageNumber="578" value="1986-07-09">
<collectingDate id="651D5062D56BFF9EFA709024B5EAF8AF" box="[841,969,1801,1827]" pageId="8" pageNumber="578" value="1986-07-09">9 July 1986</collectingDate>
</date>
<collectorName id="AC12EA9CD56BFF9EFAF69024B26FF8AF" box="[975,1100,1801,1827]" pageId="8" pageNumber="578">R.S. Miller</collectorName>
colr.”; “B.M. 1991-55”; “ 
<collectorName id="AC12EA9CD56BFF9EFC559024B76EF8CB" pageId="8" pageNumber="578">J. E. Eger Collection</collectorName>
”; “ 
<typeStatus id="DE5C31E8D56BFF9EF8539000B7CEF8CB" box="[362,493,1837,1863]" pageId="8" pageNumber="578" type="neotype">NEOTYPE</typeStatus>
<taxonomicName id="C6E7F4C9D56BFF9EF8C89003B2A2F8CB" authority="PALLIPES (DALLAS 1851) Desig. D. B. Thomas" authorityName="PALLIPES (DALLAS" authorityYear="1851" box="[497,1153,1837,1864]" class="Insecta" family="Pentatomidae" genus="Arma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="genus">
ARMA PALLIPES (
<bibRefCitation id="6576F2BBD56BFF9EFBEC9000B5A7F8C4" author="Dallas, W. S." box="[725,900,1837,1864]" pageId="8" pageNumber="578" refId="ref8744" refString="Dallas, W. S. (1851) List of the specimens of hemipterous insects in the collection of the British Museum. Part 1. Trustees of the British Museum, London, 368 pp., 11 pls." type="book" year="1851">DALLAS 1851</bibRefCitation>
) Desig. D. B. Thomas
</taxonomicName>
”; “NHMUK 010592332” (
<figureCitation id="99DC93CFD56BFF9EF9A6907CB702F8E0" box="[159,289,1873,1900]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="8" pageNumber="578">Figs 13–15</figureCitation>
). (
<collectionCode id="67F6178FD56BFF9EF804907CB789F8E7" box="[317,426,1873,1899]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="578" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="01588F4AD56BFF9FF9FE9058B46AFE79" blockId="8.[151,1436,1801,2008]" lastBlockId="9.[151,1437,150,1978]" lastPageId="9" lastPageNumber="579" pageId="8" pageNumber="578">
<materialsCitation id="B18F8517D56BFF9EF9FE9058B713F83F" ID-GBIF-Occurrence="3083796314" collectionCode="JEE" collectorName="C. W. &amp; L. B. O'Brien" country="Panama" elevation="680" location="Cerro Tute" pageId="8" pageNumber="578" specimenCount="1" specimenCount-female="1" stateProvince="Veraguas" typeStatus="paratype">
<emphasis id="33935358D56BFF9EF9FE9058B46AF803" bold="true" box="[199,585,1909,1935]" pageId="8" pageNumber="578">
<typeStatus id="DE5C31E8D56BFF9EF9FE9058B762F803" box="[199,321,1909,1935]" pageId="8" pageNumber="578" type="paratype">Paratypes</typeStatus>
. 
<collectingCountry id="79F0CFDAD56BFF9EF8779058B7EEF803" box="[334,461,1909,1935]" name="Panama" pageId="8" pageNumber="578">PANAMA</collectingCountry>
: 
<collectingRegion id="C32341A8D56BFF9EF8E39058B46AF803" box="[474,585,1909,1935]" country="Panama" name="Veraguas" pageId="8" pageNumber="578">Veraguas</collectingRegion>
</emphasis>
, 
<location id="0438D991D56BFF9EFB609058B4F6F81C" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56BFF9EFB609058B4F6F81C" box="[601,725,1909,1936]" country="Panama" name="Cerro Tute" pageId="8" pageNumber="578" stateProvince="Veraguas">Cerro Tute</location>
, 4 
<collectorName id="AC12EA9CD56BFF9EFBC99058B589F803" box="[752,938,1909,1936]" pageId="8" pageNumber="578">Km W Santa Fé</collectorName>
, 
<quantity id="C61F22AFD56BFF9EFA8E9058B5DAF803" box="[951,1017,1909,1935]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="578" unit="m" value="680.0">
<elevation id="8ACA6879D56BFF9EFA8E9058B5DAF803" box="[951,1017,1909,1935]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="578" unit="m" value="680.0">680m</elevation>
</quantity>
, 2-XIII-1995, 
<collectorName id="AC12EA9CD56BFF9EFDA79058B3BFF803" box="[1182,1436,1909,1936]" pageId="8" pageNumber="578">C.W. &amp; L. B. O’Brien</collectorName>
col., 
<specimenCount id="17E144C3D56BFF9EF9E990B7B6C6F838" box="[208,229,1946,1972]" pageId="8" pageNumber="578" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56BFF9EF9CC90B4B70BF83F" box="[245,296,1945,1971]" pageId="8" pageNumber="578">JEE</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56BFF9EF80590B4B289F83F" ID-GBIF-Occurrence="3083796331" box="[316,1194,1945,1972]" collectingDate="1951-05-10" collectionCode="MIZA" collectorName="F. Fernandes" country="Venezuela" location="Humocaro Bajo" pageId="8" pageNumber="578" specimenCount="1" specimenCount-female="1" stateProvince="Lara" typeStatus="paratype">
<emphasis id="33935358D56BFF9EF80590B4B414F83F" bold="true" box="[316,567,1945,1971]" pageId="8" pageNumber="578">
<collectingCountry id="79F0CFDAD56BFF9EF80590B4B7CCF83F" box="[316,495,1945,1971]" name="Venezuela" pageId="8" pageNumber="578">VENEZUELA</collectingCountry>
: 
<collectingRegion id="C32341A8D56BFF9EF8C590B4B414F83F" box="[508,567,1945,1971]" country="Venezuela" name="Lara" pageId="8" pageNumber="578">Lara</collectingRegion>
</emphasis>
, 
<location id="0438D991D56BFF9EFB7F90B7B4DEF83F" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56BFF9EFB7F90B7B4DEF83F" box="[582,765,1945,1972]" country="Venezuela" name="Humocaro Bajo" pageId="8" pageNumber="578" stateProvince="Lara">Humocaro Bajo</location>
, 
<date id="7559A98AD56BFF9EFA3090B4B5A7F838" box="[777,900,1945,1972]" pageId="8" pageNumber="578" value="1951-05-10">
<collectingDate id="651D5062D56BFF9EFA3090B4B5A7F838" box="[777,900,1945,1972]" pageId="8" pageNumber="578" value="1951-05-10">10-V-1951</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56BFF9EFAA990B7B200F83F" box="[912,1059,1945,1972]" pageId="8" pageNumber="578">F. Fernandes</collectorName>
, 
<specimenCount id="17E144C3D56BFF9EFD1690B7B267F838" box="[1071,1092,1946,1972]" pageId="8" pageNumber="578" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56BFF9EFD6D90B4B281F83F" box="[1108,1186,1945,1971]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="8" pageNumber="578" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="B18F8517D56BFF9EFD8F90B4B4A9F85B" ID-GBIF-Occurrence="3083796323" collectingDate="1963-09-17" collectionCode="MHNLS" collectorName="C. Gonzalez" country="Venezuela" location="Barbula" pageId="8" pageNumber="578" specimenCount="1" specimenCount-female="1" stateProvince="Carabobo" typeStatus="paratype">
<emphasis id="33935358D56BFF9EFD8F90B4B30DF83F" bold="true" box="[1206,1326,1945,1971]" pageId="8" pageNumber="578">
<collectingRegion id="C32341A8D56BFF9EFD8F90B4B30DF83F" box="[1206,1326,1945,1971]" country="Venezuela" name="Carabobo" pageId="8" pageNumber="578">Carabobo</collectingRegion>
</emphasis>
, 
<location id="0438D991D56BFF9EFC0490B4B3BBF83F" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56BFF9EFC0490B4B3BBF83F" box="[1341,1432,1945,1971]" country="Venezuela" name="Barbula" pageId="8" pageNumber="578" stateProvince="Carabobo">Barbula</location>
, 
<date id="7559A98AD56BFF9EF9AE9090B73DF85B" box="[151,286,1981,2007]" pageId="8" pageNumber="578" value="1963-09-17">
<collectingDate id="651D5062D56BFF9EF9AE9090B73DF85B" box="[151,286,1981,2007]" pageId="8" pageNumber="578" value="1963-09-17">17-IX-1963</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56BFF9EF8139090B796F85B" box="[298,437,1981,2007]" pageId="8" pageNumber="578">C. Gonzalez</collectorName>
col., 
<specimenCount id="17E144C3D56BFF9EF8CC9093B429F854" box="[501,522,1982,2008]" pageId="8" pageNumber="578" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56BFF9EFB209090B4A2F85B" box="[537,641,1981,2007]" country="Venezuela" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="8" pageNumber="578">MHNLS</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="B18F8517D56BFF9EFBAC9090B3BFF85B" ID-GBIF-Occurrence="3083796330" box="[661,1436,1981,2008]" collectorName="La Guamita &amp; Henri Pittier" country="Venezuela" location="Guamitas" pageId="8" pageNumber="578" specimenCount="1" stateProvince="Aragua" typeStatus="paratype">
<emphasis id="33935358D56BFF9EFBAC9090B4CCF85B" bold="true" box="[661,751,1981,2007]" pageId="8" pageNumber="578">
<collectingRegion id="C32341A8D56BFF9EFBAC9090B4CCF85B" box="[661,751,1981,2007]" country="Venezuela" name="Aragua" pageId="8" pageNumber="578">Aragua</collectingRegion>
</emphasis>
, 
<location id="0438D991D56BFF9EFBC49090B54BF85B" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56BFF9EFBC49090B54BF85B" box="[765,872,1981,2007]" country="Venezuela" name="Guamitas" pageId="8" pageNumber="578" stateProvince="Aragua">Guamitas</location>
(probably current 
<collectorName id="AC12EA9CD56BFF9EFD059093B2E2F85B" box="[1084,1217,1981,2008]" pageId="8" pageNumber="578">La Guamita</collectorName>
in the 
<collectorName id="AC12EA9CD56BFF9EFC299090B3BFF85B" box="[1296,1436,1981,2007]" pageId="8" pageNumber="578">Henri Pittier</collectorName>
</materialsCitation>
<materialsCitation id="B18F8517D56AFF9FF9AE97BAB54DFF3D" ID-GBIF-Occurrence="3083796301" box="[151,878,150,178]" collectionCode="MIZA" collectorName="F. Fernandes" elevation="750" location="National Park" pageId="9" pageNumber="579" specimenCount="1" specimenCount-female="1">
<location id="0438D991D56AFF9FF9AE97BAB71BFF3E" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FF9AE97BAB71BFF3E" box="[151,312,151,178]" name="National Park" pageId="9" pageNumber="579">National Park</location>
), 
<quantity id="C61F22AFD56AFF9FF87297BBB7ADFF3D" box="[331,398,150,177]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="579" unit="m" value="750.0">
<elevation id="8ACA6879D56AFF9FF87297BBB7ADFF3D" box="[331,398,150,177]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="579" unit="m" value="750.0">750m</elevation>
</quantity>
, 24-VIII-62, 
<collectorName id="AC12EA9CD56AFF9FFB1D97BAB490FF3D" box="[548,691,151,177]" pageId="9" pageNumber="579">F. Fernandes</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFBCA97BAB52BFF3D" box="[755,776,151,177]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFA2197BAB545FF3D" box="[792,870,151,177]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="579" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFA4397BAB435FF59" ID-GBIF-Occurrence="3083796326" collectingDate="1958-07-29" collectionCode="MHNLS" location="Henri Pittier National Park" municipality="Rancho Grande" pageId="9" pageNumber="579" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="E13C1530D56AFF9FFA4397BAB20AFF3D" box="[890,1065,151,177]" pageId="9" pageNumber="579">Rancho Grande</collectingMunicipality>
(currently 
<location id="0438D991D56AFF9FFD9197BAB6ECFF5A" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FFD9197BAB6ECFF5A" municipality="Rancho Grande" name="Henri Pittier National Park" pageId="9" pageNumber="579">Henri Pittier National Park</location>
), 
<date id="7559A98AD56AFF9FF9DA9797B750FF59" box="[227,371,186,213]" pageId="9" pageNumber="579" value="1958-07-29">
<collectingDate id="651D5062D56AFF9FF9DA9797B750FF59" box="[227,371,186,213]" pageId="9" pageNumber="579" value="1958-07-29">29-VII-1958</collectingDate>
</date>
, 
<specimenCount id="17E144C3D56AFF9FF8B99796B7B6FF59" box="[384,405,187,213]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FF89C9796B42EFF59" box="[421,525,187,213]" country="Venezuela" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="9" pageNumber="579">MHNLS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFB1B9796B212FF59" ID-GBIF-Occurrence="3083796307" box="[546,1073,187,213]" collectingDate="1966-01-02" collectionCode="MIZA" collectorName="F. Romero" elevation="1100" pageId="9" pageNumber="579" specimenCount="1" specimenCount-female="1">
<quantity id="C61F22AFD56AFF9FFB1B9796B451FF59" box="[546,626,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">
<elevation id="8ACA6879D56AFF9FFB1B9796B451FF59" box="[546,626,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">1100m</elevation>
</quantity>
, 
<date id="7559A98AD56AFF9FFB479796B4D2FF59" box="[638,753,187,213]" pageId="9" pageNumber="579" value="1966-01-02">
<collectingDate id="651D5062D56AFF9FFB479796B4D2FF59" box="[638,753,187,213]" pageId="9" pageNumber="579" value="1966-01-02">02-I-1966</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFBC79796B557FF59" box="[766,884,187,213]" pageId="9" pageNumber="579">F. Romero</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFA8F9796B5E8FF59" box="[950,971,187,213]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFAE29796B20AFF59" box="[987,1065,187,213]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="579" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFD049796B7DCFF75" ID-GBIF-Occurrence="3083796315" collectionCode="MIZA" collectorName="F. Fernandes &amp; C. J. Rosales" elevation="1100" pageId="9" pageNumber="579" specimenCount="1" specimenCount-female="1">
<quantity id="C61F22AFD56AFF9FFD049796B2AEFF59" box="[1085,1165,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">
<elevation id="8ACA6879D56AFF9FFD049796B2AEFF59" box="[1085,1165,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">1100m</elevation>
</quantity>
, 6-XII-56, 
<collectorName id="AC12EA9CD56AFF9FFC359796B3BFFF59" box="[1292,1436,187,213]" pageId="9" pageNumber="579">F. Fernandes</collectorName>
&amp; 
<collectorName id="AC12EA9CD56AFF9FF98F97F2B762FF75" box="[182,321,223,249]" pageId="9" pageNumber="579">C.J. Rosales</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FF8BA97F2B7BBFF75" box="[387,408,223,249]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FF89097F2B7D4FF75" box="[425,503,223,249]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="579" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFB3597F2B26DFF75" ID-GBIF-Occurrence="3083796317" box="[524,1102,222,249]" collectingDate="1952-03-12" collectionCode="MIZA" collectorName="J. R. Reguena" elevation="1100" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1">
<quantity id="C61F22AFD56AFF9FFB3597F2B478FF75" box="[524,603,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">
<elevation id="8ACA6879D56AFF9FFB3597F2B478FF75" box="[524,603,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">1100m</elevation>
</quantity>
, 
<date id="7559A98AD56AFF9FFB5197F3B4CDFF75" box="[616,750,222,249]" pageId="9" pageNumber="579" value="1952-03-12">
<collectingDate id="651D5062D56AFF9FFB5197F3B4CDFF75" box="[616,750,222,249]" pageId="9" pageNumber="579" value="1952-03-12">12-III-1952</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFBC297F2B5B2FF75" box="[763,913,223,249]" pageId="9" pageNumber="579">J.R. Reguena</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFAEA97F2B5CBFF75" box="[979,1000,223,249]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFAC197F2B265FF75" box="[1016,1094,223,249]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="579" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFD6297F2B78EFE91" ID-GBIF-Occurrence="3083796308" collectingDate="1966-04-25" collectionCode="MIZA" collectorName="J. &amp; B. Bechine" elevation="1100" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1">
<quantity id="C61F22AFD56AFF9FFD6297F2B288FF75" box="[1115,1195,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">
<elevation id="8ACA6879D56AFF9FFD6297F2B288FF75" box="[1115,1195,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="579" unit="m" value="1100.0">1100m</elevation>
</quantity>
, 
<date id="7559A98AD56AFF9FFD8197F3B31FFF75" box="[1208,1340,222,249]" pageId="9" pageNumber="579" value="1966-04-25">
<collectingDate id="651D5062D56AFF9FFD8197F3B31FFF75" box="[1208,1340,222,249]" pageId="9" pageNumber="579" value="1966-04-25">25-IV-1966</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFC7097F2B6D0FE91" pageId="9" pageNumber="579">J. &amp; B. Bechine</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FF80A962EB76BFE91" box="[307,328,259,285]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FF86E962EB786FE91" box="[343,421,259,285]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="579" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FF880962FB55AFE91" ID-GBIF-Occurrence="3083796325" box="[441,889,258,285]" collectingDate="1954-01" collectionCode="EMG" collectorName="Helga Frey" pageId="9" pageNumber="579" specimenCount="1" specimenCount-female="1">
<date id="7559A98AD56AFF9FF880962FB425FE91" box="[441,518,258,285]" pageId="9" pageNumber="579" value="1954-01">
<collectingDate id="651D5062D56AFF9FF880962FB425FE91" box="[441,518,258,285]" pageId="9" pageNumber="579" value="1954-01">I-1954</collectingDate>
</date>
, G.u. 
<collectorName id="AC12EA9CD56AFF9FFB72962EB4EBFE91" box="[587,712,259,285]" pageId="9" pageNumber="579">Helga Frey</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFA31962EB53EFE91" box="[776,797,259,285]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFA14962EB552FE91" box="[813,881,259,285]" pageId="9" pageNumber="579">EMG</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFABC962EB730FECD" ID-GBIF-Occurrence="3083796309" collectingDate="1996-05-25" collectionCode="MIZA" collectorName="F. Fernandes" elevation="450" location="El Limon" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1">
<location id="0438D991D56AFF9FFABC962EB5D7FE91" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FFABC962EB5D7FE91" box="[901,1012,259,285]" name="El Limon" pageId="9" pageNumber="579">El Limón</location>
, 
<quantity id="C61F22AFD56AFF9FFAC6962FB261FE91" box="[1023,1090,258,285]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="579" unit="m" value="450.0">
<elevation id="8ACA6879D56AFF9FFAC6962FB261FE91" box="[1023,1090,258,285]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="579" unit="m" value="450.0">450m</elevation>
</quantity>
, 
<date id="7559A98AD56AFF9FFD77962FB2EAFE91" box="[1102,1225,258,285]" pageId="9" pageNumber="579" value="1996-05-25">
<collectingDate id="651D5062D56AFF9FFD77962FB2EAFE91" box="[1102,1225,258,285]" pageId="9" pageNumber="579" value="1996-05-25">25-V-1996</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFDEC962EB347FE91" box="[1237,1380,259,285]" pageId="9" pageNumber="579">F. Fernandes</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FF9AE960AB68FFECD" box="[151,172,295,321]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
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)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FF818960AB2C4FECD" ID-GBIF-Occurrence="3083796320" box="[289,1255,294,321]" collectingDate="1982-01-09" collectionCode="AMNH" collectorName="R. T. Schuh &amp; B. M. Massie" elevation="1500" location="Barinas" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1">
<location id="0438D991D56AFF9FF818960AB75AFECD" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FF818960AB75AFECD" box="[289,377,295,321]" name="Barinas" pageId="9" pageNumber="579">Barinas</location>
, 
<collectorName id="AC12EA9CD56AFF9FF8BE960AB7CDFECD" box="[391,494,295,321]" pageId="9" pageNumber="579">El Junco</collectorName>
, 
<quantity id="C61F22AFD56AFF9FF8C5960BB46FFECD" box="[508,588,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="579" unit="m" value="1500.0">
<elevation id="8ACA6879D56AFF9FF8C5960BB46FFECD" box="[508,588,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="579" unit="m" value="1500.0">1500m</elevation>
</quantity>
, 
<date id="7559A98AD56AFF9FFB63960AB49CFECD" box="[602,703,295,321]" pageId="9" pageNumber="579" value="1982-01-09">
<collectingDate id="651D5062D56AFF9FFB63960AB49CFECD" box="[602,703,295,321]" pageId="9" pageNumber="579" value="1982-01-09">9-I-1982</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFBF5960AB56FFECD" box="[716,844,295,321]" pageId="9" pageNumber="579">R.T. Schuh</collectorName>
and 
<collectorName id="AC12EA9CD56AFF9FFABE960AB23AFECD" box="[903,1049,295,321]" pageId="9" pageNumber="579">B.M. Massie</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFD64960AB251FECD" box="[1117,1138,295,321]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFDBA960AB2FCFECD" box="[1155,1247,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="579">AMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="B18F8517D56AFF9FFDCD960BB3BAFEE9" ID-GBIF-Occurrence="3083796321" collectionCode="MPUJ" collectorName="La Almenara &amp; P. Erazo &amp; C. Lesmes &amp; L. Lopez" country="Colombia" elevation="1123" location="Santa Maria" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<emphasis id="33935358D56AFF9FFDCD960BB6CDFEE9" bold="true" pageId="9" pageNumber="579">
<collectingCountry id="79F0CFDAD56AFF9FFDCD960BB3B4FECC" box="[1268,1431,294,320]" name="Colombia" pageId="9" pageNumber="579">COLOMBIA</collectingCountry>
: 
<collectingRegion id="C32341A8D56AFF9FF9AE9667B6CDFEE9" box="[151,238,330,357]" country="Colombia" name="Boyaca" pageId="9" pageNumber="579">Boyacá</collectingRegion>
</emphasis>
, 
<location id="0438D991D56AFF9FF9C79666B7AFFEE9" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FF9C79666B7AFFEE9" box="[254,396,331,357]" country="Colombia" name="Santa Maria" pageId="9" pageNumber="579" stateProvince="Boyaca">Santa Maria</location>
, sector 
<collectorName id="AC12EA9CD56AFF9FF8DD9666B459FEE9" box="[484,634,331,357]" pageId="9" pageNumber="579">La Almenara</collectorName>
, 
<quantity id="C61F22AFD56AFF9FFBB19666B4F4FEE9" box="[648,727,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.123" pageId="9" pageNumber="579" unit="m" value="1123.0">
<elevation id="8ACA6879D56AFF9FFBB19666B4F4FEE9" box="[648,727,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.123" pageId="9" pageNumber="579" unit="m" value="1123.0">1123m</elevation>
</quantity>
, 4.87, -73.25, 
<collectorName id="AC12EA9CD56AFF9FFA479666B5FDFEE9" box="[894,990,331,357]" pageId="9" pageNumber="579">P. Erazo</collectorName>
, 
<collectorName id="AC12EA9CD56AFF9FFAD29666B245FEE9" box="[1003,1126,331,357]" pageId="9" pageNumber="579">C. Lesmes</collectorName>
, 
<collectorName id="AC12EA9CD56AFF9FFD4D9666B2F8FEE9" box="[1140,1243,331,357]" pageId="9" pageNumber="579">L. López</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFC249666B311FEE9" box="[1309,1330,331,357]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFC7A9666B3B2FEE9" box="[1347,1425,331,357]" pageId="9" pageNumber="579">MPUJ</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="B18F8517D56AFF9FF9AE9643B6DEFE21" ID-GBIF-Occurrence="3083796316" collectingDate="2012-09-15" collectionCode="UNAB" collectorName="Finca el Paraiso &amp; A. Garcia" country="Colombia" elevation="12" location="Vereda Arabia" municipality="Quipile" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca">
<emphasis id="33935358D56AFF9FF9AE9643B768FE04" bold="true" box="[151,331,366,392]" pageId="9" pageNumber="579">
<collectingRegion id="C32341A8D56AFF9FF9AE9643B768FE04" box="[151,331,366,392]" country="Colombia" name="Cundinamarca" pageId="9" pageNumber="579">Cundinamarca</collectingRegion>
</emphasis>
, 
<collectingMunicipality id="E13C1530D56AFF9FF8619642B78CFE05" box="[344,431,367,393]" pageId="9" pageNumber="579">Quipile</collectingMunicipality>
, 
<location id="0438D991D56AFF9FF8809642B479FE05" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FF8809642B479FE05" box="[441,602,367,393]" country="Colombia" municipality="Quipile" name="Vereda Arabia" pageId="9" pageNumber="579" stateProvince="Cundinamarca">Vereda Arabia</location>
, 
<collectorName id="AC12EA9CD56AFF9FFB5D9642B536FE05" box="[612,789,367,393]" pageId="9" pageNumber="579">Finca el Paraiso</collectorName>
nº 
<quantity id="C61F22AFD56AFF9FFA019642B591FE05" box="[824,946,367,393]" metricMagnitude="1" metricUnit="m" metricValue="1.22026" pageId="9" pageNumber="579" unit="m" value="12.2026">
<elevation id="8ACA6879D56AFF9FFA019642B591FE05" box="[824,946,367,393]" metricMagnitude="1" metricUnit="m" metricValue="1.22026" pageId="9" pageNumber="579" unit="m" value="12.2026">12, 2026m</elevation>
</quantity>
, 4.74, -74.56, 
<date id="7559A98AD56AFF9FFD769643B2F4FE05" box="[1103,1239,366,393]" pageId="9" pageNumber="579" value="2012-09-15">
<collectingDate id="651D5062D56AFF9FFD769643B2F4FE05" box="[1103,1239,366,393]" pageId="9" pageNumber="579" value="2012-09-15">15-IX-2012</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFDE69642B368FE05" box="[1247,1355,367,393]" pageId="9" pageNumber="579">A. Garcia</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFCBE9642B3BFFE05" box="[1415,1436,367,393]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FF9A696BEB6D6FE21" box="[159,245,403,429]" country="Colombia" httpUri="http://grbio.org/cool/hx5r-ym26" name="Universidad Nacional, Facultad de Agronomia" pageId="9" pageNumber="579">UNAB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="B18F8517D56AFF9FF83196BFB2EBFE21" ID-GBIF-Occurrence="3083796311" box="[264,1224,402,429]" collectionCode="UNAB" collectorName="Vereda El Diamante &amp; D. Rider" country="Colombia" location="Roncesvalles" pageId="9" pageNumber="579" specimenCount="1" specimenCount-female="1" stateProvince="Tolima">
<emphasis id="33935358D56AFF9FF83196BFB778FE20" bold="true" box="[264,347,402,428]" pageId="9" pageNumber="579">
<collectingRegion id="C32341A8D56AFF9FF83196BFB778FE20" box="[264,347,402,428]" country="Colombia" name="Tolima" pageId="9" pageNumber="579">Tolima</collectingRegion>
</emphasis>
, 
<location id="0438D991D56AFF9FF85096BEB423FE21" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FF85096BEB423FE21" box="[361,512,403,429]" country="Colombia" name="Roncesvalles" pageId="9" pageNumber="579" stateProvince="Tolima">Roncesvalles</location>
, 
<collectorName id="AC12EA9CD56AFF9FFB3296BEB4D3FE21" box="[523,752,403,429]" pageId="9" pageNumber="579">Vereda El Diamante</collectorName>
, 05-VI-97, M.A. &amp; 
<collectorName id="AC12EA9CD56AFF9FFAEB96BEB224FE21" box="[978,1031,403,429]" name="D. Rider" pageId="9" pageNumber="579">D.R.</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFD7F96BEB278FE21" box="[1094,1115,403,429]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFD5396BEB2E3FE21" box="[1130,1216,403,429]" country="Colombia" httpUri="http://grbio.org/cool/hx5r-ym26" name="Universidad Nacional, Facultad de Agronomia" pageId="9" pageNumber="579">UNAB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="B18F8517D56AFF9FFDEB96BFB55EFE5D" ID-GBIF-Occurrence="3083796318" collectingDate="1997-04" collectionCode="NHMUK" collectorName="Rosenberg" country="Ecuador" elevation="3500" location="Paramba" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1" stateProvince="Imbabura">
<emphasis id="33935358D56AFF9FFDEB96BFB6CDFE5D" bold="true" pageId="9" pageNumber="579">
<collectingCountry id="79F0CFDAD56AFF9FFDEB96BFB347FE20" box="[1234,1380,402,428]" name="Ecuador" pageId="9" pageNumber="579">ECUADOR</collectingCountry>
: 
<collectingRegion id="C32341A8D56AFF9FFC4896BEB6CDFE5D" country="Ecuador" name="Imbabura" pageId="9" pageNumber="579">Imbabura</collectingRegion>
</emphasis>
, 
<location id="0438D991D56AFF9FF9C2969AB743FE5D" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FF9C2969AB743FE5D" box="[251,352,439,465]" country="Ecuador" name="Paramba" pageId="9" pageNumber="579" stateProvince="Imbabura">Paramba</location>
, 
<quantity id="C61F22AFD56AFF9FF853969BB798FE5D" box="[362,443,438,465]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="579" unit="m" value="3500.0">
<elevation id="8ACA6879D56AFF9FF853969BB798FE5D" box="[362,443,438,465]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="579" unit="m" value="3500.0">3500m</elevation>
</quantity>
, 
<date id="7559A98AD56AFF9FF8FC969AB406FE5D" box="[453,549,439,465]" pageId="9" pageNumber="579" value="1997-04">
<collectingDate id="651D5062D56AFF9FF8FC969AB406FE5D" box="[453,549,439,465]" pageId="9" pageNumber="579" value="1997-04">IV-1997</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FFB16969AB485FE5D" box="[559,678,439,465]" pageId="9" pageNumber="579">Rosenberg</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FFBDB969AB4D4FE5D" box="[738,759,439,465]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFA3C969AB555FE5D" box="[773,886,439,465]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="579" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B18F8517D56AFF9FFABE969AB466FE79" ID-GBIF-Occurrence="3083796327" collectingDate="1998-08-03" collectionCode="JEE" collectorName="W. Optiz" country="Ecuador" elevation="1350" location="Maquipucuna Biol. Reserve" pageId="9" pageNumber="579" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha">
<collectingRegion id="C32341A8D56AFF9FFABE969AB5DBFE5D" box="[903,1016,439,465]" country="Ecuador" name="Pichincha" pageId="9" pageNumber="579">Pichincha</collectingRegion>
, 
<location id="0438D991D56AFF9FFD3A969AB31EFE5D" LSID="urn:lsid:plazi:treatment:894E3E5CD56BFF9AF9AE9177B4CAFE21:0438D991D56AFF9FFD3A969AB31EFE5D" box="[1027,1341,439,465]" country="Ecuador" name="Maquipucuna Biol. Reserve" pageId="9" pageNumber="579" stateProvince="Pichincha">Maquipucuna Biol. Reserve</location>
, 
<quantity id="C61F22AFD56AFF9FFC7E969BB3BBFE5D" box="[1351,1432,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="579" unit="m" value="1350.0">
<elevation id="8ACA6879D56AFF9FFC7E969BB3BBFE5D" box="[1351,1432,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="579" unit="m" value="1350.0">1350m</elevation>
</quantity>
, 2/ 
<date id="7559A98AD56AFF9FF99796F6B71BFE79" box="[174,312,475,501]" pageId="9" pageNumber="579" value="1998-08-03">
<collectingDate id="651D5062D56AFF9FF99796F6B71BFE79" box="[174,312,475,501]" pageId="9" pageNumber="579" value="1998-08-03">3-VIII-1998</collectingDate>
</date>
, 
<collectorName id="AC12EA9CD56AFF9FF87A96F6B784FE79" box="[323,423,475,501]" pageId="9" pageNumber="579">W. Optiz</collectorName>
col., 
<specimenCount id="17E144C3D56AFF9FF8DF96F6B7D8FE79" box="[486,507,475,501]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
(
<collectionCode id="67F6178FD56AFF9FFB3396F6B41EFE79" box="[522,573,475,501]" pageId="9" pageNumber="579">JEE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D56AFF9FF9FE96D2B4E3FD09" pageId="9" pageNumber="579" type="etymology">
<paragraph id="01588F4AD56AFF9FF9FE96D2B4E3FD09" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
<emphasis id="33935358D56AFF9FF9FE96D2B76DFD95" bold="true" box="[199,334,511,537]" pageId="9" pageNumber="579">Etymology.</emphasis>
We are pleased to dedicate this species to Dr Donald B. Thomas (USDA-ARS) in recognition of his remarkable work on the World 
<taxonomicName id="C6E7F4C9D56AFF9FFB1E950EB4B7FDB1" authorityName="Spinola" authorityYear="1850" box="[551,660,547,573]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="583" phylum="Arthropoda" rank="subFamily" subFamily="Asopinae">Asopinae</taxonomicName>
. The publication of his catalogues (1992, 1994) has allowed further research on the group, including this paper in which we thank him for donating to NHMUK the male specimen we are designating above as 
<typeStatus id="DE5C31E8D56AFF9FF8889546B430FD09" box="[433,531,619,645]" pageId="9" pageNumber="579" type="holotype">holotype</typeStatus>
of this species.
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D56AFF9FF9FE95A2B7FCFD25" box="[199,479,654,681]" pageId="9" pageNumber="579" type="materials_examined">
<paragraph id="01588F4AD56AFF9FF9FE95A2B7FCFD25" blockId="9.[151,1437,150,1978]" box="[199,479,654,681]" pageId="9" pageNumber="579">
<emphasis id="33935358D56AFF9FF9FE95A2B745FD24" bold="true" box="[199,358,654,681]" pageId="9" pageNumber="579">Type locality.</emphasis>
<collectingCountry id="79F0CFDAD56AFF9FF85595A2B7FCFD25" box="[364,479,655,681]" name="Venezuela" pageId="9" pageNumber="579">Venezuela</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D56AFF9FF9FE959FB2BBFCB5" pageId="9" pageNumber="579" type="diagnosis">
<paragraph id="01588F4AD56AFF9FF9FE959FB2BBFCB5" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
<emphasis id="33935358D56AFF9FF9FE959FB761FD40" bold="true" box="[199,322,690,716]" pageId="9" pageNumber="579">Diagnosis.</emphasis>
Pronotum without spots, anterolateral pronotal margins not calloused, crenulated, paler than pronotal disc; humeral angles angular elongated, without a posterior tooth, laterally directed. Apex of scutellum paler than disc (
<figureCitation id="99DC93CFD56AFF9FF9ED95D6B70FFC99" box="[212,300,763,789]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Figs 13</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FF80095D6B77BFC99" box="[313,344,763,789]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="9" pageNumber="579">22</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FF85C95D7B7A6FC99" box="[357,389,762,789]" captionStart="FIGURES 55–60" captionStartId="12.[151,264,1290,1314]" captionTargetBox="[151,1436,453,1267]" captionTargetId="figure-150@12.[151,1436,453,1267]" captionTargetPageId="12" captionText="FIGURES 55–60. Live specimens of Podisus graziae sp. n. (55–57), and Podisus thomasi sp. n. (58–60). 55–57, photos by Ricardo Brugnera, Porto Alegre, Brazil. 58–60, photos by Erika Schulz, Mindo, Pichincha, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.4692483" httpUri="https://zenodo.org/record/4692483/files/figure.png" pageId="9" pageNumber="579">58</figureCitation>
). Parameres divided into two arms, each apex acute (
<figureCitation id="99DC93CFD56AFF9FFADB95D6B214FC99" box="[994,1079,763,789]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="9" pageNumber="579">Fig. 38</figureCitation>
). Posterior margins of valvifers VIII slightly sinuous, posterior angles obtuse, lateral angles over laterotergites IX (
<figureCitation id="99DC93CFD56AFF9FFD0E9432B2A9FCB5" box="[1079,1162,798,825]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="9" pageNumber="579">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D56AFF9DF9FE946FB2C8F8D2" lastPageId="11" lastPageNumber="581" pageId="9" pageNumber="579" type="description">
<paragraph id="01588F4AD56AFF9FF9FE946FB5D1FCD1" blockId="9.[151,1437,150,1978]" box="[199,1010,834,861]" pageId="9" pageNumber="579">
<emphasis id="33935358D56AFF9FF9FE946FB77AFCD0" bold="true" box="[199,345,834,860]" pageId="9" pageNumber="579">Description.</emphasis>
Body length. 10.30–11.08 mm (
<specimenCount id="17E144C3D56AFF9FFBFE946EB4FAFCD1" box="[711,729,835,861]" pageId="9" pageNumber="579" type="female">♀</specimenCount>
) and 8.50–9.40 mm (
<specimenCount id="17E144C3D56AFF9FFAF4946EB5C7FCD1" box="[973,996,835,861]" pageId="9" pageNumber="579" type="male">♂</specimenCount>
).
</paragraph>
<paragraph id="01588F4AD56AFF9FF9FE944AB6CDFC61" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
Body castaneous to brown, anterior half of lateral margins of pronotum lighter than rest of pronotum, humeral angles slightly darker than disc, apex of scutellum pale, lighter than disc. A thin central line mostly without punctures or lighter than rest of dorsum extends from anterior margin of pronotum to posterior margin of scutellum (
<figureCitation id="99DC93CFD56AFF9FFC5D9482B696FC61" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Figs. 13</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FF9F894FEB6C2FC61" box="[193,225,979,1005]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="9" pageNumber="579">22</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD56AFF9FF9FE94DAB7E2FB2D" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
Head subrectangular, mandibular plates uniformly punctured, clypeus as long as or slightly surpassing apex of mandibular plates; ocelli lying behind an imaginary line through posterior margins of compound eyes; antennal tubercles partially visible from above, proportion of antennomeres: I&lt;II&gt;III&lt;IV&gt;V (
<figureCitation id="99DC93CFD56AFF9FFD629312B299FBD5" box="[1115,1210,1087,1113]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Figs. 13</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FFDF09312B2CAFBD5" box="[1225,1257,1087,1113]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="9" pageNumber="579">22</figureCitation>
); bucculae evanescent posteriorly, apex rounded; labium extending to posterior margin of metasternum, proportion of labiomeres: I&lt;II&gt;III&gt;IV (
<figureCitation id="99DC93CFD56AFF9FF81293AAB7ABFB2D" box="[299,392,1159,1185]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Figs. 14</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FF8AD93AAB797FB2D" box="[404,436,1159,1185]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="9" pageNumber="579">23</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD56AFF9FF9FE9386B7E7F985" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
Pronotum hexagonal, uniformly punctured; cicatrices flat; anterior and lateral margins concave in dorsal view, lateral margins crenulated and lighter than disc on anterior half; humeral angles angular, slightly produced, darker than disc. Scutellum longer than wide, reaching an imaginary line connecting fifth connexives medially, apex pale, lighter than disc; frenal margins longer than post frenal margins. Coria longer than scutellum, each attaining abdominal segment IV, uniformly punctured; hemelytral membranes well surpassing apex of abdomen, with apex spotted in brown (
<figureCitation id="99DC93CFD56AFF9FF8599272B795FAF5" box="[352,438,1375,1401]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Figs 13</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FF8FB9272B428FAF5" box="[450,523,1375,1401]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="9" pageNumber="579">22, 24</figureCitation>
). Pro-, meso- and metasterna covered by small thin setae; prosternum with a weak median carina; mesosternum carinated longitudinally; metasternum flat. Pro-, meso-, and metapleura densely punctured, evaporatoria on posterior margin of each mesopleura; metapleural evaporatoria surrounding each ostiolar peritreme; each ostiolar peritreme discal-type, elongated, with a darker spot on apical angle; each ostiolar opening directed laterally (
<figureCitation id="99DC93CFD56AFF9FF85A92C2B794F985" box="[355,439,1519,1545]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD56AFF9FF9FE913EB328F9DD" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
Abdomen sparsely punctured, posterolateral angles of each connexivum projected as small spines, longer on seventh segment. Mesial tubercle projecting from sternite III not surpassing metacoxae (
<figureCitation id="99DC93CFD56AFF9FFD4C911AB2F1F9DD" box="[1141,1234,1591,1617]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Figs. 14</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FFDE7911AB2DDF9DD" box="[1246,1278,1591,1617]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="9" pageNumber="579">23</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD56AFF9FF9FE9176B5B8F9F9" blockId="9.[151,1437,150,1978]" box="[199,923,1627,1653]" pageId="9" pageNumber="579">
<emphasis id="33935358D56AFF9FF9FE9176B728F9F9" bold="true" box="[199,267,1627,1653]" pageId="9" pageNumber="579">Male.</emphasis>
Without glandular patches on ventral surface of abdomen.
</paragraph>
<paragraph id="01588F4AD56AFF9FF9FE9152B27EF836" blockId="9.[151,1437,150,1978]" pageId="9" pageNumber="579">
Genitalia. Pygophore bowl-shaped, opened posteriorly, setose, setae denser on ventral rim (
<figureCitation id="99DC93CFD56AFF9FFDFB9152B367F915" box="[1218,1348,1663,1689]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Figs 31–33</figureCitation>
); dorsal rim concave, with 1+1 dorsal projections near segment X (
<figureCitation id="99DC93CFD56AFF9FFA0E918EB5A8F931" box="[823,907,1699,1725]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Fig. 31</figureCitation>
, dr, dp); ventral rim concave, with 1+1 inconspicuous ventral projections near posterolateral angles (
<figureCitation id="99DC93CFD56AFF9FFA3191EAB5A9F96D" box="[776,906,1735,1761]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Figs 32–33</figureCitation>
, vr, vp, pa), surface between inferior and superior layers of ventral rim excavated (
<figureCitation id="99DC93CFD56AFF9FFB0B91C6B490F889" box="[562,691,1771,1797]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Figs 32–33</figureCitation>
, il, sl, vr). Posterolateral angles acute (
<figureCitation id="99DC93CFD56AFF9FFD5E91C6B2CBF889" box="[1127,1256,1771,1797]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Figs 31–33</figureCitation>
, pa), surpassing seventh tergite (
<figureCitation id="99DC93CFD56AFF9FF8729022B7BCF8A5" box="[331,415,1807,1833]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="9" pageNumber="579">Fig. 14</figureCitation>
). Segment X tubular, ventrally directed. Head of parameres divided into two distinct arms, each apex acute, well visible posteriorly (
<figureCitation id="99DC93CFD56AFF9FFB49901EB4DAF8C1" box="[624,761,1843,1869]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Figs. 31–33</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FFA3F901EB507F8C1" box="[774,804,1843,1869]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="9" pageNumber="579">38</figureCitation>
, par). Pseudoclasper with transversal elevations, placed dorsolaterally to parameres (
<figureCitation id="99DC93CFD56AFF9FF8E3907AB441F8FD" box="[474,610,1879,1905]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="9" pageNumber="579">Figs. 31–33</figureCitation>
, 
<figureCitation id="99DC93CFD56AFF9FFB56907AB4AEF8FD" box="[623,653,1879,1905]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="9" pageNumber="579">39</figureCitation>
, pc). Phallus. Basal theca and thecal shield subequal in length, thecal shield slightly wider than basal theca (
<figureCitation id="99DC93CFD56AFF9FFB7F9056B4EEF819" box="[582,717,1915,1941]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="9" pageNumber="579">Figs. 46–48</figureCitation>
, bt, ts); vesica with two elongated arms, golf club-shaped in lateral view (
<figureCitation id="99DC93CFD56AFF9FF83790B2B743F835" box="[270,352,1951,1977]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="9" pageNumber="579">Fig. 47</figureCitation>
, v); apices of conjunctival lobes slightly sculptured (
<figureCitation id="99DC93CFD56AFF9FFA8990B2B20EF835" box="[944,1069,1951,1977]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="9" pageNumber="579">Fig. 46–48</figureCitation>
, cl).
</paragraph>
<caption id="5598DFC2D569FF9CF9AE90BAB45FF87B" ID-DOI="http://doi.org/10.5281/zenodo.4692479" ID-Zenodo-Dep="4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="10" pageNumber="580" startId="10.[151,264,1943,1967]" targetBox="[260,1315,181,1919]" targetPageId="10">
<paragraph id="01588F4AD569FF9CF9AE90BAB45FF87B" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="580">
<emphasis id="33935358D569FF9CF9AE90BAB777F823" bold="true" box="[151,340,1943,1967]" pageId="10" pageNumber="580">FIGURES 40–48.</emphasis>
Phallus of 
<taxonomicName id="C6E7F4C9D569FF9CF8FF90B5B456F83C" box="[454,629,1944,1968]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="580" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D569FF9CF8FF90B5B456F83C" box="[454,629,1944,1968]" italics="true" pageId="10" pageNumber="580">Podisus ventralis</emphasis>
</taxonomicName>
(40–42), 
<taxonomicName id="C6E7F4C9D569FF9CFBE190B5B559F823" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[728,890,1944,1968]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="580" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="33935358D569FF9CFBE190B5B559F823" box="[728,890,1944,1968]" italics="true" pageId="10" pageNumber="580">Podisus graziae</emphasis>
</taxonomicName>
<emphasis id="33935358D569FF9CFAB990B5B59AF83C" bold="true" box="[896,953,1944,1968]" pageId="10" pageNumber="580">
<taxonomicNameLabel id="28A0EE23D569FF9CFAB990B5B59AF83C" box="[896,953,1944,1968]" pageId="10" pageNumber="580" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(43–45), and 
<taxonomicName id="C6E7F4C9D569FF9CFD7F90B5B2CEF83C" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[1094,1261,1944,1968]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="580" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="33935358D569FF9CFD7F90B5B2CEF83C" box="[1094,1261,1944,1968]" italics="true" pageId="10" pageNumber="580">Podisus thomasi</emphasis>
</taxonomicName>
<emphasis id="33935358D569FF9CFDCA90B5B30FF83C" bold="true" box="[1267,1324,1944,1968]" pageId="10" pageNumber="580">
<taxonomicNameLabel id="28A0EE23D569FF9CFDCA90B5B30FF83C" box="[1267,1324,1944,1968]" pageId="10" pageNumber="580" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="5598DFC2D568FF9DF9AE93CCB55EFA04" ID-DOI="http://doi.org/10.5281/zenodo.4692481" ID-Zenodo-Dep="4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581" startId="11.[151,264,1249,1273]" subCaptionStartIDs="11.[683,831,1285,1309]" subCaptionStarts="Abbr" targetBox="[151,1435,181,1224]" targetPageId="11">
<paragraph id="01588F4AD568FF9DF9AE93CCB55EFA04" blockId="11.[151,1437,1249,1417]" pageId="11" pageNumber="581">
<emphasis id="33935358D568FF9DF9AE93CCB770FB75" bold="true" box="[151,339,1249,1273]" pageId="11" pageNumber="581">FIGURES 49–54.</emphasis>
External (49, 51, 53) and internal (50, 52, 54) female genitalia of 
<taxonomicName id="C6E7F4C9D568FF9DFAD393CCB2BAFB75" box="[1002,1177,1249,1273]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D568FF9DFAD393CCB2BAFB75" box="[1002,1177,1249,1273]" italics="true" pageId="11" pageNumber="581">Podisus ventralis</emphasis>
</taxonomicName>
(49–50), 
<taxonomicName id="C6E7F4C9D568FF9DFDC393CCB3BFFB75" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[1274,1436,1249,1273]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">Podisus graziae</taxonomicName>
<emphasis id="33935358D568FF9DF9AE9228B6F2FA91" bold="true" box="[151,209,1285,1309]" pageId="11" pageNumber="581">sp. n.</emphasis>
(51–52), and 
<taxonomicName id="C6E7F4C9D568FF9DF8589228B42BFA91" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[353,520,1285,1309]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="33935358D568FF9DF8589228B42BFA91" box="[353,520,1285,1309]" italics="true" pageId="11" pageNumber="581">Podisus thomasi</emphasis>
</taxonomicName>
<emphasis id="33935358D568FF9DFB369228B46BFA91" bold="true" box="[527,584,1285,1309]" pageId="11" pageNumber="581">
<taxonomicNameLabel id="28A0EE23D568FF9DFB369228B46BFA91" box="[527,584,1285,1309]" pageId="11" pageNumber="581" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la 
<subScript id="9D638D0FD568FF9DF8189214B70BFACB" attach="left" box="[289,296,1337,1351]" fontSize="6" pageId="11" pageNumber="581">9</subScript>
, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 
<subScript id="9D638D0FD568FF9DFB579270B456FAE7" attach="left" box="[622,629,1373,1387]" fontSize="6" pageId="11" pageNumber="581">9</subScript>
, thickening of valvulae IX; va, vesicular area; vf 
<subScript id="9D638D0FD568FF9DFD559270B250FAE7" attach="left" box="[1132,1139,1373,1387]" fontSize="6" pageId="11" pageNumber="581">8</subScript>
, valvifers VIII; vf 
<subScript id="9D638D0FD568FF9DFC169270B315FAE7" attach="left" box="[1327,1334,1373,1387]" fontSize="6" pageId="11" pageNumber="581">9</subScript>
, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm.
</paragraph>
</caption>
<paragraph id="01588F4AD568FF9DF9FE929AB2C8F8D2" blockId="11.[151,1437,1463,2030]" pageId="11" pageNumber="581">
<emphasis id="33935358D568FF9DF9FE929AB706FA5D" bold="true" box="[199,293,1463,1489]" pageId="11" pageNumber="581">Female.</emphasis>
Genitalia. Genital plates positioned ventroposteriorly (
<figureCitation id="99DC93CFD568FF9DFAA9929AB5C7FA5D" box="[912,996,1463,1489]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="11" pageNumber="581">Fig. 23</figureCitation>
); valvifers VIII oval, posterior margins sinuous, posterior angles obtuse, sutural margins straight, juxtaposed, lateral angles over laterotergites IX (
<figureCitation id="99DC93CFD568FF9DFC7C92F6B3B4FA7A" box="[1349,1431,1499,1526]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 53</figureCitation>
, vf 
<subScript id="9D638D0FD568FF9DF996913FB694F9AC" attach="right" box="[175,183,1554,1568]" fontSize="6" pageId="11" pageNumber="581">8</subScript>
). Laterotergites VIII triangular with spiracles on proximal angle (
<figureCitation id="99DC93CFD568FF9DFA9992D2B5D0F996" box="[928,1011,1535,1562]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 53</figureCitation>
, la 
<subScript id="9D638D0FD568FF9DFD2D913FB23FF9AC" attach="left" box="[1044,1052,1554,1568]" fontSize="6" pageId="11" pageNumber="581">8</subScript>
). Exposed portion of valvifers IX rectangular, wider than long (
<figureCitation id="99DC93CFD568FF9DF8E7910EB413F9B2" box="[478,560,1571,1598]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 53</figureCitation>
, vf 
<subScript id="9D638D0FD568FF9DFB6B911BB479F9C8" attach="left" box="[594,602,1590,1604]" fontSize="6" pageId="11" pageNumber="581">9</subScript>
). Exposed portion of laterotergites IX tongue-like (
<figureCitation id="99DC93CFD568FF9DFDAF910EB2CBF9B2" box="[1174,1256,1571,1598]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 53</figureCitation>
, la 
<subScript id="9D638D0FD568FF9DFC3E911BB32CF9C8" attach="left" box="[1287,1295,1590,1604]" fontSize="6" pageId="11" pageNumber="581">9</subScript>
), surpassing segment X, and almost attaining mediotergite VIII. Segment X subrectangular (
<figureCitation id="99DC93CFD568FF9DFD2F916AB24AF9EE" box="[1046,1129,1607,1634]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 53</figureCitation>
, X). Inner portion of valvifers IX projected in 1+1 straight elongated arms with acute apices (
<figureCitation id="99DC93CFD568FF9DFA449146B5EDF90A" box="[893,974,1643,1670]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, vf 
<subScript id="9D638D0FD568FF9DFAC89153B5DAF900" attach="right" box="[1009,1017,1662,1676]" fontSize="6" pageId="11" pageNumber="581">9</subScript>
). Valvulae IX with 1+1 small circular secondary thickenings. Ring sclerites inconspicuous, small, circular, placed posteriorly to thickening of vaginal intima; thickening of vaginal intima short, obtuse (
<figureCitation id="99DC93CFD568FF9DFBFE919EB53BF942" box="[711,792,1715,1742]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, tvi); arcus placed right anteriorly of thickening of vaginal intima (
<figureCitation id="99DC93CFD568FF9DF9D691FAB761F97E" box="[239,322,1751,1778]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, ar). Vesicular area almost three times the length of pars intermedialis plus capsula seminalis (
<figureCitation id="99DC93CFD568FF9DFC5691FAB694F89A" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
); Inner and median ducts of vesicular areas of uniform diameter (
<figureCitation id="99DC93CFD568FF9DFAAC91D6B5CBF89A" box="[917,1000,1787,1814]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, id, md), outer duct constricted basally (
<figureCitation id="99DC93CFD568FF9DF9999032B6D1F8B6" box="[160,242,1823,1850]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, od). Pars intermedialis tubular, shorter and narrower than capsula seminalis; capsula seminalis oval, elongated, smooth (
<figureCitation id="99DC93CFD568FF9DF87D906EB7B5F8D2" box="[324,406,1859,1886]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, pi, cs). Proximal and distal annular crests of similar width (
<figureCitation id="99DC93CFD568FF9DFD00906EB2A8F8D2" box="[1081,1163,1859,1886]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="11" pageNumber="581">Fig. 54</figureCitation>
, pa, da).
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D568FF9DF9FE904AB50CF80D" box="[199,815,1895,1922]" pageId="11" pageNumber="581" type="distribution">
<paragraph id="01588F4AD568FF9DF9FE904AB50CF80D" blockId="11.[151,1437,1463,2030]" box="[199,815,1895,1922]" pageId="11" pageNumber="581">
<emphasis id="33935358D568FF9DF9FE904AB77AF80D" bold="true" box="[199,345,1895,1921]" pageId="11" pageNumber="581">Distribution</emphasis>
: 
<collectingCountry id="79F0CFDAD568FF9DF8519045B7E7F80E" box="[360,452,1896,1922]" name="Panama" pageId="11" pageNumber="581">Panama</collectingCountry>
, 
<collectingCountry id="79F0CFDAD568FF9DF8F7904AB467F80D" box="[462,580,1895,1921]" name="Venezuela" pageId="11" pageNumber="581">Venezuela</collectingCountry>
, 
<collectingCountry id="79F0CFDAD568FF9DFB76904AB4E3F80D" box="[591,704,1895,1921]" name="Colombia" pageId="11" pageNumber="581">Colombia</collectingCountry>
, 
<collectingCountry id="79F0CFDAD568FF9DFBF5904AB508F80D" box="[716,811,1895,1921]" name="Ecuador" pageId="11" pageNumber="581">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D568FF9AF9FE90A6B4CAFE21" lastPageId="12" lastPageNumber="582" pageId="11" pageNumber="581" type="discussion">
<paragraph id="01588F4AD568FF9AF9FE90A6B7F8FF75" blockId="11.[151,1437,1463,2030]" lastBlockId="12.[151,1437,151,429]" lastPageId="12" lastPageNumber="582" pageId="11" pageNumber="581">
<emphasis id="33935358D568FF9DF9FE90A6B718F829" bold="true" box="[199,315,1931,1957]" pageId="11" pageNumber="581">Remarks.</emphasis>
<bibRefCitation id="6576F2BBD568FF9DF87B90A6B7CDF829" author="Thomas, D. B." box="[322,494,1931,1957]" pageId="11" pageNumber="581" refId="ref10128" refString="Thomas, D. B. (1992) Taxonomic Synopsis of the Asopine Pentatomidae (Heteroptera) of the Western Hemisphere. Entomological Society of America, Lanham, Maryland, 156 pp." type="book" year="1992">Thomas (1992)</bibRefCitation>
examined the 
<typeStatus id="DE5C31E8D568FF9DFBAB90A1B4CAF82A" box="[658,745,1932,1958]" pageId="11" pageNumber="581" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="C6E7F4C9D568FF9DFA3790A6B5ACF829" box="[782,911,1931,1957]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D568FF9DFA3790A6B5ACF829" box="[782,911,1931,1957]" italics="true" pageId="11" pageNumber="581">A. ventralis</emphasis>
</taxonomicName>
but was unable to find any type for 
<taxonomicName id="C6E7F4C9D568FF9DFC1D90A6B3BEF829" box="[1316,1437,1931,1957]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="33935358D568FF9DFC1D90A6B3BEF829" box="[1316,1437,1931,1957]" italics="true" pageId="11" pageNumber="581">A. pallipes</emphasis>
</taxonomicName>
and donated to NHMUK a male specimen that he designated as the 
<typeStatus id="DE5C31E8D568FF9DFABF909DB5FCF846" box="[902,991,1968,1994]" pageId="11" pageNumber="581" type="neotype">neotype</typeStatus>
of this species. While cataloguing typespecimens of 
<taxonomicName id="C6E7F4C9D568FF9DF80890FEB7BDF861" authorityName="Spinola" authorityYear="1850" box="[305,414,2003,2029]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="583" phylum="Arthropoda" rank="subFamily" subFamily="Asopinae">Asopinae</taxonomicName>
, Roell 
<emphasis id="33935358D568FF9DF8D790F9B404F861" box="[494,551,2003,2029]" italics="true" pageId="11" pageNumber="581">et al.</emphasis>
(in prep.) found an actual 
<typeStatus id="DE5C31E8D568FF9DFA6C90F9B58FF862" box="[853,940,2004,2030]" pageId="11" pageNumber="581" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="C6E7F4C9D568FF9DFAEA90FEB26FF861" box="[979,1100,2003,2029]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="33935358D568FF9DFAEA90FEB26FF861" box="[979,1100,2003,2029]" italics="true" pageId="11" pageNumber="581">A. pallipes</emphasis>
</taxonomicName>
(
<figureCitation id="99DC93CFD568FF9DFD6290FEB2E1F862" box="[1115,1218,2003,2030]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="11" pageNumber="581">Figs 4–6</figureCitation>
), the assessment of that specimen as a type is detailed in their catalogue; Thomas’s 
<typeStatus id="DE5C31E8D56FFF9AFA5E97BAB5E3FF3D" box="[871,960,151,177]" pageId="12" pageNumber="582" type="neotype">neotype</typeStatus>
is therefore hereby set aside (
<bibRefCitation id="6576F2BBD56FFF9AFC2B97BAB3BBFF3D" author="ICZN" box="[1298,1432,151,177]" pageId="12" pageNumber="582" refId="ref9299" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Art. 75.8) and we have described, in this work, a new species for this specimen, based also on the study of several other conspecific specimens.
</paragraph>
<paragraph id="01588F4AD56FFF9AF9FE962EB4CAFE21" blockId="12.[151,1437,151,429]" pageId="12" pageNumber="582">
<taxonomicName id="C6E7F4C9D56FFF9AF9FE962EB75CFE91" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[199,383,259,285]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="33935358D56FFF9AF9FE962EB75CFE91" box="[199,383,259,285]" italics="true" pageId="12" pageNumber="582">Podisus thomasi</emphasis>
</taxonomicName>
<emphasis id="33935358D56FFF9AF8BC962EB7E7FE91" bold="true" box="[389,452,259,285]" pageId="12" pageNumber="582">
<taxonomicNameLabel id="28A0EE23D56FFF9AF8BC962EB7E7FE91" box="[389,452,259,285]" pageId="12" pageNumber="582" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be differentiated from 
<taxonomicName id="C6E7F4C9D56FFF9AFBC8962EB591FE91" box="[753,946,259,285]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56FFF9AFBC8962EB591FE91" box="[753,946,259,285]" italics="true" pageId="12" pageNumber="582">Podisus ventralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="C6E7F4C9D56FFF9AFADE962EB2B9FE90" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[999,1178,259,285]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="33935358D56FFF9AFADE962EB2B9FE90" box="[999,1178,259,285]" italics="true" pageId="12" pageNumber="582">Podisus graziae</emphasis>
</taxonomicName>
<emphasis id="33935358D56FFF9AFD99962EB2FCFE91" bold="true" box="[1184,1247,259,285]" pageId="12" pageNumber="582">
<taxonomicNameLabel id="28A0EE23D56FFF9AFD99962EB2FCFE91" box="[1184,1247,259,285]" pageId="12" pageNumber="582" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the following characteristics: apex of scutellum yellowish spotted (
<figureCitation id="99DC93CFD56FFF9AFBD3960AB564FECD" box="[746,839,295,321]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="12" pageNumber="582">Figs. 13</figureCitation>
, 
<figureCitation id="99DC93CFD56FFF9AFA6D960AB550FECD" box="[852,883,295,321]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="12" pageNumber="582">22</figureCitation>
, 
<figureCitation id="99DC93CFD56FFF9AFA47960BB5E5FECD" box="[894,966,294,321]" captionStart="FIGURES 55–60" captionStartId="12.[151,264,1290,1314]" captionTargetBox="[151,1436,453,1267]" captionTargetId="figure-150@12.[151,1436,453,1267]" captionTargetPageId="12" captionText="FIGURES 55–60. Live specimens of Podisus graziae sp. n. (55–57), and Podisus thomasi sp. n. (58–60). 55–57, photos by Ricardo Brugnera, Porto Alegre, Brazil. 58–60, photos by Erika Schulz, Mindo, Pichincha, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.4692483" httpUri="https://zenodo.org/record/4692483/files/figure.png" pageId="12" pageNumber="582">58–60</figureCitation>
) [concolourous with the disc in 
<taxonomicName id="C6E7F4C9D56FFF9AFC09960AB692FEE9" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56FFF9AFC09960AB692FEE9" italics="true" pageId="12" pageNumber="582">P. ventralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="C6E7F4C9D56FFF9AF9D29666B77EFEE8" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[235,349,331,356]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56FFF9AF9D29666B77EFEE8" box="[235,349,331,356]" italics="true" pageId="12" pageNumber="582">P. graziae</emphasis>
</taxonomicName>
], apices of parameres acute (
<figureCitation id="99DC93CFD56FFF9AFB959666B4DCFEE9" box="[684,767,331,357]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="12" pageNumber="582">Fig. 38</figureCitation>
) [obtuse in 
<taxonomicName id="C6E7F4C9D56FFF9AFAB09666B22AFEE9" box="[905,1033,331,357]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56FFF9AFAB09666B22AFEE9" box="[905,1033,331,357]" italics="true" pageId="12" pageNumber="582">P. ventralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="C6E7F4C9D56FFF9AFD7B9666B297FEE8" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[1090,1204,331,356]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56FFF9AFD7B9666B297FEE8" box="[1090,1204,331,356]" italics="true" pageId="12" pageNumber="582">P. graziae</emphasis>
</taxonomicName>
], and pseudoclasper subrectangular (
<figureCitation id="99DC93CFD56FFF9AF8729642B7BEFE05" box="[331,413,367,393]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="12" pageNumber="582">Fig. 38</figureCitation>
) [suboval in 
<taxonomicName id="C6E7F4C9D56FFF9AFB089642B48CFE05" box="[561,687,367,393]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56FFF9AFB089642B48CFE05" box="[561,687,367,393]" italics="true" pageId="12" pageNumber="582">P. ventralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="C6E7F4C9D56FFF9AFBDF9642B576FE04" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[742,853,367,392]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56FFF9AFBDF9642B576FE04" box="[742,853,367,392]" italics="true" pageId="12" pageNumber="582">P. graziae</emphasis>
</taxonomicName>
]. 
<taxonomicName id="C6E7F4C9D56FFF9AFA549642B5C2FE05" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[877,993,367,393]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis id="33935358D56FFF9AFA549642B5C2FE05" box="[877,993,367,393]" italics="true" pageId="12" pageNumber="582">P. thomasi</emphasis>
</taxonomicName>
is sympatric with 
<taxonomicName id="C6E7F4C9D56FFF9AFD889642B30DFE05" box="[1201,1326,367,393]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="582" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56FFF9AFD889642B30DFE05" box="[1201,1326,367,393]" italics="true" pageId="12" pageNumber="582">P. ventralis</emphasis>
</taxonomicName>
. Both are distributed in the northern regions of South America.
</paragraph>
</subSubSection>
</treatment>
</document>