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<mods:title id="020D4BC55E466797CA451F333E5B6975">A checklist of bats (Mammalia: Chiroptera) from Lao PDR</mods:title>
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<mods:namePart id="AA35746C2E9C3D382F8B08011E3D2E6C">Thomas, Nikky M.</mods:namePart>
<mods:affiliation id="0AD077E5884CF2C9F666A23E94EAC9B2">Harrison Institute, St Botolphs Road, Sevenaoks, Kent TN 13 3 AQ, U. K. &amp; Corresponding author: E-mail: nikkythomas @ hotmail. com</mods:affiliation>
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<mods:namePart id="C57F46C98293B0F5437C7107B4AD043B">Duckworth, J. W.</mods:namePart>
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<mods:namePart id="16B34FA594EA7BBF9B517E25AA5F2C6E">Douangboubpha, Bounsavane</mods:namePart>
<mods:affiliation id="0245912FE1FCD0F07D3CAD83211C372E">Faculty of Environmental Sciences, National University of Laos, Dong Dok Campus, Xaythany District, Vientiane Capital, Lao PDR</mods:affiliation>
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<mods:affiliation id="8DFD845074C3E09FB9550D7B3910A37D">University of Greenwich School of Sciences, Room 215, Grenville Building, Medway Campus, U. K. Central Avenue, Chatham Maritime, Kent ME 4 4 TB, U. K.</mods:affiliation>
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<mods:namePart id="5838D3A1043B4687152B24A490EF6D34">Francis, Charles M.</mods:namePart>
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<treatment id="690487A5FFD1FF9D0EBAFA54FE6ED667" ID-DOI="http://doi.org/10.5281/zenodo.4342759" ID-GBIF-Taxon="165089060" ID-Zenodo-Dep="4342759" LSID="urn:lsid:plazi:treatment:690487A5FFD1FF9D0EBAFA54FE6ED667" httpUri="http://treatment.plazi.org/id/690487A5FFD1FF9D0EBAFA54FE6ED667" lastPageId="19" lastPageNumber="211" pageId="18" pageNumber="210">
<subSubSection id="A9B76538FFD1FF9C0EBAFA54FA85D6A0" box="[901,1344,1465,1491]" pageId="18" pageNumber="210" type="nomenclature">
<paragraph id="E11236B3FFD1FF9C0EBAFA54FA85D6A0" blockId="18.[816,1429,1465,1579]" box="[901,1344,1465,1491]" pageId="18" pageNumber="210">
<heading id="BA5A81DFFFD1FF9C0EBAFA54FA85D6A0" box="[901,1344,1465,1491]" pageId="18" pageNumber="210">
<taxonomicName id="26AD4D30FFD1FF9C0EBAFA54FA85D6A0" ID-CoL="6X62W" authority="Andersen, 1905" authorityName="Andersen" authorityYear="1905" box="[901,1344,1465,1491]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="210" phylum="Chordata" rank="species" species="thomasi">
<emphasis id="D3D9EAA1FFD1FF9C0EBAFA54FBBAD6A0" box="[901,1151,1465,1491]" italics="true" pageId="18" pageNumber="210">Rhinolophus thomasi</emphasis>
<bibRefCitation id="853C4B42FFD1FF9C09BAFA54FA85D6A0" author="ANDERSEN, K." box="[1157,1344,1465,1491]" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="18" pageNumber="210" pagination="75 - 145" part="2" refId="ref28355" refString="ANDERSEN, K. 1905. On some bats of the genus Rhinolophus. Proceedings of the Zoological Society of London, 2: 75 - 145." title="On some bats of the genus Rhinolophus" type="journal article" year="1905">Andersen, 1905</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="A9B76538FFD1FF9C0EEAFA37FB35D687" box="[981,1264,1498,1524]" pageId="18" pageNumber="210" type="vernacular_names">
<paragraph id="E11236B3FFD1FF9C0EEAFA37FB35D687" blockId="18.[816,1429,1465,1579]" box="[981,1264,1498,1524]" pageId="18" pageNumber="210">Thomass horseshoe bat</paragraph>
</subSubSection>
<subSubSection id="A9B76538FFD1FF9C0E0FFA17FC5AD558" pageId="18" pageNumber="210" type="reference_group">
<paragraph id="E11236B3FFD1FF9C0E0FFA17FC5AD558" blockId="18.[816,1429,1465,1579]" pageId="18" pageNumber="210">
<treatmentCitationGroup id="C1BD119DFFD1FF9C0E0FFA17FC5AD558" pageId="18" pageNumber="210">
<treatmentCitation id="600C10A2FFD1FF9C0E0FFA17FB23D57C" box="[816,1254,1530,1551]" pageId="18" pageNumber="210">
<taxonomicName id="26AD4D30FFD1FF9C0E0FFA17FB23D57C" ID-CoL="6X62W" authority="Andersen, 1905: 100" authorityName="Andersen" authorityPageNumber="100" authorityYear="1905" box="[816,1254,1530,1551]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="210" phylum="Chordata" rank="species" species="thomasi">
<emphasis id="D3D9EAA1FFD1FF9C0E0FFA17FBC7D57C" box="[816,1026,1530,1551]" italics="true" pageId="18" pageNumber="210">Rhinolophus thomasi</emphasis>
<bibRefCitation id="853C4B42FFD1FF9C0931FA17FB23D57C" author="ANDERSEN, K." box="[1038,1254,1530,1551]" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="18" pageNumber="210" pagination="75 - 145" part="2" refId="ref28355" refString="ANDERSEN, K. 1905. On some bats of the genus Rhinolophus. Proceedings of the Zoological Society of London, 2: 75 - 145." title="On some bats of the genus Rhinolophus" type="journal article" year="1905">Andersen, 1905: 100</bibRefCitation>
</taxonomicName>
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; Karin Hills, SE
<collectingCountry id="99BA7623FFD1FF9C0E69F9FBFC5FD558" box="[854,922,1558,1579]" name="Myanmar" pageId="18" pageNumber="210">Burma</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A9B76538FFD1FF9D0E0FF9BBFD95D0B2" lastPageId="19" lastPageNumber="211" pageId="18" pageNumber="210" type="distribution">
<paragraph id="E11236B3FFD1FF9C0E0FF9BBFC7AD503" blockId="18.[816,1429,1622,2017]" box="[816,959,1622,1648]" pageId="18" pageNumber="210">
<emphasis id="D3D9EAA1FFD1FF9C0E0FF9BBFC7AD503" box="[816,959,1622,1648]" italics="true" pageId="18" pageNumber="210">Distribution</emphasis>
</paragraph>
<paragraph id="E11236B3FFD1FF9C0E69F99AFAF3D5E2" blockId="18.[816,1429,1622,2017]" box="[854,1334,1655,1681]" pageId="18" pageNumber="210">
<collectingCountry id="99BA7623FFD1FF9C0E69F99AFC0FD5E2" box="[854,970,1655,1681]" name="Myanmar" pageId="18" pageNumber="210">Myanmar</collectingCountry>
,
<collectingCountry id="99BA7623FFD1FF9C0EE9F99AFBF9D5E2" box="[982,1084,1655,1681]" name="Vietnam" pageId="18" pageNumber="210">Vietnam</collectingCountry>
,
<collectingCountry id="99BA7623FFD1FF9C0977F99AFB76D5E2" box="[1096,1203,1655,1681]" name="Thailand" pageId="18" pageNumber="210">Thailand</collectingCountry>
, Lao PDR.
</paragraph>
<paragraph id="E11236B3FFD1FF9D0E69F974FD95D0B2" blockId="18.[816,1429,1622,2017]" lastBlockId="19.[159,772,223,961]" lastPageId="19" lastPageNumber="211" pageId="18" pageNumber="210">
Distribution in Lao PDR (
<figureCitation id="79962A36FFD1FF9C09B2F974FB22D5C0" box="[1165,1255,1689,1715]" captionStart="FIG" captionStartId="19.[160,173,1997,2018]" captionTargetBox="[161,770,1348,1976]" captionTargetId="graphics@19.[184,746,1358,1964]" captionTargetPageId="19" captionText="FIG. 30. Distribution of Rhinolophus thomasi" figureDoi="http://doi.org/10.5281/zenodo.3943401" httpUri="https://zenodo.org/record/3943401/files/figure.png" pageId="18" pageNumber="210">Fig. 30</figureCitation>
):
<collectingRegion id="2369F851FFD1FF9C09C4F974FCA0D5A6" country="Laos" name="Attapu" pageId="18" pageNumber="210">Attapu Province</collectingRegion>
: Ban Paam, Dong Ampham NBCA (SMF collection);
<collectingRegion id="2369F851FFD1FF9C0EA5F931FB62D585" box="[922,1191,1756,1782]" country="Laos" name="Bolikhamxai" pageId="18" pageNumber="210">Bolikhamxai Province</collectingRegion>
: limestone near Lak Sao (Ban Lak-20); Mungkon Cave (Tham Moungkon) (ROM collection); Ban Kengbit, Nam Kading NBCA (SMF collection);
<collectingRegion id="2369F851FFD1FF9C09BAF8ACFA57D428" box="[1157,1426,1857,1883]" country="Laos" name="Champasak" pageId="18" pageNumber="210">Champasak Province</collectingRegion>
: Dong Hua Sao NBCA (IEX/SMF collections);
<collectingRegion id="2369F851FFD1FF9C0E0FF869FBE3D4ED" box="[816,1062,1924,1950]" country="Laos" name="Houaphan" pageId="18" pageNumber="210">Houaphan Province</collectingRegion>
: Ban Chak, Nam Et NBCA (ROM collection); Ban Bauphath, Nam Et NBCA; Dam on Nam Et, Nam Et NBCA (EBD/ROM collection); Nam Chong, Nam Et NBCA (CMF field record);
<collectingRegion id="2369F851FFD0FF9D0C36FEECFDDFD268" box="[265,538,257,283]" country="Laos" name="Khammouan" pageId="19" pageNumber="211">Khammouan Province</collectingRegion>
: Ban Mouangkhai; Ban Vieng; Tam Houay Si (Tham Houaysy); Tam Jongchott (Tham Chongchot); Tam Khun Fart (Tham Khounfat); Tam Pha Deng (Tham Phadeng); Tam Pha Tok Camp (Tham Phatok); Tam Quaie (Tham Khoay); Tam Sompoy (Tham Sompoy) (
<bibRefCitation id="853C4B42FFD0FF9D0D98FE21FDC1D295" author="Robinson and Webber" box="[167,516,460,486]" firstAuthor="Robinson" pageId="19" pageNumber="211" pagination="21 - 45" refId="ref31811" refString="ROBINSON, M. F., and M. WEBBER. 2000. Survey of bats (Mammalia: Chiroptera) in the Khammouan Limestone National Biodiversity Area, Lao P. D. R. Natural History Bulletin of the Siam Society, 48: 21 - 45." type="journal article" year="2000">Robinson and Webber, 2000</bibRefCitation>
); Ban Gnavang (Ban Gnavay), Xe Kaman; Ban Louang; Ban Vang Manua (Ban Vangma-Nua), Hin Namno NBCA; Ban Xam-Kang, Hin Namno NBCA; Nakai Plateau (ROM collection); Ban Khankeo,
<collectingRegion id="2369F851FFD0FF9D0F57FDB9FCC1D11D" box="[616,772,596,622]" country="Laos" name="Khammouan" pageId="19" pageNumber="211">Khammouan</collectingRegion>
Limestone NBCA (SMF collection); Ban Gnommalat; Ban Houana; Ban Mouangkhai; Ban Phonsang; Ban Nathan; Xe Bangfai Cave (Tham Xebangfai), Hin Namno NBCA (CMF field record);
<collectingRegion id="2369F851FFD0FF9D0DA0FD10FE0FD064" box="[159,458,765,791]" country="Laos" name="Louangnamtha" pageId="19" pageNumber="211">Louangnamtha Province</collectingRegion>
: Ban Eng; Ban Phoulan; New Ban Namsing (Ban Namsing-mai) (ROM collection);
<collectingRegion id="2369F851FFD0FF9D0C27FCACFDA5D028" box="[280,608,833,859]" country="Laos" name="Louangphabang" pageId="19" pageNumber="211">Louangphabang Province</collectingRegion>
: Nam Khan (ROM collection); Ban Sakok, Nam Et NBCA (CMF field record);
<collectingRegion id="2369F851FFD0FF9D0C9FFC68FD4FD0EC" box="[416,650,901,927]" country="Laos" name="Viangchan" pageId="19" pageNumber="211">Vientiane Province</collectingRegion>
: Ban Nampe; Ban Nathao (ROM collection).
</paragraph>
</subSubSection>
<subSubSection id="A9B76538FFD0FF9D0DA0FC06FE6ED667" pageId="19" pageNumber="211" type="discussion">
<paragraph id="E11236B3FFD0FF9D0DA0FC06FE9CD776" blockId="19.[159,772,1003,1300]" box="[159,345,1003,1029]" pageId="19" pageNumber="211">
<emphasis id="D3D9EAA1FFD0FF9D0DA0FC06FE9CD776" box="[159,345,1003,1029]" italics="true" pageId="19" pageNumber="211">Taxonomic note</emphasis>
</paragraph>
<paragraph id="E11236B3FFD0FF9D0DFAFBE0FE6ED667" blockId="19.[159,772,1003,1300]" pageId="19" pageNumber="211">
DNA barcodes suggest several genetically distinct clusters within bats currently referred to
<taxonomicName id="26AD4D30FFD0FF9D0DA0FBBDFDD2D719" authority="(Francis et al., 2010)" baseAuthorityName="Francis" baseAuthorityYear="2010" box="[159,535,1104,1130]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="211" phylum="Chordata" rank="species" species="thomasi">
<emphasis id="D3D9EAA1FFD0FF9D0DA0FBBDFEDED719" box="[159,283,1104,1130]" italics="true" pageId="19" pageNumber="211">R. thomasi</emphasis>
(
<bibRefCitation id="853C4B42FFD0FF9D0C16FBBDFDCAD719" author="Francis" box="[297,527,1104,1130]" etAl="et al." firstAuthor="Francis" pageId="19" pageNumber="211" pagination="12575" refId="ref30303" refString="FRANCIS, C. M., A. V. BORISENKO, N. V. IVANOVA, J. L. EGER, B. K. LIM, A. GUILLEN- SERVENT, S. V. KRUSKOP, I. MACKIE, and P. D. N. HEBERT. 2010. The role of DNA barcodes in understanding and conservation of mammal diversity in Southeast Asia. PLoS ONE, 5: e 12575. doi: 10.1371 / journal. pone. 0012575." type="journal article" year="2010">
Francis
<emphasis id="D3D9EAA1FFD0FF9D0CB7FBBDFE7FD719" box="[392,442,1104,1130]" italics="true" pageId="19" pageNumber="211">et al</emphasis>
., 2010
</bibRefCitation>
)
</taxonomicName>
. Some of these have been labelled as
<taxonomicName id="26AD4D30FFD0FF9D0C4AFB9FFE24D7FF" authorityName="Temminck" authorityYear="1835" box="[373,481,1138,1164]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="211" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="D3D9EAA1FFD0FF9D0C4AFB9FFE24D7FF" box="[373,481,1138,1164]" italics="true" pageId="19" pageNumber="211">R. rouxii</emphasis>
</taxonomicName>
, but further taxonomic study comparing morphology, echolocation calls and genetics is needed to determine whether any of these represent distinct species and, if so, what are the appropriate names.
</paragraph>
</subSubSection>
</treatment>
</document>