1396 lines
151 KiB
XML
1396 lines
151 KiB
XML
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<mods:title id="BFD0D4E4917FDFC3A7090F2878A5800D">Old World Micropholcus spiders, with first records of acrocerid parasitoids in Pholcidae (Araneae)</mods:title>
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<mods:namePart id="2557E80C86AC1B8A54E4DDE077C6E322">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="A9D3B8B6F6019B08F322FBD60BF51447">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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<mods:namePart id="252BA3781478EB34BFB5BA9ECE21FCC6">Meng, Guanliang</mods:namePart>
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<mods:affiliation id="678CFB52FF22D6200D61C4E5305F4621">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany</mods:affiliation>
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<mods:title id="82F239EA3830D5712671F10D486CF8CB">ZooKeys</mods:title>
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<mods:date id="FB5989A2515D53A85B2AC27A0573F1D0">2024</mods:date>
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<mods:number id="BA00E83BB01513FA8D3B44ACE82CE632">2024-09-26</mods:number>
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<mods:number id="0F59F9A34D9DE5E88CF8A178EFF41895">1213</mods:number>
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<subSubSection id="C4A2E2A6EC3DDBDF7A946AFE777FBF73" type="nomenclature">
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<paragraph id="211FEF55E8A11DB486C491D24A5CA072">
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<heading id="E8F32436E6468D0D466ED36208E72D80" reason="label">Genus</heading>
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<taxonomicName id="15D95C6DB98FA54E6FFBAE43DADDA59C" ID-CoL="5S7N" authority="Deeleman-Reinhold & Prinsen, 1987" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
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<emphasis id="358D58452202652A5CB262360850001E" italics="true">Micropholcus</emphasis>
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<bibRefCitation id="55E238D2FD56670172E2D01718169992" author="Deeleman-Reinhold CL & Prinsen JD" issue="5" journalOrPublisher="Entomologische Berichten, Amsterdam" pagination="73-77" refId="B8" refString="Deeleman-Reinhold CL, Prinsen JD (1987) Micropholcus fauroti (Simon) n. comb., a pantropical, synanthropic spider (Araneae: Pholcidae). Entomologische Berichten, Amsterdam 47 (5): 73 - 77. https: // natuurtijdschriften. nl / pub / 1012616 / EB 1987047005003. pdf" title="Micropholcus fauroti (Simon) n. comb., a pantropical, synanthropic spider (Araneae: Pholcidae)." url="https://natuurtijdschriften.nl/pub/1012616/EB1987047005003.pdf" volume="47" year="1987">Deeleman-Reinhold & Prinsen, 1987</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="5046CBE549E6F279A4B7E1A6910C9998" type="reference_group">
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<paragraph id="C5E44D1A40E5CE5283479A1DA3ABF472">
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<treatmentCitationGroup id="A341634E4D94E49800DDD8233E70FB5F">
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<treatmentCitation id="A1896D16A48F9E5B791BA427D1634EEB" author="Deeleman-Reinhold CL & Prinsen JD" page="73" year="1987">
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<taxonomicName id="AA19DE4AD2A7567859AF73D744922B2F" ID-CoL="5S7N" authority="Deeleman-Reinhold & Prinsen, 1987: 73" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityPageNumber="73" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
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<emphasis id="36CD2246580220D8F1D2FEF7CD935FE6" italics="true">Micropholcus</emphasis>
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<comment id="11FA392AEF330C2CE516DF4A19D9A897">
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<bibRefCitation id="5EBD1496F2297897C87B98AEC5163712" author="Deeleman-Reinhold CL & Prinsen JD" issue="5" journalOrPublisher="Entomologische Berichten, Amsterdam" pagination="73-77" refId="B8" refString="Deeleman-Reinhold CL, Prinsen JD (1987) Micropholcus fauroti (Simon) n. comb., a pantropical, synanthropic spider (Araneae: Pholcidae). Entomologische Berichten, Amsterdam 47 (5): 73 - 77. https: // natuurtijdschriften. nl / pub / 1012616 / EB 1987047005003. pdf" title="Micropholcus fauroti (Simon) n. comb., a pantropical, synanthropic spider (Araneae: Pholcidae)." url="https://natuurtijdschriften.nl/pub/1012616/EB1987047005003.pdf" volume="47" year="1987">Deeleman-Reinhold & Prinsen, 1987: 73</bibRefCitation>
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; type species:
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<taxonomicName id="9258CE042DA89E8C3D35BA06D6A42F35" authority="Simon, 1887" authorityName="Simon" authorityYear="1887" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fauroti">
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<emphasis id="5A7CAB924B00B130FAC1AA642AEF3695" italics="true">Pholcus fauroti</emphasis>
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<treatmentCitation id="856EE16C4244D9E1137D35FE87EAF12A">
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<bibRefCitation id="99D5BC323FB312686611EF1935BF41E7" author="Simon E" journalOrPublisher="Bulletin de la Société Zoologique de France" pagination="452-455" refId="B45" refString="Simon E (1887) Arachnides recueillis a Obock en 1886 par M. le Dr L. Faurot. Bulletin de la Societe Zoologique de France 12: 452 - 455." title="Arachnides recueillis à Obock en 1886 par M. le Dr L. Faurot." volume="12" year="1887">Simon, 1887</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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||
.
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||
</comment>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="DC9F583803113AEFDDF3A6C501048995">
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<treatmentCitationGroup id="E6D4119D6755A97E3350FDFF86CDDFEC">
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<taxonomicName id="7A684A671FF1EA0DF7365BA41DBEF3EC" ID-CoL="5S7N" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
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<emphasis id="3353EB9CDE30FCA7861DD5ECF3C10847" italics="true">Micropholcus</emphasis>
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</taxonomicName>
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<comment id="3BAD7A21DB6A9683322DAE20EB1BAA8F">
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–
|
||
<treatmentCitation id="E3DF3BFE77C608FD66A01AC4C4429F3D" author="Deeleman-Reinhold CL & van Harten A" page="199" year="2001">
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<bibRefCitation id="67CAAC7CCC6DD348E6CFF3EC86CE2A32" author="Deeleman-Reinhold CL & van Harten A" editor="Ishwar Prakash" firstAuthor="Deeleman-Reinhold" journalOrPublisher="Scientific Publishers, India, Jodhpur" page="199" pagination="193-207" refId="B9" refString="Deeleman-Reinhold CL, van Harten A (2001) Description of some interesting, new or little known Pholcidae (Araneae) from Yemen. In: Ishwar Prakash (Ed.) Ecology of Desert Environments. Scientific Publishers, India, Jodhpur, 193–207." title="Description of some interesting, new or little known Pholcidae (Araneae) from Yemen." volumeTitle="Ecology of Desert Environments." year="2001">Deeleman-Reinhold and van Harten 2001: 199</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="0A5BEBCBB45D102DDFD9C10590697413" author="Huber BA" page="24" year="2011">
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||
<bibRefCitation id="C336E4D5EAF31ECB94DD18AF5D575D87" author="Huber BA" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" page="24" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011: 24</bibRefCitation>
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</treatmentCitation>
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.
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||
<treatmentCitation id="EFF209053204EBE4E857B56C788FDD5A" author="Huber BA & Carvalho LS & Benjamin SP" page="435" year="2014">
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<bibRefCitation id="A2B9BF3094CCB93963EA7101240149CA" DOI="10.1071/IS13050" author="Huber BA & Carvalho LS & Benjamin SP" etAl="et al." firstAuthor="Huber" journalOrPublisher="Invertebrate Systematics" page="435" pagination="432-450" refId="B28" refString="Huber BA, Carvalho LS, Benjamin SP (2014) On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae). Invertebrate Systematics 28: 432–450. https://doi.org/10.1071/IS13050" title="On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae)." volume="28" year="2014">Huber et al. 2014: 435</bibRefCitation>
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||
</treatmentCitation>
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.
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</comment>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D079F38ED30C41A64B8829F5CD7703AB">
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<treatmentCitationGroup id="EF6EE83C2F016352BCCBAA4C91968738">
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<treatmentCitation id="ABED0AD62EEEAF7B13A5F7C6EC8D2164">
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<taxonomicName id="E92D5554359D614D545BA41333F729C1" authority="Gonzalez-Sponga, 2004: 66" authorityName="Gonzalez-Sponga" authorityPageNumber="66" authorityYear="2004" class="Arachnida" family="Pholcidae" genus="Mariguitaia" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DB3245A70B2BE601DDD1EA56D726FECA" italics="true">Mariguitaia</emphasis>
|
||
<bibRefCitation id="C138B44533AF38597010D39955040D6F" author="González-Sponga MA" journalOrPublisher="Aula y Ambiente" pagination="63-76" refId="B18" refString="Gonzalez-Sponga MA (2004) Aracnidos de Venezuela. Un nuevo genero y nuevas especies de la familia Pholcidae (Araneae). Aula y Ambiente 8: 63 - 76." title="Arácnidos de Venezuela. Un nuevo género y nuevas especies de la familia Pholcidae (Araneae)." volume="8" year="2004">González-Sponga, 2004: 66</bibRefCitation>
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||
</taxonomicName>
|
||
</treatmentCitation>
|
||
; type species:
|
||
<treatmentCitation id="FFEA726149531C59A92957256C99782E">
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<taxonomicName id="A7BCACD4EE58B0730D1B145329AAE784" authority="Gonzalez-Sponga 2004" authorityName="Gonzalez-Sponga" authorityYear="2004" class="Arachnida" family="Pholcidae" genus="Mariguitaia" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="divergentis">
|
||
<emphasis id="87BA32B27E427893D692BDB409783C90" italics="true">Mariguitaia divergentis</emphasis>
|
||
<bibRefCitation id="0A1583564CD5D3347D7764E97CF9EEDD" author="González-Sponga MA" journalOrPublisher="Aula y Ambiente" pagination="63-76" refId="B18" refString="Gonzalez-Sponga MA (2004) Aracnidos de Venezuela. Un nuevo genero y nuevas especies de la familia Pholcidae (Araneae). Aula y Ambiente 8: 63 - 76." title="Arácnidos de Venezuela. Un nuevo género y nuevas especies de la familia Pholcidae (Araneae)." volume="8" year="2004">Gonzalez-Sponga 2004</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. Synonymised in
|
||
<treatmentCitation id="FDE41830A2B50BA362DFF933EDB79AAF">
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<bibRefCitation id="9E003407DA6D313B4850ED871D8706C5" DOI="10.11646/zootaxa.1970.1.3" author="Huber" firstAuthor="Huber" journalOrPublisher="Zootaxa" pagination="64-68" refId="B20" refString="Huber BA (2009) Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae). Zootaxa 1970: 64–68. https://doi.org/10.11646/zootaxa.1970.1.3" title="Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides Roewer to Filistatidae (Araneae)." volume="1970" year="2009">Huber 2009</bibRefCitation>
|
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</treatmentCitation>
|
||
.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EWEBG" type="diagnosis">
|
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<paragraph id="3C16CC105CF017C893A7A1B435139334">
|
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<heading id="073E1C7A83064B3728ED8365A161F065" reason="title">Diagnosis.</heading>
|
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</paragraph>
|
||
<paragraph id="91987B756ED63F7125B3445C695450DE">
|
||
Old World species are long-legged, eight-eyed pholcids with an oval abdomen (Figs
|
||
<figureCitation id="3CC09EAFC61C1056DBED984E01620A46" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure3" httpUri="https://binary.pensoft.net/fig/1141413">3</figureCitation>
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||
,
|
||
<figureCitation id="FA52DF6822AEF0DF3B336D4072909AF7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Oman, Morocco, and the Philippines A, B M. darbat Huber, sp. nov., male and female with egg-sac from Oman, Dhofar, near Qairoon Hairitti C M. shaat Huber, sp. nov., male from Oman, Dhofar, Shaat sinkhole D, E M. agadir (Huber), male and female with egg-sac from Morocco, Souss-Massa, Paradise Valley F M. ghar Huber, sp. nov., male from Morocco, Fès-Meknès, Kef El Ghar G M. khenifra Huber, Lecigne & Lips, sp. nov., male from Morocco, Béni Mellal-Khénifra, near Sidi Ben Daoud H M. bukidnon Huber, sp. nov., male from Philippines, Mindanao, Blue Water Cave. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure4" httpUri="https://binary.pensoft.net/fig/1141414">4</figureCitation>
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||
); New World species are more variable, sometimes without
|
||
<abbrev id="ABBRID0EEFBG" xlink_title="anterior median eye(s)">AME</abbrev>
|
||
, sometimes with elongate to worm-shaped abdomens. Most known species (except
|
||
<taxonomicName id="1AF10E1AAA0718B7C6FBA83A6A7F1F77" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="D25A912DCFC27D72039C7352BE8A6149" italics="true">M. bukidnon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C06F0F126702B0C78CF4CFD1808A2AC3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) with unique modified hair at tip of male palpal trochanter apophysis (Fig.
|
||
<figureCitation id="6FE3BF22A33B0D40E89460519E089862" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)." figureDoi="10.3897/zookeys.1213.133178.figure9" httpUri="https://binary.pensoft.net/fig/1141419">9 A – D</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="3B1B3EF6198E602CD5A85B0E62D5601B" DOI="10.1206/0003-0090(2000)254<0001:NWPSAP>2.0.CO;2" author="Huber" firstAuthor="Huber" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="1-348" refId="B19" refString="Huber BA (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1–348. https://doi.org/10.1206/0003-0090 (2000) 254 <0001: NWPSAP> 2.0. CO; 2" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." volume="254" year="2000">Huber 2000</bibRefCitation>
|
||
: figs 105, 106;
|
||
<bibRefCitation id="6408C19D2D599A7C7D6B62B31A78595C" DOI="10.1080/00222930601051196" author="Huber and Wunderlich" firstAuthor="Huber" journalOrPublisher="Journal of Natural History" pagination="2341-2360" refId="B26" refString="Huber BA, Wunderlich J (2006) Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae). Journal of Natural History 40: 2341–2360. https://doi.org/10.1080/00222930601051196" title="Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae)." volume="40" year="2006">Huber and Wunderlich 2006</bibRefCitation>
|
||
: figs 4 h, 8 d;
|
||
<bibRefCitation id="52A58D1D11ACBA1243D5E61B0DFC0E97" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
: fig. 95;
|
||
<bibRefCitation id="8A0232A1C4F01775E88B236954139280" DOI="10.1071/IS13050" author="Huber" etAl="et al." firstAuthor="Huber" journalOrPublisher="Invertebrate Systematics" pagination="432-450" refId="B28" refString="Huber BA, Carvalho LS, Benjamin SP (2014) On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae). Invertebrate Systematics 28: 432–450. https://doi.org/10.1071/IS13050" title="On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae)." volume="28" year="2014">Huber et al. 2014</bibRefCitation>
|
||
: figs 32, 57). Male chelicerae with frontal apophyses with modified hairs (Fig.
|
||
<figureCitation id="B1DC8BA83E40DE2ACE5C8FFAF2431BCF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male chelicerae: distal apophyses with modified hairs and total view (D) A, B M. alfara Huber sp. nov. C M. tanomah Huber, sp. nov. D, E M. darbat Huber, sp. nov. F, G M. ghar Huber, sp. nov. H M. bukidnon Huber, sp. nov. Scale bars: 10 µm (A – C, F – H); 100 µm (D); 2 µm (E)." figureDoi="10.3897/zookeys.1213.133178.figure6" httpUri="https://binary.pensoft.net/fig/1141416">6</figureCitation>
|
||
), i. e., similar to putative sister genus
|
||
<taxonomicName id="397877B4AB86C2030E969AEACF8B0C4B" authority="Huber" authorityName="Huber" authorityYear="2018" class="Arachnida" family="Pholcidae" genus="Cantikus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EBB5F0C5D019C7331DBADE48D4541862" italics="true">Cantikus</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
but different from other close relatives (
|
||
<taxonomicName id="404B16DDD11AC7D671CD4C046606010D" authority="Simon" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6B7783C0A7B925724CCD22525B2C0DFF" italics="true">Leptopholcus</emphasis>
|
||
Simon
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8D2450190F242B3AF00CEF8A78A11D33" authority="Deeleman-Reinhold & van Harten" authorityName="Deeleman-Reinhold & van Harten" authorityYear="2001" class="Arachnida" family="Pholcidae" genus="Pehrforsskalia" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E21FB548F683EA969C813832DA944376" italics="true">Pehrforsskalia</emphasis>
|
||
Deeleman-Reinhold & van Harten
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2C086757C34A6FB82BB8B8C05204D80E" authority="Bradley" authorityName="Bradley" authorityYear="1877" class="Arachnida" family="Pholcidae" genus="Micromerys" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F930BA56104D4D9462EBDE2C8C8354A" italics="true">Micromerys</emphasis>
|
||
Bradley
|
||
</taxonomicName>
|
||
). Procursus and bulb morphology highly variable and not diagnostic at genus level (contra
|
||
<bibRefCitation id="5BBF1181CA213257850BF50CBDE38C0C" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
). Females not diagnosable morphologically at genus level (highly variable and similar to closely related genera).
|
||
</paragraph>
|
||
<caption id="EFB7A03A6DB2BE5FC561A455C2F8C2BB" ID-DOI="10.3897/zookeys.1213.133178.figure3" ID-arpha="1B831904-D4A7-5887-8FD7-52FA5A84C675" httpUri="https://binary.pensoft.net/fig/1141413" startId="F3">
|
||
<paragraph id="2BA145FE066956AE13AB5AA577D1FA73">
|
||
<label id="7D55D48122CB87F6CF7BAFCD92C572F2">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="78C1B39787D2B9DB051218606777B416">
|
||
<taxonomicName id="324246084110929DD5298750334F6032" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="729E1721869C799BB0922100AAAD1B7C" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; live specimens from Saudi Arabia
|
||
<emphasis id="C4084F56F20A5DA4352A2E54646B740F" bold="true">A</emphasis>
|
||
<taxonomicName id="1AB54E85D0A243F68184FEDECEFD7FA4" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="dhahran" status="sp. nov.">
|
||
<emphasis id="86713486C45397FAAE7B50D4D63EA767" italics="true">M. dhahran</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="84B4530FC219F6DE8E9358E9AB8ABD9B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from ‘ Asir, W of Dhahran Al Janub
|
||
<emphasis id="7C3B763F6C0FD9F396473062B5408D95" bold="true">B</emphasis>
|
||
<taxonomicName id="D7C386628D73412F6A34BC2B1A5F6239" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="harajah" status="sp. nov.">
|
||
<emphasis id="67E11273484A2364BF17DE3319294D2B" italics="true">M. harajah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A03C8CFAD76F971AC1893CC39F85BB42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female with egg-sac from ‘ Asir, SE of Harajah
|
||
<emphasis id="463C53773D00C4370BC21D6FD8047E0C" bold="true">C, D</emphasis>
|
||
<taxonomicName id="6D2FB3C329CF4154C0EC8C07260D0FA9" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="4F3E52FF6908CDA628E7FFF648868564" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="95C55FF205560D3AB741A61ED0BCC981" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male and female with egg-sac from ‘ Asir, S of Al Fara
|
||
<emphasis id="B45226419A40F7F1390B87FFECCC151D" bold="true">E</emphasis>
|
||
<taxonomicName id="9B0A168A527FA0E74AF5A34438E7622C" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="abha" status="sp. nov.">
|
||
<emphasis id="5200ADCCAFC8DF50984D6EA925C7716B" italics="true">M. abha</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E1C1BF41A727DDD10DF23515D2826CCB" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from ‘ Asir, N of Abha
|
||
<emphasis id="E6C7F255DF9AA0AA2D1E19475E1E98A6" bold="true">F</emphasis>
|
||
<taxonomicName id="798DC1A9780CEEFD0FF4881F69FFD499" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="E4AC10863F34A3594E3F30E58201C4FD" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D9AD626AA371C512CE573CAA99C88C4B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from ‘ Asir, NW of Tanomah
|
||
<emphasis id="B03C17642C0A815D974DF363ECF3EB0B" bold="true">G</emphasis>
|
||
<taxonomicName id="AFD8D95B589255145564EC0BF4816478" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bashayer" status="sp. nov.">
|
||
<emphasis id="2408B01BF141A4D2E6000A600B903C12" italics="true">M. bashayer</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="635D53EFDDC724311A6AACCD471204ED" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female with egg-sac from ‘ Asir, NW of Al Bashayer
|
||
<emphasis id="96EF4C689A1E60C2C95EB5BE6D472009" bold="true">H</emphasis>
|
||
<taxonomicName id="B269D58ED82E7B56331C868A8F2CBDC6" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="maysaan" status="sp. nov.">
|
||
<emphasis id="79E201913EB87849421963B7FC54DDFB" italics="true">M. maysaan</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B04EF55CE8DF66FF1DD9643C9C2AFCC9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from Mecca, NW of Maysaan. Photographs BAH.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="1257718FB7675670079E25BFD92546C6" ID-DOI="10.3897/zookeys.1213.133178.figure4" ID-arpha="392B4B21-26DB-51C0-B1C9-C3BD3CC85BF9" httpUri="https://binary.pensoft.net/fig/1141414" startId="F4">
|
||
<paragraph id="31502D85E840BEEC95186EC6F31022EB">
|
||
<label id="D635388524406C0011C7B34BE62CD325">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="9C7A24649CDB502F3FC40C68D6DB4386">
|
||
<taxonomicName id="DFE46895F4259E4FC31A2BD1A0898373" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="636FA565D0FB534DE647A23B3AC3C058" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; live specimens from Oman, Morocco, and the Philippines
|
||
<emphasis id="1794AFD08231AEDA4EB9C9A4A9D9E2F8" bold="true">A, B</emphasis>
|
||
<taxonomicName id="7288CC89A3401465606A51FB7DD60883" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="90D88EBB7D293EE03621980CDE5E1A38" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A95BC93DED1EA0F033E7F59EC6979C3C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male and female with egg-sac from Oman, Dhofar, near Qairoon Hairitti
|
||
<emphasis id="0496B5246AF7EB5B2D067B448A6E7878" bold="true">C</emphasis>
|
||
<taxonomicName id="4EA0E052B78A23270A9F31D13F864FD7" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="shaat" status="sp. nov.">
|
||
<emphasis id="712EE60FBD232171CB62D4C34177560F" italics="true">M. shaat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="0E9D7AE8C42358B9D85BF20CD521F0C9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from Oman, Dhofar, Shaat sinkhole
|
||
<emphasis id="95549AB6998E36AB3D0C7C28599DF640" bold="true">D, E</emphasis>
|
||
<taxonomicName id="F7EEF8CADD5213A715E3483B672792FD" authority="(Huber)" baseAuthorityName="Huber" baseAuthorityYear="2011" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="agadir">
|
||
<emphasis id="B7B3750DE0BD2174C490F178BD7A525F" italics="true">M. agadir</emphasis>
|
||
(Huber)
|
||
</taxonomicName>
|
||
, male and female with egg-sac from Morocco, Souss-Massa, Paradise Valley
|
||
<emphasis id="492C7E469DAC0069DD251CC9A4A0BD47" bold="true">F</emphasis>
|
||
<taxonomicName id="CBEC13E0DD355D38769A009D3973875D" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="ABF1D00B91DFB567593D5D614419C808" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="514526373BB5BE9993E89CE6D800678B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from Morocco, Fès-Meknès, Kef El Ghar
|
||
<emphasis id="84A656D0A5C0B6B0D1BDD519D377BFA1" bold="true">G</emphasis>
|
||
<taxonomicName id="E80B90731F330E27CB7A19EC54C02E3E" authority="Huber, Lecigne & Lips" authorityName="Huber, Lecigne & Lips" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="khenifra" status="sp. nov.">
|
||
<emphasis id="25BE912E35AF6D784A2783A1F96EA78E" italics="true">M. khenifra</emphasis>
|
||
Huber, Lecigne & Lips
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="21A788FF104DC7BC6554CDA58F175C34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from Morocco, Béni Mellal-Khénifra, near Sidi Ben Daoud
|
||
<emphasis id="0F3D5442325A053442C2F32E271128C8" bold="true">H</emphasis>
|
||
<taxonomicName id="BE394E4D8EDEE59979A650A07564CA8B" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="F8205C8C4C4A7A25C6D0C419408BD9AB" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4ECDD7D6270B70A048D296DE8EDCE675" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male from Philippines, Mindanao, Blue Water Cave. Photographs BAH.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E3NBG" type="notes">
|
||
<paragraph id="07D69B085EF31FA51EB90D2E5339F537">
|
||
<heading id="473646A043226A7D5BCC2190C09C9E47" reason="title">Note.</heading>
|
||
</paragraph>
|
||
<paragraph id="1C919CE45A3CED53D958A7E443319037">
|
||
Most parts in this general section about
|
||
<taxonomicName id="C6B635B6907D9FAA69C8A589BEAA2758" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E07C8B3FE299A4A26E37A3821607B3A3" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
refer to the entire genus. The following description is limited to Old World taxa because they are relatively homogeneous, while some New World species (in particular those on the Caribbean islands) are superficially extremely different from South American (in particular Brazilian) and Old World species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EJOBG" type="description">
|
||
<paragraph id="E689EDFBEF7932F0DA9A170A309A1DDF">
|
||
<heading id="539B5A18F479FD8DE990F005E4086C3E" reason="title">Description</heading>
|
||
</paragraph>
|
||
<paragraph id="0FC3E7E8DAC6C2E239D1B431128313B1">
|
||
<emphasis id="E776476768DF60B764AB9D91ED0C8EC0" bold="true">
|
||
(Old World taxa). Male.
|
||
<emphasis id="085BF9D81F13587B95AB5AA77FBA0805" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Total body length ~ 2.3–3.9. Carapace width 0.8–1.5. Diameter
|
||
<abbrev id="ABBRID0EUOBG" xlink_title="posterior median eye(s)">PME</abbrev>
|
||
60–100 µm; diameter
|
||
<abbrev id="ABBRID0EYOBG" xlink_title="anterior median eye(s)">AME</abbrev>
|
||
usually 35–55 µm, in
|
||
<taxonomicName id="66A8F9993C689FE49BF467A84527CDC6" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="C30BDF2EDF098A202B72C146906331B5" italics="true">M. bukidnon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E365ADE6F94C68A736435FD2C6B0EE99" rank="species">sp. nov.</taxonomicNameLabel>
|
||
only 15 µm. Tibia 1 length 5.0–10.2. Tibia 1
|
||
<abbrev id="ABBRID0EHPBG" xlink_title="length/diameter">L / d</abbrev>
|
||
: 57–85. Leg formula 1243. Diameters of leg femora (at half length) 80–150 µm, of leg tibiae 75–120 µm.
|
||
</paragraph>
|
||
<paragraph id="856BF98068B73AB9FDAAC825070FCBBA">
|
||
<emphasis id="3AAC2EBC6E5E2149B6C5533D4552B651" bold="true" italics="true">Colour</emphasis>
|
||
(in ethanol). Prosoma and legs pale ochre-yellow to grey, carapace with dark pattern, legs with darker patellae and tibia-metatarsus joints; abdomen ochre-grey to whitish, monochromous or with whitish marks. Live specimens (Figs
|
||
<figureCitation id="D73A3D0009C9755E96FD198A0E987460" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure3" httpUri="https://binary.pensoft.net/fig/1141413">3</figureCitation>
|
||
,
|
||
<figureCitation id="AD27B0A7DC575A936D499FBDA54D4E6D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Oman, Morocco, and the Philippines A, B M. darbat Huber, sp. nov., male and female with egg-sac from Oman, Dhofar, near Qairoon Hairitti C M. shaat Huber, sp. nov., male from Oman, Dhofar, Shaat sinkhole D, E M. agadir (Huber), male and female with egg-sac from Morocco, Souss-Massa, Paradise Valley F M. ghar Huber, sp. nov., male from Morocco, Fès-Meknès, Kef El Ghar G M. khenifra Huber, Lecigne & Lips, sp. nov., male from Morocco, Béni Mellal-Khénifra, near Sidi Ben Daoud H M. bukidnon Huber, sp. nov., male from Philippines, Mindanao, Blue Water Cave. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure4" httpUri="https://binary.pensoft.net/fig/1141414">4</figureCitation>
|
||
) similar in colour but slightly darker.
|
||
</paragraph>
|
||
<paragraph id="DDD0318B04DFA5FF94CF34C9A85E87E5">
|
||
<emphasis id="DD65F47E89754BBDF023AAD17585342B" bold="true">
|
||
<emphasis id="B210A23B69FDF1D27A116D0C6095B7C0" bold="true" italics="true">Body</emphasis>
|
||
.
|
||
</emphasis>
|
||
Ocular area slightly raised (distinct in frontal view; Fig.
|
||
<figureCitation id="A93E2338BA4C36778D0E5B7A5F942472" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of prosomata (frontal views) and tips of female palps (dorsal views) A M. tanomah Huber, sp. nov.; female B M. darbat Huber, sp. nov.; male C, D M. ghar Huber, sp. nov.; male and female E, F M. bukidnon Huber, sp. nov.; male and female; note small AME G M. tanomah Huber, sp. nov., left palp H M. ghar Huber, sp. nov., right palp. Scale bars: 100 µm (A – D); 200 µm (E, F); 10 µm (G, H)." figureDoi="10.3897/zookeys.1213.133178.figure5" httpUri="https://binary.pensoft.net/fig/1141415">5</figureCitation>
|
||
). Carapace without thoracic groove. Clypeus unmodified. Sternum slightly wider than long, unmodified. Abdomen approximately twice as long as wide. Gonopore with four (rarely five) epiandrous spigots (Fig.
|
||
<figureCitation id="760A21697293B046AA69607FF3743FFD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male gonopores with epiandrous spigots and of female epigyna A, B M. tanomah Huber, sp. nov. C, D M. darbat Huber, sp. nov. E, F M. ghar Huber, sp. nov. G, H M. bukidnon Huber, sp. nov. Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (A, G); 100 µm (B, D, F, H); 20 µm (C, E)." figureDoi="10.3897/zookeys.1213.133178.figure10" httpUri="https://binary.pensoft.net/fig/1141420">10</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="197702162DAF9FB294DCB94E77E5F9AE" DOI="10.1206/0003-0090(2000)254<0001:NWPSAP>2.0.CO;2" author="Huber" firstAuthor="Huber" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="1-348" refId="B19" refString="Huber BA (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1–348. https://doi.org/10.1206/0003-0090 (2000) 254 <0001: NWPSAP> 2.0. CO; 2" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." volume="254" year="2000">Huber 2000</bibRefCitation>
|
||
: fig. 123;
|
||
<bibRefCitation id="909BD9710785CE53BFA220E5F34259F5" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
: fig. 99).
|
||
<abbrev id="ABBRID0ERQBG" xlink_title="anterior lateral spinneret(s)">ALS</abbrev>
|
||
with one strongly widened spigot, one long pointed spigot, and six cylindrical spigots (Fig.
|
||
<figureCitation id="4E872D83927C6919686B598CC7B436E4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)." figureDoi="10.3897/zookeys.1213.133178.figure9" httpUri="https://binary.pensoft.net/fig/1141419">9</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="6D97B48B304CAC8A24559ED8D0D0A1B5" DOI="10.1206/0003-0090(2000)254<0001:NWPSAP>2.0.CO;2" author="Huber" firstAuthor="Huber" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="1-348" refId="B19" refString="Huber BA (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1–348. https://doi.org/10.1206/0003-0090 (2000) 254 <0001: NWPSAP> 2.0. CO; 2" title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level." volume="254" year="2000">Huber 2000</bibRefCitation>
|
||
: figs 158, 159;
|
||
<bibRefCitation id="D9D43DC6767270F55E028DAD4770159F" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
: fig. 100);
|
||
<abbrev id="ABBRID0EBRBG" xlink_title="posterior median spinneret(s)">
|
||
<collectionCode id="C46F0D8D2BFFDC9F45B6A66AB69A29E0" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" type="">PMS</collectionCode>
|
||
</abbrev>
|
||
with two conical spigots;
|
||
<abbrev id="ABBRID0EFRBG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
|
||
without spigots.
|
||
</paragraph>
|
||
<caption id="2A287E4B6E15AF1501DA898DDA8D6529" ID-DOI="10.3897/zookeys.1213.133178.figure5" ID-arpha="487EA061-C7EB-5BA1-9C74-8A67FAF14AD7" httpUri="https://binary.pensoft.net/fig/1141415" startId="F5">
|
||
<paragraph id="091F12CFF1CBF1688528366341A1DB20">
|
||
<label id="80FD4B86F213C857C793226BC0C8C4D9">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="62186D334DDCC313506EFA91FB1C90AC">
|
||
<taxonomicName id="110A36B666DE9703EBBE3E4000EE9BA2" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4BE21879312939F636344D58AED2B069" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of prosomata (frontal views) and tips of female palps (dorsal views)
|
||
<emphasis id="DDB2C0D66D3F788953B8E70C566B843C" bold="true">A</emphasis>
|
||
<taxonomicName id="AA7665E44F3F5EDDF2D27867F935D41E" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="9A5F81FDEBB83DD515B343BEE9E261BE" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="8509E151FF095C4FC49229631F6B6326" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; female
|
||
<emphasis id="6C518ACA3AB42ACFE4054BA04D48734D" bold="true">B</emphasis>
|
||
<taxonomicName id="58E15ACC0C8652328A311FC4DF4A85A6" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="F3AE9066C24E9A80A5574911CF187DB3" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="17D8E950F90046EA43189A04DB1FBC41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; male
|
||
<emphasis id="1954773908AD33CC4A4F99D87CCF2C4B" bold="true">C, D</emphasis>
|
||
<taxonomicName id="CDF65B7EE0A17687B9E3146605794282" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="413D43FDB1FB514D9D41EFC76E85C7A6" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="03D6311FEDD1092BC422107292D9A003" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; male and female
|
||
<emphasis id="F8B9E32BC0940F8C075CA3E09E715D49" bold="true">E, F</emphasis>
|
||
<taxonomicName id="3D329740CD4A95EA4C1FE3E5C42ADE22" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="94DB38EB84079772D0364EC312813F1B" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="7D46C4B822B0AD7214D2E7F1BBC1460E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; male and female; note small
|
||
<abbrev id="ABBRID0EMTBG" xlink_title="anterior median eye(s)">AME</abbrev>
|
||
<emphasis id="1265701ADE8344CBE330DD6B56992FE0" bold="true">G</emphasis>
|
||
<taxonomicName id="B8A2D0DE6381889AD4450F7049B23ADF" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="2CE9D0E6EF79250C5D542047BE01891F" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1838985E3FABFFFF54C8277020B1412C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left palp
|
||
<emphasis id="4EE5D5E11790447C26ACE20D95434CFC" bold="true">H</emphasis>
|
||
<taxonomicName id="6B463C686CBB0EE7295B2C5609426309" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="C3C3F7FDB75F81E2B0F04A99270199A4" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A6296B90E0B22134FCD494EECE5FAB03" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, right palp. Scale bars: 100 µm (
|
||
<emphasis id="91577D90B8D4AC7FE928C129D649CC00" bold="true">A – D</emphasis>
|
||
); 200 µm (
|
||
<emphasis id="1C88BB41C0A3E76C63A808BBA9E5D094" bold="true">E, F</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="D72E819C0188ABB1F3E8EB4965CD06C9" bold="true">G, H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="47F9785821C1B5EB7E97AD39D08F2FD9">
|
||
<emphasis id="A2E644974BA099BF63F190A7DAA13AAA" bold="true">
|
||
<emphasis id="10A45B03D608543B88CB669D510845EE" bold="true" italics="true">Chelicerae</emphasis>
|
||
.
|
||
</emphasis>
|
||
Chelicerae with pair of strong frontal apophyses provided with conical or globular, strongly sculptured modified hairs (Fig.
|
||
<figureCitation id="8F12D747F3AEFADDE46CE3F3AEBF7025" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male chelicerae: distal apophyses with modified hairs and total view (D) A, B M. alfara Huber sp. nov. C M. tanomah Huber, sp. nov. D, E M. darbat Huber, sp. nov. F, G M. ghar Huber, sp. nov. H M. bukidnon Huber, sp. nov. Scale bars: 10 µm (A – C, F – H); 100 µm (D); 2 µm (E)." figureDoi="10.3897/zookeys.1213.133178.figure6" httpUri="https://binary.pensoft.net/fig/1141416">6</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="1B9213A77F09824CE3AAFD1227C9BDA8" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
: fig. 98), usually with one or two pairs of proximal processes; without stridulatory files.
|
||
</paragraph>
|
||
<caption id="402E9737FD91F5D2AA29C34ACA69F179" ID-DOI="10.3897/zookeys.1213.133178.figure6" ID-arpha="5A1128BA-627C-540B-AFD5-E8776BF5C1E4" httpUri="https://binary.pensoft.net/fig/1141416" startId="F6">
|
||
<paragraph id="933EC3D6B36CF8A7D03AF695F7E12EE9">
|
||
<label id="459BED77418D27E5E928CCC58DFF6BDA">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="E9985550ACCB3F75D8D860FAB777BDAF">
|
||
<taxonomicName id="01C970E68B661085A7251BFC6B83877F" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6A6D7745ED1D364CD2D2C7D6CC9CBD36" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of male chelicerae: distal apophyses with modified hairs and total view (
|
||
<emphasis id="9E1C894485C1297A918412A24EBEDEC1" bold="true">D</emphasis>
|
||
)
|
||
<emphasis id="8891C074408E6CCEA6551093C47F4F5C" bold="true">A, B</emphasis>
|
||
<taxonomicName id="42B6640A23772C16A89C38ACDA471758" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="676C5954F2AB4B24384D8A0E861C07B3" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="297F69F72D9137AB792543483EDA2189" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="498019A51EA7294A32897C84B53C2448" bold="true">C</emphasis>
|
||
<taxonomicName id="D811C05735CC42690335217336896E2F" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="6FF8BA85D544BC18AEB2E11963AE2386" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="8E63B35C14EBB97E74B66BAD33108200" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="35AFEC8B6A43200FAB96826142273272" bold="true">D, E</emphasis>
|
||
<taxonomicName id="A7B39EE9A789EAC0EE8E797B09D9F5F5" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="4F76851691F81A13E9CA234FD20A434F" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="00962CDA41E590FE42AEFCD2DC0BD4DC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="D86EB0330FE8AA0B9A4189557C5A05C5" bold="true">F, G</emphasis>
|
||
<taxonomicName id="3E1BF16793EF6BC6AFE98302C91860AB" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="1E869FE0CA3FBC0045805F45EEEBED7A" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F75B50D65EBA6324C291A56E028C5233" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="0C5E5B0C42AEBEA877C2ACB9ACAA4802" bold="true">H</emphasis>
|
||
<taxonomicName id="388F6750FF18BA786D650E83A0BAE726" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="2752424F8A3749F1031B3E9D1285FB97" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="9D6F4D89F109A049A01E8B99A77EC61D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scale bars: 10 µm (
|
||
<emphasis id="F01B9AB9F886331FA79868FCC62251F0" bold="true">A – C, F – H</emphasis>
|
||
); 100 µm (
|
||
<emphasis id="37960F294638B9FF3421887EDFCECCA6" bold="true">D</emphasis>
|
||
); 2 µm (
|
||
<emphasis id="12160149F08013B8476028777879F85A" bold="true">E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D98268166213A57D0E50B826735D410B">
|
||
<emphasis id="D577C78175836B930653528161BB8D12" bold="true">
|
||
<emphasis id="73C33B06041DC2C29391CF048266AFC0" bold="true" italics="true">Palps</emphasis>
|
||
.
|
||
</emphasis>
|
||
Palpal coxa unmodified. Trochanter with retrolateral-ventral apophysis usually with distinctive modified (short cylindrical) hair at tip (Fig.
|
||
<figureCitation id="E45E72AE3C05C5D458DD82A3E6AAEC79" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal trochanter tips and of spinnerets A M. alfara Huber, sp. nov. B M. tanomah Huber, sp. nov. C M. darbat Huber, sp. nov. D M. ghar Huber, sp. nov. E M. bukidnon Huber, sp. nov. F, G M. tanomah Huber, sp. nov., male ALS, and male spinnerets and anal cone (asterisk) H, I M. ghar Huber, sp. nov., male ALS and male spinnerets J M. bukidnon Huber, sp. nov., male ALS. Scale bars: 10 µm (A – F, H, J), 100 µm (G), 20 µm (I)." figureDoi="10.3897/zookeys.1213.133178.figure9" httpUri="https://binary.pensoft.net/fig/1141419">9 A – D</figureCitation>
|
||
), modified hair absent in
|
||
<taxonomicName id="74B8EB6B75ECE761F6558D22C40F8D79" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="AFE8F4E550F16E0D7DA8C4AF8C0BB43A" italics="true">M. bukidnon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="829F75C7EC380DF455737527A314CEB5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Femur variable in shape, often with rounded processes (usually ventrally and retrolaterally, sometimes also dorsally). Femur-patella joints shifted towards prolateral side. Tibia larger than femur, with two trichobothria. Tibia-tarsus joints shifted towards retrolateral side. Palpal tarsus with large capsulate tarsal organ (Fig.
|
||
<figureCitation id="6210DC078AFB48753C009A6BC162602B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 H, I</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="77F306D380DE73B68734279D3DB289F8" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
: fig. 97), outer diameter 30–35 µm, diameter of opening 15–20 µm; more open (almost exposed) in
|
||
<taxonomicName id="CB1EDB29A031E88D6340A8D6FA929340" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="803933C54BC7BFFC1A5BB9B6F7E3E654" italics="true">M. bukidnon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="47DE1A1658C25B5633F1866066308712" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Fig.
|
||
<figureCitation id="F8823035EF7AEC21B4051C645399A330" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 J</figureCitation>
|
||
). Procursus complex, often with distinct dorsal hinged process (e. g., Figs
|
||
<figureCitation id="F08FDFAE11FF113DBFA0185BD436AFE4" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Micropholcus fauroti (Simon, 1887); male from Mexico, Nuevo León, Santiago (ZFMK Ar 24655). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Abbreviations: b, genital bulb; c, coxa; f, femur; hp, dorsal hinged process; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure16" httpUri="https://binary.pensoft.net/fig/1141426">16 C</figureCitation>
|
||
,
|
||
<figureCitation id="4E23D2D4C841DE9D8F47FAEAED3D8101" captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Micropholcus bashayer Huber, sp. nov.; male from Saudi Arabia, ‘ Asir, NW of Al Bashayer (ZFMK Ar 24665). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure43" httpUri="https://binary.pensoft.net/fig/1141453">43 C</figureCitation>
|
||
,
|
||
<figureCitation id="FE9A8C54E08954990F42B3A682EE546F" captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Micropholcus darbat Huber, sp. nov.; male from Oman, Dhofar, Wadi Darbat (ZFMK Ar 24672). Left palp in prolateral (A), dorsal (B), and retrolateral (C) views. Scale bar: 0.3 mm." figureDoi="10.3897/zookeys.1213.133178.figure52" httpUri="https://binary.pensoft.net/fig/1141462">52 C</figureCitation>
|
||
), in most species with transparent prolateral membranous flap densely set with teeth (Figs
|
||
<figureCitation id="18BA720F39CAEB26CC3300DEFCB8C56B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. alfara Huber, sp. nov.; prolateral membranous flap on left procursus and right bulbal processes, prolateral view C, D M. tanomah Huber, sp. nov., tip of left procursus and prolateral membranous flap on left procursus E M. tanomah Huber, sp. nov., left procursus, dorsal view (bold arrow points at tip of procursus) F, G M. tanomah Huber, sp. nov., left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view H M. darbat Huber, sp. nov., left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (A); 100 µm (B, E, F, H); 20 µm (C, D, G)." figureDoi="10.3897/zookeys.1213.133178.figure7" httpUri="https://binary.pensoft.net/fig/1141417">7 A, D</figureCitation>
|
||
,
|
||
<figureCitation id="329D774568B78E9601140AB5B09A1BBE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. darbat Huber, sp. nov.; right bulb and procursus, prolateral view (bold arrow in A points at trochanter apophysis), and prolateral membranous flap of procursus at higher magnification C M. darbat Huber, sp. nov., left bulbal processes D M. ghar Huber, sp. nov., left procursus, retrolateral view E, F M. ghar Huber, sp. nov., right bulbal processes, prolateral and prolateral-ventral views G M. bukidnon Huber, sp. nov., left procursus, prolateral-distal view H M. bukidnon Huber, sp. nov., left bulbal processes, prolateral distal view. Abbreviations: a, putative appendix; b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; pr, procursus; ta, tarsus; u, putative uncus. Scale bars: 100 µm (A, D – H); 10 µm (B); 20 µm (C)." figureDoi="10.3897/zookeys.1213.133178.figure8" httpUri="https://binary.pensoft.net/fig/1141418">8 B</figureCitation>
|
||
). Genital bulb with distinct proximal sclerite connecting to tarsus, membranous or partly sclerotised embolus, and variably complex set of sclerotised apophyses, sometimes likely homologues of the
|
||
<taxonomicName id="7CEE2201565A72CA6EFB50EA376C70FF" authorityName="Walckenaer" authorityYear="1805" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CE0C4C72BE0537919333C276FD874560" italics="true">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
‘ appendix’ and ‘ uncus’, sometimes of uncertain homology (Figs
|
||
<figureCitation id="491F52DCFAD20C6480DC9BCE1E2B64AE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. alfara Huber, sp. nov.; prolateral membranous flap on left procursus and right bulbal processes, prolateral view C, D M. tanomah Huber, sp. nov., tip of left procursus and prolateral membranous flap on left procursus E M. tanomah Huber, sp. nov., left procursus, dorsal view (bold arrow points at tip of procursus) F, G M. tanomah Huber, sp. nov., left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view H M. darbat Huber, sp. nov., left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (A); 100 µm (B, E, F, H); 20 µm (C, D, G)." figureDoi="10.3897/zookeys.1213.133178.figure7" httpUri="https://binary.pensoft.net/fig/1141417">7 B, F</figureCitation>
|
||
,
|
||
<figureCitation id="41BA6723639B7B18DC0BA916DEDBD937" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of male palpal structures A, B M. darbat Huber, sp. nov.; right bulb and procursus, prolateral view (bold arrow in A points at trochanter apophysis), and prolateral membranous flap of procursus at higher magnification C M. darbat Huber, sp. nov., left bulbal processes D M. ghar Huber, sp. nov., left procursus, retrolateral view E, F M. ghar Huber, sp. nov., right bulbal processes, prolateral and prolateral-ventral views G M. bukidnon Huber, sp. nov., left procursus, prolateral-distal view H M. bukidnon Huber, sp. nov., left bulbal processes, prolateral distal view. Abbreviations: a, putative appendix; b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; pr, procursus; ta, tarsus; u, putative uncus. Scale bars: 100 µm (A, D – H); 10 µm (B); 20 µm (C)." figureDoi="10.3897/zookeys.1213.133178.figure8" httpUri="https://binary.pensoft.net/fig/1141418">8 C, E, F, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2C646525C4081A10612E1F63939B1D21" ID-DOI="10.3897/zookeys.1213.133178.figure7" ID-arpha="511ABB71-70F5-5E7D-922D-7D65E2C38B61" httpUri="https://binary.pensoft.net/fig/1141417" startId="F7">
|
||
<paragraph id="867FABAE466F182075B3EA1D4174A3C0">
|
||
<label id="537A9E4CB0A3F2E203D5D0867C4C23BE">Figure 7.</label>
|
||
</paragraph>
|
||
<paragraph id="17B9E9A6B9E303E220A4BC1FF003B5FA">
|
||
<taxonomicName id="31B832C2BBBF7E741FD67EEFD69957B7" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1CC70CFAB8DD80E5E1672C20F301DCE5" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of male palpal structures
|
||
<emphasis id="7B6933B08CAE2CB811A67F4C616850CF" bold="true">A, B</emphasis>
|
||
<taxonomicName id="1915CBFEA6214E54898B3FF7B538C1D7" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="3299C1E376A0321A5218D6EA84DC5518" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4E6643BC4D2BC605A77D43266118B711" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; prolateral membranous flap on left procursus and right bulbal processes, prolateral view
|
||
<emphasis id="2C742E3896C3C36137E86A4DC9B53752" bold="true">C, D</emphasis>
|
||
<taxonomicName id="1799D3D84D6410934495C0873710C077" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="690C22BE7986771060AD65DDFF8B7980" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FB1651802BB27913FA5BBEB81DFCDA12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, tip of left procursus and prolateral membranous flap on left procursus
|
||
<emphasis id="9D149345B75ED45FC8C77544174686B8" bold="true">E</emphasis>
|
||
<taxonomicName id="7A0118D8FBF6DEBB4AA4C48E38B0785A" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="71FE9C182E1FF3226E48B6663974837A" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C09210AA2674EC32D52A5BC9776F87A8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left procursus, dorsal view (bold arrow points at tip of procursus)
|
||
<emphasis id="3ED826263202CC5B10BD62F356AEDF61" bold="true">F, G</emphasis>
|
||
<taxonomicName id="A4182DD426E72DC4C463EB9A6508E0DD" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="A3758905074BDCF50FAD8BE492790CC8" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="DAE2CCFFC68B766FE0498CCB5003EC48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left bulbal processes, prolateral view, and embolus of same palp in slightly more distal view
|
||
<emphasis id="D3F6947DB1A1A73571A24163BF14EA51" bold="true">H</emphasis>
|
||
<taxonomicName id="EF40CFA98AF4C2AF8810B18AAF54B0FD" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="349443689F5CCD97C6D7EB3CB4959614" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="260197412E80E0FE2AE423E16C1F2B05" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left procursus, retrolateral view. Abbreviations: b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; rr, retrolateral ridge; ta, tarsus; tm, transparent membrane. Scale bars: 10 µm (
|
||
<emphasis id="5AAC3F77B64411CF5B5C34132E1153FD" bold="true">A</emphasis>
|
||
); 100 µm (
|
||
<emphasis id="D1821B659967A07C18C5B4BD711D429F" bold="true">B, E, F, H</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="72B3A0B5108801F36B96F41B3E8EA262" bold="true">C, D, G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="27E70854EB48FE451AB29D9012DE6FBC" ID-DOI="10.3897/zookeys.1213.133178.figure8" ID-arpha="F4392161-975D-5DC9-B4FC-08139F6485F0" httpUri="https://binary.pensoft.net/fig/1141418" startId="F8">
|
||
<paragraph id="186E1A7660DF49D369DE2417425840B2">
|
||
<label id="65D1B6D2FF46C3E25CD428BB9B82308E">Figure 8.</label>
|
||
</paragraph>
|
||
<paragraph id="D6852510A54653C057BD23B4D65BE795">
|
||
<taxonomicName id="B065F2273791773DA81ECBF6CA2F09C1" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EB1EBA7A6D2CBC3EB051DC169782D602" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of male palpal structures
|
||
<emphasis id="5EA2F2DD57322356FA63E71161DEE833" bold="true">A, B</emphasis>
|
||
<taxonomicName id="0CF1B9F03A852988995D041B68977D12" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="E6B89934B30440A4B8CC8B585E9EC5EB" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="BAA1D25BD6465FE904DCD824E2DDDEB7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; right bulb and procursus, prolateral view (bold arrow in A points at trochanter apophysis), and prolateral membranous flap of procursus at higher magnification
|
||
<emphasis id="430F96D2279B9C09FF51D47C9B987D7D" bold="true">C</emphasis>
|
||
<taxonomicName id="874D9B7DB09CE5C519931D506F9FD67D" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="B4C6E058C330619EA36B89E5466EAFD5" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6BA9E279BF6E186EB45D87BBE7CEF065" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left bulbal processes
|
||
<emphasis id="BD01D6BAD76CF00EB703F36490C43881" bold="true">D</emphasis>
|
||
<taxonomicName id="7A9E97A288300C55BEACD342F7340063" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="5F4AE011B5BF75555B3277E60D696BB5" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="5B8B56AAAA4258DA405FF854F40ABBA9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left procursus, retrolateral view
|
||
<emphasis id="00A466747C284B86A49DA750C8ABE52A" bold="true">E, F</emphasis>
|
||
<taxonomicName id="145B1978D08B4184E4447DDAEF967441" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="E1AA5496665966B02CAB76603EEB195A" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="27A2CD74362C1713D0C4588DCF3A4675" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, right bulbal processes, prolateral and prolateral-ventral views
|
||
<emphasis id="76CA404554AE7D76AACE18390054F4A0" bold="true">G</emphasis>
|
||
<taxonomicName id="3DAC2489B61A5C23ACF3FB970AFCCCFB" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="7C973806806710A729F709839CA7DF6F" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="CFCF00AFD1473F5BB961C207BABBB1B0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left procursus, prolateral-distal view
|
||
<emphasis id="406DD5A726FA67B1055BAB7F5E351029" bold="true">H</emphasis>
|
||
<taxonomicName id="F742BF4E7EFC79CA1A6278508CA8935A" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="7A3B766DF62106DD7D3A4C88F993CAC6" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="13E2A26D7BF5D80F5007010014B1CA7D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left bulbal processes, prolateral distal view. Abbreviations: a, putative appendix; b, genital bulb; e, embolus; hp, dorsal hinged process; mf, membranous prolateral flap; pr, procursus; ta, tarsus; u, putative uncus. Scale bars: 100 µm (
|
||
<emphasis id="45742A802993DF47C6F23CDA14208178" bold="true">A, D – H</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="BF316170D87C293B545FDFCCF7B214AD" bold="true">B</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="B95E6C5804207806E5C27ED3C913F799" bold="true">C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8433FBA65138F58410A22B2EB5FF18D8" ID-DOI="10.3897/zookeys.1213.133178.figure9" ID-arpha="2F7E9F26-79D1-5474-B0AB-89F379AE8041" httpUri="https://binary.pensoft.net/fig/1141419" startId="F9">
|
||
<paragraph id="A9D1ED3D18DBA2BD0875110BC1390836">
|
||
<label id="7FD1803BF028A74FC1165D4CD1C02AF2">Figure 9.</label>
|
||
</paragraph>
|
||
<paragraph id="536F59139589F33664CC6D977C1F0691">
|
||
<taxonomicName id="96F252686ADF5D99B8DF8E735CAC5D7D" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="413F4790AFC3FC7FBEDC15A693999AB5" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of male palpal trochanter tips and of spinnerets
|
||
<emphasis id="7EAE4BB1BF4D27DEB29895178BFC80A4" bold="true">A</emphasis>
|
||
<taxonomicName id="D3DA335FFCCFFBBC3B7159FE34589B14" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="C4FAE8B90FCD44ED04410FD3457A2DDE" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="0FC7101E825E172D2FBB05C583623A4E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="243A3F69086EE906A69A7EC5EC48616C" bold="true">B</emphasis>
|
||
<taxonomicName id="B42158F01A0ED9790836AE5FE1829D8B" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="E942D7ACC2A9DCDDA4317A6F127D9EA1" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FBE8F238C488ADCCDF125FE30F269BF1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="769455DBA69F319AD97D5FDF5ABD5E49" bold="true">C</emphasis>
|
||
<taxonomicName id="B7A25BC0C5F9E80604C1682C80355A99" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="5FE5629AB1E0BF567A3D832D96345BA5" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4CB5983552A10A76D7EFC97EE53BC5C6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="F747E80609B3C481C5268F6E18F66586" bold="true">D</emphasis>
|
||
<taxonomicName id="6EA290347989355DEF09A989ABA96B59" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="37D5FE6898E3BDFDEA3D653CEBA838E5" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E911F8AAF5CEAE8EB3AB79B2A6665C5B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="99BAAEF82BF37B8F831C09CABEDD48F1" bold="true">E</emphasis>
|
||
<taxonomicName id="F5C725952BC3E6C83E85EF77FD36FABA" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="208CF6F26C00D99D48037C028AB1414E" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A05262D6B0F0BD3D0C8A3843EA52600C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="CD2B57EFEF8455715FF400F2EBA770DB" bold="true">F, G</emphasis>
|
||
<taxonomicName id="581628559E4933105DC0AD087B568D0D" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="04C77C32F8A4AA274C8454B4FAE2CCF5" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="AB94CC6BE1A29AD30E58C385DC22C711" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<abbrev id="ABBRID0ESDAI" xlink_title="anterior lateral spinneret(s)">ALS</abbrev>
|
||
, and male spinnerets and anal cone (asterisk)
|
||
<emphasis id="B92F93628E7C5C147CC076E83CCE796E" bold="true">H, I</emphasis>
|
||
<taxonomicName id="B9CE0163BEC4039A20B2EDA4B07F8B57" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="BFA62632D0E9C066E23B19CE03135FCA" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="41A00B2710FB740A447228292924D040" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<abbrev id="ABBRID0EDEAI" xlink_title="anterior lateral spinneret(s)">ALS</abbrev>
|
||
and male spinnerets
|
||
<emphasis id="B18C301B13CC7D072DA7090657E71452" bold="true">J</emphasis>
|
||
<taxonomicName id="B4E37B8BF6DB310A7129F458865294BC" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="C34B274C8F3BF05A9A24A666B23C400A" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="016CCFABBF4DEACB6E8AB942797F97C2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<abbrev id="ABBRID0EUEAI" xlink_title="anterior lateral spinneret(s)">ALS</abbrev>
|
||
. Scale bars: 10 µm (
|
||
<emphasis id="AB41AFE11A401485C28C79FF205C541E" bold="true">A – F, H, J</emphasis>
|
||
), 100 µm (
|
||
<emphasis id="1467C421CCA5B2E7B9B4A5A935BEE52A" bold="true">G</emphasis>
|
||
), 20 µm (
|
||
<emphasis id="BF38E4EF1CE98B753D7A9A06FC3DF780" bold="true">I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="254C172C73B0F14BC3924BE2D8F67F96">
|
||
<emphasis id="43019373520FFCF55066152854EC3874" bold="true">
|
||
<emphasis id="6BD874F167CA62243294ED60299D31A0" bold="true" italics="true">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Without spines and curved hairs. Without slender metatarsal hairs (cf.
|
||
<bibRefCitation id="62F47A64205C2080FA5EAB7BC457ED2B" DOI="10.5852/ejt.2023.900.2301" author="Huber BA & Meng G & Král J & Ávila Herrera IM & Izquierdo MA" journalOrPublisher="European Journal of Taxonomy" pagination="32-80" refId="B31" refString="Huber BA, Meng G, Král J, Ávila Herrera IM, Izquierdo MA (2023 a) Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900: 32–80. https://doi.org/10.5852/ejt.2023.900.2301" title="Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae)." volume="900">Huber et al. 2023 a</bibRefCitation>
|
||
). Without sexually dimorphic short vertical hairs. Chemoreceptive hairs ~ 20–25 µm long, with few side branches (Fig.
|
||
<figureCitation id="0F2410BF553336379F55C80753EF7A2D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 A, H</figureCitation>
|
||
), mostly near leg tips. Retrolateral trichobothrium of tibia 1 at 5–10 % of tibia length. Prolateral trichobothrium absent on tibia 1, present on tibiae 2–4. Base of trichobothria evenly rounded, without proximal ridge (cf. Fig.
|
||
<figureCitation id="4A1B720238ACC2C3865CD67D3B914CB5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 D</figureCitation>
|
||
). Legs with roundish cuticular plates (Fig.
|
||
<figureCitation id="52F54717E8C69C171D2EE951286C6F0A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 E</figureCitation>
|
||
; diameter ~ 6–8 µm) and rimmed pores (Fig.
|
||
<figureCitation id="098DDFDFDF716567A80B89CE98DC45B3" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 B</figureCitation>
|
||
; outer diameter 2 µm, diameter of opening 0.2 µm) apparently on all leg segments. Tarsus 1 with ~ 20–30 pseudosegments, distally usually fairly distinct. Leg tarsal organs capsulate (Figs
|
||
<figureCitation id="6992F6BEB5A581E21FB0C47CE3C14440" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 E – G</figureCitation>
|
||
,
|
||
<figureCitation id="09FB4E6BB74486B1A2D0068A53781172" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 F</figureCitation>
|
||
; diameter 12–17 µm, diameter of opening 5–7 µm). Tarsus 4 with single row of comb-hairs on prolateral side (Fig.
|
||
<figureCitation id="31E86793CB2504789C490611E44433C8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 G, H</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="894265C83ED1DD2DB6D9415E79745A86" author="Huber" firstAuthor="Huber" journalOrPublisher="Bonner zoologische Monographien" pagination="1-509" refId="B21" refString="Huber BA (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien 58: 1–509. https://biostor.org/reference/137036" title="Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae)." url="https://biostor.org/reference/137036" volume="58" year="2011">Huber 2011</bibRefCitation>
|
||
: fig. 96). Main tarsal claws with ~ 10 teeth (Fig.
|
||
<figureCitation id="249F2CA14ADD67B56258711FBA178010" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Micropholcus Deeleman-Reinhold & Prinsen, SEM images of leg structures A M. tanomah Huber, sp. nov., putative chemoreceptor B M. tanomah Huber, sp. nov., rimmed pore (arrow) on left tarsus 3 C, D M. tanomah Huber, sp. nov., tarsal claws of left legs 1 and 3 E M. ghar Huber, sp. nov., cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3 F M. ghar Huber, sp. nov., pseudosegmentation (and tarsal organ) of right tarsus 3 G M. ghar Huber, sp. nov., comb-hairs on male tarsus 4 H M. bukidnon Huber, sp. nov., comb-hairs on male tarsus 4. Scale bars: 2 µm (A); 1 µm (B); 20 µm (C); 10 µm (D – H)." figureDoi="10.3897/zookeys.1213.133178.figure12" httpUri="https://binary.pensoft.net/fig/1141422">12 C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="8AE5B0F54C18452C35F30CE6CB41A390" ID-DOI="10.3897/zookeys.1213.133178.figure10" ID-arpha="80F749E8-1B7C-596C-BBAE-7B8C8B3636CF" httpUri="https://binary.pensoft.net/fig/1141420" startId="F10">
|
||
<paragraph id="0B5F588657B8175C6EBFF2F22C452BD4">
|
||
<label id="7574636F515A6F2A1694A02C394128EC">Figure 10.</label>
|
||
</paragraph>
|
||
<paragraph id="8DA316478718DCDB041736D1F714BF0A">
|
||
<taxonomicName id="14805D468A6CA4867C6D0A081ED0B103" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9570E09923914AF043FD25ABA241F563" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of male gonopores with epiandrous spigots and of female epigyna
|
||
<emphasis id="9906E4EEBFF1F8DF97F0ACBD89A0B59E" bold="true">A, B</emphasis>
|
||
<taxonomicName id="BC571CC5BA269AA702148742248E694F" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="B47EC2C23681850D1FA8C3DFDDED9997" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="145E4313E937AE8B41881BA0232F3BCB" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="CF2DBA14494A3B44D82AF9FE198666D9" bold="true">C, D</emphasis>
|
||
<taxonomicName id="6B29C2DA2A22E2769181D45BB86461C7" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="909AC65920E3FBDB6E9D99AE421F72C6" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="42916739ABD0C34747D0F2948FAD4F58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="89E53E6125E3ED4869FCD424452E4DCD" bold="true">E, F</emphasis>
|
||
<taxonomicName id="268FB3D93465CBF9DCB7F2262B4B2BAA" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="76ECACE2A5B1E5C484B05A087753884D" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FFD3550BB74536ACCEEA8E06ADCA9DA6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="DEA0E8DB925884C069BE44EE07DB5E6D" bold="true">G, H</emphasis>
|
||
<taxonomicName id="75B73133ED471ABB4A1C735FD57530E9" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="FE903E015575AE9C89379ED25CA52100" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F82AC06C3E83AF1E5259AF8B87792E23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Abbreviations: aep, anterior epigynal plate; k, epigynal ‘ knob’; pep, posterior epigynal plate. Scale bars: 10 µm (
|
||
<emphasis id="1577C343850EFE6DF35A66233CD78FF8" bold="true">A, G</emphasis>
|
||
); 100 µm (
|
||
<emphasis id="8493D6A716D7416965647F8750196D4D" bold="true">B, D, F, H</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="49E19D3B643134F97DED37D28C97F317" bold="true">C, E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89EFE91C25F16B912ECF4A6041B859F3" ID-DOI="10.3897/zookeys.1213.133178.figure11" ID-arpha="F0A9C633-B74D-57E4-9F2F-E2567A7F0496" httpUri="https://binary.pensoft.net/fig/1141421" startId="F11">
|
||
<paragraph id="1B5B24DCE051C488AA349C2CBDA5E0E1">
|
||
<label id="89A57DADE052E6E33F9FC70826900B5E">Figure 11.</label>
|
||
</paragraph>
|
||
<paragraph id="9E6846C20FD8088F1B9A6F178FACC40C">
|
||
<taxonomicName id="95D725D3DA393F13499A5DD206598A7A" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BBD350612BC1D7F6D9EB9389B054BE4" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
; SEM images of epigynal knobs (
|
||
<emphasis id="FFF706BB4831EFEDBE4A69C3C37A2FEC" bold="true">A – C</emphasis>
|
||
), trichobothria (
|
||
<emphasis id="BA2013A8D30B2EE89F1F80AB0CE19CE2" bold="true">D</emphasis>
|
||
), and tarsal organs (
|
||
<emphasis id="79A77875E9204148DED46D434B58C5F2" bold="true">E – J</emphasis>
|
||
)
|
||
<emphasis id="417B9BCE9191E2D622F17C2EE9F47364" bold="true">A</emphasis>
|
||
<taxonomicName id="7F3AD76DBE388B7023335E9ADD381C66" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="A1ADCBC683FE0DA7656A8979E53C770B" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FA01E11CEF382BA7D940EC3EE9E13A30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="474EAC097560A5247AB68A4A691E3CF1" bold="true">B</emphasis>
|
||
<taxonomicName id="653977CD5424792AC77DC2845B71440A" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="4C818C1A8EF6AA04BCC5491833117FE8" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F1DA7708B1D37C5F613E1D9C194E9E75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="9A4653B75BE7F1E62E9F240A4BD71A4C" bold="true">C</emphasis>
|
||
<taxonomicName id="B297054E20B1ECED6561F9BD1C9DD010" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="C79B9154CF056F3D12D5674B489408C0" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="93472C823FE365FBCE8454F190F82571" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="44F9569F4906262E345FE4A18FDE2662" bold="true">D</emphasis>
|
||
<taxonomicName id="D3A1F6D237624198233E6112AE0EAD64" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="B5051C12FF192A231C8053429B002BB6" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="9200CF65EADF4F983318E3932B674233" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female left palpal tibia
|
||
<emphasis id="E38A34DFEF79809A829A1DF3E035B09F" bold="true">E</emphasis>
|
||
<taxonomicName id="2184CB198265199F8625FEED09BCDB88" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="69465DDDE279C9186C46A4C5A8B70A6E" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D982A1DC10FB9F94DF6CB11BDB754BEF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male right tarsus 2
|
||
<emphasis id="F978CB1D3D5C204A1B6741AA48882C7A" bold="true">F</emphasis>
|
||
<taxonomicName id="202CC7A716D8B2DF8FBAACC4A775E263" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="FDF998767C8902EE524F633126C57D25" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="5EB72C2893648FCC9F68074B42812E4E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female left tarsus 2
|
||
<emphasis id="6F31E33F52906C035EE5FCEE9D90EF98" bold="true">G, H</emphasis>
|
||
<taxonomicName id="752BE8939D3BA792503E542C1D784287" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="darbat" status="sp. nov.">
|
||
<emphasis id="9A0C243594F94873CC397F9AB76F1D42" italics="true">M. darbat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A49B0056260031184CCF226729C9FDCF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female right tarsus 2 and male palpal tarsus
|
||
<emphasis id="0A6BE1E355ED6B895077CB45B09087D3" bold="true">I</emphasis>
|
||
<taxonomicName id="B0FBBFEF703A9F6605AD3610B195DF1D" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="730EB0F403345384BB4D346FE465B8FF" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E7F8302D7627D8C7A844CD9A06AEA160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male palpal tarsus
|
||
<emphasis id="EF7B871E78FD77829FB51F080CC5C1A5" bold="true">J</emphasis>
|
||
<taxonomicName id="53CD2961ACCADCC8AC9091D708F87361" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="9C4A5C708704F3FA22C82807670C816B" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C00D7EDE8132868864F23B84F8537CE3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male palpal tarsus. Scale bars: 20 µm (
|
||
<emphasis id="CD6033E4E5D0D23DD0FF5366ED4BC7B5" bold="true">A</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="41CE6C283F7B5FB36F1F4827679D9FDA" bold="true">B – D, H – J</emphasis>
|
||
); 2 µm (
|
||
<emphasis id="778DA645E7290E9EA2E5D0EEAE168D94" bold="true">E – G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="09960CA75A73886674666765F7F76B8A" ID-DOI="10.3897/zookeys.1213.133178.figure12" ID-arpha="A29D1DE9-72FC-5A59-BCCD-315052DE4CA7" httpUri="https://binary.pensoft.net/fig/1141422" startId="F12">
|
||
<paragraph id="16A532832D5ED98A298640FCD794C183">
|
||
<label id="687FD2DF46888F3EEE557238D7583EDB">Figure 12.</label>
|
||
</paragraph>
|
||
<paragraph id="98C33FFFD5CF0D3AC9E6CFC561369237">
|
||
<taxonomicName id="A9A8C696A55BB09B95CE7302E93FDB46" authority="Deeleman-Reinhold & Prinsen, SEM" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97ADAA2F8548E7D7FEB60C53793448A1" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen, SEM
|
||
</taxonomicName>
|
||
images of leg structures
|
||
<emphasis id="99623D3578FF1E11744AF7EB4113D21B" bold="true">A</emphasis>
|
||
<taxonomicName id="D2A4C97BE85AF82B19273EFF3605E165" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="159E7BFBBC107E443315E95000DE1AA7" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="CFC468F4AE798D162C86414BF4680330" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, putative chemoreceptor
|
||
<emphasis id="D18BD514A8C931B76D54C3D98234203D" bold="true">B</emphasis>
|
||
<taxonomicName id="9A9D0FD875B15B8DE24B0AA09532425A" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="62D9C04547D67BCE19CF74B1C714904A" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="9A6B408386A2B83A477208C1BB52D679" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, rimmed pore (arrow) on left tarsus 3
|
||
<emphasis id="C0093D6490DF64295D586623CEB1D76F" bold="true">C, D</emphasis>
|
||
<taxonomicName id="672E0E26BB36F0030BB28AAAEDD4E6E4" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="FCA7738BA59CDE429801D2F562739D10" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C353426AF372C75E22B6E49A89DF949F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, tarsal claws of left legs 1 and 3
|
||
<emphasis id="EBD415B2174C85FE6810FA0F0940CCC7" bold="true">E</emphasis>
|
||
<taxonomicName id="2F8F0D04AA84194576FEBFEA32AB7ABD" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="77C99390346FDDE85EB1F0E1CA83D3D2" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1D9CA8EB020E6E036FA45F55F3AD558D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, cuticular plate (arrow) and regular mechanoreceptor on right metatarsus 3
|
||
<emphasis id="F68BC1622E5B2BE8F3534D62B46A7AE1" bold="true">F</emphasis>
|
||
<taxonomicName id="AFFAFF7ABFE2AED80023E3CF4993DF95" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="9EFFFC6F12A765D133E296F87CE5F7B5" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B28A6B8BF490B01B32A61674A38D3FBA" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, pseudosegmentation (and tarsal organ) of right tarsus 3
|
||
<emphasis id="1FD934FC6F654E0BAF4DE460DDE22643" bold="true">G</emphasis>
|
||
<taxonomicName id="3795285C4673F8A97E04863A9E98AA3F" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="A36F54AECCC00E198EA5EAD56DB4660E" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F5E70C71930AF4F74D3590E493158303" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, comb-hairs on male tarsus 4
|
||
<emphasis id="4FAD0652D68CBDEFEE42204AD7929AB6" bold="true">H</emphasis>
|
||
<taxonomicName id="0747803289755D9D90C0FB9EC72131ED" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="4F71628B695566A6991904851679C230" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="35AB189E49097D7C19C68A1F6EAD75FB" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, comb-hairs on male tarsus 4. Scale bars: 2 µm (
|
||
<emphasis id="5DE7AD98B23A2820F7A518B0F92CFBB0" bold="true">A</emphasis>
|
||
); 1 µm (
|
||
<emphasis id="407167507C5C250D4F4A354CB7DD6CC8" bold="true">B</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="98D4473DC1D6B618EBF21464D199E32C" bold="true">C</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="D5A4704D8034B3CF777F1A28C365A09C" bold="true">D – H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DFC8B0B24F57D501DCB82A2F0261F607" ID-DOI="10.3897/zookeys.1213.133178.figure13" ID-arpha="961BF845-01EC-549A-BAF5-4865494E2863" httpUri="https://binary.pensoft.net/fig/1141423" startId="F13">
|
||
<paragraph id="3069E6E97676A2E32A19197DC47415EA">
|
||
<label id="B8B67BD0C42DBD8CD7C221D620101E1F">Figure 13.</label>
|
||
</paragraph>
|
||
<paragraph id="78BF1A794AE57E096A9D87B12288C467">
|
||
Known distribution of
|
||
<taxonomicName id="C94CA8A25F6117CB250C1C23F19AF6EB" authority="(Simon)" baseAuthorityName="Simon" baseAuthorityYear="1887" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fauroti">
|
||
<emphasis id="809B685CBCEE78A6B84D2AF6BA72A454" italics="true">Micropholcus fauroti</emphasis>
|
||
(Simon)
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="5A3BB4D9C6F9999271EF6A6BC47CCB80" bold="true">A</emphasis>
|
||
) and of Old World
|
||
<taxonomicName id="B72A3362750EE9D411CED935A846746B" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="42C302843FEB78827A23A282179EB0FF" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
in Mindanao (
|
||
<emphasis id="D1C0116FAE4CC281EF8E18C5E8287405" bold="true">B</emphasis>
|
||
), on the Arabian Peninsula (
|
||
<emphasis id="AAADD16898B7A0F92387179D2889C1FE" bold="true">C</emphasis>
|
||
) and in Morocco (
|
||
<emphasis id="E6C8DB8B50B6FD3E6ADA189254C2B317" bold="true">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="11FC67CB1CDD7A978796D2045CE66123">
|
||
<emphasis id="B198E400ECC345C11880F9BD363B32B4" bold="true">Female.</emphasis>
|
||
In general, very similar to males (Figs
|
||
<figureCitation id="BE4683F60FFC43A157FFB5DB8131CC96" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure3" httpUri="https://binary.pensoft.net/fig/1141413">3</figureCitation>
|
||
,
|
||
<figureCitation id="00A64D49CA72433C4B472C66069E5EEE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Oman, Morocco, and the Philippines A, B M. darbat Huber, sp. nov., male and female with egg-sac from Oman, Dhofar, near Qairoon Hairitti C M. shaat Huber, sp. nov., male from Oman, Dhofar, Shaat sinkhole D, E M. agadir (Huber), male and female with egg-sac from Morocco, Souss-Massa, Paradise Valley F M. ghar Huber, sp. nov., male from Morocco, Fès-Meknès, Kef El Ghar G M. khenifra Huber, Lecigne & Lips, sp. nov., male from Morocco, Béni Mellal-Khénifra, near Sidi Ben Daoud H M. bukidnon Huber, sp. nov., male from Philippines, Mindanao, Blue Water Cave. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure4" httpUri="https://binary.pensoft.net/fig/1141414">4</figureCitation>
|
||
). Legs slightly shorter than in males (male / female tibia 1 length: ~ 1.1–1.4, but sample sizes mostly small); tibia 1 length 4.3–9.1. Palpal tarsal organ smaller than in males (outer diameter: 18–25 µm, diameter of opening: 6–10 µm). Palpal tarsus ending distally in pointed tip and pair of strong dorsal hairs (Fig.
|
||
<figureCitation id="4FC27D8EC5EC7AB0FF3510DEBD9F7FFE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of prosomata (frontal views) and tips of female palps (dorsal views) A M. tanomah Huber, sp. nov.; female B M. darbat Huber, sp. nov.; male C, D M. ghar Huber, sp. nov.; male and female E, F M. bukidnon Huber, sp. nov.; male and female; note small AME G M. tanomah Huber, sp. nov., left palp H M. ghar Huber, sp. nov., right palp. Scale bars: 100 µm (A – D); 200 µm (E, F); 10 µm (G, H)." figureDoi="10.3897/zookeys.1213.133178.figure5" httpUri="https://binary.pensoft.net/fig/1141415">5 G, H</figureCitation>
|
||
). Spinnerets, leg hairs, cuticular plates, rimmed pores, comb-hairs, leg tarsal organs, and tarsal claws as in male. Epigynum anterior plate usually weakly sclerotised, with rounded process (‘ knob’; Fig.
|
||
<figureCitation id="09DAE9F5A2467A013C4B9F0B03E7605D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of epigynal knobs (A – C), trichobothria (D), and tarsal organs (E – J) A M. tanomah Huber, sp. nov. B M. darbat Huber, sp. nov. C M. ghar Huber, sp. nov. D M. alfara Huber, sp. nov., female left palpal tibia E M. alfara Huber, sp. nov., male right tarsus 2 F M. tanomah Huber, sp. nov., female left tarsus 2 G, H M. darbat Huber, sp. nov., female right tarsus 2 and male palpal tarsus I M. ghar Huber, sp. nov., male palpal tarsus J M. bukidnon Huber, sp. nov., male palpal tarsus. Scale bars: 20 µm (A); 10 µm (B – D, H – J); 2 µm (E – G)." figureDoi="10.3897/zookeys.1213.133178.figure11" httpUri="https://binary.pensoft.net/fig/1141421">11 A – C</figureCitation>
|
||
) in varying position; posterior epigynal plate short and indistinct. Internal genitalia often complex, highly variable, with distinct pair of pore plates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EHTAI" type="distribution">
|
||
<paragraph id="66095E0FDC90A6E17294B5A05F558E7E">
|
||
<heading id="678B1DFF3E673FB011CC4A245B08A365" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="B32D5C8E0AFBE20458EB781D7875869D">
|
||
The type species
|
||
<taxonomicName id="6EBCD94F7840DCB09A7870F2B1E29AF4" baseAuthorityName="Simon" baseAuthorityYear="1887" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fauroti">
|
||
<emphasis id="82EFAB27FA52082F94258F70C8D27665" italics="true">Micropholcus fauroti</emphasis>
|
||
</taxonomicName>
|
||
has attained a circumtropical distribution, with most records from between
|
||
<geoCoordinate id="92CA71D6AE911DEE37CBDF2B974A840B" degrees="25" direction="south" orientation="latitude" precision="55555" value="-25.0">25 ° S</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="B8597D61531F3D7F5431783EE8E7D382" degrees="30" direction="north" orientation="latitude" precision="55555" value="30.0">30 ° N</geoCoordinate>
|
||
(Fig.
|
||
<figureCitation id="10F8E911783851DC403D2F157A616896" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of prosomata (frontal views) and tips of female palps (dorsal views) A M. tanomah Huber, sp. nov.; female B M. darbat Huber, sp. nov.; male C, D M. ghar Huber, sp. nov.; male and female E, F M. bukidnon Huber, sp. nov.; male and female; note small AME G M. tanomah Huber, sp. nov., left palp H M. ghar Huber, sp. nov., right palp. Scale bars: 100 µm (A – D); 200 µm (E, F); 10 µm (G, H)." figureDoi="10.3897/zookeys.1213.133178.figure5" httpUri="https://binary.pensoft.net/fig/1141415">5 A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1F0F7B1810859A218BD360B889190040">
|
||
<taxonomicName id="9F76C819BFE69AED0F1808ACFFDA07BD" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AA32710BAB39FA292FDF04EB607591A8" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
is one of only two
|
||
<taxonomicName id="5B5BE3C0BC74D745DFD85DCF5DED95FB" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
|
||
genera (together with
|
||
<taxonomicName id="D90ABD4FA381A7B9E2DA324EFA67A6A5" authorityName="Walckenaer" authorityYear="1805" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="749C187D2C139316D2C755E1E407708C" italics="true">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
) with autochthonous species in both the Old and New Worlds. New World species are mostly known from the Greater Antilles and from semi-arid regions in
|
||
<collectingCountry id="939EC36839C3881DC5D8E1312B0CDB23" name="Brazil">Brazil</collectingCountry>
|
||
; the genus seems to be largely absent from the humid regions of the Amazon basin. Old World species are currently known from the Arabian Peninsula,
|
||
<collectingCountry id="2A4D4A658E1989B21A6F5319ACF4F01D" name="Morocco">Morocco</collectingCountry>
|
||
, and the
|
||
<collectingCountry id="AE13547E13E82A7594C14DE96EBCFA55" name="Philippines">Philippines</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="67707FBC807CC48B7DB5056A42DF98F2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Micropholcus Deeleman-Reinhold & Prinsen; SEM images of prosomata (frontal views) and tips of female palps (dorsal views) A M. tanomah Huber, sp. nov.; female B M. darbat Huber, sp. nov.; male C, D M. ghar Huber, sp. nov.; male and female E, F M. bukidnon Huber, sp. nov.; male and female; note small AME G M. tanomah Huber, sp. nov., left palp H M. ghar Huber, sp. nov., right palp. Scale bars: 100 µm (A – D); 200 µm (E, F); 10 µm (G, H)." figureDoi="10.3897/zookeys.1213.133178.figure5" httpUri="https://binary.pensoft.net/fig/1141415">5 B – D</figureCitation>
|
||
). Specimens have been collected from sea level to
|
||
<quantity id="935DFCEE052BE6C078709E8AAA7C7FCD" metricMagnitude="3" metricUnit="m" metricValue="2.37" unit="m" value="2370.0">2370 m</quantity>
|
||
. In
|
||
<collectingCountry id="8C4ABA811FE156544F46BDA04DCFD0B4" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
, all new species described herein were collected above
|
||
<quantity id="CFB5F799949DC383BF1666B17CD635D7" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
|
||
. Several of the localities visited below
|
||
<quantity id="3FE85150B701B080689FE6B311B7B8FF" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</quantity>
|
||
had suitable habitats but no
|
||
<taxonomicName id="27A9CE0B3BD1FC5E35CD77B6AF3C224B" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C57446CE689D84E977B19F59011ACDB2" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E3UAI" type="natural history">
|
||
<paragraph id="F105684282FFE4AA53CDA40326217EA6">
|
||
<heading id="C1C86A658B6640A91DE59E640ACB6FB2" reason="title">Natural history.</heading>
|
||
</paragraph>
|
||
<paragraph id="E6E7438A80D42B1A3EFE82D04402B6FB">
|
||
Old World
|
||
<taxonomicName id="E9E0C8081EF5AFC060EA8AB79B0738A0" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="585867F900226318245F50D2B6981AFD" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
seem to be very homogeneous with respect to their preferred microhabitats. Most species have been collected from rocks: in caves and at cave entrances, in small caverns of rock walls, and on the undersides of large boulders (Fig.
|
||
<figureCitation id="68B5845DB62BD04571112D4E77825F8F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Typical habitats of Micropholcus Deeleman-Reinhold & Prinsen in the Old World A Saudi Arabia, ‘ Asir, SE of Harajah (type locality of M. harajah Huber, sp. nov.) B Saudi Arabia, ‘ Asir, S of Al Fara (type locality of M. alfara Huber, sp. nov.) C Saudi Arabia, ‘ Asir, N of Abha (type locality of M. abha Huber, sp. nov.) D Saudi Arabia, Mecca, NW of Maysaan (type locality of M. maysaan Huber, sp. nov.) E Oman, Dhofar, near Shaat sinkhole (type locality of M. shaat Huber, sp. nov.) F Morocco, Fès-Meknès, Kef El Ghar (type locality of M. ghar Huber, sp. nov.) G Morocco, Béni Mellal-Khénifra, Imi n’Ifri (M. khenifra Huber, Lecigne & Lips, sp. nov.) H Philippines, Mindanao, Kabyaw Cave (M. bukidnon Huber, sp. nov.). Photos BAH." figureDoi="10.3897/zookeys.1213.133178.figure14" httpUri="https://binary.pensoft.net/fig/1141424">14</figureCitation>
|
||
). They share this type of microhabitat with most South American species, and with the majority of species of the putative sister genus (
|
||
<taxonomicName id="36B4A0EED1033E6A376B982AFD62C429" authorityName="Huber" authorityYear="2018" class="Arachnida" family="Pholcidae" genus="Cantikus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D68A8CE3978FB07D52CE69CA2BF02275" italics="true">Cantikus</emphasis>
|
||
</taxonomicName>
|
||
), suggesting that this might be the plesiomorphic microhabitat. The unusual microhabitat reported for
|
||
<taxonomicName id="5DC2120BDCF15C153B695A034409CC9F" authorityName="Deeleman-Reinhold & van Harten" authorityYear="2001" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="jacominae">
|
||
<emphasis id="C97F39E8C9034185B47748B2FA4D7424" italics="true">M. jacominae</emphasis>
|
||
</taxonomicName>
|
||
(dry plant debris in an irrigated banana plantation) needs confirmation. The synanthropic
|
||
<taxonomicName id="70FBF5E8E186CAE90397CBD0BA9D79D3" baseAuthorityName="Simon" baseAuthorityYear="1887" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fauroti">
|
||
<emphasis id="A3AE7EDC2EF726754F4AD8E3F8720178" italics="true">M. fauroti</emphasis>
|
||
</taxonomicName>
|
||
is usually found in the upper corners and edges of rooms. Caribbean species have shifted to the undersides of leaves, probably explaining their significantly different general body shape and colour (see
|
||
<bibRefCitation id="2278007BA4C52F20789DCB53B4BD57AC" DOI="10.1080/00222930601051196" author="Huber and Wunderlich" firstAuthor="Huber" journalOrPublisher="Journal of Natural History" pagination="2341-2360" refId="B26" refString="Huber BA, Wunderlich J (2006) Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae). Journal of Natural History 40: 2341–2360. https://doi.org/10.1080/00222930601051196" title="Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae)." volume="40" year="2006">Huber and Wunderlich 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9DB800558B15D10A71C70F6E7690B47A" DOI="10.1071/IS13050" author="Huber" etAl="et al." firstAuthor="Huber" journalOrPublisher="Invertebrate Systematics" pagination="432-450" refId="B28" refString="Huber BA, Carvalho LS, Benjamin SP (2014) On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae). Invertebrate Systematics 28: 432–450. https://doi.org/10.1071/IS13050" title="On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae)." volume="28" year="2014">Huber et al. 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="F6DF6E208ABBA746E84F2BEE7C4A4E00" ID-DOI="10.3897/zookeys.1213.133178.figure14" ID-arpha="E74B2949-B7B7-5433-B625-C8AA25646E89" httpUri="https://binary.pensoft.net/fig/1141424" startId="F14">
|
||
<paragraph id="91B8810EB4FBC2CB76F5C033D135CE79">
|
||
<label id="A0AA57400FA656B2B4107EC6128AABEE">Figure 14.</label>
|
||
</paragraph>
|
||
<paragraph id="283E05472294D3D312CD653F31110F08">
|
||
Typical habitats of
|
||
<taxonomicName id="B002C25AFD372850389C0A2A0C77DD70" authority="Deeleman-Reinhold & Prinsen" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01128DDC4246C20DACBB8EFD59FC0508" italics="true">Micropholcus</emphasis>
|
||
Deeleman-Reinhold & Prinsen
|
||
</taxonomicName>
|
||
in the Old World
|
||
<emphasis id="F5894164A1DE5BDEC7F5607EA7829F92" bold="true">A</emphasis>
|
||
Saudi Arabia, ‘ Asir, SE of Harajah (type locality of
|
||
<taxonomicName id="578C006674BA78EDB02F567CC80DAE97" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="harajah" status="sp. nov.">
|
||
<emphasis id="523925C992FA9216D1F0B6E662875069" italics="true">M. harajah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="44047C870EF40B97470A9E21FB262421" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="E371D883F2726A0CB41964DDE51FA400" bold="true">B</emphasis>
|
||
Saudi Arabia, ‘ Asir, S of Al Fara (type locality of
|
||
<taxonomicName id="3E49AA8374A587AE7C7D8274D3997DFE" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alfara" status="sp. nov.">
|
||
<emphasis id="2CBB98D3441D2EEDE864BE553D181E94" italics="true">M. alfara</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F2CDE2293BB6ECE1BB96DF25677CC7D4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="C2EE86926E731078F67A35C03976098A" bold="true">C</emphasis>
|
||
Saudi Arabia, ‘ Asir, N of Abha (type locality of
|
||
<taxonomicName id="E81DF11C1ED91DAF3DDD1252BC1C7613" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="abha" status="sp. nov.">
|
||
<emphasis id="5E231800627BD4576456F72933331C1B" italics="true">M. abha</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="BAD505B440A6919630AE7846656C6694" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="6CE6A878BF1B01637F468C654382F64A" bold="true">D</emphasis>
|
||
Saudi Arabia, Mecca, NW of Maysaan (type locality of
|
||
<taxonomicName id="97C9DEE22B50D0A47FF162295E1B0159" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="maysaan" status="sp. nov.">
|
||
<emphasis id="E0E60CD109394BEAB32416C028DD81EB" italics="true">M. maysaan</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EB2DBA05E3168D8EEB24605A1E3B33F5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="DA123A42612D5697E9D883FDC4698CDA" bold="true">E</emphasis>
|
||
Oman, Dhofar, near Shaat sinkhole (type locality of
|
||
<taxonomicName id="FEF6C65B13406CC4A50E6B02445CB745" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="shaat" status="sp. nov.">
|
||
<emphasis id="AF81018B1029F739E51AAA786C2A0636" italics="true">M. shaat</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C47D2FE607F0B93E734D7EC82504CD09" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="B1EAC6CB8E13415CE859AF820D1BCB3B" bold="true">F</emphasis>
|
||
Morocco, Fès-Meknès, Kef El Ghar (type locality of
|
||
<taxonomicName id="5319633AF21819069A3086C769864036" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="EF4946F09B3D09C20C812777EE3E344A" italics="true">M. ghar</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="E85BED491481AB20FD76BDAB7A58B88E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="23E19A54A954F8AF578D3D6A08F7F5DB" bold="true">G</emphasis>
|
||
Morocco, Béni Mellal-Khénifra, Imi n’Ifri (
|
||
<taxonomicName id="432EA772DEC9E00E9DE2AE1CC83A15B7" authority="Huber, Lecigne & Lips" authorityName="Huber, Lecigne & Lips" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="khenifra" status="sp. nov.">
|
||
<emphasis id="530133AD601638994E8EEC3B32172576" italics="true">M. khenifra</emphasis>
|
||
Huber, Lecigne & Lips
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="78FC4C61118733BDF32EA0546892E056" rank="species">sp. nov.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="D6560AF6E1F2AC3F954922C25BE26398" bold="true">H</emphasis>
|
||
Philippines, Mindanao, Kabyaw Cave (
|
||
<taxonomicName id="AAD6D8F83E43EA06D41A627140ACF432" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="AC8D39CCFF1C3CCD079BBFD3AC3F17BD" italics="true">M. bukidnon</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D82A36F66AD1C5D0EE7B0B3E8DFA1DB0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Photos BAH.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="95A1D2465B17E7CA721004DE61AFE964">
|
||
Old World and Brazilian
|
||
<taxonomicName id="07F5F683EDEB745330CBE9EC5338A91E" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5DB11B9038CB5E74A93FE980920B76FF" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
spiders build fine dome-shaped webs but during the day, most species (except those deeper in caves, e. g.,
|
||
<taxonomicName id="03E0FBBC4076695F9C62E2BF24869B60" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="C75D8BA95D83C727AA8FEA8A647A98D6" italics="true">M. ghar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="602AFC0082E451AAC047331F82191E55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) sit flat on the rock surface (Fig.
|
||
<figureCitation id="20DE98D7EF3611ED4AD30370177F5DE7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Section of cave ceiling (16 × 11 cm), showing nine adult specimens of M. tanomah Huber, sp. nov.; from NW of Tanomah, ‘ Asir, Saudi Arabia." figureDoi="10.3897/zookeys.1213.133178.figure15" httpUri="https://binary.pensoft.net/fig/1141425">15</figureCitation>
|
||
). The webs of leaf-dwelling Caribbean species have not yet been described. Upon disturbance, the spiders show a range of reactions, from refusing to move, bouncing, walking or running away, to dropping out of the web. Egg sacs are round (in Caribbean species elongated), covered by a barely visible sparse layer of silk (Figs
|
||
<figureCitation id="908CF1CAE7E81EF806265B96EEEFA22D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Saudi Arabia A M. dhahran Huber, sp. nov., male from ‘ Asir, W of Dhahran Al Janub B M. harajah Huber, sp. nov., female with egg-sac from ‘ Asir, SE of Harajah C, D M. alfara Huber, sp. nov., male and female with egg-sac from ‘ Asir, S of Al Fara E M. abha Huber, sp. nov., male from ‘ Asir, N of Abha F M. tanomah Huber, sp. nov., male from ‘ Asir, NW of Tanomah G M. bashayer Huber, sp. nov., female with egg-sac from ‘ Asir, NW of Al Bashayer H M. maysaan Huber, sp. nov., male from Mecca, NW of Maysaan. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure3" httpUri="https://binary.pensoft.net/fig/1141413">3</figureCitation>
|
||
,
|
||
<figureCitation id="C445C2630ACF053EA9F0932FCC0311B6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Micropholcus Deeleman-Reinhold & Prinsen; live specimens from Oman, Morocco, and the Philippines A, B M. darbat Huber, sp. nov., male and female with egg-sac from Oman, Dhofar, near Qairoon Hairitti C M. shaat Huber, sp. nov., male from Oman, Dhofar, Shaat sinkhole D, E M. agadir (Huber), male and female with egg-sac from Morocco, Souss-Massa, Paradise Valley F M. ghar Huber, sp. nov., male from Morocco, Fès-Meknès, Kef El Ghar G M. khenifra Huber, Lecigne & Lips, sp. nov., male from Morocco, Béni Mellal-Khénifra, near Sidi Ben Daoud H M. bukidnon Huber, sp. nov., male from Philippines, Mindanao, Blue Water Cave. Photographs BAH." figureDoi="10.3897/zookeys.1213.133178.figure4" httpUri="https://binary.pensoft.net/fig/1141414">4</figureCitation>
|
||
), and contain up to ~
|
||
<specimenCount id="941278568F12E7F3F01A0BE7D9A6F584" count="35" type="egg">35 eggs</specimenCount>
|
||
; egg diameters range from
|
||
<quantity id="AD22234D297145896AAD2D67F59B1946" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.1" metricValueMin="5.4" unit="mm" value="0.625" valueMax="0.71" valueMin="0.54">0.54 to 0.71 mm</quantity>
|
||
. For more detailed observations on Brazilian species, see
|
||
<bibRefCitation id="8A6E0C5B555418A5B43DD5B5446471EA" DOI="10.1071/IS13050" author="Huber BA & Carvalho LS & Benjamin SP" etAl="et al." firstAuthor="Huber" journalOrPublisher="Invertebrate Systematics" pagination="432-450" refId="B28" refString="Huber BA, Carvalho LS, Benjamin SP (2014) On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae). Invertebrate Systematics 28: 432–450. https://doi.org/10.1071/IS13050" title="On the New World spiders previously misplaced in Leptopholcus: molecular and morphological analyses and descriptions of four new species (Araneae, Pholcidae)." volume="28" year="2014">Huber et al. (2014)</bibRefCitation>
|
||
. For detailed life history data of
|
||
<taxonomicName id="8FBC741940DA5AFEA5BC5F7A396B783B" baseAuthorityName="Simon" baseAuthorityYear="1887" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fauroti">
|
||
<emphasis id="D627101BA75FA61758CB2560C7CC3E8B" italics="true">M. fauroti</emphasis>
|
||
</taxonomicName>
|
||
under lab conditions see
|
||
<bibRefCitation id="6F54174A93449FEB426EC3B01D6191CD" DOI="10.21608/ajesa.2017.164166" author="Ahmad NFR & Abou-Setta MM" firstAuthor="Ahmad" journalOrPublisher="Acarines" pagination="31-35" refId="B2" refString="Ahmad NFR, Abou-Setta MM (2017) Life history of the pholcid spider, Micropholcus fauroti (Simon, 1887) (Araneae: Pholcidae) in Egypt. Acarines 11: 31–35. https://doi.org/10.21608/ajesa.2017.164166" title="Life history of the pholcid spider, Micropholcus fauroti (Simon, 1887) (Araneae: Pholcidae) in Egypt." volume="11" year="2017">Ahmad and Abou-Setta (2017)</bibRefCitation>
|
||
. For anecdotal observations on further Old World species, see individual species descriptions below.
|
||
</paragraph>
|
||
<caption id="1E5DC66E14B8CA26E62188ADBC1CC713" ID-DOI="10.3897/zookeys.1213.133178.figure15" ID-arpha="3F506B7C-0B62-5A64-991D-FF4192E3BB2D" httpUri="https://binary.pensoft.net/fig/1141425" startId="F15">
|
||
<paragraph id="4A4F455F7FC757A6CF9B133E072DEB2B">
|
||
<label id="F1AE6A57827EBFFFC079233B45B41CC1">Figure 15.</label>
|
||
</paragraph>
|
||
<paragraph id="6ABFAFB862E881CF4D2137A920E94EDE">
|
||
Section of cave ceiling (16 × 11 cm), showing nine adult specimens of
|
||
<taxonomicName id="954DD803786EDB55E3A9B786AC594D01" authority="Huber" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tanomah" status="sp. nov.">
|
||
<emphasis id="E16A6C87F12D0842570571577BC99316" italics="true">M. tanomah</emphasis>
|
||
Huber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="7B92076B3FADA9719D8DC8CD99644682" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; from NW of Tanomah, ‘ Asir, Saudi Arabia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZ3AI" type="Relationships">
|
||
<paragraph id="461D714459AE24A4A744C56BAA374F74">
|
||
<heading id="D7AEAE5B2745AFE3EC35ABB9F32D6BE3" reason="title">Relationships.</heading>
|
||
</paragraph>
|
||
<paragraph id="EEC9A87FF2CC96A08D7CD410958AF351">
|
||
The molecular analysis of
|
||
<bibRefCitation id="ADD7A674DCC6851B59ECEC4E56DF048D" DOI="10.1186/s12862-018-1244-8" author="Eberle J & Dimitrov D & Valdez-Mondragón A & Huber BA" etAl="et al." firstAuthor="Eberle" refId="B12" refString="Eberle J, Dimitrov D, Valdez-Mondragón A, Huber BA (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology 18: 141. https://doi.org/10.1186/s12862-018-1244-8" year="2018">Eberle et al. (2018)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="BD3D6AF9289C6CFD78B021587601CA13" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F7AE2735224F46B7E2F4779480E0177" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
as sister to the South-East Asian genus
|
||
<taxonomicName id="E8610DA66F94681D3471AB9FD5460880" authority="Huber, 2018" authorityName="Huber" authorityYear="2018" class="Arachnida" family="Pholcidae" genus="Cantikus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9D5DBFB918137852E3AAA19A1A291A8A" italics="true">Cantikus</emphasis>
|
||
<bibRefCitation id="65E028B81B4A0300F485657D4DF8EB45" DOI="10.3897/zookeys.789.22781" author="Huber BA & Eberle J & Dimitrov D" journalOrPublisher="ZooKeys" pagination="51-101" refId="B30" refString="Huber BA, Eberle J, Dimitrov D (2018) The phylogeny of pholcid spiders: a critical evaluation of relationships suggested by molecular data (Araneae, Pholcidae). ZooKeys 789: 51 - 101. https: // doi. org / 10.3897 / zookeys. 789.22781" title="The phylogeny of pholcid spiders: a critical evaluation of relationships suggested by molecular data (Araneae, Pholcidae)." volume="789" year="2018">Huber, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
, within a clade that included three further Old World genera:
|
||
<taxonomicName id="72592FB5C4DD523163D337DCDB095744" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Pholcidae" genus="Leptopholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BDB06BC23C63FEDF99848CEFEB4774E9" italics="true">Leptopholcus</emphasis>
|
||
Simon, 1893
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="EA51DC3BB37C8FA82162D9DF179A395E" authority="Deeleman-Reinhold & van Harten, 2001" authorityName="Deeleman-Reinhold & van Harten" authorityYear="2001" class="Arachnida" family="Pholcidae" genus="Pehrforsskalia" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23AA61254E2B85190453DE0C23663806" italics="true">Pehrforsskalia</emphasis>
|
||
<bibRefCitation id="249FB8BB8A8F1FE39C604F724F8CCFA5" author="Deeleman-Reinhold CL & van Harten A" editor="Ishwar Prakash" journalOrPublisher="Scientific Publishers, India, Jodhpur" pagination="193-207" refId="B9" refString="Deeleman-Reinhold CL, van Harten A (2001) Description of some interesting, new or little known Pholcidae (Araneae) from Yemen. In: Ishwar Prakash (Ed.) Ecology of Desert Environments. Scientific Publishers, India, Jodhpur, 193 - 207." title="Description of some interesting, new or little known Pholcidae (Araneae) from Yemen." volumeTitle="Ecology of Desert Environments." year="2001">Deeleman-Reinhold & van Harten, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="3AFCCBE27CB514EC3DECE338DC0A25FC" authority="Bradley, 1877" authorityName="Bradley" authorityYear="1877" class="Arachnida" family="Pholcidae" genus="Micromerys" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8C7D77FFF37FC67DC0F70DD6846E2BD3" italics="true">Micromerys</emphasis>
|
||
Bradley, 1877
|
||
</taxonomicName>
|
||
. That analysis included ten species of
|
||
<taxonomicName id="A000EE65601507F7B0F14C350312F580" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B8403F3CC303E7EC86978DCCC92488A7" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
: the type species
|
||
<taxonomicName id="545497C5CC6269D9BBBE49EACE207A75" baseAuthorityName="Simon" baseAuthorityYear="1887" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="fauroti">
|
||
<emphasis id="ADF1D5109E18EA09C1362263184402C0" italics="true">M. fauroti</emphasis>
|
||
</taxonomicName>
|
||
, two further Old World species (
|
||
<taxonomicName id="1FC9BCD88EC742FA43B8495BF9E92022" baseAuthorityName="Huber" baseAuthorityYear="2011" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="agadir">
|
||
<emphasis id="1A13C2F5CBA3B284EA5E2876B3B53704" italics="true">M. agadir</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2E3A87F7C5D6080E0CDFB4B949B9095F" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bukidnon" status="sp. nov.">
|
||
<emphasis id="CE956EB43DF299F304D062211DD61BBB" italics="true">M. bukidnon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="23A6E1660A09553C1D86E79A564934A1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and seven New World species. The monophyly of the genus received maximum support. New World species were nested within a paraphyletic Old World group.
|
||
</paragraph>
|
||
<paragraph id="AA34E2FA93178E5E49CCCFB247ACD8E9">
|
||
Our
|
||
<abbrev id="ABBRID0EQ6AI" xlink_title="neighbor-joining">NJ</abbrev>
|
||
tree (Fig.
|
||
<figureCitation id="30F30992CEAAD06B0FE6BB68E4DC491E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Neighbor-joining tree of analysed CO 1 sequences using the Kimura 2 - parameter model; numbers on the branches are bootstrap supports from 5000 replications (%). The labels North and South refer to two geographically and morphologically distinct species groups within Saudi Arabia." figureDoi="10.3897/zookeys.1213.133178.figure1" httpUri="https://binary.pensoft.net/fig/1141411">1</figureCitation>
|
||
) is not expected to reliably reflect phylogenetic relationships. However, some clades that receive reasonable to high support are either congruent with the results of
|
||
<bibRefCitation id="5389827025D0A136091B81AE1E21CFB5" DOI="10.1186/s12862-018-1244-8" author="Eberle J & Dimitrov D & Valdez-Mondragón A & Huber BA" etAl="et al." firstAuthor="Eberle" refId="B12" refString="Eberle J, Dimitrov D, Valdez-Mondragón A, Huber BA (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology 18: 141. https://doi.org/10.1186/s12862-018-1244-8" year="2018">Eberle et al. (2018)</bibRefCitation>
|
||
(Caribbean clade) or include geographically neighbouring species: the Moroccan clade, the southern Saudi Arabian clade, and the northern Saudi Arabian clade. The latter two are also supported by several morphological similarities each, but it is not clear which of these are synapomorphies and which not. Thus, relationships within
|
||
<taxonomicName id="4CB10076D6B9C9A66C74ED94D9908C9A" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F468F0E42E60440B407B600BF411C500" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
are largely unresolved and Fig.
|
||
<figureCitation id="34445F002B03588D5A25BED1217F3704" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Neighbor-joining tree of analysed CO 1 sequences using the Kimura 2 - parameter model; numbers on the branches are bootstrap supports from 5000 replications (%). The labels North and South refer to two geographically and morphologically distinct species groups within Saudi Arabia." figureDoi="10.3897/zookeys.1213.133178.figure1" httpUri="https://binary.pensoft.net/fig/1141411">1</figureCitation>
|
||
should not be misinterpreted in a phylogenetic context.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EHABI" type="Composition">
|
||
<paragraph id="E1D05CED3CFA40BCA27806B35C8909EE">
|
||
<heading id="0BA94096F2EC76D65C5857D0EB9E135A" reason="title">Composition.</heading>
|
||
</paragraph>
|
||
<paragraph id="9019B98E58632A413D1CA91AEE0B983E">
|
||
The genus now includes 30 described species:
|
||
<collectingCountry id="37B3A38F6E531899EE4CE2909DFECE7A" name="Dominican Republic">the Dominican</collectingCountry>
|
||
amber fossil
|
||
<taxonomicName id="A6CBBB2A7FBC871D6FDCCE52A215935A" authority="(Huber & Wunderlich, 2006)" baseAuthorityName="Huber & Wunderlich" baseAuthorityYear="2006" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kiskeya">
|
||
<emphasis id="6CF1AF3D97D2F0EC7B75E5E18C37ABDE" italics="true">M. kiskeya</emphasis>
|
||
(
|
||
<bibRefCitation id="9BC3A797819A3EC0EBD624A0A738C3AD" DOI="10.1080/00222930601051196" author="Huber BA & Wunderlich J" journalOrPublisher="Journal of Natural History" pagination="2341-2360" refId="B26" refString="Huber BA, Wunderlich J (2006) Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae). Journal of Natural History 40: 2341 - 2360. https: // doi. org / 10.1080 / 00222930601051196" title="Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first case of egg-parasitism in pholcid spiders (Araneae: Pholcidae)." volume="40" year="2006">Huber & Wunderlich, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and 29 extant species. Of the latter, seven occur in South America, six on the Caribbean islands, and
|
||
<quantity id="AE184DC432B48AEDD9243359B170AC4E" metricMagnitude="-1" metricUnit="m" metricValue="4.064" unit="in" value="16.0">16 in</quantity>
|
||
the Old World. All Old World species are treated below except for
|
||
<taxonomicName id="DDAE353972168194419727648A189E57" authority="Barrientos, 2019" authorityName="Barrientos" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tegulifer">
|
||
<emphasis id="C33BC1B524AA60E534C7E36E973FBC7C" italics="true">M. tegulifer</emphasis>
|
||
Barrientos, 2019
|
||
</taxonomicName>
|
||
(a loan request was denied by the curator of arthropods, Museu de Ciències Naturals de Barcelona). Numerous undescribed New World species are available in collections, in particular from
|
||
<collectingCountry id="480DDE136F8D6922DDB588178D9DB848" name="Brazil">Brazil</collectingCountry>
|
||
(L. S. Carvalho, pers. comm.
|
||
<date id="61B7B696E538C8DE28F9E8199508D6F3" value="2020-07-02">2 July 2020</date>
|
||
). At least one further undescribed species is known to occur in
|
||
<collectingCountry id="97656DE19D8312E571AAF27D30BB6BC4" name="Morocco">Morocco</collectingCountry>
|
||
, represented by a single male specimen deposited in the Muséum d’histoire naturelle,
|
||
<collectingRegion id="F4570ED7E359393AC22B5180D0AB1950" country="Switzerland" name="Geneve">Genève</collectingRegion>
|
||
,
|
||
<collectingCountry id="5737DCA61C20325F4705DE23871CBC16" name="Switzerland">Switzerland</collectingCountry>
|
||
(“ sp. Gen 377 ” in Fig.
|
||
<figureCitation id="B21173707ABF290979F813DAB8C08743" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Known distribution of Micropholcus fauroti (Simon) (A) and of Old World Micropholcus in Mindanao (B), on the Arabian Peninsula (C) and in Morocco (D)." figureDoi="10.3897/zookeys.1213.133178.figure13" httpUri="https://binary.pensoft.net/fig/1141423">13 D</figureCitation>
|
||
). It resembles
|
||
<taxonomicName id="F2CF26AB9C47B9D45C027C44BE6A57A4" authorityName="Barrientos" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tegulifer">
|
||
<emphasis id="2897E2C604A6163911A9FDF99A21A4EF" italics="true">M. tegulifer</emphasis>
|
||
</taxonomicName>
|
||
but has a very different uncus. It originates from the Gorges du Dades area in the Drâa-Tafilalet Region, ~
|
||
<named-content id="BA8739515B6D665A9B1D407F2CA96E3B" content-type="dwc:verbatimCoordinates">
|
||
<geoCoordinate id="825F4237B6DA12B5D623B274B7F1228B" degrees="31.535" direction="north" orientation="latitude" precision="55" value="31.535">31.535 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="797330921D84EA1F3529F64A040ACDDD" degrees="5.918" direction="west" orientation="longitude" precision="55" value="-5.918">5.918 ° W</geoCoordinate>
|
||
</named-content>
|
||
. Our molecular data indicate that the “
|
||
<taxonomicName id="BE5A0CE6A426BFC1EF07D9FEDE40240F" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="461EA35B052882B9C9159A3CBB3338B4" italics="true">Pholcus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
” CO 1 barcode published in
|
||
<bibRefCitation id="603A7A9DDD1BFDFFB41DEC3A0ED2CCE0" DOI="10.1016/j.ympev.2008.04.027" author="Dimitrov D & Arnedo MA & Ribera C" etAl="et al." firstAuthor="Dimitrov" journalOrPublisher="Molecular Phylogenetics and Evolution" pagination="596-614" refId="B10" refString="Dimitrov D, Arnedo MA, Ribera C (2008) Colonization and diversification of the spider genus Pholcus Walckenaer, 1805 (Araneae, Pholcidae) in the Macaronesian archipelagos: Evidence for long-term occupancy yet rapid speciation. Molecular Phylogenetics and Evolution 48: 596–614. https://doi.org/10.1016/j.ympev.2008.04.027" title="Colonization and diversification of the spider genus Pholcus Walckenaer, 1805 (Araneae, Pholcidae) in the Macaronesian archipelagos: Evidence for long-term occupancy yet rapid speciation." volume="48" year="2008">Dimitrov et al. (2008)</bibRefCitation>
|
||
is also from a
|
||
<taxonomicName id="30DB6345A2FF31943B2E41E6A3B0FC2E" authorityName="Deeleman-Reinhold & Prinsen, SEM" authorityYear="1987" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B0104B97E50D4F7A03A1495B360827A6" italics="true">Micropholcus</emphasis>
|
||
</taxonomicName>
|
||
, different from
|
||
<taxonomicName id="9A8288880415C9FCB555DD126C455416" baseAuthorityName="Huber" baseAuthorityYear="2011" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="agadir">
|
||
<emphasis id="AA489FDB811E7F6D383DDA3E3EA38D93" italics="true">M. agadir</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0B6006048094BBEFF8E614D9664EF012" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="ghar" status="sp. nov.">
|
||
<emphasis id="DBBCD09ADE9C96A0655F8D694648564C" italics="true">M. ghar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3477B076547EA7FF08043EE564A298BE" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="1053000BD141B44BD845846B2125A9A8" authorityName="Huber & Meng" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="khenifra" status="sp. nov.">
|
||
<emphasis id="2C1BA290F9CAB132BC4AD80DC1D716AA" italics="true">M. khenifra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12C0C93DD634D10AC88E7A5C0C72A4D6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Figs
|
||
<figureCitation id="4BFDD49C85E4A9AC32D7F11B0A658D84" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Neighbor-joining tree of analysed CO 1 sequences using the Kimura 2 - parameter model; numbers on the branches are bootstrap supports from 5000 replications (%). The labels North and South refer to two geographically and morphologically distinct species groups within Saudi Arabia." figureDoi="10.3897/zookeys.1213.133178.figure1" httpUri="https://binary.pensoft.net/fig/1141411">1</figureCitation>
|
||
,
|
||
<figureCitation id="7F56E13043E2725C3B4DE523C978FAC7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. ASAP species delimitation analysis based on CO 1 data. The analysis was performed on the web server https://bioinfo.mnhn.fr/abi/public/asap/. The Kimura 2 - parameter model was applied. The ten best partitions are shown. The numbers for each column (partition) are: (1) Total number of species as identified by ASAP in the corresponding partition. (2) Score, an indicator of how good a partition is (the lower the score, the better the partition). (3) Rank of the scores. Note that the partition with the lowest (best) score splits M. maysaan sp. nov. into two species but joins M. alfara sp. nov. and M. dhahran sp. nov." figureDoi="10.3897/zookeys.1213.133178.figure2" httpUri="https://binary.pensoft.net/fig/1141412">2</figureCitation>
|
||
). It could be
|
||
<taxonomicName id="F39BFF2C2494A37ED24E6CD34CABB007" authorityName="Barrientos" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Micropholcus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tegulifer">
|
||
<emphasis id="CEB46DD15DEF209AE6EDA3964025858D" italics="true">M. tegulifer</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="1224C5E76EF644FBE69AA05A592947EC" italics="true">M.</emphasis>
|
||
sp. Gen 377, or a different new species. Its geographic origin cannot be reconstructed, and the only available (juvenile) specimen is lost (D. Dimitrov and C. Ribera, pers. comm.
|
||
<date id="0B70786B37D0D342FA882A0439D3C6F3" value="2024-03-20" valueMax="2024-04-19" valueMin="2024-03-20">20 Mar. and 19 Apr. 2024</date>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |